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<document id="41AC344D863C4B4E3E7C1581F652FA96" ID-DOI="10.11646/zootaxa.4801.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3898162" ID-ZooBank="14C5CC99-A3DF-4363-82DC-68BDF2FA849D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1592376217238" checkinUser="plazi" docAuthor="Porta, Andrés O., Michalik, Peter, Franchi, Ezequiel &amp; Proud, Daniel N." docDate="2020" docId="0383AA189753A85476CBFF172A6497FA" docLanguage="en" docName="zootaxa.4801.1.6.pdf" docOrigin="Zootaxa 4801 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Procheiridium judsoni Porta, Michalik &amp; Proud 2020, n. sp." docType="treatment" docUuid="66629322-6B08-414B-92C8-2288885860D6" docUuidSource="ZooBank" docVersion="4" lastPageNumber="147" masterDocId="FFBAD2609757A851765CFF8E287A9701" masterDocTitle="The first fossil pycnocheiridiine pseudoscorpion (Pseudoscorpiones: Cheiridiidae Procheiridium gen. nov.) from Cretaceous Burmese amber" masterLastPageNumber="150" masterPageNumber="142" pageNumber="146" updateTime="1700276736462" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="D0EA484F3BFACC09E18D65C550737DBB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2DF81031D798A764D004D7BDE569016E">
<mods:title id="426B2BDF4118B262CCB926EA68770AAC">The first fossil pycnocheiridiine pseudoscorpion (Pseudoscorpiones: Cheiridiidae Procheiridium gen. nov.) from Cretaceous Burmese amber</mods:title>
</mods:titleInfo>
<mods:name id="D8F70FA2C822B026164470E0F34430E4" type="personal">
<mods:role id="43796044A372CFBB90095C6B41EFE983">
<mods:roleTerm id="25DC57421A2BDFD350FE74DFD5A519A7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F8698FC8D5A17C0B894A24658BF918CA">Porta, Andrés O.</mods:namePart>
<mods:affiliation id="DB3B338153789790EB3D7277066CB33C">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” - Av. Ángel Gallardo 470 C 1405 DJR, Buenos Aires, Argentina.</mods:affiliation>
</mods:name>
<mods:name id="03F79C72D12CB12127336DDB72F2E7B0" type="personal">
<mods:role id="65FE27665368ACD9955EF3DAE3588F41">
<mods:roleTerm id="34B420B99CB6C846DB93587266DDDBAF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="86936B6B00715DA138914DC6865F9BB1">Michalik, Peter</mods:namePart>
<mods:affiliation id="AF2CC0DB0006333421634B1998EB81AE">Zoological Museum, University of Greifswald, Loitzer Str. 26, 17489, Greifswald, Mecklenburg-Vorpommern, Germany. michalik @ uni-greifswald. de; https: // orcid. org / 0000 - 0003 - 2459 - 9153</mods:affiliation>
</mods:name>
<mods:name id="DD462D8C89ECE44A08CB9DFDA39C3D27" type="personal">
<mods:role id="29BCBDD682DE893A7BDA42981D0C13AD">
<mods:roleTerm id="8AD5236DFB3B24503A2D9A22F9CBCA2E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0F227B0713A781BB936818FADC9A1BF1">Franchi, Ezequiel</mods:namePart>
<mods:affiliation id="5A373FAE1D3C28C41EE30FFB2260B08E">División Icnología, Museo Argentino de Ciencias Naturales, Av. Ángel Gallardo 470, 1405, CABA, Argentina. ezequielfranchi @ hotmail. com; https: // orcid. org / 0000 - 0003 - 0337 - 3403</mods:affiliation>
</mods:name>
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<mods:roleTerm id="5170FB84C0D73B9D3D70DFCEEE059367">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="807DE495DF5E2C3DD7673BD0BBA3C2AA">Proud, Daniel N.</mods:namePart>
<mods:affiliation id="4273BCE6C90787E1275DCA1378BC3651">Department of Biological Sciences, Moravian College, 1200 Main Street, Bethlehem, Pennsylvania, 18018, USA. proudd @ moravian. edu; https: // orcid. org / 0000 - 0003 - 1842 - 6395</mods:affiliation>
</mods:name>
<mods:typeOfResource id="9D3AE32A7E79262ACCC08FC6F3954E31">text</mods:typeOfResource>
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<mods:title id="490E23E3BA2D99529E4286A10C51EE8A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5BFA2008841AA2409D5AA853DD53786F">
<mods:date id="65EA18C9B5DC6182A39D57A979767FB4">2020</mods:date>
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<mods:number id="8ED49822B1653F9DF04E1AFE334A95E2">2020-06-17</mods:number>
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<mods:number id="639BC959CE2195C927361556F1C0DFCE">4801</mods:number>
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<mods:number id="05FE2495A169C646F262D0A84BAFD646">1</mods:number>
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<mods:classification id="C57F055A4B8CF4BEABB89C4C7A93F6FC">journal article</mods:classification>
<mods:identifier id="166CF373152E06F26711B622822C582A" type="DOI">10.11646/zootaxa.4801.1.6</mods:identifier>
<mods:identifier id="768E43655FD66122576FE155511E94D0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="58DDAFDCC2A7EC0095AAD0F603EDDB51" type="Zenodo-Dep">3898162</mods:identifier>
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<treatment id="0383AA189753A85476CBFF172A6497FA" LSID="urn:lsid:zoobank.org:act:66629322-6B08-414B-92C8-2288885860D6" httpUri="http://treatment.plazi.org/id/0383AA189753A85476CBFF172A6497FA" lastPageId="5" lastPageNumber="147" pageId="4" pageNumber="146">
<subSubSection id="C33048859753A85576CBFF172B3897B5" box="[151,834,153,180]" pageId="4" pageNumber="146" type="nomenclature">
<paragraph id="8B951B0E9753A85576CBFF172B3897B5" blockId="4.[151,834,153,216]" box="[151,834,153,180]" pageId="4" pageNumber="146">
<heading id="D0DDAC629753A85576CBFF172B3897B5" bold="true" box="[151,834,153,180]" fontSize="11" level="1" pageId="4" pageNumber="146" reason="1">
<emphasis id="B95EC71C9753A85576CBFF172B3897B5" bold="true" box="[151,834,153,180]" pageId="4" pageNumber="146">
†
<taxonomicName id="4C2A608D9753A85576FAFF172A8297B5" authority="Porta, Michalik &amp; Proud" authorityName="Porta, Michalik &amp; Proud" authorityYear="2020" box="[166,760,153,180]" class="Arachnida" family="Cheiridiidae" genus="Procheiridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="146" phylum="Arthropoda" rank="species" species="judsoni" status="sp. nov.">
<emphasis id="B95EC71C9753A85576FAFF1729CD97B5" bold="true" box="[166,439,153,180]" italics="true" pageId="4" pageNumber="146">Procheiridium judsoni</emphasis>
Porta, Michalik &amp; Proud
</taxonomicName>
<taxonomicNameLabel id="A26D7A679753A85574A3FF142B3897B5" box="[767,834,154,180]" pageId="4" pageNumber="146" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33048859753A85576CBFF33288297D9" box="[151,248,189,216]" pageId="4" pageNumber="146" type="description">
<paragraph id="8B951B0E9753A85576CBFF33288297D9" blockId="4.[151,834,153,216]" box="[151,248,189,216]" pageId="4" pageNumber="146">
<figureCitation id="1311078B9753A85576CBFF33288297D9" box="[151,248,189,216]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1768,1793]" captionStartId-1="3.[151,250,1767,1792]" captionStartId-2="4.[151,250,1880,1905]" captionStartId-3="5.[151,250,1820,1845]" captionStartId-4="6.[151,250,1862,1887]" captionTargetBox-0="[151,1436,179,1740]" captionTargetBox-1="[151,1436,597,1741]" captionTargetBox-2="[251,1323,481,1844]" captionTargetBox-3="[178,1403,283,1786]" captionTargetBox-4="[151,1436,741,1837]" captionTargetId-0="figure@2.[151,1436,179,1740]" captionTargetId-1="figure@3.[151,1436,597,1741]" captionTargetId-2="figure@4.[236,1352,453,1854]" captionTargetId-3="figure@5.[177,1411,273,1795]" captionTargetId-4="figure@6.[151,1436,741,1837]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1: Procheiridium judsoni n. gen. et n. sp., holotype (MLP 36246): A. Carapace; B. Tergites 1–3 showing details of the surface; C. Right palpus, femur, dorsal view; D. Habitus, dorsal; E. Detail of the lateral setae of the last tergites; F. Right palpus, femur, ventral view, with details of the setae; G. Left chela, ventral view of the hand; H. Same view of the moveable digit; I. Same view of the articulation. Scale bars: A, 0.2 mm; B, 0.05 mm; C, 0.2 mm; D, 0.5 mm; E–G, 0.05 mm; H, 0.1 mm, I, 0.05 mm." captionText-1="FIGURE 2: Procheiridium judsoni n. gen. et n. sp., holotype (MLP 36246): A. Habitus, ventral; B. Right coxae; C. Manducatory process; D. Anal plate and last sternites; E. Left leg IV, junction between femur and patella. Scale bars: A, 0.5 mm; B, 0.2 mm; C, 0.05 mm; D–E, 0.1 mm." captionText-2="FIGURE 3: Procheiridium judsoni n. sp., holotype (MLP 36246): A. Carapax; B. Right chelicera; C. Rallum; D. Right Coxae, E. Right leg I; legs not belonging to the holotype in the same amber sample: F. Right leg III; G. Right leg IV. Scale bars: A–B, 0.2 mm; C, 0.01 mm; D–G, 0.2 mm." captionText-3="FIGURE 4: Procheiridium judsoni n. gen. et n. sp., holotype (MLP 36246): A. Right chela, dorsal view; B. Left chela, ventral view; C. Right palpus, trochanter to patella, ventrolateral view. Scale bars: A–C, 0.20 mm." captionText-4="FIGURE 5: Procheiridium judsoni n. gen. et n. sp., holotype (MLP 36246): A. Right leg I, ventral view; B. Right tarsus III; C. Right tarsus II distal portion; D. Right chelicera and detail of the galea; E. Left chelicera, ventral view, showing rallum (arrow); F. Same, detail of rallum (arrow). Scale bars: A, 0.2 mm; B–D, 0.05 mm; E, 0.1 mm; F, 0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3898164" figureDoi-1="http://doi.org/10.5281/zenodo.3898166" figureDoi-2="http://doi.org/10.5281/zenodo.3898168" figureDoi-3="http://doi.org/10.5281/zenodo.3898170" figureDoi-4="http://doi.org/10.5281/zenodo.3898172" httpUri-0="https://zenodo.org/record/3898164/files/figure.png" httpUri-1="https://zenodo.org/record/3898166/files/figure.png" httpUri-2="https://zenodo.org/record/3898168/files/figure.png" httpUri-3="https://zenodo.org/record/3898170/files/figure.png" httpUri-4="https://zenodo.org/record/3898172/files/figure.png" pageId="4" pageNumber="146">Figs 1–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33048859753A855769BFE882ADF9642" pageId="4" pageNumber="146" type="etymology">
<paragraph id="8B951B0E9753A855769BFE882ADF9642" blockId="4.[151,1436,261,432]" pageId="4" pageNumber="146">
<emphasis id="B95EC71C9753A855769BFE8829329621" bold="true" box="[199,328,262,288]" pageId="4" pageNumber="146">Etymology</emphasis>
. The new speccies is named in honor of Dr. Mark L. I. Judson who contributed to the phylogeny and paleobiology of the superfamily
<taxonomicName id="4C2A608D9753A8557459FEA72ADB9642" authorityName="Hansen" authorityYear="1894" box="[517,673,297,323]" order="Null" pageId="4" pageNumber="146" rank="superFamily" superFamily="Cheiridioidea">Cheiridioidea</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33048859753A854769BFEC02A1B97D6" lastPageId="5" lastPageNumber="147" pageId="4" pageNumber="146" type="materials_examined">
<paragraph id="8B951B0E9753A855769BFEC02D0296AE" blockId="4.[151,1436,261,432]" pageId="4" pageNumber="146">
<emphasis id="B95EC71C9753A855769BFEC029169669" bold="true" box="[199,364,333,360]" pageId="4" pageNumber="146">Type material</emphasis>
.
<typeStatus id="5491A5AC9753A8557725FEC3299B9666" box="[377,481,333,359]" pageId="4" pageNumber="146" type="holotype">Holotype</typeStatus>
(
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36246), apparently adult based on the sclerotization, and probably a male based on the morphology of the galea. Preserved in a single amber sample collected from the Hukawng Valley of
<collectingRegion id="49EED5EC9753A8557317FEFF288796AE" country="Myanmar" name="Kachin" pageId="4" pageNumber="146">Kachin Province</collectingRegion>
, northern
<collectingCountry id="F33D5B9E9753A8557733FE18299A96B1" box="[367,480,406,432]" name="Myanmar" pageId="4" pageNumber="146">Myanmar</collectingCountry>
; lowermost Cenomanian, mid-Cretaceous (
<bibRefCitation id="EFBB66FF9753A8557599FE1B2CC896AE" author="Grimaldi, D. A. &amp; Engel, M. S. &amp; Nascimbene, P. C." box="[965,1202,405,432]" pageId="4" pageNumber="146" pagination="1 - 7" refId="ref3317" refString="Grimaldi, D. A., Engel, M. S. &amp; Nascimbene, P. C. (2002) Fossiliferous Cretaceous amber from Myanmar (Burma): its rediscovery, biotic diversity, and paleontological significance. American Museum Novitates, 3361, 1 - 7. https: // doi. org / 10.1206 / 0003 - 0082 (2002) 361 &lt;0001: fcafmb&gt; 2.0. co; 2" type="journal article" year="2002">
Grimaldi et
<emphasis id="B95EC71C9753A8557210FE1B2C1396AE" box="[1100,1129,405,431]" italics="true" pageId="4" pageNumber="146">al.</emphasis>
, 2002
</bibRefCitation>
,
<bibRefCitation id="EFBB66FF9753A85572E1FE1B2D1696AE" author="Shi, G. &amp; Grimaldi, D. A. &amp; Harlow, G. E. &amp; Wang, J. &amp; Wang, J. &amp; Yang, M. &amp; Lei, W. &amp; Li, Q. &amp; Li, X." box="[1213,1388,405,432]" pageId="4" pageNumber="146" pagination="155 - 163" refId="ref4225" refString="Shi, G., Grimaldi, D. A., Harlow, G. E., Wang, J., Wang, J., Yang, M., Lei, W., Li, Q. &amp; Li, X. (2012) Age constraint on Burmese amber based on UePb dating of zircons. Cretaceous Research, 37, 155 - 163. https: // doi. org / 10.1016 / j. cretres. 2012.03.014" type="journal article" year="2012">
Shi et
<emphasis id="B95EC71C9753A8557358FE1B2D5B96AE" box="[1284,1313,405,431]" italics="true" pageId="4" pageNumber="146">al.</emphasis>
, 2012
</bibRefCitation>
).
</paragraph>
<caption id="DF554B869753A85576CBF8D62A7690B9" ID-DOI="http://doi.org/10.5281/zenodo.3898168" ID-Zenodo-Dep="3898168" httpUri="https://zenodo.org/record/3898168/files/figure.png" pageId="4" pageNumber="146" startId="4.[151,250,1880,1905]" targetBox="[251,1323,481,1844]" targetPageId="4">
<paragraph id="8B951B0E9753A85576CBF8D62A7690B9" blockId="4.[151,1436,1880,1977]" pageId="4" pageNumber="146">
<emphasis id="B95EC71C9753A85576CBF8D6296F9070" bold="true" box="[151,277,1880,1905]" pageId="4" pageNumber="146">FIGURE 3:</emphasis>
<taxonomicName id="4C2A608D9753A8557747F8D629859071" authority="Porta &amp; Michalik &amp; Franchi &amp; Proud, 2020" authorityName="Porta &amp; Michalik &amp; Franchi &amp; Proud" authorityYear="2020" box="[283,511,1880,1904]" class="Arachnida" family="Cheiridiidae" genus="Procheiridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="146" phylum="Arthropoda" rank="species" species="judsoni" status="sp. nov.">
<emphasis id="B95EC71C9753A8557747F8D629859071" box="[283,511,1880,1904]" italics="true" pageId="4" pageNumber="146">Procheiridium judsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D7A679753A8557459F8D72A479071" box="[517,573,1881,1904]" pageId="4" pageNumber="146" rank="species">n. sp.</taxonomicNameLabel>
, holotype (MLP 36246):
<emphasis id="B95EC71C9753A8557561F8D62B359070" bold="true" box="[829,847,1880,1905]" pageId="4" pageNumber="146">A</emphasis>
. Carapax;
<emphasis id="B95EC71C9753A85575E1F8D62BB49071" bold="true" box="[957,974,1880,1904]" pageId="4" pageNumber="146">B</emphasis>
. Right chelicera;
<emphasis id="B95EC71C9753A85572DDF8D62CE99070" bold="true" box="[1153,1171,1880,1905]" pageId="4" pageNumber="146">C</emphasis>
. Rallum;
<emphasis id="B95EC71C9753A85572A4F8D62D709071" bold="true" box="[1272,1290,1880,1904]" pageId="4" pageNumber="146">D</emphasis>
. Right Coxae,
<emphasis id="B95EC71C9753A85576CBF8F228D49095" bold="true" box="[151,174,1916,1940]" pageId="4" pageNumber="146">E.</emphasis>
Right leg I; legs not belonging to the holotype in the same amber sample:
<emphasis id="B95EC71C9753A85575F8F8F22BCD9095" bold="true" box="[932,951,1916,1940]" pageId="4" pageNumber="146">F.</emphasis>
Right leg III;
<emphasis id="B95EC71C9753A8557214F8F22C199094" bold="true" box="[1096,1123,1916,1941]" pageId="4" pageNumber="146">G.</emphasis>
Right leg IV. Scale bars:
<emphasis id="B95EC71C9753A855733AF8F22DEC9095" bold="true" box="[1382,1430,1916,1941]" pageId="4" pageNumber="146">A–B</emphasis>
, 0.2 mm;
<emphasis id="B95EC71C9753A85576AEF82E297E90B8" bold="true" box="[242,260,1952,1977]" pageId="4" pageNumber="146">C</emphasis>
, 0.01 mm;
<emphasis id="B95EC71C9753A8557724F82E29D190B8" bold="true" box="[376,427,1952,1977]" pageId="4" pageNumber="146">D–G</emphasis>
, 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B951B0E9752A854769BFF172BB697B2" blockId="5.[199,972,152,251]" box="[199,972,152,179]" pageId="5" pageNumber="147">
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<emphasis id="B95EC71C9752A854769BFF17291C97B2" bold="true" box="[199,358,152,179]" pageId="5" pageNumber="147">
<typeStatus id="5491A5AC9752A854769BFF17297897B2" box="[199,258,153,179]" pageId="5" pageNumber="147">Type</typeStatus>
locality.
</emphasis>
<collectingMunicipality id="6BF181749752A8547730FF172A5297B2" box="[364,552,153,179]" pageId="5" pageNumber="147">Hukawng Valley</collectingMunicipality>
southwest of
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in
<collectingRegion id="49EED5EC9752A8547527FF172BB697B2" box="[891,972,153,179]" country="Myanmar" name="Kachin" pageId="5" pageNumber="147">Kachin</collectingRegion>
</materialsCitation>
</paragraph>
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State (
<geoCoordinate id="EE1E7DC99752A8547752FF33290B97D6" box="[270,369,189,215]" degrees="26" direction="north" minutes="20" orientation="latitude" pageId="5" pageNumber="147" precision="925" value="26.333334">26º20’N</geoCoordinate>
,
<geoCoordinate id="EE1E7DC99752A8547720FF3329A097D9" box="[380,474,189,216]" degrees="96" direction="east" minutes="36" orientation="longitude" pageId="5" pageNumber="147" precision="925" value="96.6">96º36’E</geoCoordinate>
),
<collectingCountry id="F33D5B9E9752A85477B2FF332A2797D6" box="[494,605,189,215]" name="Myanmar" pageId="5" pageNumber="147">Myanmar</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33048859752A854769BFF6E2A6497FA" box="[199,542,224,251]" pageId="5" pageNumber="147" type="diagnosis">
<paragraph id="8B951B0E9752A854769BFF6E2A6497FA" blockId="5.[199,972,152,251]" box="[199,542,224,251]" pageId="5" pageNumber="147">
<emphasis id="B95EC71C9752A854769BFF6E293897FA" bold="true" box="[199,322,224,251]" pageId="5" pageNumber="147">Diagnosis.</emphasis>
Same as the genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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