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<document id="E310F989E6AD2E81C1C90AF693315988" ID-DOI="10.11646/zootaxa.5372.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10146342" ID-ZooBank="1300D562-3E0B-4F72-933F-8FA7D28F6853" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700186674367" checkinUser="plazi" docAuthor="Mendonça, Maria Thayane Da Silva, Silva, Valéria Juliete Da &amp; Fernandes, José Antônio Marin" docDate="2023" docId="414B87B5FFC09859FF63DED0711BF88F" docLanguage="en" docName="zootaxa.5372.1.1.pdf" docOrigin="Zootaxa 5372 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5372.1.1/52272" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Edessa (Edessa) brevihumeralis Mendonça &amp; Silva &amp; Fernandes 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="20" masterDocId="BD72FFCDFFD3984AFFF4DE477513FFC1" masterDocTitle="Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="20" updateTime="1700188109802" updateUser="ExternalLinkService">
<mods:mods id="74F2A4B11226A8C19FE68BCE82A2446D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="44ECF7A86B94886ADF287D18CD702F9F">
<mods:title id="FAF25B5AD26B16E4B5EA72F542E8C984">Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)</mods:title>
</mods:titleInfo>
<mods:name id="243505868B9C6466DC338B0B71B1D113" type="personal">
<mods:role id="3C8F0FEAAEDD8ACE637923120854A4E0">
<mods:roleTerm id="27ECE702F8266D34C0F43822649E0925">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="915C801DDDCABE6ADA5BFE1169932775">Mendonça, Maria Thayane Da Silva</mods:namePart>
<mods:affiliation id="DBE05F7A8C2073D43F302B4837E59A3A">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Programa de Pós-Graduação em Zoologia UFPA / MPEG</mods:affiliation>
</mods:name>
<mods:name id="0F35F8A9EFB14D331B01CADB311ACD34" type="personal">
<mods:role id="71115C754756C562D73A23CCC1DF1B96">
<mods:roleTerm id="49A078F7CFC99320D57069C7CC2EDB7B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A84169ACFA7FDC58253978BD2E3FF5C7">Silva, Valéria Juliete Da</mods:namePart>
<mods:nameIdentifier id="DBC476AE49DE618B9B10D227CF76C0BC" type="ORCID">0000-0002-1845-5228</mods:nameIdentifier>
<mods:affiliation id="B7A36B5D5D1B324E37C06FE3D0BE92C6">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; valeriajuliete @ hotmail. com; https: // orcid. org / 0000 - 0002 - 1845 - 5228</mods:affiliation>
<mods:nameIdentifier id="AE0EDE19EE63EF01625FC5E6FA07EA57" type="email">valeriajuliete@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="3CADDBF48607372610D9794342D116CA" type="personal">
<mods:role id="28792C556DD5A70C9D89E8385FFF05A9">
<mods:roleTerm id="8B130A0FFFE0B2AFB26C9C3D955E36D2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C274C060C455BEF564E0245EA598E14D">Fernandes, José Antônio Marin</mods:namePart>
<mods:nameIdentifier id="E9E013CEF22215BF570BA33A9F7A01DF" type="ORCID">0000-0001-7450-5296</mods:nameIdentifier>
<mods:affiliation id="A173BFEFCE90B46E7640918673CD6395">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Corresponding author: joseamf @ ufpa. br; https: // orcid. org / 0000 - 0001 - 7450 - 5296</mods:affiliation>
<mods:nameIdentifier id="8FB89C7718137FBEB825D2F1B329B1E8" type="email">joseamf@ufpa.br</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="371F2D2720EA1B3C0A4E40B48F774B31">text</mods:typeOfResource>
<mods:relatedItem id="6F3D79777E687DA1829B3A3E411ED97C" type="host">
<mods:titleInfo id="10300A1B39E45ED1C09FA1C33BF3AF0E">
<mods:title id="2495D3AD40B8C7635AB8E79034858318">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="44E1A72968D25CD723C66C7FD24C726B">
<mods:date id="95157A0BBFAE80E9108E90E50FD11738">2023</mods:date>
<mods:detail id="3C13792401CFFDB086423ACBB5E9221E" type="pubDate">
<mods:number id="1215629108FEC35FE5B87B6113851E15">2023-11-14</mods:number>
</mods:detail>
<mods:detail id="B20D1457B7F22FF0394E40E20FBDF00A" type="volume">
<mods:number id="0FE396966A1F7405057EAF75F80F3219">5372</mods:number>
</mods:detail>
<mods:detail id="33E3295836CFEBC15E9ED0A7F901C1D5" type="issue">
<mods:number id="C1D6076666CCAD74905FC69A039AB807">1</mods:number>
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<mods:start id="2C9208358C454056DBFEDFA284ED31D9">1</mods:start>
<mods:end id="805132617332C4E707A3B27F24DB50D6">128</mods:end>
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<mods:location id="7F626A6ADA571090C872869BFC47414F">
<mods:url id="C49C1624F15F34988CF03999D8C7D109">https://www.mapress.com/zt/article/download/zootaxa.5372.1.1/52272</mods:url>
</mods:location>
<mods:classification id="C5BF7931BB2353E24BFC9D477B46AF33">journal article</mods:classification>
<mods:identifier id="BF73A9A2B767E116F75B989C70D44EDE" type="DOI">10.11646/zootaxa.5372.1.1</mods:identifier>
<mods:identifier id="41F75172C2B2F6F99621912804CF0C54" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="2550296045DFDA0E37C8F9392D362D07" type="Zenodo-Dep">10146342</mods:identifier>
<mods:identifier id="01F95311AF42ADF86B256C1FF5F5C12E" type="ZooBank">1300D562-3E0B-4F72-933F-8FA7D28F6853</mods:identifier>
</mods:mods>
<treatment id="414B87B5FFC09859FF63DED0711BF88F" LSID="urn:lsid:plazi:treatment:414B87B5FFC09859FF63DED0711BF88F" httpUri="http://treatment.plazi.org/id/414B87B5FFC09859FF63DED0711BF88F" lastPageNumber="20" pageId="19" pageNumber="20">
<subSubSection id="81F86528FFC09859FF63DED07770FF70" box="[151,611,151,178]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="C95D36A3FFC09859FF63DED07770FF70" blockId="19.[151,611,151,214]" box="[151,611,151,178]" pageId="19" pageNumber="20">
<heading id="921581CFFFC09859FF63DED07770FF70" bold="true" box="[151,611,151,178]" fontSize="11" level="2" pageId="19" pageNumber="20" reason="4">
<emphasis id="FB96EAB1FFC09859FF63DED07770FF70" bold="true" box="[151,611,151,178]" pageId="19" pageNumber="20">
<taxonomicName id="0EE24D20FFC09859FF63DED07709FF73" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[151,538,151,178]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="brevihumeralis" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FF63DED07709FF73" bold="true" box="[151,538,151,178]" italics="true" pageId="19" pageNumber="20">Edessa (Edessa) brevihumeralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0A557CAFFC09859FDD5DED07770FF70" box="[545,611,151,177]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFC09859FF63DEFC746DFF14" box="[151,382,187,214]" pageId="19" pageNumber="20" type="description">
<paragraph id="C95D36A3FFC09859FF63DEFC746DFF14" blockId="19.[151,611,151,214]" box="[151,382,187,214]" pageId="19" pageNumber="20">
(
<figureCitation id="51D92A26FFC09859FF6BDEFC75FCFF14" box="[159,239,187,213]" captionStart="FIGURE 6" captionStartId="20.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-22@20.[185,1402,181,1919]" captionTargetPageId="20" captionText="FIGURE 6. A–G. Edessa (E.) brevihumeralis sp. n.:A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146358" httpUri="https://zenodo.org/record/10146358/files/figure.png" pageId="19" pageNumber="20">
Figs.
<geoCoordinate id="ACD65064FFC09859FF2BDEFC75FCFF14" box="[223,239,187,213]" degrees="6, 55" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="555" value="6.55">6</geoCoordinate>
</figureCitation>
,
<figureCitation id="51D92A26FFC09859FF0FDEFC745DFF17" box="[251,334,187,214]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">55 E–F</figureCitation>
,
<figureCitation id="51D92A26FFC09859FEACDEFC7466FF14" box="[344,373,187,213]" captionStart="FIGURE 72" captionStartId="125.[152,255,1071,1095]" captionTargetBox="[195,1391,181,1040]" captionTargetId="figure-20@125.[195,1391,181,1050]" captionTargetPageId="125" captionText="FIGURE 72. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146495" httpUri="https://zenodo.org/record/10146495/files/figure.png" pageId="19" pageNumber="20">72</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFC09859FF63DF44703DFEDC" box="[151,1326,259,286]" pageId="19" pageNumber="20" type="etymology">
<paragraph id="C95D36A3FFC09859FF63DF44703DFEDC" blockId="19.[151,1437,259,1870]" box="[151,1326,259,286]" pageId="19" pageNumber="20">
<emphasis id="FB96EAB1FFC09859FF63DF44740BFEDC" bold="true" box="[151,280,259,285]" pageId="19" pageNumber="20">Etymology</emphasis>
. The name refers to the length of the humeral angles (L.
<emphasis id="FB96EAB1FFC09859FC63DF4476C9FEDC" box="[919,986,259,285]" italics="true" pageId="19" pageNumber="20">brevis</emphasis>
, short; L.
<emphasis id="FB96EAB1FFC09859FBBADF4471BDFEDC" box="[1102,1198,259,285]" italics="true" pageId="19" pageNumber="20">humerus</emphasis>
, shoulder).
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFC09859FF33DF6076DAFE10" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="C95D36A3FFC09859FF33DF607716FEA4" blockId="19.[151,1437,259,1870]" pageId="19" pageNumber="20">
<emphasis id="FB96EAB1FFC09859FF33DF6074A0FE80" bold="true" box="[199,435,295,321]" pageId="19" pageNumber="20">Material examined.</emphasis>
<materialsCitation id="798A3CFEFFC09859FE49DF607716FEA4" collectingDate="1979-03" collectionCode="MNRJ" collectorName="O. Roppa &amp; A. Domingos" country="Brazil" location="Mato Grosso" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Mato Grosso" typeStatus="holotype">
<typeStatus id="16598801FFC09859FE49DF607736FE80" box="[445,549,295,321]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
male.
<emphasis id="FB96EAB1FFC09859FD82DF60768BFE80" bold="true" box="[630,920,295,321]" pageId="19" pageNumber="20">
<collectingCountry id="B1F57633FFC09859FD82DF6077F2FE80" box="[630,737,295,321]" name="Brazil" pageId="19" pageNumber="20">BRAZIL</collectingCountry>
,
<collectingRegion id="0B26F841FFC09859FD06DF607680FE80" box="[754,915,295,321]" country="Brazil" name="Mato Grosso" pageId="19" pageNumber="20">Mato Grosso</collectingRegion>
:
</emphasis>
<taxonomicName id="0EE24D20FFC09859FC55DF6071B8FE80" authority=", Fazenda S" authorityName="Fazenda S" box="[929,1195,295,321]" class="Magnoliopsida" family="Podostemaceae" genus="Diamantina" kingdom="Plantae" order="Malpighiales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="genus">Diamantina, Fazenda S</taxonomicName>
„o Jo„o,
<date id="BD5C1063FFC09859FAE7DF607060FE80" box="[1299,1395,295,321]" pageId="19" pageNumber="20" value="1979-03">
<collectingDate id="AD18E98BFFC09859FAE7DF607060FE80" box="[1299,1395,295,321]" pageId="19" pageNumber="20" value="1979-03">III-1979</collectingDate>
</date>
,
<collectorName id="64175375FFC09859FA75DF6075F3FEA7" pageId="19" pageNumber="20">O. Roppa</collectorName>
&amp;
<collectorName id="64175375FFC09859FEF7DF0C7489FEA4" box="[259,410,331,357]" pageId="19" pageNumber="20">A. Domingos</collectorName>
(
<collectionCode id="AFF3AE66FFC09859FE5CDF0C74EAFEA4" box="[424,505,331,357]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="19" pageNumber="20">MNRJ</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="C95D36A3FFC09859FF33DF3776DAFE10" blockId="19.[151,1437,259,1870]" pageId="19" pageNumber="20">
<materialsCitation id="798A3CFEFFC09859FF33DF3776E6FE48" box="[199,1013,367,394]" collectingDate="1979-03" collectionCode="MNRJ" collectorName="O. Roppa &amp; A. Domingos" country="Brazil" location="Mato Grosso" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Mato Grosso" typeStatus="paratype">
<typeStatus id="16598801FFC09859FF33DF37742BFE4B" box="[199,312,368,394]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
.
<emphasis id="FB96EAB1FFC09859FEB6DF28774CFE48" bold="true" box="[322,607,367,393]" pageId="19" pageNumber="20">
<collectingCountry id="B1F57633FFC09859FEB6DF2874BEFE48" box="[322,429,367,393]" name="Brazil" pageId="19" pageNumber="20">BRAZIL</collectingCountry>
,
<collectingRegion id="0B26F841FFC09859FE4FDF287749FE48" box="[443,602,367,393]" country="Brazil" name="Mato Grosso" pageId="19" pageNumber="20">Mato Grosso</collectingRegion>
:
</emphasis>
<specimenCount id="DFE4FD2AFFC09859FD92DF28779AFE48" box="[614,649,367,393]" count="1" pageId="19" pageNumber="20" type="female">1♀</specimenCount>
, same data as holotype (
<collectionCode id="AFF3AE66FFC09859FC68DF2876FEFE48" box="[924,1005,367,393]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="19" pageNumber="20">MNRJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="798A3CFEFFC09859FBF4DF36770BFE6C" collectingDate="1966-11-01" collectingDateMax="1966-11-12" collectingDateMin="1966-11-01" collectionCode="MZUSP" collectorName="Lenko &amp; Pereira" country="Brazil" location="Rio Papagaio" municipality="Utiariti" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Mato Grosso" typeStatus="paratype">
1♁,
<collectingMunicipality id="2939ACD9FFC09859FBC5DF287196FE48" box="[1073,1157,367,393]" pageId="19" pageNumber="20">Utiariti</collectingMunicipality>
,
<location id="CC3D6078FFC09859FB64DF28703AFE48" LSID="urn:lsid:plazi:treatment:414B87B5FFC09859FF63DED0711BF88F:CC3D6078FFC09859FB64DF28703AFE48" box="[1168,1321,367,393]" country="Brazil" municipality="Utiariti" name="Rio Papagaio" pageId="19" pageNumber="20" stateProvince="Mato Grosso">Rio Papagaio</location>
,
<date id="BD5C1063FFC09859FAC0DF2875C1FE6C" pageId="19" pageNumber="20" value="1966-11-01" valueMax="1966-11-12" valueMin="1966-11-01">
<collectingDate id="AD18E98BFFC09859FAC0DF2875C1FE6C" pageId="19" pageNumber="20" value="1966-11-01" valueMax="1966-11-12" valueMin="1966-11-01">1–12-XI-1966</collectingDate>
</date>
,
<collectorName id="64175375FFC09859FF2ADFD47435FE6C" box="[222,294,403,429]" pageId="19" pageNumber="20">Lenko</collectorName>
&amp;
<collectorName id="64175375FFC09859FEB8DFD47488FE6C" box="[332,411,403,429]" pageId="19" pageNumber="20">Pereira</collectorName>
(
<collectionCode id="AFF3AE66FFC09859FE58DFD47703FE6C" box="[428,528,403,429]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="20">MZUSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="798A3CFEFFC09859FDD0DFD475C9FE10" collectingDate="1997-11-04" collectingDateMax="1997-11-06" collectingDateMin="1997-11-04" collectionCode="JEE" country="Brazil" location="Fazenda Rancho Grande" municipality="Ariquemes" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Rondonia" typeStatus="paratype">
<emphasis id="FB96EAB1FFC09859FDD0DFD477B1FE6C" bold="true" box="[548,674,403,429]" pageId="19" pageNumber="20">
<collectingRegion id="0B26F841FFC09859FDD0DFD4778EFE6C" box="[548,669,403,429]" country="Brazil" name="Rondonia" pageId="19" pageNumber="20">Rondônia</collectingRegion>
:
</emphasis>
<specimenCount id="DFE4FD2AFFC09859FD5EDFD477DEFE6C" box="[682,717,403,429]" count="1" pageId="19" pageNumber="20" type="female">1♀</specimenCount>
,
<collectingMunicipality id="2939ACD9FFC09859FD2FDFD4764AFE6C" box="[731,857,403,429]" pageId="19" pageNumber="20">Ariquemes</collectingMunicipality>
,
<location id="CC3D6078FFC09859FC91DFD47169FE6C" LSID="urn:lsid:plazi:treatment:414B87B5FFC09859FF63DED0711BF88F:CC3D6078FFC09859FC91DFD47169FE6C" box="[869,1146,403,429]" country="Brazil" municipality="Ariquemes" name="Fazenda Rancho Grande" pageId="19" pageNumber="20" stateProvince="Rondonia">Fazenda Rancho Grande</location>
<quantity id="0E1A9B46FFC09859FB77DFD471D8FE6C" box="[1155,1227,403,429]" metricMagnitude="4" metricUnit="m" metricValue="6.2" pageId="19" pageNumber="20" unit="km" value="62.0">62 km</quantity>
SW,
<date id="BD5C1063FFC09859FAFEDFD4708FFE6C" box="[1290,1436,403,429]" pageId="19" pageNumber="20" value="1997-11-04" valueMax="1997-11-06" valueMin="1997-11-04">
<collectingDate id="AD18E98BFFC09859FAFEDFD4708FFE6C" box="[1290,1436,403,429]" pageId="19" pageNumber="20" value="1997-11-04" valueMax="1997-11-06" valueMin="1997-11-04">4–6-XI-1997</collectingDate>
</date>
(
<collectionCode id="AFF3AE66FFC09859FF6BDFF075C1FE10" box="[159,210,439,465]" pageId="19" pageNumber="20">JEE</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="798A3CFEFFC09859FF11DFF076D6FE10" box="[229,965,439,468]" collectingDate="1929-04" collectionCode="USNM" collectorName="H. S. Parigh" country="Peru" location="Yurimaguas" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Loreto" typeStatus="paratype">
<emphasis id="FB96EAB1FFC09859FF11DFF074BDFE10" bold="true" box="[229,430,439,465]" pageId="19" pageNumber="20">
<collectingCountry id="B1F57633FFC09859FF11DFF07422FE10" box="[229,305,439,465]" name="Peru" pageId="19" pageNumber="20">PERU</collectingCountry>
,
<collectingRegion id="0B26F841FFC09859FECBDFF07487FE10" box="[319,404,439,465]" country="Peru" name="Loreto" pageId="19" pageNumber="20">Loreto</collectingRegion>
: 1
</emphasis>
♁,
<location id="CC3D6078FFC09859FE24DFF0774AFE10" LSID="urn:lsid:plazi:treatment:414B87B5FFC09859FF63DED0711BF88F:CC3D6078FFC09859FE24DFF0774AFE10" box="[464,601,439,465]" country="Peru" name="Yurimaguas" pageId="19" pageNumber="20" stateProvince="Loreto">Yurimaguas</location>
,
<date id="BD5C1063FFC09859FD90DFF077D0FE10" box="[612,707,439,465]" pageId="19" pageNumber="20" value="1929-04">
<collectingDate id="AD18E98BFFC09859FD90DFF077D0FE10" box="[612,707,439,465]" pageId="19" pageNumber="20" value="1929-04">IV-1929</collectingDate>
</date>
,
<collectorName id="64175375FFC09859FD3ADFFF7644FE10" box="[718,855,439,466]" pageId="19" pageNumber="20">H. S. Parigh</collectorName>
(
<collectionCode id="AFF3AE66FFC09859FC92DFF076AEFE10" box="[870,957,439,465]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="20" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFC09859FF33DF9C7137FDFC" pageId="19" pageNumber="20" type="description">
<paragraph id="C95D36A3FFC09859FF33DF9C7137FDFC" blockId="19.[151,1437,259,1870]" pageId="19" pageNumber="20">
<emphasis id="FB96EAB1FFC09859FF33DF9C7465FE34" bold="true" box="[199,374,475,501]" pageId="19" pageNumber="20">Measurements</emphasis>
(n= 5). Total length: 16.8–19.1; head length: 1.5; head width: 2.9–3.4; pronotum length: 3.3– 3.5; pronotum width: 11.1–12.0; scutellum length: 7.9–9.0; scutellum width: 5.8–6.5; abdominal width: 9.2–10.0; length antennomeres: I: 0.9–1.0; II: 1.5–1.6; III: 1.6–2.0; IV: 3.8–3.9; V: 4.0–4.2.
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFC09859FF33DC0071F3F9C8" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="C95D36A3FFC09859FF33DC0071F3F9C8" blockId="19.[151,1437,259,1870]" pageId="19" pageNumber="20">
<emphasis id="FB96EAB1FFC09859FF33DC007451FDA0" bold="true" box="[199,322,583,609]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
Large (
<quantity id="0E1A9B46FFC09859FE68DC007754FDA0" box="[412,583,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="1.7950000000000004" metricValueMax="1.9100000000000001" metricValueMin="1.6800000000000002" pageId="19" pageNumber="20" unit="mm" value="17.95" valueMax="19.1" valueMin="16.8">16.8–19.1 mm</quantity>
). Dorsal body surface olive green (
<figureCitation id="51D92A26FFC09859FC2FDC00715FFDA0" box="[987,1100,583,610]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 E</figureCitation>
). Ventral surface dark yellow with transversal black lines on thorax and bands on abdomen (
<figureCitation id="51D92A26FFC09859FC9ADC2C76CDFD44" box="[878,990,619,646]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 F</figureCitation>
). Antennae reddish brown (
<figureCitation id="51D92A26FFC09859FAEADC2C7083FD44" box="[1310,1424,619,646]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 E</figureCitation>
). Pronotum densely punctured, with brown punctures (
<figureCitation id="51D92A26FFC09859FCF4DCC87660FD68" box="[768,883,655,682]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 E</figureCitation>
); anterolateral margin and cicatrices with black punctures. Humeral angles short (1.25 times wider than long); apex with black spot restricted to the angles in dorsal and ventral views; slightly bent posteriorly (
<figureCitation id="51D92A26FFC09859FD7FDC907604FD30" box="[651,791,727,754]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 E–F</figureCitation>
). Scutellum with brown punctures; apex not reaching the end of coria (
<figureCitation id="51D92A26FFC09859FEDBDCBC748EFCD4" box="[303,413,763,790]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 E</figureCitation>
). Coria with all veins lighter than surrounding area (
<figureCitation id="51D92A26FFC09859FC1BDCBC714EFCD4" box="[1007,1117,763,790]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 E</figureCitation>
). Connexival segments with concavities almost entirely covered by subrectangular black spots separated by a large yellow median spot (
<figureCitation id="51D92A26FFC09859FAB8DD5875B8FC9C" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 E</figureCitation>
); black spots extending ventrally, triangular (
<figureCitation id="51D92A26FFC09859FD5BDD04760FFC9C" box="[687,796,835,862]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 F</figureCitation>
). Posterolateral angles of connexivum with apices black (
<figureCitation id="51D92A26FFC09859FF6BDD20741FFC40" box="[159,268,871,898]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 E</figureCitation>
). Ventral surface, thorax with black lines; dark line of propleura covering 2/3 of the width of the sclerite (
<figureCitation id="51D92A26FFC09859FF6BDDCC741EFC64" box="[159,269,907,934]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 F</figureCitation>
). Evaporatorium concolorous with the surrounding surface (
<figureCitation id="51D92A26FFC09859FC49DDCC7139FC64" box="[957,1066,907,934]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 F</figureCitation>
). Metasternal process (
<figureCitation id="51D92A26FFC09859FAC6DDCC7080FC64" box="[1330,1427,907,933]" captionStart="FIGURE 6" captionStartId="20.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-22@20.[185,1402,181,1919]" captionTargetPageId="20" captionText="FIGURE 6. A–G. Edessa (E.) brevihumeralis sp. n.:A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146358" httpUri="https://zenodo.org/record/10146358/files/figure.png" pageId="19" pageNumber="20">Fig. 6 G</figureCitation>
) with arms of anterior bifurcation straight and laterally barely expanded at apex; anterior bifurcation somewhat excavated receiving fourth and a small part of the third rostral segment. Legs with tibiae and tarsi reddish, remaining yellow to orange (
<figureCitation id="51D92A26FFC09859FE99DDB074CDFBD0" box="[365,478,1015,1042]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 F</figureCitation>
). Abdomen with spine of segment III acuminated (
<figureCitation id="51D92A26FFC09859FBC7DDB07184FBD0" box="[1075,1175,1015,1041]" captionStart="FIGURE 6" captionStartId="20.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-22@20.[185,1402,181,1919]" captionTargetPageId="20" captionText="FIGURE 6. A–G. Edessa (E.) brevihumeralis sp. n.:A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146358" httpUri="https://zenodo.org/record/10146358/files/figure.png" pageId="19" pageNumber="20">Fig. 6 G</figureCitation>
). Intersegmental areas covered by narrow solid black bands with well-defined margins, reaching the lateral spots (
<figureCitation id="51D92A26FFC09859FB76DA5C71FAFBF4" box="[1154,1257,1051,1078]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 F</figureCitation>
). Pseudosutures concolorous with the surrounding surface (
<figureCitation id="51D92A26FFC09859FD73DA7877E7FB9B" box="[647,756,1087,1114]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 F</figureCitation>
, dark lines are a deterioration of the exoskeleton). Median longitudinal brown band incomplete, restricted to the segment VII (
<figureCitation id="51D92A26FFC09859FC89DA2476F6FBBC" box="[893,997,1123,1150]" captionStart="FIGURE 55" captionStartId="108.[152,255,1943,1967]" captionTargetBox="[260,1328,186,1913]" captionTargetId="figure-22@108.[254,1334,181,1919]" captionTargetPageId="108" captionText="FIGURE 55. A–F. Dorsal and ventral surface: A–B, Edessa (E.) atricornis sp. n.; C–D, Edessa (E.) brevicornis sp. n.; E–F, Edessa (E.) brevihumeralis sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146459" httpUri="https://zenodo.org/record/10146459/files/figure.png" pageId="19" pageNumber="20">Fig. 55 F</figureCitation>
). Trichobothria one in line with spiracle and the other laterad. Posterolateral angles of segment VII exceed the level of apices of laterotergites IX (
<figureCitation id="51D92A26FFC09859FAACDAC075B9FB04" captionStart="FIGURE 6" captionStartId="20.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-22@20.[185,1402,181,1919]" captionTargetPageId="20" captionText="FIGURE 6. A–G. Edessa (E.) brevihumeralis sp. n.:A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146358" httpUri="https://zenodo.org/record/10146358/files/figure.png" pageId="19" pageNumber="20">Fig. 6 F</figureCitation>
). Male genitalia, posterolateral angles of the pygophore developed, projected laterally (
<figureCitation id="51D92A26FFC09859FB71DAEC701BFB04" box="[1157,1288,1195,1221]" captionStart="FIGURE 6" captionStartId="20.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-22@20.[185,1402,181,1919]" captionTargetPageId="20" captionText="FIGURE 6. A–G. Edessa (E.) brevihumeralis sp. n.:A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146358" httpUri="https://zenodo.org/record/10146358/files/figure.png" pageId="19" pageNumber="20">Fig. 6 A–B</figureCitation>
); apices with spot black (
<figureCitation id="51D92A26FFC09859FEE9DA8874F4FB28" box="[285,487,1231,1257]" captionStart="FIGURE 6" captionStartId="20.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-22@20.[185,1402,181,1919]" captionTargetPageId="20" captionText="FIGURE 6. A–G. Edessa (E.) brevihumeralis sp. n.:A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146358" httpUri="https://zenodo.org/record/10146358/files/figure.png" pageId="19" pageNumber="20">Fig. 6 A–B, D–E</figureCitation>
). Superior processes of the genital cup rectangular, narrow, flattened and coarse in posterior view, continuing ventrally in a crenulate high carina with a dentiform projection (
<figureCitation id="51D92A26FFC09859FB52DAB4703BFACF" box="[1190,1320,1267,1294]" captionStart="FIGURE 6" captionStartId="20.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-22@20.[185,1402,181,1919]" captionTargetPageId="20" captionText="FIGURE 6. A–G. Edessa (E.) brevihumeralis sp. n.:A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146358" httpUri="https://zenodo.org/record/10146358/files/figure.png" pageId="19" pageNumber="20">Fig. 6 B, E</figureCitation>
, crenulate carina barely visible). Parameres (
<figureCitation id="51D92A26FFC09859FDD0DB5077BFFAF0" box="[548,684,1303,1329]" captionStart="FIGURE 6" captionStartId="20.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-22@20.[185,1402,181,1919]" captionTargetPageId="20" captionText="FIGURE 6. A–G. Edessa (E.) brevihumeralis sp. n.:A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146358" httpUri="https://zenodo.org/record/10146358/files/figure.png" pageId="19" pageNumber="20">Fig. 6 C–D</figureCitation>
) dark brown; anterior lobe large and subrectangular; dorsal lobe large, laterally curved and subtriangular; posterior lobe large and rounded. Proctiger, posterior face triangular (
<figureCitation id="51D92A26FFC09859FA9BDB7C75F5FAB8" captionStart="FIGURE 6" captionStartId="20.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-22@20.[185,1402,181,1919]" captionTargetPageId="20" captionText="FIGURE 6. A–G. Edessa (E.) brevihumeralis sp. n.:A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146358" httpUri="https://zenodo.org/record/10146358/files/figure.png" pageId="19" pageNumber="20">Fig. 6 D–E</figureCitation>
). Ventral surface densely punctured; points distributed in “V”. Ventral rim setose, with median tuft (
<figureCitation id="51D92A26FFC09859FAADDB1875DAFA5C" captionStart="FIGURE 6" captionStartId="20.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-22@20.[185,1402,181,1919]" captionTargetPageId="20" captionText="FIGURE 6. A–G. Edessa (E.) brevihumeralis sp. n.:A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146358" httpUri="https://zenodo.org/record/10146358/files/figure.png" pageId="19" pageNumber="20">Fig. 6 A,C</figureCitation>
); with expansions little developed, rounded and concolorous with the surrounding surface (
<figureCitation id="51D92A26FFC09859FB27DBC47026FA5C" box="[1235,1333,1411,1437]" captionStart="FIGURE 6" captionStartId="20.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-22@20.[185,1402,181,1919]" captionTargetPageId="20" captionText="FIGURE 6. A–G. Edessa (E.) brevihumeralis sp. n.:A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146358" httpUri="https://zenodo.org/record/10146358/files/figure.png" pageId="19" pageNumber="20">Fig. 6 C</figureCitation>
). Female genitalia, plates with dark and deep punctures. Valvifers VIII, sutural margins contiguous and brown, distally with small U-shaped concavity; posterior margin sinuous. Laterotergites VIII with one small dark spot adjacent to outer lateral margins. Laterotergites IX with apices rounded, exceeding the mediotergite VIII (
<figureCitation id="51D92A26FFC09859FB83DBA871C0F9C8" box="[1143,1235,1519,1545]" captionStart="FIGURE 6" captionStartId="20.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-22@20.[185,1402,181,1919]" captionTargetPageId="20" captionText="FIGURE 6. A–G. Edessa (E.) brevihumeralis sp. n.:A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146358" httpUri="https://zenodo.org/record/10146358/files/figure.png" pageId="19" pageNumber="20">Fig. 6 F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFC09859FF33D85477DBF8E8" pageId="19" pageNumber="20" type="discussion">
<paragraph id="C95D36A3FFC09859FF33D85477DBF8E8" blockId="19.[151,1437,259,1870]" pageId="19" pageNumber="20">
<emphasis id="FB96EAB1FFC09859FF33D854745CF9EC" bold="true" box="[199,335,1555,1581]" pageId="19" pageNumber="20">Comments.</emphasis>
<taxonomicName id="0EE24D20FFC09859FEA2D854779BF9EC" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[342,648,1555,1581]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="brevihumeralis" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FEA2D854779BF9EC" box="[342,648,1555,1581]" italics="true" pageId="19" pageNumber="20">Edessa (E.) brevihumeralis</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFC09859FD7BD85477DDF9EC" bold="true" box="[655,718,1555,1581]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="E0A557CAFFC09859FD7BD85477DDF9EC" box="[655,718,1555,1581]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="0EE24D20FFC09859FCB8D8547134F9EC" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[844,1063,1555,1581]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="imitans" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FCB8D8547134F9EC" box="[844,1063,1555,1581]" italics="true" pageId="19" pageNumber="20">Edessa (E.) imitans</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFC09859FBDBD854717DF9EC" bold="true" box="[1071,1134,1555,1581]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="E0A557CAFFC09859FBDBD854717DF9EC" box="[1071,1134,1555,1581]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="0EE24D20FFC09859FB88D8547067F9EC" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[1148,1396,1555,1581]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="indiscreta" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FB88D8547067F9EC" box="[1148,1396,1555,1581]" italics="true" pageId="19" pageNumber="20">Edessa (E.) indiscreta</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFC09859FA8FD85475BDF990" bold="true" pageId="19" pageNumber="20">
<taxonomicNameLabel id="E0A557CAFFC09859FA8FD85475BDF990" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="0EE24D20FFC09859FF48D87074D7F990" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[188,452,1591,1617]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tenuivittata" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FF48D87074D7F990" box="[188,452,1591,1617]" italics="true" pageId="19" pageNumber="20">Edessa (E.) tenuivittata</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFC09859FE3FD8707719F990" bold="true" box="[459,522,1591,1617]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="E0A557CAFFC09859FE3FD8707719F990" box="[459,522,1591,1617]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="0EE24D20FFC09859FDBCD87076AEF990" authority="Breddin, 1903" authorityName="Breddin" authorityYear="1903" box="[584,957,1591,1617]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lavata" subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FDBCD8707605F990" box="[584,790,1591,1617]" italics="true" pageId="19" pageNumber="20">Edessa (E.) lavata</emphasis>
Breddin, 1903
</taxonomicName>
(see
<bibRefCitation id="AD734B52FFC09859FC0CD87071E4F990" author="Mendonca, M. T. S. &amp; Silva, V. J. &amp; Fernandes, J. A. M." box="[1016,1271,1591,1618]" pageId="19" pageNumber="20" pagination="1 - 63" refId="ref74748" refString="Mendonca, M. T. S., Silva, V. J. &amp; Fernandes, J. A. M. (2023) Diagnose of the nominal subgenus of Edessa and description of the E. sexdens group (Heteroptera, Pentatomidae, Edessinae). Zootaxa, 5240 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 5240.1.1" type="journal article" year="2023">
Mendonça
<emphasis id="FB96EAB1FFC09859FB82D87F71B4F990" box="[1142,1191,1591,1617]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
., 2023
</bibRefCitation>
,
<figureCitation id="51D92A26FFC09859FAF6D8707086F990" box="[1282,1429,1591,1618]" captionStart="FIGURE 10" captionStartId="27.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-20@27.[185,1402,181,1919]" captionTargetPageId="27" captionText="FIGURE 10. A–G. Edessa (E.) decolorata sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146367" httpUri="https://zenodo.org/record/10146367/files/figure.png" pageId="19" pageNumber="20">Figs. 10 F–G</figureCitation>
,
<figureCitation id="51D92A26FFC09859FF63D81C75DFF9B4" box="[151,204,1627,1653]" captionStart="FIGURE 11" captionStartId="29.[152,255,1943,1967]" captionTargetBox="[189,1398,192,1911]" captionTargetId="figure-20@29.[189,1398,179,1917]" captionTargetPageId="29" captionText="FIGURE 11. A–G. Edessa (E.) echyra sp. n.: A–E, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146369" httpUri="https://zenodo.org/record/10146369/files/figure.png" pageId="19" pageNumber="20">11 F</figureCitation>
). The five species have a dark spot on the humeral angles restricted to the angles in dorsal view, and ventrally black intersegmental areas on the abdomen. Species can be separated by some characteristics, pronotum with black punctures in cicatrices occurs in
<taxonomicName id="0EE24D20FFC09859FDF8D8E3761BF97C" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[524,776,1699,1725]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="brevihumeralis" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FDF8D8E3761BF97C" box="[524,776,1699,1725]" italics="true" pageId="19" pageNumber="20">E. (E.) brevihumeralis</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFC09859FCE4D8E47642F97C" bold="true" box="[784,849,1699,1725]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="E0A557CAFFC09859FCE4D8E47642F97C" box="[784,849,1699,1725]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="0EE24D20FFC09859FC94D8E37120F97C" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[864,1075,1699,1725]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tenuivittata" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FC94D8E37120F97C" box="[864,1075,1699,1725]" italics="true" pageId="19" pageNumber="20">E. (E.) tenuivittata</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFC09859FBCFD8E4716FF97C" bold="true" box="[1083,1148,1699,1725]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="E0A557CAFFC09859FBCFD8E4716FF97C" box="[1083,1148,1699,1725]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="0EE24D20FFC09859FB40D8E3705DF97C" authorityName="Breddin" authorityYear="1903" box="[1204,1358,1699,1725]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lavata" subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FB40D8E3705DF97C" box="[1204,1358,1699,1725]" italics="true" pageId="19" pageNumber="20">E. (E.) lavata</emphasis>
</taxonomicName>
; while genital plates punctured with black punctures are found in
<taxonomicName id="0EE24D20FFC09859FCDBD88F7139F920" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[815,1066,1735,1761]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="brevihumeralis" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FCDBD88F7139F920" box="[815,1066,1735,1761]" italics="true" pageId="19" pageNumber="20">E. (E.) brevihumeralis</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFC09859FBC5D8807162F920" bold="true" box="[1073,1137,1735,1761]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="E0A557CAFFC09859FBC5D8807162F920" box="[1073,1137,1735,1761]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="0EE24D20FFC09859FB74D88F7037F920" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[1152,1316,1735,1761]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="imitans" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FB74D88F7037F920" box="[1152,1316,1735,1761]" italics="true" pageId="19" pageNumber="20">E. (E.) imitans</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFC09859FAD8D880707FF920" bold="true" box="[1324,1388,1735,1761]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="E0A557CAFFC09859FAD8D880707FF920" box="[1324,1388,1735,1761]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="0EE24D20FFC09859FF63D8AB743EF8C4" authorityName="Breddin" authorityYear="1903" box="[151,301,1771,1797]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lavata" subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FF63D8AB743EF8C4" box="[151,301,1771,1797]" italics="true" pageId="19" pageNumber="20">E. (E.) lavata</emphasis>
</taxonomicName>
, but the shape of the apices of laterotergites IX are different (
<taxonomicName id="0EE24D20FFC09859FC2CD8AB71C3F8C4" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[984,1232,1771,1797]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="brevihumeralis" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FC2CD8AB71C3F8C4" box="[984,1232,1771,1797]" italics="true" pageId="19" pageNumber="20">E. (E.) brevihumeralis</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFC09859FB22D8AC7006F8C4" bold="true" box="[1238,1301,1771,1797]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="E0A557CAFFC09859FB22D8AC7006F8C4" box="[1238,1301,1771,1797]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
rounded,
<taxonomicName id="0EE24D20FFC09859FA70D8AB7408F8E8" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="imitans" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FA70D8AB7408F8E8" italics="true" pageId="19" pageNumber="20">E. (E.) imitans</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFC09859FED6D9487471F8E8" bold="true" box="[290,354,1807,1833]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="E0A557CAFFC09859FED6D9487471F8E8" box="[290,354,1807,1833]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="0EE24D20FFC09859FE6CD957773CF8E8" authorityName="Breddin" authorityYear="1903" box="[408,559,1807,1833]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lavata" subGenus="Edessa">
<emphasis id="FB96EAB1FFC09859FE6CD957773CF8E8" box="[408,559,1807,1833]" italics="true" pageId="19" pageNumber="20">E. (E.) lavata</emphasis>
</taxonomicName>
acuminated).
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFC09859FF33D974711BF88F" box="[199,1032,1843,1870]" pageId="19" pageNumber="20" type="distribution">
<paragraph id="C95D36A3FFC09859FF33D974711BF88F" blockId="19.[151,1437,259,1870]" box="[199,1032,1843,1870]" pageId="19" pageNumber="20">
<emphasis id="FB96EAB1FFC09859FF33D974744AF88C" bold="true" box="[199,345,1843,1869]" pageId="19" pageNumber="20">Distribution</emphasis>
(
<figureCitation id="51D92A26FFC09859FE9DD97474A9F88C" box="[361,442,1843,1869]" captionStart="FIGURE 72" captionStartId="125.[152,255,1071,1095]" captionTargetBox="[195,1391,181,1040]" captionTargetId="figure-20@125.[195,1391,181,1050]" captionTargetPageId="125" captionText="FIGURE 72. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146495" httpUri="https://zenodo.org/record/10146495/files/figure.png" pageId="19" pageNumber="20">Fig. 72</figureCitation>
).
<collectingCountry id="B1F57633FFC09859FE24D9747729F88C" box="[464,570,1843,1869]" name="Brazil" pageId="19" pageNumber="20">BRAZIL</collectingCountry>
:
<collectingRegion id="0B26F841FFC09859FDB0D97377CBF88C" box="[580,728,1843,1870]" country="Brazil" name="Mato Grosso" pageId="19" pageNumber="20">Mato Grosso</collectingRegion>
,
<collectingRegion id="0B26F841FFC09859FD10D9747646F88C" box="[740,853,1843,1869]" country="Brazil" name="Rondonia" pageId="19" pageNumber="20">Rondônia</collectingRegion>
;
<collectingCountry id="B1F57633FFC09859FCABD97376B8F88F" box="[863,939,1844,1870]" name="Peru" pageId="19" pageNumber="20">PERU</collectingCountry>
:
<collectingRegion id="0B26F841FFC09859FC42D9737117F88F" box="[950,1028,1844,1870]" country="Peru" name="Loreto" pageId="19" pageNumber="20">Loreto</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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