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<document id="E19E896BC4C724C57451B81CD022B90B" ID-DOI="10.11646/zootaxa.5372.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10146342" ID-ZooBank="1300D562-3E0B-4F72-933F-8FA7D28F6853" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700186674367" checkinUser="plazi" docAuthor="Mendonça, Maria Thayane Da Silva, Silva, Valéria Juliete Da &amp; Fernandes, José Antônio Marin" docDate="2023" docId="414B87B5FFCB9852FF63DED0776AFA5F" docLanguage="en" docName="zootaxa.5372.1.1.pdf" docOrigin="Zootaxa 5372 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5372.1.1/52272" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Edessa (Edessa) cornicula Mendonça &amp; Silva &amp; Fernandes 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="25" masterDocId="BD72FFCDFFD3984AFFF4DE477513FFC1" masterDocTitle="Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="25" updateTime="1700188109802" updateUser="ExternalLinkService">
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<mods:title id="8E1D0571B52F26EBA5CBC4D15C690475">Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)</mods:title>
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<mods:namePart id="103B695761BC36D50574B8D6C87DD19A">Mendonça, Maria Thayane Da Silva</mods:namePart>
<mods:affiliation id="206F92356E9D04FBE760155285F3AE4F">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Programa de Pós-Graduação em Zoologia UFPA / MPEG</mods:affiliation>
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<mods:namePart id="320D60C8DD438063611E23ECF1F0F0E5">Silva, Valéria Juliete Da</mods:namePart>
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<mods:affiliation id="C83EE47D1354939695FBACE176E85090">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; valeriajuliete @ hotmail. com; https: // orcid. org / 0000 - 0002 - 1845 - 5228</mods:affiliation>
<mods:nameIdentifier id="DC5C7975153107903A63584C9EBFA48B" type="email">valeriajuliete@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="B8404C3264AD6960BB735BC97686D08D">Fernandes, José Antônio Marin</mods:namePart>
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<mods:affiliation id="BB8E5B374B950CF0FCDA82BD635C8388">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Corresponding author: joseamf @ ufpa. br; https: // orcid. org / 0000 - 0001 - 7450 - 5296</mods:affiliation>
<mods:nameIdentifier id="E30CB29AB343C1FCDDE16A89E6B8FE95" type="email">joseamf@ufpa.br</mods:nameIdentifier>
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<mods:title id="66A73688C6FEAB45F9897A9D740C3182">Zootaxa</mods:title>
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<mods:date id="872CCBB895064C57032534932AB88839">2023</mods:date>
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<mods:number id="49E02D96E2D0573FA7FF52650EEF7E48">2023-11-14</mods:number>
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<mods:number id="1E2A01102075EC65CB03C90AEA81B0DF">5372</mods:number>
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<mods:number id="AD4B5654594734B04B813376C820B833">1</mods:number>
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<treatment id="414B87B5FFCB9852FF63DED0776AFA5F" LSID="urn:lsid:plazi:treatment:414B87B5FFCB9852FF63DED0776AFA5F" httpUri="http://treatment.plazi.org/id/414B87B5FFCB9852FF63DED0776AFA5F" lastPageNumber="25" pageId="24" pageNumber="25">
<subSubSection id="81F86528FFCB9852FF63DED07733FF70" box="[151,544,151,178]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="C95D36A3FFCB9852FF63DED07733FF70" blockId="24.[151,544,151,214]" box="[151,544,151,178]" pageId="24" pageNumber="25">
<heading id="921581CFFFCB9852FF63DED07733FF70" bold="true" box="[151,544,151,178]" fontSize="11" level="2" pageId="24" pageNumber="25" reason="4">
<emphasis id="FB96EAB1FFCB9852FF63DED07733FF70" bold="true" box="[151,544,151,178]" pageId="24" pageNumber="25">
<taxonomicName id="0EE24D20FFCB9852FF63DED074C4FF73" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[151,471,151,178]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cornicula" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFCB9852FF63DED074C4FF73" bold="true" box="[151,471,151,178]" italics="true" pageId="24" pageNumber="25">Edessa (Edessa) cornicula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0A557CAFFCB9852FE2ADED07733FF70" box="[478,544,151,177]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="81F86528FFCB9852FF63DEFC7495FF14" box="[151,390,187,214]" pageId="24" pageNumber="25" type="description">
<paragraph id="C95D36A3FFCB9852FF63DEFC7495FF14" blockId="24.[151,544,151,214]" box="[151,390,187,214]" pageId="24" pageNumber="25">
(
<figureCitation id="51D92A26FFCB9852FF6BDEFC75FDFF14" box="[159,238,187,213]" captionStart="FIGURE 8" captionStartId="23.[152,255,1943,1967]" captionTargetBox="[186,1401,188,1913]" captionTargetId="figure-20@23.[179,1408,181,1919]" captionTargetPageId="23" captionText="FIGURE 8. AB. Edessa (E.) cornicula sp. n.: A, genital plates; B, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146362" httpUri="https://zenodo.org/record/10146362/files/figure.png" pageId="24" pageNumber="25">Figs. 8</figureCitation>
,
<figureCitation id="51D92A26FFCB9852FF0FDEFC7446FF14" box="[251,341,187,214]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">56 CD</figureCitation>
,
<figureCitation id="51D92A26FFCB9852FE95DEFC746DFF14" box="[353,382,187,213]" captionStart="FIGURE 73" captionStartId="125.[152,255,2007,2031]" captionTargetBox="[183,1404,1119,1983]" captionTargetId="figure-33@125.[183,1404,1119,1983]" captionTargetPageId="125" captionText="FIGURE 73. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146497" httpUri="https://zenodo.org/record/10146497/files/figure.png" pageId="24" pageNumber="25">73</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFCB9852FF63DF447098FEDC" box="[151,1419,259,286]" pageId="24" pageNumber="25" type="etymology">
<paragraph id="C95D36A3FFCB9852FF63DF447098FEDC" blockId="24.[151,1437,259,1438]" box="[151,1419,259,286]" pageId="24" pageNumber="25">
<emphasis id="FB96EAB1FFCB9852FF63DF44740BFEDC" bold="true" box="[151,280,259,285]" pageId="24" pageNumber="25">Etymology</emphasis>
. The name refers to the size and development of the humeral angles (L.
<emphasis id="FB96EAB1FFCB9852FBB7DF437197FEDC" box="[1091,1156,260,285]" italics="true" pageId="24" pageNumber="25">cornu</emphasis>
, horn;
<taxonomicName id="0EE24D20FFCB9852FB21DF447053FEDC" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[1237,1344,259,285]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cornicula">
<emphasis id="FB96EAB1FFCB9852FB21DF447053FEDC" box="[1237,1344,259,285]" italics="true" pageId="24" pageNumber="25">cornicula</emphasis>
</taxonomicName>
, dim).
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFCB9852FF33DF60741CFEA4" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="C95D36A3FFCB9852FF33DF60741CFEA4" blockId="24.[151,1437,259,1438]" pageId="24" pageNumber="25">
<emphasis id="FB96EAB1FFCB9852FF33DF6074A6FE80" bold="true" box="[199,437,295,321]" pageId="24" pageNumber="25">Material examined.</emphasis>
<materialsCitation id="798A3CFEFFCB9852FE34DF60741CFEA4" collectingDate="1966-05-15" collectingDateMax="1966-05-22" collectingDateMin="1966-05-15" collectionCode="MZUSP" collectorName="Malkin" country="Peru" location="Rio Ampiacu" municipality="Estiron" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Loreto" typeStatus="holotype">
<typeStatus id="16598801FFCB9852FE34DF60773BFE80" box="[448,552,295,321]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
female.
<emphasis id="FB96EAB1FFCB9852FD65DF60765AFE80" bold="true" box="[657,841,295,321]" pageId="24" pageNumber="25">
<collectingCountry id="B1F57633FFCB9852FD65DF6077CEFE80" box="[657,733,295,321]" name="Peru" pageId="24" pageNumber="25">PERU</collectingCountry>
,
<collectingRegion id="0B26F841FFCB9852FD04DF607656FE80" box="[752,837,295,321]" country="Peru" name="Loreto" pageId="24" pageNumber="25">Loreto</collectingRegion>
:
</emphasis>
<collectingMunicipality id="2939ACD9FFCB9852FCA1DF6076BAFE80" box="[853,937,295,321]" pageId="24" pageNumber="25">Estiron</collectingMunicipality>
,
<location id="CC3D6078FFCB9852FC4CDF607147FE80" LSID="urn:lsid:plazi:treatment:414B87B5FFCB9852FF63DED0776AFA5F:CC3D6078FFCB9852FC4CDF607147FE80" box="[952,1108,295,321]" country="Peru" municipality="Estiron" name="Rio Ampiacu" pageId="24" pageNumber="25" stateProvince="Loreto">Rio Ampiacu</location>
,
<date id="BD5C1063FFCB9852FB97DF607014FE83" box="[1123,1287,295,322]" pageId="24" pageNumber="25" value="1966-05-15" valueMax="1966-05-22" valueMin="1966-05-15">
<collectingDate id="AD18E98BFFCB9852FB97DF607014FE83" box="[1123,1287,295,322]" pageId="24" pageNumber="25" value="1966-05-15" valueMax="1966-05-22" valueMin="1966-05-15">1522-V-1966</collectingDate>
</date>
,
<collectorName id="64175375FFCB9852FAE3DF60707BFE80" box="[1303,1384,295,321]" pageId="24" pageNumber="25">Malkin</collectorName>
leg. (
<collectionCode id="AFF3AE66FFCB9852FF6BDF0C7410FEA4" box="[159,259,331,357]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="24" pageNumber="25">MZUSP</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFCB9852FF33DF28746AFE10" pageId="24" pageNumber="25" type="description">
<paragraph id="C95D36A3FFCB9852FF33DF28746AFE10" blockId="24.[151,1437,259,1438]" pageId="24" pageNumber="25">
<emphasis id="FB96EAB1FFCB9852FF33DF287465FE48" bold="true" box="[199,374,367,393]" pageId="24" pageNumber="25">Measurements</emphasis>
(n= 1). Total length: 22.3; head length: 1.5; head width: 3.5; pronotum length: 4.5; pronotum width: 14.2; scutellum length: 10.3; scutellum width: 7.5; abdominal width: 12.8; length antennomeres: I: 1.5; II: 2.0; III: 2.3; IV: 4.8.
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFCB9852FF33DF9C76B2FB60" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="C95D36A3FFCB9852FF33DF9C76B2FB60" blockId="24.[151,1437,259,1438]" pageId="24" pageNumber="25">
<emphasis id="FB96EAB1FFCB9852FF33DF9C7451FE34" bold="true" box="[199,322,475,501]" pageId="24" pageNumber="25">Diagnosis.</emphasis>
Large (
<quantity id="0E1A9B46FFCB9852FE62DF9C74ECFE34" box="[406,511,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" pageId="24" pageNumber="25" unit="mm" value="22.3">22.3 mm</quantity>
). Dorsal body surface olive green (
<figureCitation id="51D92A26FFCB9852FC76DF9C76FDFE34" box="[898,1006,475,502]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 C</figureCitation>
). Ventral surface dark yellow to brown with transversal brown bands on thorax and abdomen (
<figureCitation id="51D92A26FFCB9852FCF8DFB8766CFDD8" box="[780,895,511,538]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 D</figureCitation>
). Antennae yellowish (
<figureCitation id="51D92A26FFCB9852FB71DFB8700AFDD8" box="[1157,1305,511,538]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 CD</figureCitation>
). Pronotum with brown punctures (
<figureCitation id="51D92A26FFCB9852FE6BDC64771CFDFC" box="[415,527,547,574]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 C</figureCitation>
); anterolateral margin with a yellow band extending from the humeral angles to the head; anterolateral margin and cicatrices with brown punctures. Humeral angles very short (2.0 times wider than long); apex concolorous with the surrounding surface, slightly acuminate and curved backwards (
<figureCitation id="51D92A26FFCB9852FAD4DC2C709CFD44" box="[1312,1423,619,646]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 C</figureCitation>
). Scutellum with brown punctures; apex not reaching the end of coria (
<figureCitation id="51D92A26FFCB9852FC5DDCC8710BFD68" box="[937,1048,655,682]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 C</figureCitation>
). Coria with all veins concolorous with the surrounding surface (
<figureCitation id="51D92A26FFCB9852FE12DCF47741FD0C" box="[486,594,691,718]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 C</figureCitation>
). Connexival segments with concavities entirely covered by subrectangular black spots separated by a large dark yellow median spot (
<figureCitation id="51D92A26FFCB9852FCE2DC907693FD30" box="[790,896,727,754]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 C</figureCitation>
); black spots extending ventrally, subrectangular (
<figureCitation id="51D92A26FFCB9852FF6BDCBC741CFCD4" box="[159,271,763,790]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 D</figureCitation>
). Posterolateral angles of connexivum with apices black (
<figureCitation id="51D92A26FFCB9852FC61DCBC7111FCD4" box="[917,1026,763,790]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 C</figureCitation>
). Ventral surface, thorax with brown bands; dark band of the propleura covering 2/3 of the width of the sclerite (
<figureCitation id="51D92A26FFCB9852FC15DD587142FCF8" box="[993,1105,799,826]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 D</figureCitation>
). Evaporatorium concolorous with the surrounding surface; peritreme straight (
<figureCitation id="51D92A26FFCB9852FD30DD047625FC9C" box="[708,822,835,862]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 D</figureCitation>
). Metasternal process (
<figureCitation id="51D92A26FFCB9852FBC8DD047188FC9C" box="[1084,1179,835,861]" captionStart="FIGURE 8" captionStartId="23.[152,255,1943,1967]" captionTargetBox="[186,1401,188,1913]" captionTargetId="figure-20@23.[179,1408,181,1919]" captionTargetPageId="23" captionText="FIGURE 8. AB. Edessa (E.) cornicula sp. n.: A, genital plates; B, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146362" httpUri="https://zenodo.org/record/10146362/files/figure.png" pageId="24" pageNumber="25">Fig. 8 B</figureCitation>
) with arms of anterior bifurcation straight, laterally well expanded at apex; anterior bifurcation broadly excavated receiving fourth rostral segment. Legs yellowish (
<figureCitation id="51D92A26FFCB9852FE4EDDCC773BFC64" box="[442,552,907,934]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 D</figureCitation>
). Abdomen with spine of segment III rounded (
<figureCitation id="51D92A26FFCB9852FBC1DDCC718EFC67" box="[1077,1181,907,934]" captionStart="FIGURE 8" captionStartId="23.[152,255,1943,1967]" captionTargetBox="[186,1401,188,1913]" captionTargetId="figure-20@23.[179,1408,181,1919]" captionTargetPageId="23" captionText="FIGURE 8. AB. Edessa (E.) cornicula sp. n.: A, genital plates; B, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146362" httpUri="https://zenodo.org/record/10146362/files/figure.png" pageId="24" pageNumber="25">Figs. 8 B</figureCitation>
;
<figureCitation id="51D92A26FFCB9852FB5DDDCC71F0FC64" box="[1193,1251,907,934]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">56 D</figureCitation>
). Intersegmental areas covered by narrow smudged brown bands with well-defined margins, reaching the lateral spots (
<figureCitation id="51D92A26FFCB9852FAEADDE87083FC08" box="[1310,1424,943,970]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 D</figureCitation>
). Pseudosutures concolorous with the surrounding surface (
<figureCitation id="51D92A26FFCB9852FCD9DD9476B2FC2C" box="[813,929,979,1006]" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 D</figureCitation>
). Median longitudinal area spotless (
<figureCitation id="51D92A26FFCB9852FABDDD9475BCFBD0" captionStart="FIGURE 56" captionStartId="109.[152,255,1943,1967]" captionTargetBox="[258,1329,187,1913]" captionTargetId="figure-20@109.[252,1335,181,1919]" captionTargetPageId="109" captionText="FIGURE 56. AF. Dorsal and ventral surface:AB, Edessa (E.) chloroicterica sp. n.; CD, Edessa (E.) cornicula sp. n.; EF, Edessa (E.) crassicornis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146461" httpUri="https://zenodo.org/record/10146461/files/figure.png" pageId="24" pageNumber="25">Fig. 56 D</figureCitation>
). Trichobothria both parallel to the spiracle. Posterolateral angles of segment VII exceeding the level of apices of laterotergites IX (
<figureCitation id="51D92A26FFCB9852FE94DA5C74D1FBF4" box="[352,450,1051,1077]" captionStart="FIGURE 8" captionStartId="23.[152,255,1943,1967]" captionTargetBox="[186,1401,188,1913]" captionTargetId="figure-20@23.[179,1408,181,1919]" captionTargetPageId="23" captionText="FIGURE 8. AB. Edessa (E.) cornicula sp. n.: A, genital plates; B, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146362" httpUri="https://zenodo.org/record/10146362/files/figure.png" pageId="24" pageNumber="25">Fig. 8 A</figureCitation>
). Male genitalia. Unknown. Female genitalia, valvifers VIII with concolorous with the surrounding surface punctures; sutural margins contiguous; posterior margin subrectangular with small and shallow U-shaped excavation delimited by small rounded projections. Laterotergites VIII without dark band. Laterotergites IX with apices acuminate and level with mediotergite VIII (
<figureCitation id="51D92A26FFCB9852FCC0DAC07687FB60" box="[820,916,1159,1185]" captionStart="FIGURE 8" captionStartId="23.[152,255,1943,1967]" captionTargetBox="[186,1401,188,1913]" captionTargetId="figure-20@23.[179,1408,181,1919]" captionTargetPageId="23" captionText="FIGURE 8. AB. Edessa (E.) cornicula sp. n.: A, genital plates; B, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146362" httpUri="https://zenodo.org/record/10146362/files/figure.png" pageId="24" pageNumber="25">Fig. 8 A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFCB9852FF33DAEC7125FABB" pageId="24" pageNumber="25" type="discussion">
<paragraph id="C95D36A3FFCB9852FF33DAEC7125FABB" blockId="24.[151,1437,259,1438]" pageId="24" pageNumber="25">
<emphasis id="FB96EAB1FFCB9852FF33DAEC745CFB04" bold="true" box="[199,335,1195,1221]" pageId="24" pageNumber="25">Comments.</emphasis>
<taxonomicName id="0EE24D20FFCB9852FEADDAEC7741FB04" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[345,594,1195,1221]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cornicula" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFCB9852FEADDAEC7741FB04" box="[345,594,1195,1221]" italics="true" pageId="24" pageNumber="25">Edessa (E.) cornicula</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFCB9852FDA9DAEC778CFB04" bold="true" box="[605,671,1195,1221]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="E0A557CAFFCB9852FDA9DAEC778CFB04" box="[605,671,1195,1221]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="0EE24D20FFCB9852FCD7DAEC718AFB04" authority="Erichson, 1848" authorityName="Erichson" authorityYear="1848" box="[803,1177,1195,1221]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="urus" subGenus="Edessa">
<emphasis id="FB96EAB1FFCB9852FCD7DAEC76F0FB04" box="[803,995,1195,1221]" italics="true" pageId="24" pageNumber="25">Edessa (E.) urus</emphasis>
Erichson, 1848
</taxonomicName>
(see
<bibRefCitation id="AD734B52FFCB9852FB2EDAEC75C1FB28" author="Mendonca, M. T. S. &amp; Silva, V. J. &amp; Fernandes, J. A. M." pageId="24" pageNumber="25" pagination="1 - 63" refId="ref74748" refString="Mendonca, M. T. S., Silva, V. J. &amp; Fernandes, J. A. M. (2023) Diagnose of the nominal subgenus of Edessa and description of the E. sexdens group (Heteroptera, Pentatomidae, Edessinae). Zootaxa, 5240 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 5240.1.1" type="journal article" year="2023">
Mendonça
<emphasis id="FB96EAB1FFCB9852FAAEDAEB7085FB04" box="[1370,1430,1195,1221]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
, 2023
</bibRefCitation>
,
<figureCitation id="51D92A26FFCB9852FF2ADA887442FB2B" box="[222,337,1231,1258]" captionStart="FIGURE 23" captionStartId="52.[152,255,1943,1967]" captionTargetBox="[180,1408,188,1912]" captionTargetId="figure-22@52.[173,1415,181,1919]" captionTargetPageId="52" captionText="FIGURE 23. AB. Edessa (E.) illuminocornis sp. n.: A, genital plates; B, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146393" httpUri="https://zenodo.org/record/10146393/files/figure.png" pageId="24" pageNumber="25">Figs. 23 F</figureCitation>
,
<figureCitation id="51D92A26FFCB9852FEAADA8874A6FB28" box="[350,437,1231,1257]" captionStart="FIGURE 29" captionStartId="63.[152,255,1943,1967]" captionTargetBox="[190,1397,188,1913]" captionTargetId="figure-20@63.[190,1397,181,1919]" captionTargetPageId="63" captionText="FIGURE 29. AG. Edessa (E.) limerai sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146405" httpUri="https://zenodo.org/record/10146405/files/figure.png" pageId="24" pageNumber="25">29 EF</figureCitation>
). Both species share the humeral angles concolorous with the surrounding surface, short and curved; body ventrally with intersegmental areas brown. Both species can be separated by characteristics of the genitalia, in
<taxonomicName id="0EE24D20FFCB9852FEA4DB5F771CFAF0" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[336,527,1303,1329]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cornicula" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFCB9852FEA4DB5F771CFAF0" box="[336,527,1303,1329]" italics="true" pageId="24" pageNumber="25">E. (E.) cornicula</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFCB9852FDECDB50774AFAF0" bold="true" box="[536,601,1303,1329]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="E0A557CAFFCB9852FDECDB50774AFAF0" box="[536,601,1303,1329]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
the posterior margin of the valvifers VIII is subrectangular and apices of the laterotergites IX level with mediotergite VIII, while in
<taxonomicName id="0EE24D20FFCB9852FCDFDB7B76BCFA94" authorityName="Erichson" authorityYear="1848" box="[811,943,1339,1365]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="urus" subGenus="Edessa">
<emphasis id="FB96EAB1FFCB9852FCDFDB7B76BCFA94" box="[811,943,1339,1365]" italics="true" pageId="24" pageNumber="25">E. (E.) urus</emphasis>
</taxonomicName>
the posterior margin of the valvifers VIII is clearly arched and the apices of the laterotergites IX passing the mediotergite VIII.
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFCB9852FF33DBC4776AFA5F" box="[199,633,1411,1438]" pageId="24" pageNumber="25" type="distribution">
<paragraph id="C95D36A3FFCB9852FF33DBC4776AFA5F" blockId="24.[151,1437,259,1438]" box="[199,633,1411,1438]" pageId="24" pageNumber="25">
<emphasis id="FB96EAB1FFCB9852FF33DBC4744AFA5C" bold="true" box="[199,345,1411,1437]" pageId="24" pageNumber="25">Distribution</emphasis>
(
<figureCitation id="51D92A26FFCB9852FE9DDBC474A9FA5C" box="[361,442,1411,1437]" captionStart="FIGURE 73" captionStartId="125.[152,255,2007,2031]" captionTargetBox="[183,1404,1119,1983]" captionTargetId="figure-33@125.[183,1404,1119,1983]" captionTargetPageId="125" captionText="FIGURE 73. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146497" httpUri="https://zenodo.org/record/10146497/files/figure.png" pageId="24" pageNumber="25">Fig. 73</figureCitation>
).
<collectingCountry id="B1F57633FFCB9852FE24DBC3770FFA5F" box="[464,540,1412,1438]" name="Peru" pageId="24" pageNumber="25">PERU</collectingCountry>
:
<collectingRegion id="0B26F841FFCB9852FDD3DBC37766FA5F" box="[551,629,1412,1438]" country="Peru" name="Loreto" pageId="24" pageNumber="25">Loreto</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>