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<document id="DF88A28E66C813D008D604A3C32DDFA2" ID-DOI="10.11646/zootaxa.5397.2.9" ID-ISSN="1175-5326" ID-ZooBank="E1DBCA3A-65EF-4DE3-999D-FBC16BA492E0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1704707603527" checkinUser="plazi" docAuthor="Shear, William A." docDate="2024" docId="038387B4B129FFC9A88C7AADFE48FD54" docLanguage="en" docName="zootaxa.5397.2.9.pdf" docOrigin="Zootaxa 5397 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5397.2.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lineagona Shear 2024, new genus" docType="treatment" docVersion="1" lastPageNumber="287" masterDocId="FFBAFFCCB12BFFCDA81B7F54FF92FFFD" masterDocTitle="Lineagona, a new millipede genus (Diplopoda, Chordeumatida, Cleidogonidae) from Texas, USA, and Nuevo León, México" masterLastPageNumber="290" masterPageNumber="283" pageNumber="285" updateTime="1704707685572" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="FC6F1FFECD5F0C18CDFA647F3BB5AA67">Lineagona, a new millipede genus (Diplopoda, Chordeumatida, Cleidogonidae) from Texas, USA, and Nuevo León, México</mods:title>
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<mods:namePart id="4EF6274102ECBD1ACD20F938969E84AD">Shear, William A.</mods:namePart>
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<mods:date id="9A3FFA2340C6470A6FB6FAF12F7D2466">2024</mods:date>
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<mods:identifier id="1BDC02B3BCCBF4565AD2CBBFCFB98EC7" type="DOI">10.11646/zootaxa.5397.2.9</mods:identifier>
<mods:identifier id="7C7341DE210710484556AF56F61F5A15" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038387B4B129FFC9A88C7AADFE48FD54" LSID="urn:lsid:plazi:treatment:038387B4B129FFC9A88C7AADFE48FD54" httpUri="http://treatment.plazi.org/id/038387B4B129FFC9A88C7AADFE48FD54" lastPageId="4" lastPageNumber="287" pageId="2" pageNumber="285">
<subSubSection id="C3306529B129FFCFA88C7AADFE38F9EE" box="[151,426,1529,1555]" pageId="2" pageNumber="285" type="nomenclature">
<paragraph id="8B9536A2B129FFCFA88C7AADFE38F9EE" blockId="2.[151,426,1529,1555]" box="[151,426,1529,1555]" pageId="2" pageNumber="285">
<heading id="D0DD81CEB129FFCFA88C7AADFE38F9EE" bold="true" box="[151,426,1529,1555]" fontSize="11" level="1" pageId="2" pageNumber="285" reason="1">
<emphasis id="B95EEAB0B129FFCFA88C7AADFE38F9EE" bold="true" box="[151,426,1529,1555]" pageId="2" pageNumber="285">
<taxonomicName id="4C2A4D21B129FFCFA88C7AADFE8BF9EE" authority="Shear, 2024" authorityName="Shear" authorityYear="2024" box="[151,281,1529,1555]" class="Diplopoda" family="Cleidogonidae" genus="Lineagona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95EEAB0B129FFCFA88C7AADFE8BF9EE" bold="true" box="[151,281,1529,1555]" italics="true" pageId="2" pageNumber="285">Lineagona</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26D57CBB129FFCFA9337AADFE38F9EE" box="[296,426,1529,1555]" pageId="2" pageNumber="285" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3306529B129FFCFA88C796CFC04F993" pageId="2" pageNumber="285" type="materials_examined">
<paragraph id="8B9536A2B129FFCFA88C796CFCC2F9B2" blockId="2.[151,918,1591,1646]" box="[151,848,1591,1615]" pageId="2" pageNumber="285">
<emphasis id="B95EEAB0B129FFCFA88C796CFE84F9B2" box="[151,278,1592,1615]" italics="true" pageId="2" pageNumber="285">
<typeStatus id="54918800B129FFCFA88C796CFF54F9B2" box="[151,198,1592,1615]" pageId="2" pageNumber="285">Type</typeStatus>
species
</emphasis>
:
<emphasis id="B95EEAB0B129FFCFA938796CFD8AF9B2" box="[291,536,1591,1615]" italics="true" pageId="2" pageNumber="285">
<taxonomicName id="4C2A4D21B129FFCFA938796CFD86F9B2" authority="Shear, 2024" authorityName="Shear" authorityYear="2024" box="[291,532,1591,1615]" class="Diplopoda" family="Cleidogonidae" genus="Lineagona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="culmenicola" status="sp. nov.">Lineagona culmenicola</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A26D57CBB129FFCFAA047963FDC5F9B2" box="[543,599,1591,1615]" pageId="2" pageNumber="285" rank="species">
<emphasis id="B95EEAB0B129FFCFAA047963FDC3F9B2" bold="true" box="[543,593,1591,1615]" pageId="2" pageNumber="285">n. sp</emphasis>
.
</taxonomicNameLabel>
, by present designation.
</paragraph>
<paragraph id="8B9536A2B129FFCFA88C7902FC04F993" blockId="2.[151,918,1591,1646]" box="[151,918,1622,1646]" pageId="2" pageNumber="285">
<emphasis id="B95EEAB0B129FFCFA88C7902FEADF990" box="[151,319,1622,1646]" italics="true" pageId="2" pageNumber="285">Included species</emphasis>
: In addition to the
<typeStatus id="54918800B129FFCFAA1A7902FDBDF993" box="[513,559,1622,1646]" pageId="2" pageNumber="285">type</typeStatus>
,
<taxonomicName id="4C2A4D21B129FFCFAA227902FC00F993" authority="Loomis, 1963" authorityName="Loomis" authorityYear="1963" box="[569,914,1622,1646]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="chisosi">
<emphasis id="B95EEAB0B129FFCFAA227902FD6EF993" box="[569,764,1622,1646]" italics="true" pageId="2" pageNumber="285">Cleidogona chisosi</emphasis>
<bibRefCitation id="EFBB4B53B129FFCFAB197902FC00F993" author="Loomis, H. F." box="[770,914,1622,1646]" pageId="2" pageNumber="285" pagination="118 - 126" refId="ref4866" refString="Loomis, H. F. (1963) Millipedes from states immediately north and south of the Mexican boundary. Journal of the Kansas Entomological Society, 36, 118 - 126." type="journal article" year="1963">Loomis, 1963</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3306529B129FFCFA88C79C8FDF4F873" pageId="2" pageNumber="285" type="diagnosis">
<paragraph id="8B9536A2B129FFCFA88C79C8FDF4F873" blockId="2.[151,1437,1692,2043]" pageId="2" pageNumber="285">
<emphasis id="B95EEAB0B129FFCFA88C79C8FE86F94B" bold="true" box="[151,276,1692,1718]" pageId="2" pageNumber="285">Diagnosis:</emphasis>
Closely resembling
<taxonomicName id="4C2A4D21B129FFCFA9E679C8FD4AF94B" box="[509,728,1692,1718]" pageId="2" pageNumber="285">
<emphasis id="B95EEAB0B129FFCFA9E679C8FD12F94B" box="[509,640,1692,1718]" italics="true" pageId="2" pageNumber="285">Cleidogona</emphasis>
species
</taxonomicName>
in nonsexual characters. Differing from that genus in the deep division of the gonopod angiocoxite into two branches, sometimes with the posterior or lateral branch subdivided, and in the reduction of the ninth legpair to four or five short articles, the coxae enlarged and lobed, but the femora not swollen and broadly subtriangular as in
<emphasis id="B95EEAB0B129FFCFAA90785CFC87F8DF" box="[651,789,1800,1826]" italics="true" pageId="2" pageNumber="285">
<taxonomicName id="4C2A4D21B129FFCFAA90785CFC83F8DF" box="[651,785,1800,1826]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">Cleidogona</taxonomicName>
.
</emphasis>
Differing from
<taxonomicName id="4C2A4D21B129FFCFABD0785CFBF4F8DF" authorityName="Cope" authorityYear="1869" box="[971,1126,1800,1826]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAB0B129FFCFABD0785CFBF4F8DF" box="[971,1126,1800,1826]" italics="true" pageId="2" pageNumber="285">Pseudotremia</emphasis>
</taxonomicName>
and
<emphasis id="B95EEAB0B129FFCFAC84785CFAAAF8DF" box="[1183,1336,1800,1826]" italics="true" pageId="2" pageNumber="285">
<taxonomicName id="4C2A4D21B129FFCFAC84785CFAA6F8DF" authorityName="Hoffman" authorityYear="1950" box="[1183,1332,1800,1826]" class="Diplopoda" family="Cleidogonidae" genus="Solaenogona" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">Solaenogona</taxonomicName>
,
</emphasis>
which it resembles in the divided angiocoxites, in not having the colpocoxites fused basally, and in lacking median processes between the coxites. Differing from
<taxonomicName id="4C2A4D21B129FFCFAA237804FD07F897" authorityName="Loomis" authorityYear="1964" box="[568,661,1872,1898]" class="Diplopoda" family="Cleidogonidae" genus="Dybasia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAB0B129FFCFAA237804FD07F897" box="[568,661,1872,1898]" italics="true" pageId="2" pageNumber="285">Dybasia</emphasis>
</taxonomicName>
in lacking a sternal process on the gonopods, and from
<taxonomicName id="4C2A4D21B129FFCFAD077804FAECF897" authorityName="Shear" authorityYear="1982" box="[1308,1406,1872,1898]" class="Diplopoda" family="Cleidogonidae" genus="Cabraca" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAB0B129FFCFAD077804FAECF897" box="[1308,1406,1872,1898]" italics="true" pageId="2" pageNumber="285">Cabraca</emphasis>
</taxonomicName>
in having telopodites of the male ninth legs.
</paragraph>
</subSubSection>
<subSubSection id="C3306529B129FFC9A8DC78CCFE48FD54" lastPageId="4" lastPageNumber="287" pageId="2" pageNumber="285" type="etymology">
<paragraph id="8B9536A2B129FFCFA8DC78CCFDBFF807" blockId="2.[151,1437,1692,2043]" pageId="2" pageNumber="285">
<emphasis id="B95EEAB0B129FFCFA8DC78CCFEC0F84F" bold="true" box="[199,338,1944,1970]" pageId="2" pageNumber="285">Etymology:</emphasis>
From Latin,
<emphasis id="B95EEAB0B129FFCFA9FC78CCFDB7F84F" box="[487,549,1944,1970]" italics="true" pageId="2" pageNumber="285">linea,</emphasis>
borderline, combined with the stem –
<emphasis id="B95EEAB0B129FFCFABC978CDFB82F84F" box="[978,1040,1945,1970]" italics="true" pageId="2" pageNumber="285">gona,</emphasis>
used to form generic names in this family. The name refers to the occurrence of the genus on both sides of the border between the
<collectingCountry id="F33D7632B129FFCFACCA78E8FA95F82B" box="[1233,1287,1980,2006]" name="United States of America" pageId="2" pageNumber="285">USA</collectingCountry>
and
<collectingCountry id="F33D7632B129FFCFAD5B78E8FA0BF82B" box="[1344,1433,1980,2006]" name="Mexico" pageId="2" pageNumber="285">México</collectingCountry>
. The gender of the name is feminine.
</paragraph>
<caption id="DF55662AB128FFCEA88C78C3FE18F80A" pageId="3" pageNumber="286" startId="3.[151,264,1943,1967]" targetBox="[162,1425,181,1920]" targetPageId="3" targetType="figure">
<paragraph id="8B9536A2B128FFCEA88C78C3FE18F80A" blockId="3.[151,1437,1943,2040]" pageId="3" pageNumber="286">
<emphasis id="B95EEAB0B128FFCEA88C78C3FEADF852" bold="true" box="[151,319,1943,1967]" pageId="3" pageNumber="286">FIGURES 1–2.</emphasis>
<emphasis id="B95EEAB0B128FFCEA95178CCFDD1F84D" box="[330,579,1944,1968]" italics="true" pageId="3" pageNumber="286">
<taxonomicName id="4C2A4D21B128FFCEA95178CCFDADF84D" authority="Shear, 2024" authorityName="Shear" authorityYear="2024" box="[330,575,1944,1968]" class="Diplopoda" family="Cleidogonidae" genus="Lineagona" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="culmenicola" status="sp. nov.">Lineagona culmenicola</taxonomicName>
,
</emphasis>
<emphasis id="B95EEAB0B128FFCEAA5578CCFD19F84D" bold="true" box="[590,651,1944,1968]" pageId="3" pageNumber="286">
<taxonomicNameLabel id="A26D57CBB128FFCEAA5578CCFD19F84D" box="[590,651,1944,1968]" pageId="3" pageNumber="286" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male.
<emphasis id="B95EEAB0B128FFCEAB5978C3FCDDF852" bold="true" box="[834,847,1943,1967]" pageId="3" pageNumber="286">1</emphasis>
. Gonopods, ventral view.
<emphasis id="B95EEAB0B128FFCEAC7078C3FBEAF852" bold="true" box="[1131,1144,1943,1967]" pageId="3" pageNumber="286">2</emphasis>
. Gonopods, posterior view.
<emphasis id="B95EEAB0B128FFCEA88C78EFFEA2F82E" bold="true" box="[151,304,1979,2003]" pageId="3" pageNumber="286">Abbreviations</emphasis>
:
<emphasis id="B95EEAB0B128FFCEA95878E8FEFAF829" bold="true" box="[323,360,1980,2004]" pageId="3" pageNumber="286">aac</emphasis>
, anterior branch of angiocoxite;
<emphasis id="B95EEAB0B128FFCEAAD578E8FD76F829" bold="true" box="[718,740,1980,2004]" pageId="3" pageNumber="286">cc</emphasis>
, colpocoxite;
<emphasis id="B95EEAB0B128FFCEAB6478E8FC0AF829" bold="true" box="[895,920,1980,2004]" pageId="3" pageNumber="286">cp</emphasis>
, coxal pore;
<emphasis id="B95EEAB0B128FFCEAC3678E8FBD7F829" bold="true" box="[1069,1093,1980,2004]" pageId="3" pageNumber="286">cx</emphasis>
, coxa;
<emphasis id="B95EEAB0B128FFCEAC8278E8FB2DF829" bold="true" box="[1177,1215,1980,2004]" pageId="3" pageNumber="286">pac</emphasis>
, posterior branch of angiocoxite;
<emphasis id="B95EEAB0B128FFCEA90078B4FEB7F805" bold="true" box="[283,293,2016,2040]" pageId="3" pageNumber="286">s</emphasis>
, sternum.
</paragraph>
</caption>
<paragraph id="8B9536A2B12FFFC9A8DC7FC3FE48FD54" blockId="4.[151,1437,151,681]" pageId="4" pageNumber="287">
<emphasis id="B95EEAB0B12FFFC9A8DC7FC3FE98FF4C" bold="true" box="[199,266,151,177]" pageId="4" pageNumber="287">Notes</emphasis>
: Upon initial dissection the male
<typeStatus id="54918800B12FFFC9AA977FC3FD7CFF4C" box="[652,750,151,177]" pageId="4" pageNumber="287" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2A4D21B12FFFC9AB0D7FC3FC2FFF4C" authority="Shear, 2024" authorityName="Shear" authorityYear="2024" box="[790,957,151,177]" class="Diplopoda" family="Cleidogonidae" genus="Lineagona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="culmenicola" status="sp. nov.">
<emphasis id="B95EEAB0B12FFFC9AB0D7FC3FC2FFF4C" box="[790,957,151,177]" italics="true" pageId="4" pageNumber="287">L. culmenicola</emphasis>
</taxonomicName>
<emphasis id="B95EEAB0B12FFFC9ABDD7FC3FB94FF4C" bold="true" box="[966,1030,151,177]" pageId="4" pageNumber="287">
<taxonomicNameLabel id="A26D57CBB12FFFC9ABDD7FC3FB94FF4C" box="[966,1030,151,177]" pageId="4" pageNumber="287" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
struck me as being nearly a perfect intermediate between species of
<taxonomicName id="4C2A4D21B12FFFC9AA127FEFFD1EFF28" box="[521,652,187,213]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAB0B12FFFC9AA127FEFFD1EFF28" box="[521,652,187,213]" italics="true" pageId="4" pageNumber="287">Cleidogona</emphasis>
</taxonomicName>
and
<emphasis id="B95EEAB0B12FFFC9AADE7FEFFCF4FF28" box="[709,870,187,213]" italics="true" pageId="4" pageNumber="287">
<taxonomicName id="4C2A4D21B12FFFC9AADE7FEFFCF0FF28" authorityName="Cope" authorityYear="1869" box="[709,866,187,213]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">Pseudotremia</taxonomicName>
,
</emphasis>
albeit with the soma of
<emphasis id="B95EEAB0B12FFFC9AC677FEFFA95FF28" box="[1148,1287,187,213]" italics="true" pageId="4" pageNumber="287">
<taxonomicName id="4C2A4D21B12FFFC9AC677FEFFA90FF28" box="[1148,1282,187,213]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">Cleidogona</taxonomicName>
.
</emphasis>
Closer study revealed that the colpocoxites were not fused as they originally appeared, and that the gonopods resembled a more elaborate version of those of
<emphasis id="B95EEAB0B12FFFC9AA3A7E57FD07FEE0" box="[545,661,259,285]" italics="true" pageId="4" pageNumber="287">
<taxonomicName id="4C2A4D21B12FFFC9AA3A7E57FD03FEE0" authorityName="Loomis" authorityYear="1963" box="[545,657,259,285]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="chisosi">L. chisosi</taxonomicName>
.
</emphasis>
Nevertheless, the presence of this genus geographically between the Mexican center of diversity for
<taxonomicName id="4C2A4D21B12FFFC9AA1D7E73FD1BFEBC" box="[518,649,295,321]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAB0B12FFFC9AA1D7E73FD1BFEBC" box="[518,649,295,321]" italics="true" pageId="4" pageNumber="287">Cleidogona</emphasis>
</taxonomicName>
and the more northerly distribution of
<taxonomicName id="4C2A4D21B12FFFC9AC567E73FB7AFEBC" authorityName="Cope" authorityYear="1869" box="[1101,1256,295,321]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAB0B12FFFC9AC567E73FB7AFEBC" box="[1101,1256,295,321]" italics="true" pageId="4" pageNumber="287">Pseudotremia</emphasis>
</taxonomicName>
is suggestive. I (
<bibRefCitation id="EFBB4B53B12FFFC9A8847E1FFEB6FE98" author="Shear, W. A." box="[159,292,331,357]" pageId="4" pageNumber="287" pagination="151 - 352" refId="ref5035" refString="Shear, W. A. (1972) Studies in the milliped order Chordeumida (Diplopoda): a revision of the family Cleidogonidae and a reclassification of the order Chordeumida in the New World. Bulletin of the Museum of Comparative Zoology, 144, 151 - 352." type="journal article" year="1972">Shear 1972</bibRefCitation>
,
<bibRefCitation id="EFBB4B53B12FFFC9A92A7E1FFEFEFE98" author="Shear, W. A." box="[305,364,331,357]" pageId="4" pageNumber="287" pagination="1 - 36" refId="ref5275" refString="Shear, W. A. (2003) The milliped family Trichopetalidae, Part 1: Introduction and genera Trigenotyla Causey, Nannopetalum n. gen., and Causeylla n. gen. (Diplopoda, Chordeumatida, Cleidogonoidea) Zootaxa, 321 (1), 1 - 36. https: // doi. org / 10.11646 / zootaxa. 321.1.1" type="journal article" year="2003">2003</bibRefCitation>
,
<bibRefCitation id="EFBB4B53B12FFFC9A9627E1FFE21FE98" author="Shear, W. A." box="[377,435,331,357]" pageId="4" pageNumber="287" pagination="1 - 62" refId="ref5340" refString="Shear, W. A. (2010) The milliped family Trichopetalidae, Part 2: The genera Trichopetalum, Zygonopus and Scoterpes. Zootaxa, 2385 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 2385.1.1" type="journal article" year="2010">2010</bibRefCitation>
) emphasized the close relationship between
<taxonomicName id="4C2A4D21B12FFFC9ABA87E1FFBC4FE98" authorityName="Cook" authorityYear="1896" box="[947,1110,331,357]" class="Diplopoda" family="Cleidogonidae" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="family">Cleidogonidae</taxonomicName>
and
<taxonomicName id="4C2A4D21B12FFFC9AC8B7E1FFAD0FE98" authorityName="Verhoeff" authorityYear="1914" box="[1168,1346,331,357]" class="Diplopoda" family="Trichopetalidae" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="family">Trichopetalidae</taxonomicName>
, and on that basis postulated that
<emphasis id="B95EEAB0B12FFFC9A9AA7E3BFDC0FE74" box="[433,594,367,393]" italics="true" pageId="4" pageNumber="287">
<taxonomicName id="4C2A4D21B12FFFC9A9AA7E3BFDDCFE74" authorityName="Cope" authorityYear="1869" box="[433,590,367,393]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">Pseudotremia</taxonomicName>
,
</emphasis>
which more closely resembles trichopetalids, represented the basic form of North American cleidogonid. This basal stock may have been extinguished in central
<collectingRegion id="49EEF840B12FFFC9AC487EC7FB59FE50" box="[1107,1227,403,429]" country="Mexico" name="Distrito Federal" pageId="4" pageNumber="287">México by</collectingRegion>
the opening of the Cretaceous seaway, after having invaded south to Central America (
<taxonomicName id="4C2A4D21B12FFFC9AB8E7EE3FBB5FE2C" authorityName="Hoffman" authorityYear="1950" box="[917,1063,439,465]" class="Diplopoda" family="Cleidogonidae" genus="Solaenogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAB0B12FFFC9AB8E7EE3FBB5FE2C" box="[917,1063,439,465]" italics="true" pageId="4" pageNumber="287">Solaenogona</emphasis>
</taxonomicName>
) and expanding north into North America (
<taxonomicName id="4C2A4D21B12FFFC9A9107E8FFE37FE08" authorityName="Cope" authorityYear="1869" box="[267,421,475,501]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAB0B12FFFC9A9107E8FFE37FE08" box="[267,421,475,501]" italics="true" pageId="4" pageNumber="287">Pseudotremia</emphasis>
</taxonomicName>
). In this model,
<taxonomicName id="4C2A4D21B12FFFC9AA777E8FFD7DFE08" box="[620,751,475,501]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAB0B12FFFC9AA777E8FFD7DFE08" box="[620,751,475,501]" italics="true" pageId="4" pageNumber="287">Cleidogona</emphasis>
</taxonomicName>
originated in the Paleogene in Central
<collectingRegion id="49EEF840B12FFFC9ACA07E8FFA83FE08" box="[1211,1297,475,501]" country="Mexico" name="Mexico" pageId="4" pageNumber="287">México</collectingRegion>
and rapidly spread both north and south (
<bibRefCitation id="EFBB4B53B12FFFC9A9C57EABFDF1FDE4" author="Shear, W. A." box="[478,611,511,537]" pageId="4" pageNumber="287" pagination="151 - 352" refId="ref5035" refString="Shear, W. A. (1972) Studies in the milliped order Chordeumida (Diplopoda): a revision of the family Cleidogonidae and a reclassification of the order Chordeumida in the New World. Bulletin of the Museum of Comparative Zoology, 144, 151 - 352." type="journal article" year="1972">Shear 1972</bibRefCitation>
). Now I would add that
<taxonomicName id="4C2A4D21B12FFFC9AB697EABFC78FDE5" authority="Shear, 2024" authorityName="Shear" authorityYear="2024" box="[882,1002,511,536]" class="Diplopoda" family="Cleidogonidae" genus="Lineagona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95EEAB0B12FFFC9AB697EABFC78FDE5" box="[882,1002,511,536]" italics="true" pageId="4" pageNumber="287">Lineagona</emphasis>
</taxonomicName>
<emphasis id="B95EEAB0B12FFFC9ABEA7EABFBC9FDE4" bold="true" box="[1009,1115,511,537]" pageId="4" pageNumber="287">
<taxonomicNameLabel id="A26D57CBB12FFFC9ABEA7EABFBC9FDE4" box="[1009,1115,511,537]" pageId="4" pageNumber="287" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
may represent a relict of the pathway from a
<taxonomicName id="4C2A4D21B12FFFC9A9547D77FE7BFDC0" authorityName="Cope" authorityYear="1869" box="[335,489,547,573]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAB0B12FFFC9A9547D77FE7BFDC0" box="[335,489,547,573]" italics="true" pageId="4" pageNumber="287">Pseudotremia</emphasis>
</taxonomicName>
-like ancestor to
<emphasis id="B95EEAB0B12FFFC9AABC7D77FCA2FDC0" box="[679,816,547,573]" italics="true" pageId="4" pageNumber="287">
<taxonomicName id="4C2A4D21B12FFFC9AABC7D77FCBEFDC0" box="[679,812,547,573]" class="Diplopoda" family="Cleidogonidae" genus="Cleidogona" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="287" phylum="Arthropoda" rank="genus">Cleidogona</taxonomicName>
,
</emphasis>
or perhaps a relict of their common ancestral stock. At some point in cleidogonid studies, it is hoped that mtDNA data amenable to the application of a molecular clock might be obtained. Along with information from nuclear genes, both a phylogeny and time line for evolution of the family might be constructed.
</paragraph>
</subSubSection>
</treatment>
</document>
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