treatments-xml/data/41/4B/87/414B87B5FF839818FF63DDCD7615FC64.xml
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<document id="584CF6D57D0516356C34C13D0044F8F9" ID-DOI="10.11646/zootaxa.5372.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10146342" ID-ZooBank="1300D562-3E0B-4F72-933F-8FA7D28F6853" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700186674367" checkinUser="plazi" docAuthor="Mendonça, Maria Thayane Da Silva, Silva, Valéria Juliete Da &amp; Fernandes, José Antônio Marin" docDate="2023" docId="414B87B5FF839818FF63DDCD7615FC64" docLanguage="en" docName="zootaxa.5372.1.1.pdf" docOrigin="Zootaxa 5372 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5372.1.1/52272" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Edessa (Edessa) pseudonigroangulata Mendonça &amp; Silva &amp; Fernandes 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="83" masterDocId="BD72FFCDFFD3984AFFF4DE477513FFC1" masterDocTitle="Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="81" updateTime="1700188109802" updateUser="ExternalLinkService">
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<mods:title id="43502EB15D4C797C610712117FA890B5">Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)</mods:title>
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<mods:namePart id="01512262B1F8ED0757E1BBA89FC69928">Mendonça, Maria Thayane Da Silva</mods:namePart>
<mods:affiliation id="3C56D23AFB5F5226A919BFD8D80052E3">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Programa de Pós-Graduação em Zoologia UFPA / MPEG</mods:affiliation>
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<mods:namePart id="0DECB9F1A0B45D1299CAEF8F7881BB2C">Silva, Valéria Juliete Da</mods:namePart>
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<mods:affiliation id="F136C0817BBF5039BBB1422C7CF68C1F">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; valeriajuliete @ hotmail. com; https: // orcid. org / 0000 - 0002 - 1845 - 5228</mods:affiliation>
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<mods:namePart id="33E8D73A032D3BE49DD22167A829282D">Fernandes, José Antônio Marin</mods:namePart>
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<mods:affiliation id="A1BB2AEFA1F4013861963250D25C7F7F">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Corresponding author: joseamf @ ufpa. br; https: // orcid. org / 0000 - 0001 - 7450 - 5296</mods:affiliation>
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<emphasis id="FB96EAB1FF83981AFF63DDCD77BDFC64" bold="true" box="[151,686,906,933]" pageId="80" pageNumber="81">
<taxonomicName id="0EE24D20FF83981AFF63DDCD7776FC64" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[151,613,906,933]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pseudonigroangulata" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FF83981AFF63DDCD7776FC64" bold="true" box="[151,613,906,933]" italics="true" pageId="80" pageNumber="81">Edessa (Edessa) pseudonigroangulata</emphasis>
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(
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Figs.
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<figureCitation id="51D92A26FF83981AFEFDDDE8744FFC08" box="[265,348,943,969]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">66 EF</figureCitation>
,
<figureCitation id="51D92A26FF83981AFE92DDE87490FC08" box="[358,387,943,969]" captionStart="FIGURE 74" captionStartId="126.[152,255,1633,1657]" captionTargetBox="[183,1404,741,1606]" captionTargetId="figure-270@126.[183,1404,741,1613]" captionTargetPageId="126" captionText="FIGURE 74. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146499" httpUri="https://zenodo.org/record/10146499/files/figure.png" pageId="80" pageNumber="81">74</figureCitation>
)
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<emphasis id="FB96EAB1FF83981AFF63DDB7740BFBCB" bold="true" box="[151,280,1008,1034]" pageId="80" pageNumber="81">Etymology</emphasis>
. The name refers to the similarity with
<taxonomicName id="0EE24D20FF83981AFD20DDB671F6FBCA" authority="Fernandes &amp; Silva, 2015" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[724,1253,1008,1035]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="nigroangulata" subGenus="Edessa">
<emphasis id="FB96EAB1FF83981AFD20DDB676D7FBCB" box="[724,964,1008,1034]" italics="true" pageId="80" pageNumber="81">E. (E.) nigroangulata</emphasis>
<bibRefCitation id="AD734B52FF83981AFC3FDDB771F6FBCA" author="Fernandes, J. A. &amp; Silva, V. J. &amp; Correia, A. O. &amp; Nunes, B. M." box="[971,1253,1008,1035]" pageId="80" pageNumber="81" pagination="511 - 536" refId="ref74519" refString="Fernandes, J. A. M, Silva, V. J., Correia, A. O. &amp; Nunes, B. M. (2015) New species of Edessa Fabricius, 1803 (Hemiptera: Pentatomidae) from Costa Rica. Zootaxa, 3999 (4), 511 - 536. https: // doi. org / 10.11646 / zootaxa. 3999.4.3" type="journal article" year="2015">Fernandes &amp; Silva, 2015</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<emphasis id="FB96EAB1FF83981AFF33DA5374A7FBEF" bold="true" box="[199,436,1044,1070]" pageId="80" pageNumber="81">Material examined.</emphasis>
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:
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1♁,
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:
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</materialsCitation>
;
<materialsCitation id="798A3CFEFF83981AFDADDAE374BDFB23" collectingDate="1980-03-13" collectingDateMax="1980-03-22" collectingDateMin="1980-03-13" collectionCode="UFPA" collectorName="D. H. Jenzen" country="Costa Rica" location="Corcovado National Park" municipality="Sirena" pageId="80" pageNumber="81" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
<specimenCount id="DFE4FD2AFF83981AFDADDAE3776FFB7F" box="[601,636,1188,1214]" count="1" pageId="80" pageNumber="81" type="female">1♀</specimenCount>
,
<collectingMunicipality id="2939ACD9FF83981AFD7FDAE377C6FB7F" box="[651,725,1188,1214]" pageId="80" pageNumber="81">Sirena</collectingMunicipality>
,
<location id="CC3D6078FF83981AFD15DAE37111FB7F" LSID="urn:lsid:plazi:treatment:414B87B5FF839818FF63DDCD7615FC64:CC3D6078FF83981AFD15DAE37111FB7F" box="[737,1026,1188,1214]" country="Costa Rica" municipality="Sirena" name="Corcovado National Park" pageId="80" pageNumber="81" stateProvince="Puntarenas">Corcovado National Park</location>
,
<date id="BD5C1063FF83981AFBFBDAE371ADFB7F" box="[1039,1214,1188,1214]" pageId="80" pageNumber="81" value="1980-03-13" valueMax="1980-03-22" valueMin="1980-03-13">
<collectingDate id="AD18E98BFF83981AFBFBDAE371ADFB7F" box="[1039,1214,1188,1214]" pageId="80" pageNumber="81" value="1980-03-13" valueMax="1980-03-22" valueMin="1980-03-13">1322-III-1980</collectingDate>
</date>
,
<collectorName id="64175375FF83981AFB3EDAE2704FFB7E" box="[1226,1372,1189,1215]" pageId="80" pageNumber="81">D. H. Jenzen</collectorName>
coll., CRI001 715426 (
<collectionCode id="AFF3AE66FF83981AFEADDA8F74B6FB23" box="[345,421,1224,1250]" pageId="80" pageNumber="81">UFPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="81F86528FF83981AFF33DAAB76E7FA8E" pageId="80" pageNumber="81" type="description">
<paragraph id="C95D36A3FF83981AFF33DAAB76E7FA8E" blockId="80.[151,1437,1008,2043]" pageId="80" pageNumber="81">
<emphasis id="FB96EAB1FF83981AFF33DAAB7465FAC7" bold="true" box="[199,374,1260,1286]" pageId="80" pageNumber="81">Measurements</emphasis>
(n= 4). Total length: 15.416.9; head length: 1.11.9; head width: 3.03.1; pronotum length: 3.23.3; pronotum width: 10.510.7; scutellum length: 7.47.7; scutellum width: 5.86.0; abdominal width: 8.9 9.1; length antennomeres: I: 0.81.0; II: 1.41.5; III: 1.5; IV: 3.03.6; V: 3.5.
</paragraph>
</subSubSection>
<subSubSection id="81F86528FF839818FF33DB1F713FFE10" lastPageId="82" lastPageNumber="83" pageId="80" pageNumber="81" type="diagnosis">
<paragraph id="C95D36A3FF839818FF33DB1F713FFE10" blockId="80.[151,1437,1008,2043]" lastBlockId="82.[151,1437,151,933]" lastPageId="82" lastPageNumber="83" pageId="80" pageNumber="81">
<emphasis id="FB96EAB1FF83981AFF33DB1F7451FAB3" bold="true" box="[199,322,1368,1394]" pageId="80" pageNumber="81">Diagnosis.</emphasis>
Large (
<quantity id="0E1A9B46FF83981AFE6ADB1F775AFAB3" box="[414,585,1368,1395]" metricMagnitude="-2" metricUnit="m" metricValue="1.6149999999999998" metricValueMax="1.69" metricValueMin="1.54" pageId="80" pageNumber="81" unit="mm" value="16.15" valueMax="16.9" valueMin="15.4">15.416.9 mm</quantity>
). Dorsal body surface olive green, densely punctured (
<figureCitation id="51D92A26FF83981AFB37DB1F7026FAB3" box="[1219,1333,1368,1394]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 E</figureCitation>
). Ventral surface dark yellow to reddish brown with transversal black bands on thorax and abdomen (
<figureCitation id="51D92A26FF83981AFB22DB3B705AFA57" box="[1238,1353,1404,1430]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 F</figureCitation>
). Head with black punctures. Antennae yellow (
<figureCitation id="51D92A26FF83981AFD91DBE777E7FA7B" box="[613,756,1440,1466]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 EF</figureCitation>
). Pronotum with brown punctures (
<figureCitation id="51D92A26FF83981AFB7EDBE771EAFA7B" box="[1162,1273,1440,1466]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 E</figureCitation>
); anterolateral margin and cicatrices with black punctures. Humeral angles short (1.5 times wider than long); apex with black spot restricted to the angles in dorsal and ventral views; bent backward (
<figureCitation id="51D92A26FF83981AFC36DBAF713CF9C3" box="[962,1071,1512,1538]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 E</figureCitation>
). Scutellum with black to brown punctures; posterior part excavated medially; apex at the same level as the end of coria (
<figureCitation id="51D92A26FF83981AFB6FD84B701FF9E7" box="[1179,1292,1548,1574]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 E</figureCitation>
). Coria with all veins concolorous with the surrounding surface (
<figureCitation id="51D92A26FF83981AFD03D8777678F98B" box="[759,875,1584,1610]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 E</figureCitation>
). Connexival segments with concavities entirely covered by rectangular black spots separated by a large yellow median spot (
<figureCitation id="51D92A26FF83981AFC0FD8137174F9AF" box="[1019,1127,1620,1646]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 E</figureCitation>
), spots extending ventrally, subrectangular (
<figureCitation id="51D92A26FF83981AFEBED83F74A3F953" box="[330,432,1656,1682]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 F</figureCitation>
). Posterolateral angles of connexivum with apices black (
<figureCitation id="51D92A26FF83981AFBD3D83F7183F953" box="[1063,1168,1656,1682]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 E</figureCitation>
). Ventral surface, thorax with black bands; dark band of the propleura covering 2/3 of the width of the sclerite (
<figureCitation id="51D92A26FF83981AFBB3D8DB71BCF977" box="[1095,1199,1692,1718]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 F</figureCitation>
). Proepisternum with dark band (
<figureCitation id="51D92A26FF83981AFEEDD8877494F91B" box="[281,391,1728,1754]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 F</figureCitation>
). Evaporatorium concolorous with the surrounding surface (
<figureCitation id="51D92A26FF83981AFBC3D88771B5F91B" box="[1079,1190,1728,1754]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 F</figureCitation>
). Metasternal process (
<figureCitation id="51D92A26FF83981AFF6BD8A37407F93F" box="[159,276,1764,1790]" captionStart="FIGURE 39" captionStartId="81.[152,255,1943,1967]" captionTargetBox="[186,1402,187,1913]" captionTargetId="figure-20@81.[186,1402,181,1919]" captionTargetPageId="81" captionText="FIGURE 39. AG. Edessa (E.) pseudonigroangulata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146426" httpUri="https://zenodo.org/record/10146426/files/figure.png" pageId="80" pageNumber="81">Figs. 39 G</figureCitation>
;
<figureCitation id="51D92A26FF83981AFED6D8A37445F93F" box="[290,342,1764,1790]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">66 F</figureCitation>
) with arms of anterior bifurcation straight and laterally well expanded at apex; anterior bifurcation broadly excavated receiving fourth rostral segment. Legs with tibiae and tarsi reddish, remaining dark yellow (
<figureCitation id="51D92A26FF83981AFA9BD94F75C3F887" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 F</figureCitation>
). Abdomen with spine of segment III rounded (
<figureCitation id="51D92A26FF83981AFD03D96B7661F887" box="[759,882,1836,1862]" captionStart="FIGURE 39" captionStartId="81.[152,255,1943,1967]" captionTargetBox="[186,1402,187,1913]" captionTargetId="figure-20@81.[186,1402,181,1919]" captionTargetPageId="81" captionText="FIGURE 39. AG. Edessa (E.) pseudonigroangulata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146426" httpUri="https://zenodo.org/record/10146426/files/figure.png" pageId="80" pageNumber="81">Figs. 39 G</figureCitation>
;
<figureCitation id="51D92A26FF83981AFC70D96B76AFF887" box="[900,956,1836,1862]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">66 F</figureCitation>
). Intersegmental areas covered by narrow black bands with smudged margins, reaching the lateral spots (
<figureCitation id="51D92A26FF83981AFCA3D91776D1F8AB" box="[855,962,1872,1898]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 F</figureCitation>
). Pseudosutures solid black with smudged margins (
<figureCitation id="51D92A26FF83981AFEF4D9337479F84F" box="[256,362,1908,1934]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 F</figureCitation>
). Median longitudinal black band incomplete (
<figureCitation id="51D92A26FF83981AFC80D93376CEF84F" box="[884,989,1908,1934]" captionStart="FIGURE 66" captionStartId="119.[152,255,1943,1967]" captionTargetBox="[264,1323,189,1915]" captionTargetId="figure-20@119.[258,1329,181,1919]" captionTargetPageId="119" captionText="FIGURE 66. AF. Dorsal and ventral surface:AB, Edessa (E.) omopalla sp. n.; CD, Edessa (E.) proxima sp. n.; EF, Edessa (E.) pseudonigroangulata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146483" httpUri="https://zenodo.org/record/10146483/files/figure.png" pageId="80" pageNumber="81">Fig. 66 F</figureCitation>
). Trichobothria one in line with spiracle and the other laterad. Posterolateral angles of segment VII not reaching the level of apices of laterotergites IX (
<figureCitation id="51D92A26FF83981AFA9BD9DF75DFF817" captionStart="FIGURE 39" captionStartId="81.[152,255,1943,1967]" captionTargetBox="[186,1402,187,1913]" captionTargetId="figure-20@81.[186,1402,181,1919]" captionTargetPageId="81" captionText="FIGURE 39. AG. Edessa (E.) pseudonigroangulata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146426" httpUri="https://zenodo.org/record/10146426/files/figure.png" pageId="80" pageNumber="81">Fig. 39 F</figureCitation>
). Male genitalia, dorsal side of the pygophore with a suffused brown area occupying 2/3 of the surface (
<figureCitation id="51D92A26FF83981AFAB8D9FB75DCF83B" captionStart="FIGURE 39" captionStartId="81.[152,255,1943,1967]" captionTargetBox="[186,1402,187,1913]" captionTargetId="figure-20@81.[186,1402,181,1919]" captionTargetPageId="81" captionText="FIGURE 39. AG. Edessa (E.) pseudonigroangulata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146426" httpUri="https://zenodo.org/record/10146426/files/figure.png" pageId="80" pageNumber="81">Fig. 39 AB</figureCitation>
). Posterolateral angles of the pygophore developed; well projected laterally, apices with spots black (
<figureCitation id="51D92A26FF839818FAB3D9A775E1FF70" captionStart="FIGURE 39" captionStartId="81.[152,255,1943,1967]" captionTargetBox="[186,1402,187,1913]" captionTargetId="figure-20@81.[186,1402,181,1919]" captionTargetPageId="81" captionText="FIGURE 39. AG. Edessa (E.) pseudonigroangulata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146426" httpUri="https://zenodo.org/record/10146426/files/figure.png" lastPageId="82" lastPageNumber="83" pageId="80" pageNumber="81">Fig. 39 AB, D</figureCitation>
). Superior processes of the genital cup rectangular, thick, in flattened and coarse posterior view, continuing ventrally in a crenulate, long and high carina (
<figureCitation id="51D92A26FF819818FD61DEFC760FFF14" box="[661,796,187,213]" captionStart="FIGURE 39" captionStartId="81.[152,255,1943,1967]" captionTargetBox="[186,1402,187,1913]" captionTargetId="figure-20@81.[186,1402,181,1919]" captionTargetPageId="81" captionText="FIGURE 39. AG. Edessa (E.) pseudonigroangulata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146426" httpUri="https://zenodo.org/record/10146426/files/figure.png" pageId="82" pageNumber="83">Fig. 39 B, E</figureCitation>
crenulate carina barely visible). Parameres (
<figureCitation id="51D92A26FF819818FAFEDEFC7086FF14" box="[1290,1429,187,213]" captionStart="FIGURE 39" captionStartId="81.[152,255,1943,1967]" captionTargetBox="[186,1402,187,1913]" captionTargetId="figure-20@81.[186,1402,181,1919]" captionTargetPageId="81" captionText="FIGURE 39. AG. Edessa (E.) pseudonigroangulata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146426" httpUri="https://zenodo.org/record/10146426/files/figure.png" pageId="82" pageNumber="83">Fig. 39 DE</figureCitation>
) with brown margins; anterior lobe subrectangular; dorsal lobe rounded, with strongly curved apex; posterior lobe subrectangular. Proctiger, posterior face subtriangular (
<figureCitation id="51D92A26FF819818FCF0DF447686FEDC" box="[772,917,259,285]" captionStart="FIGURE 39" captionStartId="81.[152,255,1943,1967]" captionTargetBox="[186,1402,187,1913]" captionTargetId="figure-20@81.[186,1402,181,1919]" captionTargetPageId="81" captionText="FIGURE 39. AG. Edessa (E.) pseudonigroangulata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146426" httpUri="https://zenodo.org/record/10146426/files/figure.png" pageId="82" pageNumber="83">Fig. 39 DE</figureCitation>
). Surface ventral with dark spots laterally and medially (
<figureCitation id="51D92A26FF819818FEFDDF607465FE80" box="[265,374,295,321]" captionStart="FIGURE 39" captionStartId="81.[152,255,1943,1967]" captionTargetBox="[186,1402,187,1913]" captionTargetId="figure-20@81.[186,1402,181,1919]" captionTargetPageId="81" captionText="FIGURE 39. AG. Edessa (E.) pseudonigroangulata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146426" httpUri="https://zenodo.org/record/10146426/files/figure.png" pageId="82" pageNumber="83">Fig. 39 C</figureCitation>
). Ventral rim setose, with medial tuft (
<figureCitation id="51D92A26FF819818FCD1DF6076A7FE80" box="[805,948,295,321]" captionStart="FIGURE 39" captionStartId="81.[152,255,1943,1967]" captionTargetBox="[186,1402,187,1913]" captionTargetId="figure-20@81.[186,1402,181,1919]" captionTargetPageId="81" captionText="FIGURE 39. AG. Edessa (E.) pseudonigroangulata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146426" httpUri="https://zenodo.org/record/10146426/files/figure.png" pageId="82" pageNumber="83">Fig. 39 AC</figureCitation>
); with expansions little developed, rounded and black (
<figureCitation id="51D92A26FF819818FEEEDF0C749DFEA4" box="[282,398,331,357]" captionStart="FIGURE 39" captionStartId="81.[152,255,1943,1967]" captionTargetBox="[186,1402,187,1913]" captionTargetId="figure-20@81.[186,1402,181,1919]" captionTargetPageId="81" captionText="FIGURE 39. AG. Edessa (E.) pseudonigroangulata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146426" httpUri="https://zenodo.org/record/10146426/files/figure.png" pageId="82" pageNumber="83">Fig. 39 C</figureCitation>
). Female genitalia, valvifers VIII with dark punctures; sutural margins contiguous brown and not divergent; posterior margin in open V-shaped excavation, brown and slightly arched. Valvifers IX not carinated, large. Laterotergites VIII with two dark spots on lateral and a large yellow spot in middle of the margins. Laterotergites IX with apices acuminate passing the mediotergite VIII (
<figureCitation id="51D92A26FF819818FC41DFF0710CFE10" box="[949,1055,439,465]" captionStart="FIGURE 39" captionStartId="81.[152,255,1943,1967]" captionTargetBox="[186,1402,187,1913]" captionTargetId="figure-20@81.[186,1402,181,1919]" captionTargetPageId="81" captionText="FIGURE 39. AG. Edessa (E.) pseudonigroangulata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146426" httpUri="https://zenodo.org/record/10146426/files/figure.png" pageId="82" pageNumber="83">Fig. 39 F</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="9D9D662BFF82981BFF6CD9D07130F812" ID-DOI="http://doi.org/10.5281/zenodo.10146426" ID-Zenodo-Dep="10146426" httpUri="https://zenodo.org/record/10146426/files/figure.png" pageId="81" pageNumber="82" startId="81.[152,255,1943,1967]" targetBox="[186,1402,187,1913]" targetPageId="81" targetType="figure">
<paragraph id="C95D36A3FF82981BFF6CD9D07130F812" blockId="81.[151,1437,1943,2004]" pageId="81" pageNumber="82">
<emphasis id="FB96EAB1FF82981BFF6CD9D0743BF86E" bold="true" box="[152,296,1943,1967]" pageId="81" pageNumber="82">FIGURE 39.</emphasis>
AG.
<taxonomicName id="0EE24D20FF82981BFE84D9DF77DBF871" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[368,712,1944,1968]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="pseudonigroangulata" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FF82981BFE84D9DF77DBF871" box="[368,712,1944,1968]" italics="true" pageId="81" pageNumber="82">Edessa (E.) pseudonigroangulata</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FF82981BFD25D9DF761EF871" bold="true" box="[721,781,1944,1968]" pageId="81" pageNumber="82">
<taxonomicNameLabel id="E0A557CAFF82981BFD25D9DF761EF871" box="[721,781,1944,1968]" pageId="81" pageNumber="82" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm.
</paragraph>
</caption>
<subSubSection id="81F86528FF819818FF33DF9D7064FC40" pageId="82" pageNumber="83" type="discussion">
<paragraph id="C95D36A3FF819818FF33DF9D7064FC40" blockId="82.[151,1437,151,933]" pageId="82" pageNumber="83">
<emphasis id="FB96EAB1FF819818FF33DF9D745CFE35" bold="true" box="[199,335,474,500]" pageId="82" pageNumber="83">Comments.</emphasis>
<taxonomicName id="0EE24D20FF819818FEA2DF9C77DEFE34" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[342,717,475,501]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="pseudonigroangulata" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FF819818FEA2DF9C77DEFE34" box="[342,717,475,501]" italics="true" pageId="82" pageNumber="83">Edessa (E.) pseudonigroangulata</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FF819818FD27DF9C7600FE34" bold="true" box="[723,787,475,501]" pageId="82" pageNumber="83">
<taxonomicNameLabel id="E0A557CAFF819818FD27DF9C7600FE34" box="[723,787,475,501]" pageId="82" pageNumber="83" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="0EE24D20FF819818FC64DF9C708FFE34" authority="Fernandes &amp; Silva, 2015" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[912,1436,474,501]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nigroangulata" subGenus="Edessa">
<emphasis id="FB96EAB1FF819818FC64DF9C716CFE34" box="[912,1151,475,501]" italics="true" pageId="82" pageNumber="83">E. (E.) nigroangulata</emphasis>
<bibRefCitation id="AD734B52FF819818FB72DF9C708FFE34" author="Fernandes, J. A. &amp; Silva, V. J. &amp; Correia, A. O. &amp; Nunes, B. M." box="[1158,1436,474,501]" pageId="82" pageNumber="83" pagination="511 - 536" refId="ref74519" refString="Fernandes, J. A. M, Silva, V. J., Correia, A. O. &amp; Nunes, B. M. (2015) New species of Edessa Fabricius, 1803 (Hemiptera: Pentatomidae) from Costa Rica. Zootaxa, 3999 (4), 511 - 536. https: // doi. org / 10.11646 / zootaxa. 3999.4.3" type="journal article" year="2015">Fernandes &amp; Silva, 2015</bibRefCitation>
</taxonomicName>
(see description in
<bibRefCitation id="AD734B52FF819818FE9DDFB8774CFDD8" author="Fernandes, J. A. &amp; Silva, V. J. &amp; Correia, A. O. &amp; Nunes, B. M." box="[361,607,510,537]" pageId="82" pageNumber="83" pagination="511 - 536" refId="ref74519" refString="Fernandes, J. A. M, Silva, V. J., Correia, A. O. &amp; Nunes, B. M. (2015) New species of Edessa Fabricius, 1803 (Hemiptera: Pentatomidae) from Costa Rica. Zootaxa, 3999 (4), 511 - 536. https: // doi. org / 10.11646 / zootaxa. 3999.4.3" type="journal article" year="2015">
Fernandes
<emphasis id="FB96EAB1FF819818FE16DFB87709FDD8" box="[482,538,511,537]" italics="true" pageId="82" pageNumber="83">et al.</emphasis>
, 2015
</bibRefCitation>
and images in
<bibRefCitation id="AD734B52FF819818FCF1DFB87112FDD8" author="Mendonca, M. T. S. &amp; Silva, V. J. &amp; Fernandes, J. A. M." box="[773,1025,511,537]" pageId="82" pageNumber="83" pagination="1 - 63" refId="ref74748" refString="Mendonca, M. T. S., Silva, V. J. &amp; Fernandes, J. A. M. (2023) Diagnose of the nominal subgenus of Edessa and description of the E. sexdens group (Heteroptera, Pentatomidae, Edessinae). Zootaxa, 5240 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 5240.1.1" type="journal article" year="2023">
Mendonça
<emphasis id="FB96EAB1FF819818FC76DFB876AAFDD8" box="[898,953,511,537]" italics="true" pageId="82" pageNumber="83">et al.</emphasis>
, 2023
</bibRefCitation>
,
<figureCitation id="51D92A26FF819818FBF8DFB871B1FDD8" box="[1036,1186,511,537]" captionStart="FIGURE 17" captionStartId="41.[152,255,1943,1967]" captionTargetBox="[195,1392,187,1914]" captionTargetId="figure-20@41.[195,1392,181,1919]" captionTargetPageId="41" captionText="FIGURE 17.AG. Edessa (E.) fuliginocornis sp. n.:AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146381" httpUri="https://zenodo.org/record/10146381/files/figure.png" pageId="82" pageNumber="83">Figs. 17 AC</figureCitation>
,
<figureCitation id="51D92A26FF819818FB58DFB87012FDD8" box="[1196,1281,511,537]" captionStart="FIGURE 28" captionStartId="61.[152,255,1943,1967]" captionTargetBox="[185,1402,187,1913]" captionTargetId="figure-20@61.[185,1402,181,1919]" captionTargetPageId="61" captionText="FIGURE 28. AG. Edessa (E.) infuscata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146403" httpUri="https://zenodo.org/record/10146403/files/figure.png" pageId="82" pageNumber="83">28 EF</figureCitation>
), both species share: antennae yellow; coria with all veins concolorous with the surrounding surface; ventrally abdomen with intersegmental areas black; and ventral rim of the pygophore with lateral subdistal tuft of setae. On the other hand, both species can be easily separated by: anterolateral margin of the pronotum coarse, crenulate and emarginated in
<taxonomicName id="0EE24D20FF819818FF40DCC874B6FD68" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[180,421,655,681]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nigroangulata" subGenus="Edessa">
<emphasis id="FB96EAB1FF819818FF40DCC874B6FD68" box="[180,421,655,681]" italics="true" pageId="82" pageNumber="83">E. (E.) nigroangulata</emphasis>
</taxonomicName>
(in
<taxonomicName id="0EE24D20FF819818FE20DCC87607FD68" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[468,788,655,681]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="pseudonigroangulata" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FF819818FE20DCC87607FD68" box="[468,788,655,681]" italics="true" pageId="82" pageNumber="83">E. (E.) pseudonigroangulata</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FF819818FCEFDCC87648FD68" bold="true" box="[795,859,655,681]" pageId="82" pageNumber="83">
<taxonomicNameLabel id="E0A557CAFF819818FCEFDCC87648FD68" box="[795,859,655,681]" pageId="82" pageNumber="83" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
anterolateral margin smooth and not emarginated); width of the humeral angles (wider in
<taxonomicName id="0EE24D20FF819818FDA3DCF4765FFD0C" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[599,844,691,717]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nigroangulata" subGenus="Edessa">
<emphasis id="FB96EAB1FF819818FDA3DCF4765FFD0C" box="[599,844,691,717]" italics="true" pageId="82" pageNumber="83">E. (E.) nigroangulata</emphasis>
</taxonomicName>
); color of the legs (in
<taxonomicName id="0EE24D20FF819818FBA3DCF4708EFD0C" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[1111,1437,691,717]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="pseudonigroangulata" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FF819818FBA3DCF4708EFD0C" box="[1111,1437,691,717]" italics="true" pageId="82" pageNumber="83">E. (E.) pseudonigroangulata</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FF819818FF63DC9075C8FD30" bold="true" box="[151,219,727,753]" pageId="82" pageNumber="83">
<taxonomicNameLabel id="E0A557CAFF819818FF63DC9075C8FD30" box="[151,219,727,753]" pageId="82" pageNumber="83" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
legs with tibiae and tarsi more red than the other articles, all articles yellow in
<taxonomicName id="0EE24D20FF819818FB67DC907098FD30" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[1171,1419,727,753]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nigroangulata" subGenus="Edessa">
<emphasis id="FB96EAB1FF819818FB67DC907098FD30" box="[1171,1419,727,753]" italics="true" pageId="82" pageNumber="83">E. (E.) nigroangulata</emphasis>
</taxonomicName>
); connexival segments with rectangular spots (
<taxonomicName id="0EE24D20FF819818FD7BDCBC766DFCD4" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[655,894,763,789]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nigroangulata" subGenus="Edessa">
<emphasis id="FB96EAB1FF819818FD7BDCBC766DFCD4" box="[655,894,763,789]" italics="true" pageId="82" pageNumber="83">E. (E.) nigroangulata</emphasis>
</taxonomicName>
with trapezoidal spots); ventrally abdomen with pseudosutures black (
<taxonomicName id="0EE24D20FF819818FE73DD587766FCF8" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[391,629,799,825]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nigroangulata" subGenus="Edessa">
<emphasis id="FB96EAB1FF819818FE73DD587766FCF8" box="[391,629,799,825]" italics="true" pageId="82" pageNumber="83">E. (E.) nigroangulata</emphasis>
</taxonomicName>
with pseudosutures brown); dorsal side of the pygophore with a suffused brown area occupying 2/3 of the surface (
<taxonomicName id="0EE24D20FF819818FD9FDD047648FC9C" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[619,859,835,861]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nigroangulata" subGenus="Edessa">
<emphasis id="FB96EAB1FF819818FD9FDD047648FC9C" box="[619,859,835,861]" italics="true" pageId="82" pageNumber="83">E. (E.) nigroangulata</emphasis>
</taxonomicName>
with dorsal side of the pygophore without suffused brown area); and superior processes of the genital cup with high carina (
<taxonomicName id="0EE24D20FF819818FC35DD2071A2FC40" authorityName="Fernandes &amp; Silva" authorityYear="2015" box="[961,1201,871,897]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="nigroangulata" subGenus="Edessa">
<emphasis id="FB96EAB1FF819818FC35DD2071A2FC40" box="[961,1201,871,897]" italics="true" pageId="82" pageNumber="83">E. (E.) nigroangulata</emphasis>
</taxonomicName>
with low carina).
</paragraph>
</subSubSection>
<subSubSection id="81F86528FF819818FF33DDCD7615FC64" box="[199,774,906,933]" pageId="82" pageNumber="83" type="distribution">
<paragraph id="C95D36A3FF819818FF33DDCD7615FC64" blockId="82.[151,1437,151,933]" box="[199,774,906,933]" pageId="82" pageNumber="83">
<emphasis id="FB96EAB1FF819818FF33DDCD744AFC65" bold="true" box="[199,345,906,932]" pageId="82" pageNumber="83">Distribution</emphasis>
(
<figureCitation id="51D92A26FF819818FE9DDDCC74A9FC64" box="[361,442,907,933]" captionStart="FIGURE 74" captionStartId="126.[152,255,1633,1657]" captionTargetBox="[183,1404,741,1606]" captionTargetId="figure-270@126.[183,1404,741,1613]" captionTargetPageId="126" captionText="FIGURE 74. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146499" httpUri="https://zenodo.org/record/10146499/files/figure.png" pageId="82" pageNumber="83">Fig. 74</figureCitation>
):
<collectingCountry id="B1F57633FF819818FE27DDCC776AFC64" box="[467,633,907,933]" name="Costa Rica" pageId="82" pageNumber="83">COSTA RICA</collectingCountry>
:
<collectingRegion id="0B26F841FF819818FD77DDCC7611FC64" box="[643,770,907,933]" country="Costa Rica" name="Puntarenas" pageId="82" pageNumber="83">Puntarenas</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>