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<mods:title id="49C0BC9E40A7B6E9C8BC5B78CD938C9C">Integrative taxonomy of the primitively segmented spider genus Ganthela (Araneae: Mesothelae: Liphistiidae): DNA barcoding gap agrees with morphology</mods:title>
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<mods:namePart id="98E6E4CCB6DF38DA35704D974E429484">Xu, Xin</mods:namePart>
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<heading id="D0DD819AFFBF6B37FC68FB844F0BFBCB" box="[909,1358,1098,1120]" centered="true" fontSize="9" level="2" pageId="14" pageNumber="302" reason="4">
<taxonomicName id="4C2A4D75FFBF6B37FC68FB844EC9FBF5" authority="Xu" authorityName="Xu &amp; Liu &amp; Chen &amp; Li &amp; Kuntner" authorityYear="2015" box="[909,1164,1098,1119]" class="Arachnida" family="Liphistiidae" genus="Ganthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="303" phylum="Arthropoda" rank="species" species="venus" status="sp. nov.">
<emphasis id="B95EEAE4FFBF6B37FC68FB844E27FBF5" bold="true" box="[909,1122,1098,1119]" italics="true" pageId="14" pageNumber="302">GANTHELA VENUS</emphasis>
XU
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<emphasis id="B95EEAE4FFBF6B37FB70FB834EB5FBF4" bold="true" box="[1173,1264,1098,1119]" pageId="14" pageNumber="302">SP. NOV.</emphasis>
(
<figureCitation id="13112A73FFBF6B37FAE4FB844F03FBCB" box="[1281,1350,1098,1120]" captionStart="Figure 9" captionStartId="15.[144,219,547,566]" captionTargetBox="[367,1199,198,517]" captionTargetId="figure-539@15.[367,1199,198,517]" captionTargetPageId="15" captionText="Figure 9. Ganthela venus Xu sp. nov. A, B, female genitalia (XUX-2013-160): A, dorsal view; B, ventral view. RC, receptacular cluster. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5338482" httpUri="https://zenodo.org/record/5338482/files/figure.png" pageId="14" pageNumber="302">FIG. 9</figureCitation>
)
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<typeStatus id="54918854FFBF6B37FCDFFBBE49D8FB2E" box="[826,925,1136,1157]" pageId="14" pageNumber="302" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
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<materialsCitation id="3B423CABFFBF6B37FCDFFB414EE8FAB4" collectingDate="2013-07-11" collectionCode="T" collectorName="F. X. Liu &amp; Liu, X &amp; Zhang. No" country="China" county="Putian City" elevation="459" latitude="25.66" location="Female" longLatPrecision="747" longitude="118.88" municipality="Hanjiang District" pageId="14" pageNumber="302" specimenCount="1" stateProvince="Fujian" typeStatus="holotype">
<location id="8EF5602DFFBF6B37FCDFFB4149CEFB0E" LSID="urn:lsid:plazi:treatment:038387E0FFBF6B36FC68FB8448FDFADB:8EF5602DFFBF6B37FCDFFB4149CEFB0E" box="[826,907,1167,1189]" country="China" county="Putian City" latitude="25.66" longLatPrecision="747" longitude="118.88" municipality="Hanjiang District" name="Female" pageId="14" pageNumber="302" stateProvince="Fujian">Female</location>
(XUX-2013-160),
<location id="8EF5602DFFBF6B37FBB5FB414F53FB0E" LSID="urn:lsid:plazi:treatment:038387E0FFBF6B36FC68FB8448FDFADB:8EF5602DFFBF6B37FBB5FB414F53FB0E" box="[1104,1302,1167,1189]" country="China" county="Putian City" latitude="25.66" longLatPrecision="747" longitude="118.88" municipality="Hanjiang District" name="Mount Wangjiang" pageId="14" pageNumber="302" stateProvince="Fujian">Mount Wangjiang</location>
,
<location id="8EF5602DFFBF6B37FAC4FB4149C9FB6F" LSID="urn:lsid:plazi:treatment:038387E0FFBF6B36FC68FB8448FDFADB:8EF5602DFFBF6B37FAC4FB4149C9FB6F" country="China" county="Putian City" latitude="25.66" longLatPrecision="747" longitude="118.88" municipality="Hanjiang District" name="Huangyang Village" pageId="14" pageNumber="302" stateProvince="Fujian">Huangyang Village</location>
,
<location id="8EF5602DFFBF6B37FC7CFB604E29FB68" LSID="urn:lsid:plazi:treatment:038387E0FFBF6B36FC68FB8448FDFADB:8EF5602DFFBF6B37FC7CFB604E29FB68" box="[921,1132,1198,1220]" country="China" county="Putian City" latitude="25.66" longLatPrecision="747" longitude="118.88" municipality="Hanjiang District" name="Zhuangbian Town" pageId="14" pageNumber="302" stateProvince="Fujian">Zhuangbian Town</location>
,
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,
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,
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,
<collectingCountry id="F33D7666FFBF6B37FBBFFB024EE3FB49" box="[1114,1190,1228,1250]" name="China" pageId="14" pageNumber="302">China</collectingCountry>
,
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,
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,
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<quantity id="4CD29B13FFBF6B37FCDFFB2549C5FAAB" box="[826,896,1259,1281]" metricMagnitude="2" metricUnit="m" metricValue="4.59" pageId="14" pageNumber="302" unit="m" value="459.0">459 m</quantity>
a.s.l.
</elevation>
,
<date id="FF941036FFBF6B37FC2FFB254E1EFAAB" box="[970,1115,1259,1281]" pageId="14" pageNumber="302" value="2013-07-11">
<collectingDate id="EFD0E9DEFFBF6B37FC2FFB254E1EFAAB" box="[970,1115,1259,1281]" pageId="14" pageNumber="302" value="2013-07-11">11 July 2013</collectingDate>
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, collected by
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, X. Xu, and Z.
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.
<collectorName id="26DF5320FFBF6B37FC44FAC74E5CFAB4" box="[929,1049,1289,1311]" pageId="14" pageNumber="302">Zhang. No</collectorName>
male found.
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</paragraph>
<paragraph id="8B9536F6FFBF6B37FCDFFA8449F4FAF4" blockId="14.[826,1376,1354,1406]" box="[826,945,1354,1375]" pageId="14" pageNumber="302">
<emphasis id="B95EEAE4FFBF6B37FCDFFA8449F4FAF4" box="[826,945,1354,1375]" italics="true" pageId="14" pageNumber="302">Etymology</emphasis>
</paragraph>
<paragraph id="8B9536F6FFBF6B37FCDFFAA64F25FAD5" blockId="14.[826,1376,1354,1406]" box="[826,1376,1384,1406]" pageId="14" pageNumber="302">Named after the Greek goddess of love, Venus.</paragraph>
<paragraph id="8B9536F6FFBF6B37FCDFFA6449EFFA14" blockId="14.[826,1442,1450,1716]" box="[826,938,1450,1471]" pageId="14" pageNumber="302">
<emphasis id="B95EEAE4FFBF6B37FCDFFA6449EFFA14" box="[826,938,1450,1471]" italics="true" pageId="14" pageNumber="302">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B9536F6FFBF6B37FCDFFA064E9BF91F" blockId="14.[826,1442,1450,1716]" pageId="14" pageNumber="302">
Females of
<taxonomicName id="4C2A4D75FFBF6B37FC59FA064E59FA76" authority="Xu &amp; Liu &amp; Chen &amp; Li &amp; Kuntner, 2015" authorityName="Xu &amp; Liu &amp; Chen &amp; Li &amp; Kuntner" authorityYear="2015" box="[956,1052,1480,1501]" class="Arachnida" family="Liphistiidae" genus="Ganthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="302" phylum="Arthropoda" rank="species" species="venus" status="sp. nov.">
<emphasis id="B95EEAE4FFBF6B37FC59FA064E59FA76" box="[956,1052,1480,1501]" italics="true" pageId="14" pageNumber="302">G. venus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D579FFFBF6B37FBC6FA064E33FA76" box="[1059,1142,1480,1501]" pageId="14" pageNumber="302" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="4C2A4D75FFBF6B37FCDFFA294E4EFA57" authority="Xu &amp; Liu &amp; Chen &amp; Li &amp; Kuntner, 2015" authorityName="Xu &amp; Liu &amp; Chen &amp; Li &amp; Kuntner" authorityYear="2015" box="[826,1035,1511,1532]" class="Arachnida" family="Liphistiidae" genus="Ganthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="302" phylum="Arthropoda" rank="species" species="wangjiangensis" status="sp. nov.">
<emphasis id="B95EEAE4FFBF6B37FCDFFA294E4EFA57" box="[826,1035,1511,1532]" italics="true" pageId="14" pageNumber="302">G. wangjiangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D579FFFBF6B37FBF7FA294E23FA57" box="[1042,1126,1511,1532]" pageId="14" pageNumber="302" rank="species">sp. nov.</taxonomicNameLabel>
by details in the genitalia, which have separated stalks on the anterior part (
<figureCitation id="13112A73FFBF6B37FCA7F9ED49F7F992" box="[834,946,1571,1593]" captionStart="Figure 9" captionStartId="15.[144,219,547,566]" captionTargetBox="[367,1199,198,517]" captionTargetId="figure-539@15.[367,1199,198,517]" captionTargetPageId="15" captionText="Figure 9. Ganthela venus Xu sp. nov. A, B, female genitalia (XUX-2013-160): A, dorsal view; B, ventral view. RC, receptacular cluster. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5338482" httpUri="https://zenodo.org/record/5338482/files/figure.png" pageId="14" pageNumber="302">Fig. 9A, B</figureCitation>
).
<taxonomicName id="4C2A4D75FFBF6B37FC23F9ED4E37F992" authority="Xu &amp; Liu &amp; Chen &amp; Li &amp; Kuntner, 2015" authorityName="Xu &amp; Liu &amp; Chen &amp; Li &amp; Kuntner" authorityYear="2015" box="[966,1138,1571,1593]" class="Arachnida" family="Liphistiidae" genus="Ganthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="302" phylum="Arthropoda" rank="species" species="venus" status="sp. nov.">
<emphasis id="B95EEAE4FFBF6B37FC23F9ED4E37F992" box="[966,1138,1571,1593]" italics="true" pageId="14" pageNumber="302">Ganthela venus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D579FFFBF6B37FB92F9EA4E8DF992" box="[1143,1224,1572,1593]" pageId="14" pageNumber="302" rank="species">sp. nov.</taxonomicNameLabel>
furthermore differs from all other
<taxonomicName id="4C2A4D75FFBF6B37FC3FF98C4ED0F9F3" box="[986,1173,1602,1624]" class="Arachnida" family="Liphistiidae" genus="Ganthela" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="302" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B95EEAE4FFBF6B37FC3FF98C4E05F9FC" box="[986,1088,1602,1623]" italics="true" pageId="14" pageNumber="302">Ganthela</emphasis>
species
</taxonomicName>
by the following unique nucleotide substitutions in the standard DNA barcode alignment: C (36), A (80),
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(227), C (333), C (341), G (347),
<collectionCode id="ED3BAE33FFBF6B37FC61F95049D1F918" box="[900,916,1694,1715]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="302">T</collectionCode>
(354), C (431), and
<collectionCode id="ED3BAE33FFBF6B37FB66F9504ED6F918" box="[1155,1171,1694,1715]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="302">T</collectionCode>
(539).
</paragraph>
<paragraph id="8B9536F6FFBF6B37FCDFF91149FEF95F" blockId="14.[826,1441,1759,1903]" box="[826,955,1759,1780]" pageId="14" pageNumber="302">
<emphasis id="B95EEAE4FFBF6B37FCDFF91149FEF95F" box="[826,955,1759,1780]" italics="true" pageId="14" pageNumber="302">Description</emphasis>
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<materialsCitation id="3B423CABFFBF6B36FCDFF9334808FC68" collectionCode="OW, ALE, PLE, PME, AME" lastPageId="15" lastPageNumber="303" pageId="14" pageNumber="302" specimenCode="BL 9.80, CL 4.38, CW 4.20, OL 5.6" specimenCount="1" typeStatus="holotype">
Female (
<typeStatus id="54918854FFBF6B37FC46F9334E4FF8B8" box="[931,1034,1789,1811]" pageId="14" pageNumber="302" type="holotype">holotype</typeStatus>
). Carapace and opisthosoma light brown; tergites slightly dark brown; sternum narrow, length more or less twice of width; a few long pointed hairs running over ocular mound in a longitudinal row; chelicerae robust, with promargin of cheliceral groove with ten strong denticles of variable size; legs with strong hairs and spines; opisthosoma with 12 tergites, with tergites 26 larger than others, and with tergite 4 being largest; seven spinnerets. Measurements:
<specimenCode id="DB8C9E8DFFBE6B36FD79FCDA48B6FC82" box="[668,755,788,809]" collectionCode="BL" pageId="15" pageNumber="303">BL 9.80</specimenCode>
,
<specimenCode id="DB8C9E8DFFBE6B36FF75FCFC4AAFFCEC" box="[144,234,818,839]" collectionCode="CL" pageId="15" pageNumber="303">CL 4.38</specimenCode>
,
<specimenCode id="DB8C9E8DFFBE6B36FF12FCFC4B1CFCEC" box="[247,345,818,839]" collectionCode="CW" pageId="15" pageNumber="303">CW 4.20</specimenCode>
,
<specimenCode id="DB8C9E8DFFBE6B36FE80FCFC4BF6FCE3" box="[357,435,818,840]" collectionCode="OL" pageId="15" pageNumber="303">OL 5.6</specimenCode>
, and
<collectionCode id="ED3BAE33FFBE6B36FE16FCFC485BFCEC" box="[499,542,818,839]" pageId="15" pageNumber="303">OW</collectionCode>
5.00;
<collectionCode id="ED3BAE33FFBE6B36FD87FCFC48D3FCEC" box="[610,662,818,839]" pageId="15" pageNumber="303">ALE</collectionCode>
&gt;
<collectionCode id="ED3BAE33FFBE6B36FD55FCFC48A7FCEC" box="[688,738,818,839]" pageId="15" pageNumber="303">PLE</collectionCode>
&gt;
<collectionCode id="ED3BAE33FFBE6B36FF75FC9F4A8CFCCD" box="[144,201,849,870]" pageId="15" pageNumber="303">PME</collectionCode>
&gt;
<collectionCode id="ED3BAE33FFBE6B36FF07FC9F4B5AFCCD" box="[226,287,849,870]" pageId="15" pageNumber="303">AME</collectionCode>
; palp 7.63 (2.50 + 1.31 + 1.65 + 2.17), leg I missing, leg II 8.49 (2.71 + 1.50 + 1.68 + 1.50 + 1.10), leg III 9.34 (2.52 + 1.58 + 1.56 + 2.30 + 1.38), and leg IV 13.70 (3.75 + 1.60 + 2.55 + 3.68 + 2.12)
</materialsCitation>
.
</paragraph>
<caption id="DF55667EFFBE6B36FF75FDED4BEFFDFF" ID-DOI="http://doi.org/10.5281/zenodo.5338482" ID-Zenodo-Dep="5338482" httpUri="https://zenodo.org/record/5338482/files/figure.png" pageId="15" pageNumber="303" startId="15.[144,219,547,566]" targetBox="[367,1199,198,517]" targetPageId="15">
<paragraph id="8B9536F6FFBE6B36FF75FDED4BEFFDFF" blockId="15.[144,1422,546,596]" pageId="15" pageNumber="303">
<emphasis id="B95EEAE4FFBE6B36FF75FDED4AB7FD9D" bold="true" box="[144,242,547,566]" pageId="15" pageNumber="303">Figure 9.</emphasis>
<taxonomicName id="4C2A4D75FFBE6B36FF19FDEC4B89FD9D" authority="Xu" authorityName="Xu &amp; Liu &amp; Chen &amp; Li &amp; Kuntner" authorityYear="2015" box="[252,460,546,566]" class="Arachnida" family="Liphistiidae" genus="Ganthela" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="303" phylum="Arthropoda" rank="species" species="venus" status="sp. nov.">
<emphasis id="B95EEAE4FFBE6B36FF19FDEC4BECFD9D" bold="true" box="[252,425,546,566]" italics="true" pageId="15" pageNumber="303">Ganthela venus</emphasis>
Xu
</taxonomicName>
<emphasis id="B95EEAE4FFBE6B36FE37FDED4861FD9D" bold="true" box="[466,548,547,566]" pageId="15" pageNumber="303">
<taxonomicNameLabel id="A26D579FFFBE6B36FE37FDED4861FD9D" box="[466,548,547,566]" pageId="15" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, B, female genitalia (XUX-2013-160): A, dorsal view; B, ventral view. RC, receptacular cluster. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B9536F6FFBE6B36FF75FC304B0AFBB8" blockId="15.[144,759,1022,1167]" box="[144,335,1022,1043]" pageId="15" pageNumber="303">
<emphasis id="B95EEAE4FFBE6B36FF75FC304B0AFBB8" box="[144,335,1022,1043]" italics="true" pageId="15" pageNumber="303">Female genitalia</emphasis>
</paragraph>
<paragraph id="8B9536F6FFBE6B36FF75FBD34B59FB25" blockId="15.[144,759,1022,1167]" pageId="15" pageNumber="303">
Posterior part of genital area W-shaped (
<figureCitation id="13112A73FFBE6B36FD97FBD348ACFB98" box="[626,745,1053,1075]" captionStart="Figure 9" captionStartId="15.[144,219,547,566]" captionTargetBox="[367,1199,198,517]" captionTargetId="figure-539@15.[367,1199,198,517]" captionTargetPageId="15" captionText="Figure 9. Ganthela venus Xu sp. nov. A, B, female genitalia (XUX-2013-160): A, dorsal view; B, ventral view. RC, receptacular cluster. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5338482" httpUri="https://zenodo.org/record/5338482/files/figure.png" pageId="15" pageNumber="303">Fig. 9A, B</figureCitation>
), with pair of receptacular clusters with short stalks on the anterior part of genital separated from one another (
<figureCitation id="13112A73FFBE6B36FF7DFBB74B4AFB25" box="[152,271,1145,1167]" captionStart="Figure 9" captionStartId="15.[144,219,547,566]" captionTargetBox="[367,1199,198,517]" captionTargetId="figure-539@15.[367,1199,198,517]" captionTargetPageId="15" captionText="Figure 9. Ganthela venus Xu sp. nov. A, B, female genitalia (XUX-2013-160): A, dorsal view; B, ventral view. RC, receptacular cluster. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5338482" httpUri="https://zenodo.org/record/5338482/files/figure.png" pageId="15" pageNumber="303">Fig. 9A, B</figureCitation>
).
</paragraph>
<paragraph id="8B9536F6FFBE6B36FF75FB054B5EFB4B" blockId="15.[144,525,1227,1279]" box="[144,283,1227,1248]" pageId="15" pageNumber="303">
<emphasis id="B95EEAE4FFBE6B36FF75FB054B5EFB4B" box="[144,283,1227,1248]" italics="true" pageId="15" pageNumber="303">Distribution</emphasis>
</paragraph>
<paragraph id="8B9536F6FFBE6B36FF75FB274848FB54" blockId="15.[144,525,1227,1279]" box="[144,525,1257,1279]" pageId="15" pageNumber="303">
<collectingRegion id="49EEF814FFBE6B36FF75FB274A9EFB54" box="[144,219,1257,1279]" country="China" name="Fujian" pageId="15" pageNumber="303">Fujian</collectingRegion>
(Putian) Province,
<collectingCountry id="F33D7666FFBE6B36FE25FB27484CFB54" box="[448,521,1257,1279]" name="China" pageId="15" pageNumber="303">China</collectingCountry>
.
</paragraph>
<paragraph id="8B9536F6FFBE6B36FF75FAF24AB0FAFA" blockId="15.[144,696,1340,1392]" box="[144,245,1340,1361]" pageId="15" pageNumber="303">
<emphasis id="B95EEAE4FFBE6B36FF75FAF24AB0FAFA" box="[144,245,1340,1361]" italics="true" pageId="15" pageNumber="303">Remarks</emphasis>
</paragraph>
<paragraph id="8B9536F6FFBE6B36FF75FA9448FDFADB" blockId="15.[144,696,1340,1392]" box="[144,696,1370,1392]" pageId="15" pageNumber="303">
See comments under
<taxonomicName id="4C2A4D75FFBE6B36FE69FA954818FADB" authority="Xu &amp; Liu &amp; Chen &amp; Li &amp; Kuntner, 2015" authorityName="Xu &amp; Liu &amp; Chen &amp; Li &amp; Kuntner" authorityYear="2015" box="[396,605,1371,1392]" class="Arachnida" family="Liphistiidae" genus="Ganthela" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="303" phylum="Arthropoda" rank="species" species="wangjiangensis" status="sp. nov.">
<emphasis id="B95EEAE4FFBE6B36FE69FA954818FADB" box="[396,605,1371,1392]" italics="true" pageId="15" pageNumber="303">G. wangjiangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D579FFFBE6B36FD81FA9548FDFADB" box="[612,696,1371,1392]" pageId="15" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>