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<document id="E7238014773C83AC66881A8D0872BFD8" ID-DOI="10.11646/zootaxa.3640.4.8" ID-GBIF-Dataset="6e7dc9c0-25c8-460c-a7b8-3af0d49dfc77" ID-ISSN="1175-5326" ID-Zenodo-Dep="221490" ID-ZooBank="E7E9E0AB-D80A-47BE-8ACB-DEF05BC78D4B" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460594722954" checkinUser="plazi" docAuthor="Lima, Lucas R. C., Cruz, Paulo Vilela &amp; Pinheiro, Ulisses" docDate="2013" docId="0391103BFF99FF9FFF54FB23FC962820" docLanguage="en" docName="zt03640p600.pdf" docOrigin="Zootaxa 3640 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paracloeodes charrua Emmerich &amp; Nieto 2009" docType="treatment" docVersion="8" lastPageNumber="599" masterDocId="FFA86843FF99FF9DFFC3FF90F8432D4F" masterDocTitle="The adult stage of Paracloeodes charrua Emmerich &amp; Nieto, 2009 (Ephemeroptera: Baetidae)" masterLastPageNumber="600" masterPageNumber="597" pageNumber="597" updateTime="1698326540175" updateUser="plazi">
<mods:mods id="F77FC2F9A54A0C20686AFD6B70A3F8D7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E57519FE67FDD80A2F9B6C0D274E52C6">
<mods:title id="2F3FE38164381E6C803A560182916464">The adult stage of Paracloeodes charrua Emmerich &amp; Nieto, 2009 (Ephemeroptera: Baetidae)</mods:title>
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<mods:namePart id="E95C78265EE2F41009D73FE15FE11CD1">Lima, Lucas R. C.</mods:namePart>
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<mods:name id="F9969FB2E42DAD09597E9F00E7E7378A" type="personal">
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<mods:namePart id="B139EF1A45BCDE590CD29596BFFC68CE">Cruz, Paulo Vilela</mods:namePart>
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<mods:name id="465B92B4AFB396C755A29646B2FF18A3" type="personal">
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<mods:namePart id="4054EEC5A29E751A821F9F6A3E72D7BA">Pinheiro, Ulisses</mods:namePart>
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<mods:title id="01B3002CCAA0DF80986B770A3D1D214B">Zootaxa</mods:title>
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<mods:part id="30AE00AA9E4C54085B283B07DBB13F43">
<mods:date id="96992DE55271951B4FF7E8E060BBE994">2013</mods:date>
<mods:detail id="DC3D4FF8D64A3654DFBAA7F7F024BF4A" type="volume">
<mods:number id="4D874B81543CDFF5CF52BB3452EECD92">3640</mods:number>
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<mods:start id="79BF05B8EA7F790037A4046BE7B405FB">597</mods:start>
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<mods:classification id="DFAFDF6527F566388D7DE5B21E0F214D">journal article</mods:classification>
<mods:identifier id="F3CCDF61CD85D6D169F84BA5F434B1FA" type="DOI">10.11646/zootaxa.3640.4.8</mods:identifier>
<mods:identifier id="3FA20F16FD30AB2A58929AA97CEA0F52" type="GBIF-Dataset">6e7dc9c0-25c8-460c-a7b8-3af0d49dfc77</mods:identifier>
<mods:identifier id="A0FA33B7AE8BE90F10B12743303361C1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6703FD45C72F1B76DB1759D421E96549" type="Zenodo-Dep">221490</mods:identifier>
<mods:identifier id="CFCACA5C7EC6858A22CC306AEC9D8D7A" type="ZooBank">E7E9E0AB-D80A-47BE-8ACB-DEF05BC78D4B</mods:identifier>
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<treatment id="0391103BFF99FF9FFF54FB23FC962820" ID-DOI="http://doi.org/10.5281/zenodo.6164880" ID-GBIF-Taxon="119566348" ID-Zenodo-Dep="6164880" LSID="urn:lsid:plazi:treatment:0391103BFF99FF9FFF54FB23FC962820" httpUri="http://treatment.plazi.org/id/0391103BFF99FF9FFF54FB23FC962820" lastPageId="2" lastPageNumber="599" pageId="0" pageNumber="597">
<subSubSection id="C322F2A6FF99FF9DFF54FB23F95029A3" pageId="0" pageNumber="597" type="nomenclature">
<paragraph id="8B87A12DFF99FF9DFF54FB23FA9D2983" blockId="0.[151,734,1202,1260]" box="[151,734,1202,1229]" pageId="0" pageNumber="597">
<heading id="D0CF1641FF99FF9DFF54FB23FA9D2983" bold="true" box="[151,734,1202,1229]" fontSize="11" level="1" pageId="0" pageNumber="597" reason="1">
<taxonomicName id="4C38DAAEFF99FF9DFF54FB23FA9D2983" ID-CoL="4CQKT" authority="Emmerich &amp; Nieto, 2009" authorityName="Emmerich &amp; Nieto" authorityYear="2009" box="[151,734,1202,1229]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="species" species="charrua">
<emphasis id="B94C7D3FFF99FF9DFF54FB23FA9D2983" bold="true" box="[151,734,1202,1229]" pageId="0" pageNumber="597">
<emphasis id="B94C7D3FFF99FF9DFF54FB23F9E22982" bold="true" box="[151,417,1203,1229]" italics="true" pageId="0" pageNumber="597">Paracloeodes charrua</emphasis>
Emmerich &amp; Nieto, 2009
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B87A12DFF99FF9DFF5EFB45F95029A3" blockId="0.[151,734,1202,1260]" box="[157,275,1237,1260]" pageId="0" pageNumber="597">
(
<figureCitation id="1303BDA8FF99FF9DFF66FB45F94929A3" box="[165,266,1237,1260]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,257,1980,2001]" captionTargetBox="[151,1415,333,1949]" captionTargetId="figure@1.[151,1436,333,1953]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Adults of Paracloeodes charrua. Emmerich &amp; Nieto, 2009. 1, male imago (dorsal view). 2, male imago (lateral view). 3, forewing. 4 a, hind wing. 4 b, hind wing (enlarged). 5, genitalia. 6, female imago (dorsal view) (SP = small projection)." httpUri="https://zenodo.org/record/221491/files/figure.png" pageId="0" pageNumber="597">Figs. 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C322F2A6FF99FF9DFF54FA8BF95E28AF" pageId="0" pageNumber="597" type="diagnosis">
<paragraph id="8B87A12DFF99FF9DFF54FA8BF9D028D5" blockId="0.[151,1436,1307,1434]" pageId="0" pageNumber="597">
<emphasis id="B94C7D3FFF99FF9DFF54FA8BF946287C" bold="true" box="[151,261,1307,1331]" pageId="0" pageNumber="597">Diagnosis</emphasis>
. The male imagos of
<emphasis id="B94C7D3FFF99FF9DFDC7FA8BFA36287D" box="[516,629,1307,1330]" italics="true" pageId="0" pageNumber="597">
<taxonomicName id="4C38DAAEFF99FF9DFDC7FA8BFA05287D" box="[516,582,1307,1330]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="species" species="cha">P. cha</taxonomicName>
rr ua
</emphasis>
can be distinguished from the other species of genus by the following combination of characters: (1) hind wings present; (2) tibia of foreleg almost twice the length of femur; (3) abdominal terga light brown, submedially with a pair of small red spots; (
<docIssue id="CF5BDEC2FF99FF9DFCECFAF0FB7D2838" box="[815,830,1376,1399]" pageId="0" pageNumber="597">4</docIssue>
) posterior margin of subgenital plate convex, posteriorly with a small projection.
</paragraph>
<paragraph id="8B87A12DFF99FF9DFF54FA58F95E28AF" blockId="0.[151,285,1480,1504]" box="[151,285,1480,1504]" pageId="0" pageNumber="597">
<emphasis id="B94C7D3FFF99FF9DFF54FA58F95E28AF" bold="true" box="[151,285,1480,1504]" pageId="0" pageNumber="597">Male imago</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C322F2A6FF99FF9FFF54F99DFABC2FD8" lastPageId="2" lastPageNumber="599" pageId="0" pageNumber="597" type="description">
<paragraph id="8B87A12DFF99FF9DFF54F99DFA472B08" blockId="0.[151,1437,1549,2026]" pageId="0" pageNumber="597">
<emphasis id="B94C7D3FFF99FF9DFF54F99DF9172B6A" bold="true" box="[151,340,1549,1573]" pageId="0" pageNumber="597">Maximal length.</emphasis>
Body:
<quantity id="4CC00CC8FF99FF9DFE65F99DFA6F2B6B" box="[422,556,1549,1572]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.8" metricValueMin="3.3" pageId="0" pageNumber="597" unit="mm" value="3.55" valueMax="3.8" valueMin="3.3">3.3–3.8 mm</quantity>
; cerci broken; forewing:
<quantity id="4CC00CC8FF99FF9DFC83F99DFB852B6B" box="[832,966,1549,1572]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.2" metricValueMin="3.3" pageId="0" pageNumber="597" unit="mm" value="3.75" valueMax="4.2" valueMin="3.3">3.3–4.2 mm</quantity>
; hind wing:
<quantity id="4CC00CC8FF99FF9DFB93F99DFCA02B6B" box="[1104,1251,1549,1572]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" metricValueMax="8.0" metricValueMin="5.4" pageId="0" pageNumber="597" unit="mm" value="0.67" valueMax="0.8" valueMin="0.54">0.54–0.8 mm</quantity>
; tibia I:
<quantity id="4CC00CC8FF99FF9DFA82F99DFDDB2B6B" box="[1345,1432,1549,1572]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="0" pageNumber="597" unit="mm" value="1.3">1.3 mm</quantity>
; tibia II:
<quantity id="4CC00CC8FF99FF9DFF2EF9A0F9012B08" box="[237,322,1584,1607]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="0" pageNumber="597" unit="mm" value="0.8">0.8 mm</quantity>
; tibia III:
<quantity id="4CC00CC8FF99FF9DFE68F9A0FA432B08" box="[427,512,1584,1607]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="0" pageNumber="597" unit="mm" value="0.7">0.7 mm</quantity>
.
</paragraph>
<paragraph id="8B87A12DFF99FF9DFF04F9C3FBD12BE0" blockId="0.[151,1437,1549,2026]" pageId="0" pageNumber="597">
<emphasis id="B94C7D3FFF99FF9DFF04F9C3F9472B24" bold="true" box="[199,260,1619,1643]" pageId="0" pageNumber="597">Head</emphasis>
(
<figureCitation id="1303BDA8FF99FF9DFED7F9C3F93E2B25" box="[276,381,1619,1642]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,257,1980,2001]" captionTargetBox="[151,1415,333,1949]" captionTargetId="figure@1.[151,1436,333,1953]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Adults of Paracloeodes charrua. Emmerich &amp; Nieto, 2009. 1, male imago (dorsal view). 2, male imago (lateral view). 3, forewing. 4 a, hind wing. 4 b, hind wing (enlarged). 5, genitalia. 6, female imago (dorsal view) (SP = small projection)." httpUri="https://zenodo.org/record/221491/files/figure.png" pageId="0" pageNumber="597">Figs. 1–2</figureCitation>
). Coloration brown; compound eyes blackish, turbinate portion light brown. Antenna with scape and pedicel light brown, flagellum lighter. Dorsal portion of turbinate eyes oval; length 1.8x width; stalk height 1.1x width of dorsal portion; inner margins not parallel, divergent anteriorly.
</paragraph>
<paragraph id="8B87A12DFF99FF9DFF04F92BFD302AEB" blockId="0.[151,1437,1549,2026]" pageId="0" pageNumber="597">
<emphasis id="B94C7D3FFF99FF9DFF04F92BF9582B9C" bold="true" box="[199,283,1723,1747]" pageId="0" pageNumber="597">Thorax</emphasis>
(
<figureCitation id="1303BDA8FF99FF9DFEEDF92BF9D92B9D" box="[302,410,1723,1746]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,257,1980,2001]" captionTargetBox="[151,1415,333,1949]" captionTargetId="figure@1.[151,1436,333,1953]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Adults of Paracloeodes charrua. Emmerich &amp; Nieto, 2009. 1, male imago (dorsal view). 2, male imago (lateral view). 3, forewing. 4 a, hind wing. 4 b, hind wing (enlarged). 5, genitalia. 6, female imago (dorsal view) (SP = small projection)." httpUri="https://zenodo.org/record/221491/files/figure.png" pageId="0" pageNumber="597">Figs. 1–2</figureCitation>
). Pronotum brown; mesonotum brown with medioparapsidal suture darker, sublateroscutum brown, posterior scutal protuberance brown; metanotum brown; prosternum brown. Anteronotal protuberance rounded. Legs yellowish with a blackish line on posterior face. Leg I: tibia 1.9× length of femur; tarsi 1.7× length of femur. Leg II tibia 1.4× length of femur; tarsi 0.5× length of femur. Leg III tibia 1.4× length of femur; tarsi 0.5× length of femur. Forewing (
<figureCitation id="1303BDA8FF99FF9DFECDF8D5F9122A13" box="[270,337,1861,1884]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,257,1980,2001]" captionTargetBox="[151,1415,333,1949]" captionTargetId="figure@1.[151,1436,333,1953]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Adults of Paracloeodes charrua. Emmerich &amp; Nieto, 2009. 1, male imago (dorsal view). 2, male imago (lateral view). 3, forewing. 4 a, hind wing. 4 b, hind wing (enlarged). 5, genitalia. 6, female imago (dorsal view) (SP = small projection)." httpUri="https://zenodo.org/record/221491/files/figure.png" pageId="0" pageNumber="597">Fig. 3</figureCitation>
): membrane hyaline, veins light yellow, stigmatic area with five cross veins, not touching subcostal vein. Marginal intercalary veins paired, except between veins ICu2 and A; length of forewing about 2.2× width.
<collectingCountry id="F32FE1BDFF99FF9DFAA4F8F8FDDF2A30" box="[1383,1436,1896,1919]" name="India" pageId="0" pageNumber="597">Hind</collectingCountry>
wing hyaline with two complete longitudinal veins; costal process quadrangular located on basal third (
<figureCitation id="1303BDA8FF99FF9DFB26F81DFD6C2AEB" box="[1253,1327,1933,1956]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,257,1980,2001]" captionTargetBox="[151,1415,333,1949]" captionTargetId="figure@1.[151,1436,333,1953]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Adults of Paracloeodes charrua. Emmerich &amp; Nieto, 2009. 1, male imago (dorsal view). 2, male imago (lateral view). 3, forewing. 4 a, hind wing. 4 b, hind wing (enlarged). 5, genitalia. 6, female imago (dorsal view) (SP = small projection)." httpUri="https://zenodo.org/record/221491/files/figure.png" pageId="0" pageNumber="597">Figs. 4</figureCitation>
a–4b).
</paragraph>
<paragraph id="8B87A12DFF99FF9CFF04F820FC012C58" blockId="0.[151,1437,1549,2026]" lastBlockId="1.[151,1437,152,279]" lastPageId="1" lastPageNumber="598" pageId="0" pageNumber="597">
<emphasis id="B94C7D3FFF99FF9DFF04F820F9752A87" bold="true" box="[199,310,1968,1992]" pageId="0" pageNumber="597">Abdomen</emphasis>
(
<figureCitation id="1303BDA8FF99FF9DFE85F820F9EC2A88" box="[326,431,1968,1991]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,257,1980,2001]" captionTargetBox="[151,1415,333,1949]" captionTargetId="figure@1.[151,1436,333,1953]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Adults of Paracloeodes charrua. Emmerich &amp; Nieto, 2009. 1, male imago (dorsal view). 2, male imago (lateral view). 3, forewing. 4 a, hind wing. 4 b, hind wing (enlarged). 5, genitalia. 6, female imago (dorsal view) (SP = small projection)." httpUri="https://zenodo.org/record/221491/files/figure.png" pageId="0" pageNumber="597">Figs. 1–2</figureCitation>
). Terga light brown, submedially with a pair of small red spots; segments VII–X darker and segments II–VI lighter. Tracheation black. Sterna yellowish, medially with two pairs of small red spots. Genitalia (
<figureCitation id="1303BDA8FF99FF9DFA96F843FDD72AA5" box="[1365,1428,2003,2026]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,257,1980,2001]" captionTargetBox="[151,1415,333,1949]" captionTargetId="figure@1.[151,1436,333,1953]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Adults of Paracloeodes charrua. Emmerich &amp; Nieto, 2009. 1, male imago (dorsal view). 2, male imago (lateral view). 3, forewing. 4 a, hind wing. 4 b, hind wing (enlarged). 5, genitalia. 6, female imago (dorsal view) (SP = small projection)." httpUri="https://zenodo.org/record/221491/files/figure.png" pageId="0" pageNumber="597">Fig. 5</figureCitation>
) with segments of forceps whitish, basally washed with brown. Forceps segment I cylindrical, without distomedial projection; 0.3× length of segment II; distance between base of forceps 0.4× distance between lateral margins of forceps. Forceps segment II narrow submedially. Forceps segment III elongate, length 2.9× the width; 0.2× length of segment II. Posterior margin of subgenital plate convex, posteriorly with a small projection (
<figureCitation id="1303BDA8FF98FF9CFC30FE90FC772C58" box="[1011,1076,256,279]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,257,1980,2001]" captionTargetBox="[151,1415,333,1949]" captionTargetId="figure@1.[151,1436,333,1953]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Adults of Paracloeodes charrua. Emmerich &amp; Nieto, 2009. 1, male imago (dorsal view). 2, male imago (lateral view). 3, forewing. 4 a, hind wing. 4 b, hind wing (enlarged). 5, genitalia. 6, female imago (dorsal view) (SP = small projection)." httpUri="https://zenodo.org/record/221491/files/figure.png" pageId="1" pageNumber="598">Fig. 5</figureCitation>
).
</paragraph>
<caption id="DF47F1A5FF98FF9CFF54F82CFD1E2ABD" httpUri="https://zenodo.org/record/221491/files/figure.png" pageId="1" pageNumber="598" targetBox="[151,1415,333,1949]" targetPageId="1">
<paragraph id="8B87A12DFF98FF9CFF54F82CFD1E2ABD" blockId="1.[151,1436,1978,2034]" pageId="1" pageNumber="598">
<emphasis id="B94C7D3FFF98FF9CFF54F82CF9772A9E" bold="true" box="[151,308,1980,2001]" pageId="1" pageNumber="598">FIGURES 1–6.</emphasis>
Adults of
<taxonomicName id="4C38DAAEFF98FF9CFE60F82CFA3E2A9E" box="[419,637,1980,2001]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="charrua">
<emphasis id="B94C7D3FFF98FF9CFE60F82CFA3E2A9E" box="[419,637,1980,2001]" italics="true" pageId="1" pageNumber="598">Paracloeodes charrua</emphasis>
</taxonomicName>
. Emmerich &amp; Nieto, 2009. 1, male imago (dorsal view). 2, male imago (lateral view). 3, forewing. 4a, hind wing. 4b, hind wing (enlarged). 5, genitalia. 6, female imago (dorsal view) (SP = small projection).
</paragraph>
</caption>
<paragraph id="8B87A12DFF9BFF9FFF54FF07F9752DE0" blockId="2.[151,310,151,175]" box="[151,310,151,175]" pageId="2" pageNumber="599">
<emphasis id="B94C7D3FFF9BFF9FFF54FF07F9752DE0" bold="true" box="[151,310,151,175]" pageId="2" pageNumber="599">Female imago</emphasis>
</paragraph>
<paragraph id="8B87A12DFF9BFF9FFF54FF4CFA472C58" blockId="2.[151,1437,220,1391]" pageId="2" pageNumber="599">
<emphasis id="B94C7D3FFF9BFF9FFF54FF4CF9102DBB" bold="true" box="[151,339,220,244]" pageId="2" pageNumber="599">Maximal length.</emphasis>
Body: 3.6–4.0 mm; cerci: broken; forewing:
<quantity id="4CC00CC8FF9BFF9FFCFDFF4DFB812DBB" box="[830,962,221,244]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.8" metricValueMin="3.4" pageId="2" pageNumber="599" unit="mm" value="3.5999999999999996" valueMax="3.8" valueMin="3.4">3.4–3.8 mm</quantity>
; hind wing:
<quantity id="4CC00CC8FF9BFF9FFB8AFF4DFCAB2DBB" box="[1097,1256,221,244]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.2" metricValueMin="4.5" pageId="2" pageNumber="599" unit="mm" value="0.485" valueMax="0.52" valueMin="0.45">0.45–0.52 mm</quantity>
; tibia I:
<quantity id="4CC00CC8FF9BFF9FFA80FF4DFDDB2DBB" box="[1347,1432,221,244]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="599" unit="mm" value="0.6">0.6 mm</quantity>
; tibia II:
<quantity id="4CC00CC8FF9BFF9FFF2EFE90F9012C58" box="[237,322,256,279]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="599" unit="mm" value="0.7">0.7 mm</quantity>
; tibia III:
<quantity id="4CC00CC8FF9BFF9FFE68FE90FA432C58" box="[427,512,256,279]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="599" unit="mm" value="0.6">0.6 mm</quantity>
.
</paragraph>
<paragraph id="8B87A12DFF9BFF9FFF04FEB2F8A12C13" blockId="2.[151,1437,220,1391]" pageId="2" pageNumber="599">
<emphasis id="B94C7D3FFF9BFF9FFF04FEB2F9472C75" bold="true" box="[199,260,290,314]" pageId="2" pageNumber="599">Head</emphasis>
(
<figureCitation id="1303BDA8FF9BFF9FFED1FEB2F9172C76" box="[274,340,290,313]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,257,1980,2001]" captionTargetBox="[151,1415,333,1949]" captionTargetId="figure@1.[151,1436,333,1953]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Adults of Paracloeodes charrua. Emmerich &amp; Nieto, 2009. 1, male imago (dorsal view). 2, male imago (lateral view). 3, forewing. 4 a, hind wing. 4 b, hind wing (enlarged). 5, genitalia. 6, female imago (dorsal view) (SP = small projection)." httpUri="https://zenodo.org/record/221491/files/figure.png" pageId="2" pageNumber="599">Fig. 6</figureCitation>
). Coloration light brown; compound eyes blackish. Antenna with scape, pedicel and flagellum light brown.
</paragraph>
<paragraph id="8B87A12DFF9BFF9FFF04FEF7FA552F1E" blockId="2.[151,1437,220,1391]" pageId="2" pageNumber="599">
<emphasis id="B94C7D3FFF9BFF9FFF04FEF7F9582C30" bold="true" box="[199,283,359,383]" pageId="2" pageNumber="599">Thorax</emphasis>
(
<figureCitation id="1303BDA8FF9BFF9FFEEFFEF8F9322C30" box="[300,369,360,383]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,257,1980,2001]" captionTargetBox="[151,1415,333,1949]" captionTargetId="figure@1.[151,1436,333,1953]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Adults of Paracloeodes charrua. Emmerich &amp; Nieto, 2009. 1, male imago (dorsal view). 2, male imago (lateral view). 3, forewing. 4 a, hind wing. 4 b, hind wing (enlarged). 5, genitalia. 6, female imago (dorsal view) (SP = small projection)." httpUri="https://zenodo.org/record/221491/files/figure.png" pageId="2" pageNumber="599">Fig. 6</figureCitation>
). Pronotum light brown; mesonotum brown with medioparapsidal suture darker, sublateroscutum brown, posterior scutal protuberance brown; metanotum brown; prosternum brown. Anteronotal protuberance rounded. Leg I: tibia 1.2× length of femur; tarsi 0.4× length of femur. Leg II tibia 1.4× length of femur; tarsi 0.4× length of femur. Leg III tibia 1.2× length of femur; tarsi 0.4× length of femur. Forewing hyaline,veins light yellow, stigmatic area with five cross veins, two touching subcostal vein. Marginal intercalary veins paired, except between veins MP2 and ICu2; length of forewing about 2.6× width.
<collectingCountry id="F32FE1BDFF9BFF9FFD94FD88FACF2F60" box="[599,652,536,559]" name="India" pageId="2" pageNumber="599">Hind</collectingCountry>
wing hyaline with two complete longitudinal veins; costal process quadrangular located on basal third.
</paragraph>
<paragraph id="8B87A12DFF9BFF9FFF04FDCCFABC2FD8" blockId="2.[151,1437,220,1391]" pageId="2" pageNumber="599">
<emphasis id="B94C7D3FFF9BFF9FFF04FDCCF9752F3B" bold="true" box="[199,310,604,628]" pageId="2" pageNumber="599">Abdomen</emphasis>
(
<figureCitation id="1303BDA8FF9BFF9FFE84FDCDF9CF2F3B" box="[327,396,605,628]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,257,1980,2001]" captionTargetBox="[151,1415,333,1949]" captionTargetId="figure@1.[151,1436,333,1953]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Adults of Paracloeodes charrua. Emmerich &amp; Nieto, 2009. 1, male imago (dorsal view). 2, male imago (lateral view). 3, forewing. 4 a, hind wing. 4 b, hind wing (enlarged). 5, genitalia. 6, female imago (dorsal view) (SP = small projection)." httpUri="https://zenodo.org/record/221491/files/figure.png" pageId="2" pageNumber="599">Fig. 6</figureCitation>
). Terga light brown, submedially with a pair of small reddish spots. Tracheation black. Sterna yellowish, medially with two pairs of small reddish spots.
</paragraph>
</subSubSection>
<subSubSection id="C322F2A6FF9BFF9FFF04FD32FA522E80" pageId="2" pageNumber="599" type="materials_examined">
<paragraph id="8B87A12DFF9BFF9FFF04FD32FA522E80" blockId="2.[151,1437,220,1391]" pageId="2" pageNumber="599">
<emphasis id="B94C7D3FFF9BFF9FFF04FD32F9EA2FF5" bold="true" box="[199,425,674,698]" pageId="2" pageNumber="599">Material examined.</emphasis>
<collectingCountry id="F32FE1BDFF9BFF9FFE70FD32FA552FF6" box="[435,534,674,697]" name="Brazil" pageId="2" pageNumber="599">BRAZIL</collectingCountry>
, Pernambuco State, Amaraji, Rio Amaraji,
<geoCoordinate id="EE0CC7EAFF9BFF9FFC39FD32FC382FF6" box="[1018,1147,674,697]" direction="south" orientation="latitude" pageId="2" pageNumber="599" precision="1" value="-8.363584">8°21'48.9&quot;S</geoCoordinate>
,
<geoCoordinate id="EE0CC7EAFF9BFF9FFB48FD32FD622FF6" box="[1163,1313,674,697]" direction="west" orientation="longitude" pageId="2" pageNumber="599" precision="1" value="-35.480278">35°28'49.0&quot;W</geoCoordinate>
,
<date id="FF8687EDFF9BFF9FFAF3FD32FDDB2FF6" box="[1328,1432,674,697]" pageId="2" pageNumber="599" value="2011-01-20">
20/i/
<quantity id="4CC00CC8FF9BFF9FFAA0FD32FDDB2FF6" box="[1379,1432,674,697]" metricMagnitude="6" metricUnit="m" metricValue="2.01132" pageId="2" pageNumber="599" unit="m" value="2011320.0">2011</quantity>
</date>
, 320m, Lima L.R.C., Nicacio G. cols, three male imagos (light trap) (UFPE); same data as preceding except
<date id="FF8687EDFF9BFF9FFAE1FD55FDDB2F93" box="[1314,1432,709,732]" pageId="2" pageNumber="599" value="2012-04-21">21/iv/2012</date>
, one male imago (CZNC);
<collectingCountry id="F32FE1BDFF9BFF9FFE74FD78FA592FB0" box="[439,538,744,767]" name="Brazil" pageId="2" pageNumber="599">BRAZIL</collectingCountry>
, Pernambuco State, Amaraji, Poço do Carreiro,
<geoCoordinate id="EE0CC7EAFF9BFF9FFBE9FD78FCE82FB0" box="[1066,1195,744,767]" direction="south" orientation="latitude" pageId="2" pageNumber="599" precision="1" value="-8.367206">8°22'1.94&quot;S</geoCoordinate>
,
<geoCoordinate id="EE0CC7EAFF9BFF9FFB7AFD78FD122FB0" box="[1209,1361,744,767]" direction="west" orientation="longitude" pageId="2" pageNumber="599" precision="1" value="-35.469093">35°28'8.73&quot;W</geoCoordinate>
,
<date id="FF8687EDFF9BFF9FFA9DFD78F88C2E6E" pageId="2" pageNumber="599" value="2012-04-28">
28/iv/
<quantity id="4CC00CC8FF9BFF9FFF54FC9AF88C2E6E" box="[151,207,778,801]" metricMagnitude="6" metricUnit="m" metricValue="2.012269" pageId="2" pageNumber="599" unit="m" value="2012269.0">2012</quantity>
</date>
, 269m, Lima L.R.C. coll., one male imago, (light trap) (INPA);
<collectingCountry id="F32FE1BDFF9BFF9FFCBAFC9AFB9F2E6E" box="[889,988,778,801]" name="Brazil" pageId="2" pageNumber="599">BRAZIL</collectingCountry>
, Pernambuco State, Jaqueira, Rio Pirangi,
<geoCoordinate id="EE0CC7EAFF9BFF9FFF54FCBDF95A2E0B" box="[151,281,813,836]" direction="south" orientation="latitude" pageId="2" pageNumber="599" precision="1" value="-8.748055">8°44'53.0&quot;S</geoCoordinate>
,
<geoCoordinate id="EE0CC7EAFF9BFF9FFEEBFCBDF9832E0B" box="[296,448,813,836]" direction="west" orientation="longitude" pageId="2" pageNumber="599" precision="1" value="-35.814194">35°48'51.1&quot;W</geoCoordinate>
,
<date id="FF8687EDFF9BFF9FFE0DFCBDFA7E2E0B" box="[462,573,813,836]" pageId="2" pageNumber="599" value="2012-05-23">
23/v/
<quantity id="4CC00CC8FF9BFF9FFDC5FCBDFA7E2E0B" box="[518,573,813,836]" metricMagnitude="6" metricUnit="m" metricValue="2.012189" pageId="2" pageNumber="599" unit="m" value="2012189.0">2012</quantity>
</date>
, 189m, Lima L.R.C. coll., two male imagos (INPA); same data as preceding except
<date id="FF8687EDFF9BFF9FFF2DFCC0F9282E28" box="[238,363,848,871]" pageId="2" pageNumber="599" value="2012-07-13">13/vii/2012</date>
, two male imagos (reared) (CZNC);
<collectingCountry id="F32FE1BDFF9BFF9FFCF5FCC0FBDA2E28" box="[822,921,848,871]" name="Brazil" pageId="2" pageNumber="599">BRAZIL</collectingCountry>
, Pernambuco State, Ribeirão, Rio Amaraji,
<geoCoordinate id="EE0CC7EAFF9BFF9FFF54FCE2F95B2EC6" box="[151,280,882,905]" direction="south" orientation="latitude" pageId="2" pageNumber="599" precision="1" value="-8.442805">8°26'34.1&quot;S</geoCoordinate>
,
<geoCoordinate id="EE0CC7EAFF9BFF9FFEE9FCE2F9812EC6" box="[298,450,882,905]" direction="west" orientation="longitude" pageId="2" pageNumber="599" precision="1" value="-35.40614">35°24'22.1&quot;W</geoCoordinate>
,
<date id="FF8687EDFF9BFF9FFE17FCE2FA782EC6" box="[468,571,882,905]" pageId="2" pageNumber="599" value="2011-01-21">
21/i/
<quantity id="4CC00CC8FF9BFF9FFDC6FCE2FA782EC6" box="[517,571,882,905]" metricMagnitude="6" metricUnit="m" metricValue="2.0111239999999997" pageId="2" pageNumber="599" unit="m" value="2011124.0">2011</quantity>
</date>
, 124m, Lima LRC, Nicacio G. cols., one female imago (UFPE);
<collectingCountry id="F32FE1BDFF9BFF9FFAF6FCE2FDDB2EC6" box="[1333,1432,882,905]" name="Brazil" pageId="2" pageNumber="599">BRAZIL</collectingCountry>
, Pernambuco State,
<collectingCountry id="F32FE1BDFF9BFF9FFEACFC05F9F72EE3" box="[367,436,917,940]" name="Indonesia" pageId="2" pageNumber="599">Flores</collectingCountry>
, Rio Pajeú,
<geoCoordinate id="EE0CC7EAFF9BFF9FFD86FC05FA842EE3" box="[581,711,917,940]" direction="south" orientation="latitude" pageId="2" pageNumber="599" precision="1" value="-7.8765836">7°52'35.7&quot;S</geoCoordinate>
,
<geoCoordinate id="EE0CC7EAFF9BFF9FFD14FC05FB2D2EE3" box="[727,878,917,940]" direction="west" orientation="longitude" pageId="2" pageNumber="599" precision="1" value="-38.00111">38°00'04.0&quot;W</geoCoordinate>
,
<date id="FF8687EDFF9BFF9FFCBCFC05FBB82EE3" box="[895,1019,917,940]" pageId="2" pageNumber="599" value="2011-07-19">
19/vii/
<quantity id="4CC00CC8FF9BFF9FFC06FC05FBB82EE3" box="[965,1019,917,940]" metricMagnitude="6" metricUnit="m" metricValue="2.011473" pageId="2" pageNumber="599" unit="m" value="2011473.0">2011</quantity>
</date>
, 473m, Lima LRC coll., two female imagos (reared) (1 INPA, 1 UFPE).
</paragraph>
</subSubSection>
<subSubSection id="C322F2A6FF9BFF9FFF04FC4AFC962820" pageId="2" pageNumber="599" type="discussion">
<paragraph id="8B87A12DFF9BFF9FFF04FC4AFC962820" blockId="2.[151,1437,220,1391]" pageId="2" pageNumber="599">
<emphasis id="B94C7D3FFF9BFF9FFF04FC4AF90A2EBD" bold="true" box="[199,329,986,1010]" pageId="2" pageNumber="599">Comments.</emphasis>
The male imago of
<taxonomicName id="4C38DAAEFF9BFF9FFDE6FC4AFAD42EBE" box="[549,663,986,1009]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="charrua">
<emphasis id="B94C7D3FFF9BFF9FFDE6FC4AFAD42EBE" box="[549,663,986,1009]" italics="true" pageId="2" pageNumber="599">P. charrua</emphasis>
</taxonomicName>
can be distinguished from other male imagos described by: the ratio of tibia/femur of foreleg which is almost twice longer in
<taxonomicName id="4C38DAAEFF9BFF9FFD15FC6DFB05295B" box="[726,838,1021,1044]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="charrua">
<emphasis id="B94C7D3FFF9BFF9FFD15FC6DFB05295B" box="[726,838,1021,1044]" italics="true" pageId="2" pageNumber="599">P. charrua</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C38DAAEFF9BFF9FFC5FFC6DFC48295B" box="[924,1035,1021,1044]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B94C7D3FFF9BFF9FFC5FFC6DFC48295B" box="[924,1035,1021,1044]" italics="true" pageId="2" pageNumber="599">P. minutus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C38DAAEFF9BFF9FFBFDFC6DFCA6295B" box="[1086,1253,1021,1044]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="B94C7D3FFF9BFF9FFBFDFC6DFCA6295B" box="[1086,1253,1021,1044]" italics="true" pageId="2" pageNumber="599">P. portoricensis</emphasis>
</taxonomicName>
(about 1.4x); the abdominal color pattern; the presence of hind wings; and one conspicuous characteristic, the presence of a small pointed projection on the posterior margin of the subgenital plate (
<figureCitation id="1303BDA8FF9BFF9FFCD9FBD2FB1E2916" box="[794,861,1090,1113]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,257,1980,2001]" captionTargetBox="[151,1415,333,1949]" captionTargetId="figure@1.[151,1436,333,1953]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Adults of Paracloeodes charrua. Emmerich &amp; Nieto, 2009. 1, male imago (dorsal view). 2, male imago (lateral view). 3, forewing. 4 a, hind wing. 4 b, hind wing (enlarged). 5, genitalia. 6, female imago (dorsal view) (SP = small projection)." httpUri="https://zenodo.org/record/221491/files/figure.png" pageId="2" pageNumber="599">Fig. 5</figureCitation>
). This projection is for the first time reported to the genus, since it is not present in male imagos of
<taxonomicName id="4C38DAAEFF9BFF9FFD48FBF5FABA2933" box="[651,761,1125,1148]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B94C7D3FFF9BFF9FFD48FBF5FABA2933" box="[651,761,1125,1148]" italics="true" pageId="2" pageNumber="599">P. minutus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C38DAAEFF9BFF9FFCE8FBF5FB912933" box="[811,978,1125,1148]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="B94C7D3FFF9BFF9FFCE8FBF5FB912933" box="[811,978,1125,1148]" italics="true" pageId="2" pageNumber="599">P. portoricensis</emphasis>
</taxonomicName>
. Although uncommon, a similar projection has also been reported for male adults of
<taxonomicName id="4C38DAAEFF9BFF9FFD98FB18FCFB29D0" authority="Lugo-Ortiz and McCafferty, 1998" authorityName="Lugo-Ortiz and McCafferty" authorityYear="1998" box="[603,1208,1160,1183]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="minantenna">
<emphasis id="B94C7D3FFF9BFF9FFD98FB18FB0329D0" box="[603,832,1160,1183]" italics="true" pageId="2" pageNumber="599">Rivudiva minantenna</emphasis>
Lugo-Ortiz and McCafferty, 1998
</taxonomicName>
, and because of that, two species described based only on adults were transferred to this genus:
<taxonomicName id="4C38DAAEFF9BFF9FFC18FB3BFD09298E" authority="Traver, 1971" authorityName="Traver" authorityYear="1971" box="[987,1354,1194,1218]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="coveloae">
<emphasis id="B94C7D3FFF9BFF9FFC18FB3BFCE0298D" box="[987,1187,1195,1218]" italics="true" pageId="2" pageNumber="599">Rivudiva coveloae</emphasis>
(Traver, 1971)
</taxonomicName>
and
<taxonomicName id="4C38DAAEFF9BFF9FFA46FB3BFB4F29AB" authority="(Traver, 1943) (Salles &amp; Nascimento, 2011)" authorityName="(Traver, 1943) (Salles &amp; Nascimento" authorityYear="2011" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B94C7D3FFF9BFF9FFA46FB3BF96B29AB" italics="true" pageId="2" pageNumber="599">R. venezuelensis</emphasis>
(Traver, 1943) (Salles &amp; Nascimento, 2011)
</taxonomicName>
. Other similarities occurs between the subgenital plates of
<taxonomicName id="4C38DAAEFF9BFF9FFA4BFB5DF9672848" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="B94C7D3FFF9BFF9FFA4BFB5DF9672848" italics="true" pageId="2" pageNumber="599">P. portoricensis</emphasis>
</taxonomicName>
and the recently described male imago of
<taxonomicName id="4C38DAAEFF9BFF9FFD25FB60F89B2866" authority="Lugo-Ortiz &amp; McCafferty, 1998 (Cruz et al. 2011)" authorityName="Lugo-Ortiz &amp; McCafferty, 1998 (Cruz et al." authorityYear="2011" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="trichobasis">
<emphasis id="B94C7D3FFF9BFF9FFD25FB60FB822848" box="[742,961,1264,1287]" italics="true" pageId="2" pageNumber="599">Rivudiva trichobasis</emphasis>
Lugo-Ortiz &amp; McCafferty, 1998 (Cruz
<emphasis id="B94C7D3FFF9BFF9FFAA4FB60FDD62848" box="[1383,1429,1264,1287]" italics="true" pageId="2" pageNumber="599">et al</emphasis>
. 2011)
</taxonomicName>
: both possess the subgenital plate concave, though much deeper in
<taxonomicName id="4C38DAAEFF9BFF9FFC77FA83FC0A2865" box="[948,1097,1299,1322]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="trichobasis">
<emphasis id="B94C7D3FFF9BFF9FFC77FA83FC0A2865" box="[948,1097,1299,1322]" italics="true" pageId="2" pageNumber="599">R. trichobasis</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C38DAAEFF9BFF9FFB67FA83FD0F2865" box="[1188,1356,1299,1322]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="portoricensis">
<emphasis id="B94C7D3FFF9BFF9FFB67FA83FD0F2865" box="[1188,1356,1299,1322]" italics="true" pageId="2" pageNumber="599">P. portoricensis</emphasis>
</taxonomicName>
. Given the few number of species described based on nymphs and adults, it is difficult to ascertain at this time if these similarities are an example of convergence or an indicative of close relationship between both genera.
</paragraph>
</subSubSection>
</treatment>
</document>
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