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<document id="804F243D1317A243B3A295FE787FA4F6" ID-CLB-Dataset="7049" ID-DOI="10.1080/00222933.2012.763060" ID-GBIF-Dataset="eb5297a4-9107-4eef-8c81-109ee8948145" ID-ISSN="1464-5262" ID-Zenodo-Dep="4631916" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616524360025" checkinUser="carolina" docAuthor="Heyer, Jacob den, Ueckermann, Edward A. &amp; Khanjani, Mohammad" docDate="2013" docId="038387B2CC0AFFA7FE10C6E81120BB3F" docLanguage="en" docName="JNATHIST.47.31-32.2049-2070.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 47 (31 - 32)" docSource="http://dx.doi.org/10.1080/00222933.2012.763060" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Pulaeus razanensis Den Heyer 2013, sp. nov." docType="treatment" docUuid="306AF22C-8B27-47A0-A02D-6DB41E94AE17" docUuidSource="ZooBank" docVersion="8" lastPageNumber="2061" masterDocId="FFBAFFCACC06FFA9FFCEC64F150DB95C" masterDocTitle="Iranian Cunaxidae (Acari: Prostigmata: Bdelloidea). Part III. Subfamily Cunaxoidinae" masterLastPageNumber="2070" masterPageNumber="2049" pageNumber="2059" updateTime="1698934678465" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4F131EFA0BC72E8CFA41C4DFC9F98B16">Iranian Cunaxidae (Acari: Prostigmata: Bdelloidea). Part III. Subfamily Cunaxoidinae</mods:title>
</mods:titleInfo>
<mods:name id="7991895E82B9BEF3CBF13CDB6639B283" type="personal">
<mods:role id="98EF51E0FC517F0EF39B0DB7C3EDC884">
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<mods:namePart id="A937A3AEA37A63A4C7E3FF76834B81BA">Heyer, Jacob den</mods:namePart>
<mods:affiliation id="926E6165C307367519E2CFD5229DB4A9">Department of Zoology and Entomology, University of the Free State, Bloemfontein, South Africa;</mods:affiliation>
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<mods:namePart id="75EDB21B4DD2CFCC63525E15789E3C1E">Ueckermann, Edward A.</mods:namePart>
<mods:affiliation id="3A6D609AB7EC11053D1FD93763CC5D87">ARC-Plant Protection Research Institute, Pretoria, South Africa; &amp; School of Environmental Sciences and Development, North-West University, Potchefstroom, South Africa;</mods:affiliation>
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<mods:namePart id="1C2116A8A8C1BC8F90DCE261A9509DEB">Khanjani, Mohammad</mods:namePart>
<mods:affiliation id="9E96FB5D0D4963ACF24DB6E7E2759AB6">Department of Plant Protection, College of Agriculture, Bu Ali-Sina University, Hamadan, Iran</mods:affiliation>
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<mods:title id="8CE3F80C9555568A8CE06597F93A3647">Journal of Natural History</mods:title>
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<mods:part id="C4B14857AFE6362B34DF6BD1441F5113">
<mods:date id="2CE21B7616191756FD03E2AA5B720094">2013</mods:date>
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<mods:title id="3181D99E672BB0D77BFEF8147FFA6DAD">J. Nat. Hist.</mods:title>
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<mods:number id="01CADF74F7ECE7502D312A40DC2A75BF">2013-05-14</mods:number>
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<mods:detail id="E8FEAB746FF85549908BBE8DCCA7A095" type="volume">
<mods:number id="81474DFE18EE50A1F8FE155AF58F2DF7">47</mods:number>
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<mods:number id="5E5C60F97C7DEC8C8831A3DFB20BD4A2">31 - 32</mods:number>
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<mods:start id="E2D23D9A26789BBC30E34B93388A644F">2049</mods:start>
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<mods:identifier id="6609ADB2676076C1136A33A4307AB538" type="DOI">10.1080/00222933.2012.763060</mods:identifier>
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<treatment id="038387B2CC0AFFA7FE10C6E81120BB3F" ID-DOI="http://doi.org/10.5281/zenodo.4742838" ID-GBIF-Taxon="181882103" ID-Zenodo-Dep="4742838" LSID="urn:lsid:zoobank.org:act:306AF22C-8B27-47A0-A02D-6DB41E94AE17" httpUri="http://treatment.plazi.org/id/038387B2CC0AFFA7FE10C6E81120BB3F" lastPageId="14" lastPageNumber="2061" pageId="12" pageNumber="2059">
<subSubSection id="C330652FCC0AFFA5FE10C6E81684B9E3" box="[478,905,167,192]" pageId="12" pageNumber="2059" type="nomenclature">
<paragraph id="8B9536A4CC0AFFA5FE10C6E81684B9E3" blockId="12.[478,905,167,223]" box="[478,905,167,192]" pageId="12" pageNumber="2059">
<heading id="D0DD81C8CC0AFFA5FE10C6E81684B9E3" box="[478,905,167,192]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="2059" reason="2">
<taxonomicName id="4C2A4D27CC0AFFA5FE10C6E81623B99C" authority="Den Heyer" authorityName="Den Heyer" authorityYear="2013" box="[478,814,167,192]" class="Arachnida" family="Cunaxidae" genus="Pulaeus" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="2059" phylum="Arthropoda" rank="species" species="razanensis" status="sp. nov.">
<emphasis id="B95EEAB6CC0AFFA5FE10C6E817A1B99C" bold="true" box="[478,684,167,192]" italics="true" pageId="12" pageNumber="2059">Pulaeus razanensis</emphasis>
Den Heyer
</taxonomicName>
<taxonomicNameLabel id="A26D57CDCC0AFFA5FCFBC6E71684B9E3" box="[821,905,168,191]" pageId="12" pageNumber="2059" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330652FCC0AFFA5FD9DC688161EB983" box="[595,787,199,223]" pageId="12" pageNumber="2059" type="description">
<paragraph id="8B9536A4CC0AFFA5FD9DC688161EB983" blockId="12.[478,905,167,223]" box="[595,787,199,223]" pageId="12" pageNumber="2059">
(
<figureCitation id="13112A21CC0AFFA5FD95C68817D1B983" box="[603,732,199,223]" captionStart="Figure 5" captionStartId="11.[143,210,1512,1534]" captionTargetBox="[146,1102,162,1477]" captionTargetId="figure-5@11.[145,1102,162,1478]" captionTargetPageId="11" captionText="Figure 5. Pulaeus krama (Chaudhri, 1977), male. (A) Leg II; (B) leg III; (C) leg IV. Pulaeus razanensis Den Heyer sp. nov. female. (D) Dorsum; (E) sensilla vi; (F) sensilla sce; (G) venter." figureDoi="http://doi.org/10.5281/zenodo.4631929" httpUri="https://zenodo.org/record/4631929/files/figure.png" pageId="12" pageNumber="2059">Figures 5D</figureCitation>
,
<figureCitation id="13112A21CC0AFFA5FD29C6881606B983" box="[743,779,199,223]" captionStart="Figure 6" captionStartId="13.[143,210,1577,1599]" captionTargetBox="[145,1102,169,1544]" captionTargetId="figure-5@13.[145,1102,169,1544]" captionTargetPageId="13" captionText="Figure 6. Pulaeus razanensis Den Heyer sp. nov. female. (A) Hypognathum; (B) palp; (C) chelicera; (D) leg I; (E) leg II; (F) leg III; (G) leg IV." figureDoi="http://doi.org/10.5281/zenodo.4631931" httpUri="https://zenodo.org/record/4631931/files/figure.png" pageId="12" pageNumber="2059">6G</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C330652FCC0AFFA5FF1DC76B144BBB5F" pageId="12" pageNumber="2059" type="materials_examined">
<paragraph id="8B9536A4CC0AFFA5FF1DC76B1487B860" blockId="12.[211,1171,292,515]" box="[211,394,292,316]" pageId="12" pageNumber="2059">
<heading id="D0DD81C8CC0AFFA5FF1DC76B1487B860" box="[211,394,292,316]" fontSize="10" level="3" pageId="12" pageNumber="2059" reason="5">
<emphasis id="B95EEAB6CC0AFFA5FF1DC76B1487B860" box="[211,394,292,316]" italics="true" pageId="12" pageNumber="2059">Material studied</emphasis>
</heading>
</paragraph>
<paragraph id="8B9536A4CC0AFFA5FF1DC704144BBB5F" blockId="12.[211,1171,292,515]" pageId="12" pageNumber="2059">
<materialsCitation id="3B423CF9CC0AFFA5FF1DC704144BBB5F" ID-GBIF-Occurrence="3107838301" collectingDate="1998-08-27" collectingDateMax="2006-06-30" collectingDateMin="1998-08-27" country="Iran" location="Hamadan" pageId="12" pageNumber="2059" specimenCount="3" specimenCount-female="3" stateProvince="Hamadan" typeStatus="holotype">
<typeStatus id="54918806CC0AFFA5FF1DC7041433B83F" box="[211,318,331,355]" pageId="12" pageNumber="2059" type="holotype">Holotype</typeStatus>
female, collected from underneath
<taxonomicName id="4C2A4D27CC0AFFA5FCCBC7031454B8DF" authority="Fisch., Razan" authorityName="Fisch., Razan" class="Magnoliopsida" family="Fabaceae" genus="Alhagi" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="2059" phylum="Tracheophyta" rank="species" species="camelorum">
<emphasis id="B95EEAB6CC0AFFA5FCCBC70316D7B838" box="[773,986,332,356]" italics="true" pageId="12" pageNumber="2059">Alhagi camelorum</emphasis>
Fisch., Razan (
<collectingRegion id="49EEF846CC0AFFA5FF15C724145DB8DF" box="[219,336,363,387]" country="Iran" name="Hamadan" pageId="12" pageNumber="2059">Hamadan</collectingRegion>
)
</taxonomicName>
,
<collectingCountry id="F33D7634CC0AFFA5FEAAC723149AB8DF" box="[356,407,364,387]" name="Iran" pageId="12" pageNumber="2059">Iran</collectingCountry>
,
<date id="FF941064CC0AFFA5FE6DC7241759B8DF" box="[419,596,363,387]" pageId="12" pageNumber="2059" value="1998-08-27">
<collectingDate id="EFD0E98CCC0AFFA5FE6DC7241759B8DF" box="[419,596,363,387]" pageId="12" pageNumber="2059" value="1998-08-27">27 August 1998</collectingDate>
</date>
,
<collectorName id="26DF5372CC0AFFA5FDAEC72317F3B8DF" box="[608,766,363,387]" pageId="12" pageNumber="2059">M. Khanjani.</collectorName>
;
<specimenCount id="9D2CFD2DCC0AFFA5FCC7C72416D9B8DF" box="[777,980,363,388]" pageId="12" pageNumber="2059" type="female" typeStatus="paratype">1 paratype female</specimenCount>
,
<taxonomicName id="4C2A4D27CC0AFFA5FC11C723119EB8DF" box="[991,1171,364,388]" class="Magnoliopsida" family="Fabaceae" genus="Medicago" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="2059" phylum="Tracheophyta" rank="species" species="sativa">
<emphasis id="B95EEAB6CC0AFFA5FC11C723119EB8DF" box="[991,1171,364,388]" italics="true" pageId="12" pageNumber="2059">Medicago sativa</emphasis>
</taxonomicName>
L, Shabestar (
<collectingRegion id="49EEF846CC0AFFA5FEBDC7C41722B8FF" box="[371,559,395,419]" country="Iran" name="Azarbayjan-e Sharqi" pageId="12" pageNumber="2059">East Azerbaijan</collectingRegion>
),
<collectingCountry id="F33D7634CC0AFFA5FD8DC7C3177BB8FF" box="[579,630,396,419]" name="Iran" pageId="12" pageNumber="2059">Iran</collectingCountry>
,
<date id="FF941064CC0AFFA5FD4DC7C41616B8FF" box="[643,795,395,419]" pageId="12" pageNumber="2059" value="2006-06-30">
<collectingDate id="EFD0E98CCC0AFFA5FD4DC7C41616B8FF" box="[643,795,395,419]" pageId="12" pageNumber="2059" value="2006-06-30">30 June 2006</collectingDate>
</date>
.
<collectorName id="26DF5372CC0AFFA5FCE8C7C316BBB8FF" box="[806,950,395,419]" pageId="12" pageNumber="2059">P. Lotfollahi</collectorName>
;
<specimenCount id="9D2CFD2DCC0AFFA5FC0CC7C41182B8FF" box="[962,1167,395,420]" pageId="12" pageNumber="2059" type="female" typeStatus="paratype">1 paratype female</specimenCount>
, soil, Najafābād (
<collectingRegion id="49EEF846CC0AFFA5FE54C7E414F1B89F" box="[410,508,427,451]" country="Iran" name="Esfahan" pageId="12" pageNumber="2059">Esfahan</collectingRegion>
),
<collectingCountry id="F33D7634CC0AFFA5FDDAC7E3174AB89F" box="[532,583,428,451]" name="Iran" pageId="12" pageNumber="2059">Iran</collectingCountry>
,
<date id="FF941064CC0AFFA5FD96C7E41603B89F" box="[600,782,427,451]" pageId="12" pageNumber="2059" value="2002-03-30">
<collectingDate id="EFD0E98CCC0AFFA5FD96C7E41603B89F" box="[600,782,427,451]" pageId="12" pageNumber="2059" value="2002-03-30">30 March 2002</collectingDate>
</date>
,
<collectorName id="26DF5372CC0AFFA5FCD1C7E316A7B89F" box="[799,938,427,451]" pageId="12" pageNumber="2059">M. Jalaeian</collectorName>
;
<specimenCount id="9D2CFD2DCC0AFFA5FC75C7E41182B89F" box="[955,1167,427,452]" pageId="12" pageNumber="2059" type="female" typeStatus="paratype">1 paratype female</specimenCount>
,
<taxonomicName id="4C2A4D27CC0AFFA5FF1DC7841481B8BF" box="[211,396,459,483]" class="Magnoliopsida" family="Fabaceae" genus="Medicago" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="2059" phylum="Tracheophyta" rank="species" species="sativa">
<emphasis id="B95EEAB6CC0AFFA5FF1DC7841481B8BF" box="[211,396,459,483]" italics="true" pageId="12" pageNumber="2059">Medicago sativa</emphasis>
</taxonomicName>
L (Alfalfa), Sonqor (
<collectingRegion id="49EEF846CC0AFFA5FD41C784162BB8BF" box="[655,806,459,483]" country="Iran" name="Kermanshah" pageId="12" pageNumber="2059">Kermānshāh</collectingRegion>
),
<collectingCountry id="F33D7634CC0AFFA5FCF0C783167CB8BF" box="[830,881,460,483]" name="Iran" pageId="12" pageNumber="2059">Iran</collectingCountry>
,
<date id="FF941064CC0AFFA5FC4EC784116FB8BF" box="[896,1122,459,483]" pageId="12" pageNumber="2059" value="2003-09-12">
<collectingDate id="EFD0E98CCC0AFFA5FC4EC784116FB8BF" box="[896,1122,459,483]" pageId="12" pageNumber="2059" value="2003-09-12">12 September 2003</collectingDate>
</date>
,
<collectorName id="26DF5372CC0AFFA5FBBCC783144BBB5F" pageId="12" pageNumber="2059">M. Khanjani.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C330652FCC0AFFA7FF1DC4071498B882" lastPageId="14" lastPageNumber="2061" pageId="12" pageNumber="2059" type="diagnosis">
<paragraph id="8B9536A4CC0AFFA5FF1DC4071432BB03" blockId="12.[211,1172,584,775]" box="[211,319,584,607]" pageId="12" pageNumber="2059">
<heading id="D0DD81C8CC0AFFA5FF1DC4071432BB03" box="[211,319,584,607]" fontSize="10" level="3" pageId="12" pageNumber="2059" reason="5">
<emphasis id="B95EEAB6CC0AFFA5FF1DC4071432BB03" box="[211,319,584,607]" italics="true" pageId="12" pageNumber="2059">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9536A4CC0AFFA5FF1DC42017EBBA5B" blockId="12.[211,1172,584,775]" pageId="12" pageNumber="2059">
This species very strongly resembles
<taxonomicName id="4C2A4D27CC0AFFA5FDA4C43F16CFBBDB" authority="Den Heyer, 1979" authorityName="Den Heyer" authorityYear="1979" box="[618,962,623,648]" class="Arachnida" family="Cunaxidae" genus="Pulaeus" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="2059" phylum="Arthropoda" rank="species" species="glebulentus">
<emphasis id="B95EEAB6CC0AFFA5FDA4C43F160DBBD4" box="[618,768,624,648]" italics="true" pageId="12" pageNumber="2059">P. glebulentus</emphasis>
<bibRefCitation id="EFBB4B55CC0AFFA5FCC9C43F16CFBBDB" author="Den Heyer J" box="[775,962,623,648]" pageId="12" pageNumber="2059" pagination="18 - 31" refId="ref7710" refString="Den Heyer J. 1979. Pulaeus, a new cunaxid genus (Prostigmata: Acari). Acarologia. 21; 1: 18 - 31." type="journal article" year="1979">Den Heyer, 1979</bibRefCitation>
</taxonomicName>
by also possessing a dorsal shield with transverse lobes. It differs, however, in the following aspects: ventral hypognathal ornamention of papillae and striae; genua I 3 asl (on one leg and 2 asl on the other leg),(1 asl,1 sts), 4 sts: genua II 2 asl, 5 sts; tarsus I 4 asl (on left leg) {3asl (on right leg)}, 1 famulus pit,1 dtsl, 2 tsl, 22 sts.
</paragraph>
<paragraph id="8B9536A4CC0AFFA5FF1DC5041745BA38" blockId="12.[211,1172,843,1003]" box="[211,584,843,868]" pageId="12" pageNumber="2059">
<heading id="D0DD81C8CC0AFFA5FF1DC5041745BA38" box="[211,584,843,868]" fontSize="10" level="3" pageId="12" pageNumber="2059" reason="5">
<emphasis id="B95EEAB6CC0AFFA5FF1DC5041706BA3F" box="[211,523,843,867]" italics="true" pageId="12" pageNumber="2059">
Female (
<figureCitation id="13112A21CC0AFFA5FEFFC50314BCBA3F" box="[305,433,844,867]" captionStart="Figure 5" captionStartId="11.[143,210,1512,1534]" captionTargetBox="[146,1102,162,1477]" captionTargetId="figure-5@11.[145,1102,162,1478]" captionTargetPageId="11" captionText="Figure 5. Pulaeus krama (Chaudhri, 1977), male. (A) Leg II; (B) leg III; (C) leg IV. Pulaeus razanensis Den Heyer sp. nov. female. (D) Dorsum; (E) sensilla vi; (F) sensilla sce; (G) venter." figureDoi="http://doi.org/10.5281/zenodo.4631929" httpUri="https://zenodo.org/record/4631929/files/figure.png" pageId="12" pageNumber="2059">Figures 5D</figureCitation>
,
<figureCitation id="13112A21CC0AFFA5FE72C50414EDBA3F" box="[444,480,843,867]" captionStart="Figure 6" captionStartId="13.[143,210,1577,1599]" captionTargetBox="[145,1102,169,1544]" captionTargetId="figure-5@13.[145,1102,169,1544]" captionTargetPageId="13" captionText="Figure 6. Pulaeus razanensis Den Heyer sp. nov. female. (A) Hypognathum; (B) palp; (C) chelicera; (D) leg I; (E) leg II; (F) leg III; (G) leg IV." figureDoi="http://doi.org/10.5281/zenodo.4631931" httpUri="https://zenodo.org/record/4631931/files/figure.png" pageId="12" pageNumber="2059">6G</figureCitation>
) (n
</emphasis>
=
<emphasis id="B95EEAB6CC0AFFA5FDE3C5031745BA38" box="[557,584,844,868]" italics="true" pageId="12" pageNumber="2059">4)</emphasis>
</heading>
</paragraph>
<paragraph id="8B9536A4CC0AFFA5FF1DC53B164CBAB7" blockId="12.[211,1172,843,1003]" pageId="12" pageNumber="2059">
<emphasis id="B95EEAB6CC0AFFA5FF1DC53B1704BAD7" box="[211,521,883,907]" italics="true" pageId="12" pageNumber="2059">Dimensions and description</emphasis>
. Idiosoma: length 323 (319–327); width 204 (193–212); hypognathum: length 144 (139–154); width 117(92–142); length palp 96 (93–98); chelicera 139 (131–146); legs I 206 (193–212); II 179(166–193); III 201(193–216); IV 223 (212–235); sensillae vi, 119(115–123), sce 107(100–112).
</paragraph>
<paragraph id="8B9536A4CC0AFFA5FF1DC25B17C2BDF6" blockId="12.[211,1172,1042,1194]" pageId="12" pageNumber="2059">
<emphasis id="B95EEAB6CC0AFFA5FF1DC25B14D2BD76" box="[211,479,1043,1067]" italics="true" pageId="12" pageNumber="2059">
Dorsum (
<figureCitation id="13112A21CC0AFFA5FEF3C25C14ACBD76" box="[317,417,1043,1066]" captionStart="Figure 5" captionStartId="11.[143,210,1512,1534]" captionTargetBox="[146,1102,162,1477]" captionTargetId="figure-5@11.[145,1102,162,1478]" captionTargetPageId="11" captionText="Figure 5. Pulaeus krama (Chaudhri, 1977), male. (A) Leg II; (B) leg III; (C) leg IV. Pulaeus razanensis Den Heyer sp. nov. female. (D) Dorsum; (E) sensilla vi; (F) sensilla sce; (G) venter." figureDoi="http://doi.org/10.5281/zenodo.4631929" httpUri="https://zenodo.org/record/4631929/files/figure.png" pageId="12" pageNumber="2059">Figure 5</figureCitation>
D–F)
</emphasis>
. As in
<taxonomicName id="4C2A4D27CC0AFFA5FDF4C25B17DEBD77" authorityName="Den Heyer" authorityYear="1979" box="[570,723,1043,1067]" class="Arachnida" family="Cunaxidae" genus="Pulaeus" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="2059" phylum="Arthropoda" rank="species" species="glebulentus">
<emphasis id="B95EEAB6CC0AFFA5FDF4C25B17DEBD77" box="[570,723,1043,1067]" italics="true" pageId="12" pageNumber="2059">P. glebulentus</emphasis>
</taxonomicName>
, entire idiosomal shield provided with nearly transverse lobes (
<figureCitation id="13112A21CC0AFFA5FE2EC27D1754BD16" box="[480,601,1074,1098]" captionStart="Figure 5" captionStartId="11.[143,210,1512,1534]" captionTargetBox="[146,1102,162,1477]" captionTargetId="figure-5@11.[145,1102,162,1478]" captionTargetPageId="11" captionText="Figure 5. Pulaeus krama (Chaudhri, 1977), male. (A) Leg II; (B) leg III; (C) leg IV. Pulaeus razanensis Den Heyer sp. nov. female. (D) Dorsum; (E) sensilla vi; (F) sensilla sce; (G) venter." figureDoi="http://doi.org/10.5281/zenodo.4631929" httpUri="https://zenodo.org/record/4631929/files/figure.png" pageId="12" pageNumber="2059">Figure 5D</figureCitation>
). Shield carries coarse setose sensillum pairs
<emphasis id="B95EEAB6CC0AFFA5FB81C27C116FBD16" box="[1103,1122,1075,1098]" italics="true" pageId="12" pageNumber="2059">vi</emphasis>
and
<emphasis id="B95EEAB6CC0AFFA5FF1DC21C15F9BD36" box="[211,244,1107,1130]" italics="true" pageId="12" pageNumber="2059">sce</emphasis>
(
<figureCitation id="13112A21CC0AFFA5FECFC21D1497BD36" box="[257,410,1106,1130]" captionStart="Figure 5" captionStartId="11.[143,210,1512,1534]" captionTargetBox="[146,1102,162,1477]" captionTargetId="figure-5@11.[145,1102,162,1478]" captionTargetPageId="11" captionText="Figure 5. Pulaeus krama (Chaudhri, 1977), male. (A) Leg II; (B) leg III; (C) leg IV. Pulaeus razanensis Den Heyer sp. nov. female. (D) Dorsum; (E) sensilla vi; (F) sensilla sce; (G) venter." figureDoi="http://doi.org/10.5281/zenodo.4631929" httpUri="https://zenodo.org/record/4631929/files/figure.png" pageId="12" pageNumber="2059">Figures 5E, F</figureCitation>
) as well as setae pairs
<emphasis id="B95EEAB6CC0AFFA5FD42C21C17AEBD36" box="[652,675,1107,1130]" italics="true" pageId="12" pageNumber="2059">ve</emphasis>
,
<emphasis id="B95EEAB6CC0AFFA5FD61C21C17C1BD36" box="[687,716,1107,1130]" italics="true" pageId="12" pageNumber="2059">sci</emphasis>
,
<emphasis id="B95EEAB6CC0AFFA5FD17C21C17FFBD37" box="[729,754,1107,1131]" italics="true" pageId="12" pageNumber="2059">c1</emphasis>
,
<emphasis id="B95EEAB6CC0AFFA5FD30C21C161ABD37" box="[766,791,1107,1131]" italics="true" pageId="12" pageNumber="2059">c2</emphasis>
,
<emphasis id="B95EEAB6CC0AFFA5FCEDC21C1630BD37" box="[803,829,1107,1131]" italics="true" pageId="12" pageNumber="2059">d1</emphasis>
,
<emphasis id="B95EEAB6CC0AFFA5FC84C21C166EBD37" box="[842,867,1107,1131]" italics="true" pageId="12" pageNumber="2059">e1</emphasis>
. Lyrifissures about halfway between setae
<emphasis id="B95EEAB6CC0AFFA5FEB4C23C149FBDD7" box="[378,402,1139,1163]" italics="true" pageId="12" pageNumber="2059">e1</emphasis>
and
<emphasis id="B95EEAB6CC0AFFA5FE1EC23C14EABDD7" box="[464,487,1139,1163]" italics="true" pageId="12" pageNumber="2059">f2</emphasis>
. Setae
<emphasis id="B95EEAB6CC0AFFA5FDF2C23C175EBDD6" box="[572,595,1139,1162]" italics="true" pageId="12" pageNumber="2059">ve</emphasis>
,
<emphasis id="B95EEAB6CC0AFFA5FDAAC23C178CBDD6" box="[612,641,1139,1162]" italics="true" pageId="12" pageNumber="2059">sci</emphasis>
,
<emphasis id="B95EEAB6CC0AFFA5FD5CC23C17A6BDD7" box="[658,683,1139,1163]" italics="true" pageId="12" pageNumber="2059">e1</emphasis>
and
<emphasis id="B95EEAB6CC0AFFA5FD27C23D1609BDD6" box="[745,772,1138,1162]" italics="true" pageId="12" pageNumber="2059">h1</emphasis>
of about equal length; other setae somewhat shorter. Integumental striae lobed.
</paragraph>
<paragraph id="8B9536A4CC0AFFA5FF1DC29C1632BC75" blockId="12.[211,1172,1234,1321]" pageId="12" pageNumber="2059">
<emphasis id="B95EEAB6CC0AFFA5FF1DC29C14CDBDB5" box="[211,448,1234,1258]" italics="true" pageId="12" pageNumber="2059">
Venter (
<figureCitation id="13112A21CC0AFFA5FEFBC29D14BABDB5" box="[309,439,1234,1257]" captionStart="Figure 5" captionStartId="11.[143,210,1512,1534]" captionTargetBox="[146,1102,162,1477]" captionTargetId="figure-5@11.[145,1102,162,1478]" captionTargetPageId="11" captionText="Figure 5. Pulaeus krama (Chaudhri, 1977), male. (A) Leg II; (B) leg III; (C) leg IV. Pulaeus razanensis Den Heyer sp. nov. female. (D) Dorsum; (E) sensilla vi; (F) sensilla sce; (G) venter." figureDoi="http://doi.org/10.5281/zenodo.4631929" httpUri="https://zenodo.org/record/4631929/files/figure.png" pageId="12" pageNumber="2059">Figure 5G</figureCitation>
)
</emphasis>
. Coxae II carry on their posteromedian edges a pair of propodogastral setae while coxae IV bear a pair of paracoxal setae on their median edges. Integumental striae consist of long broken lobes.
</paragraph>
<paragraph id="8B9536A4CC0AFFA5FF1DC31D1706BCD5" blockId="12.[211,1171,1361,1417]" pageId="12" pageNumber="2059">
<emphasis id="B95EEAB6CC0AFFA5FF1DC31D1711BC35" box="[211,540,1361,1386]" italics="true" pageId="12" pageNumber="2059">
<taxonomicName id="4C2A4D27CC0AFFA5FF1DC31D1450BC36" authorityName="Krefft" authorityYear="1871" box="[211,349,1362,1386]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="12" pageNumber="2059" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
(
<figureCitation id="13112A21CC0AFFA5FEBEC31D14DBBC35" box="[368,470,1361,1385]" captionStart="Figure 6" captionStartId="13.[143,210,1577,1599]" captionTargetBox="[145,1102,169,1544]" captionTargetId="figure-5@13.[145,1102,169,1544]" captionTargetPageId="13" captionText="Figure 6. Pulaeus razanensis Den Heyer sp. nov. female. (A) Hypognathum; (B) palp; (C) chelicera; (D) leg I; (E) leg II; (F) leg III; (G) leg IV." figureDoi="http://doi.org/10.5281/zenodo.4631931" httpUri="https://zenodo.org/record/4631931/files/figure.png" pageId="12" pageNumber="2059">Figure 6</figureCitation>
A–C).
</emphasis>
Very well sclerotized. Length ratio gnathosomal coxal region: hypostome 1 or less.
</paragraph>
<paragraph id="8B9536A4CC0AFFA5FF1DC3FE168FBF55" blockId="12.[211,1172,1457,1545]" pageId="12" pageNumber="2059">
<emphasis id="B95EEAB6CC0AFFA5FF1DC3FE1700BC95" box="[211,525,1457,1481]" italics="true" pageId="12" pageNumber="2059">
Hypognathum (
<figureCitation id="13112A21CC0AFFA5FE4CC3FD14F2BC95" box="[386,511,1457,1481]" captionStart="Figure 6" captionStartId="13.[143,210,1577,1599]" captionTargetBox="[145,1102,169,1544]" captionTargetId="figure-5@13.[145,1102,169,1544]" captionTargetPageId="13" captionText="Figure 6. Pulaeus razanensis Den Heyer sp. nov. female. (A) Hypognathum; (B) palp; (C) chelicera; (D) leg I; (E) leg II; (F) leg III; (G) leg IV." figureDoi="http://doi.org/10.5281/zenodo.4631931" httpUri="https://zenodo.org/record/4631931/files/figure.png" pageId="12" pageNumber="2059">Figure 6A</figureCitation>
).
</emphasis>
Ventral hypognathal coxal region posteriad to
<emphasis id="B95EEAB6CC0AFFA5FBFEC3FE1155BC95" box="[1072,1112,1457,1481]" italics="true" pageId="12" pageNumber="2059">hg4</emphasis>
with striae; laterad and anteriad, to latter setae, with papillae. Long setae
<emphasis id="B95EEAB6CC0AFFA5FBDCC39E1137BCB5" box="[1042,1082,1489,1513]" italics="true" pageId="12" pageNumber="2059">hg3</emphasis>
, nearly reaching
<emphasis id="B95EEAB6CC0AFFA5FEF4C3BE146FBF55" box="[314,354,1521,1545]" italics="true" pageId="12" pageNumber="2059">hg2</emphasis>
, placed posteriad off edge of palpal attachment.
</paragraph>
<paragraph id="8B9536A4CC0AFFA5FF1DC07E14E9BF34" blockId="12.[211,1172,1584,1640]" pageId="12" pageNumber="2059">
<emphasis id="B95EEAB6CC0AFFA5FF1DC07E1493BF15" box="[211,414,1585,1609]" italics="true" pageId="12" pageNumber="2059">
Palp (
<figureCitation id="13112A21CC0AFFA5FED8C07E149DBF15" box="[278,400,1585,1609]" captionStart="Figure 6" captionStartId="13.[143,210,1577,1599]" captionTargetBox="[145,1102,169,1544]" captionTargetId="figure-5@13.[145,1102,169,1544]" captionTargetPageId="13" captionText="Figure 6. Pulaeus razanensis Den Heyer sp. nov. female. (A) Hypognathum; (B) palp; (C) chelicera; (D) leg I; (E) leg II; (F) leg III; (G) leg IV." figureDoi="http://doi.org/10.5281/zenodo.4631931" httpUri="https://zenodo.org/record/4631931/files/figure.png" pageId="12" pageNumber="2059">Figure 6B</figureCitation>
).
</emphasis>
Resembles that of
<taxonomicName id="4C2A4D27CC0AFFA5FDB4C07D16C7BF14" authority="Den Heyer, 1981" authorityName="Den Heyer" authorityYear="1981" box="[634,970,1584,1609]" class="Arachnida" family="Cunaxidae" genus="Pulaeus" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="2059" phylum="Arthropoda" rank="species" species="franciscae">
<emphasis id="B95EEAB6CC0AFFA5FDB4C07D160BBF15" box="[634,774,1585,1609]" italics="true" pageId="12" pageNumber="2059">P. franciscae</emphasis>
<bibRefCitation id="EFBB4B55CC0AFFA5FCC1C07E16C7BF14" author="Den Heyer J" box="[783,970,1584,1608]" pageId="12" pageNumber="2059" pagination="87 - 99" refId="ref7769" refString="Den Heyer J. 1981. Three new Afrotropical species of the genus Pulaeus (Cunaxidae: Acarida). Phytophylactica. 13: 87 - 99." type="journal article" year="1981">Den Heyer,1981</bibRefCitation>
</taxonomicName>
. Proximal part of medial surface papillate.
</paragraph>
<caption id="DF55662CCC0BFFA4FF41C0661775BF00" ID-DOI="http://doi.org/10.5281/zenodo.4631931" ID-Zenodo-Dep="4631931" httpUri="https://zenodo.org/record/4631931/files/figure.png" pageId="13" pageNumber="2060" startId="13.[143,210,1577,1599]" targetBox="[145,1102,169,1544]" targetPageId="13">
<paragraph id="8B9536A4CC0BFFA4FF41C0661775BF00" blockId="13.[143,1103,1577,1628]" pageId="13" pageNumber="2060">
Figure 6.
<taxonomicName id="4C2A4D27CC0BFFA4FF39C0661724BF63" authority="Den Heyer" authorityName="Den Heyer" authorityYear="2013" box="[247,553,1577,1599]" class="Arachnida" family="Cunaxidae" genus="Pulaeus" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="2060" phylum="Arthropoda" rank="species" species="razanensis" status="sp. nov.">
<emphasis id="B95EEAB6CC0BFFA4FF39C06614BFBF63" box="[247,434,1577,1599]" italics="true" pageId="13" pageNumber="2060">Pulaeus razanensis</emphasis>
Den Heyer
</taxonomicName>
<taxonomicNameLabel id="A26D57CDCC0BFFA4FDFEC0651770BF63" box="[560,637,1578,1599]" pageId="13" pageNumber="2060" rank="species">sp. nov.</taxonomicNameLabel>
female. (A) Hypognathum; (B) palp; (C) chelicera; (D) leg I; (E) leg II; (F) leg III; (G) leg IV.
</paragraph>
</caption>
<paragraph id="8B9536A4CC08FFA7FF1DC6E71170B9E3" blockId="14.[211,1149,167,192]" box="[211,1149,167,192]" pageId="14" pageNumber="2061">
<heading id="D0DD81C8CC08FFA7FF1DC6E71170B9E3" box="[211,1149,167,192]" centered="true" fontSize="10" level="3" pageId="14" pageNumber="2061" reason="5">
<emphasis id="B95EEAB6CC08FFA7FF1DC6E714D3B99C" box="[211,478,168,192]" italics="true" pageId="14" pageNumber="2061">
Chelicerae (
<figureCitation id="13112A21CC08FFA7FE98C6E714DDB99C" box="[342,464,168,192]" captionStart="Figure 6" captionStartId="13.[143,210,1577,1599]" captionTargetBox="[145,1102,169,1544]" captionTargetId="figure-5@13.[145,1102,169,1544]" captionTargetPageId="13" captionText="Figure 6. Pulaeus razanensis Den Heyer sp. nov. female. (A) Hypognathum; (B) palp; (C) chelicera; (D) leg I; (E) leg II; (F) leg III; (G) leg IV." figureDoi="http://doi.org/10.5281/zenodo.4631931" httpUri="https://zenodo.org/record/4631931/files/figure.png" pageId="14" pageNumber="2061">Figure 6C</figureCitation>
).
</emphasis>
Typical for genus. Proximal part of dorsal surface papillate.
</heading>
</paragraph>
<paragraph id="8B9536A4CC08FFA7FF1DC6A7160BB8C2" blockId="14.[211,1172,231,414]" pageId="14" pageNumber="2061">
<emphasis id="B95EEAB6CC08FFA7FF1DC6A714B2B9A3" box="[211,447,231,255]" italics="true" pageId="14" pageNumber="2061">
Legs (
<figureCitation id="13112A21CC08FFA7FED8C6A7147AB9A3" box="[278,375,231,255]" captionStart="Figure 6" captionStartId="13.[143,210,1577,1599]" captionTargetBox="[145,1102,169,1544]" captionTargetId="figure-5@13.[145,1102,169,1544]" captionTargetPageId="13" captionText="Figure 6. Pulaeus razanensis Den Heyer sp. nov. female. (A) Hypognathum; (B) palp; (C) chelicera; (D) leg I; (E) leg II; (F) leg III; (G) leg IV." figureDoi="http://doi.org/10.5281/zenodo.4631931" httpUri="https://zenodo.org/record/4631931/files/figure.png" pageId="14" pageNumber="2061">Figure 6</figureCitation>
D–G).
</emphasis>
Leg chaetotaxy: coxae I–IV 3 sts, 1 peg – 2 sts (+ 1 ppgs) – 3 sts – 2 sts (+ 1 pcs); trochanters I–IV 1–1–2–1 sts; basifemora I–V 4–6–3–2 sts; telofemora I–IV 5sts –5sts –3sts, 1 ms –2 sts, 1 ms; genua I–V 3 or 2 asl, (1 asl,1 sts), 4 sts–2 asl, 5 sts –1 asl, 5 sts –1 asl, 5 sts; tibiae I–IV 1 asl, 5 sts–1bsl, 5sts –1 bsl, 5 sts – 4 sts, 1 long T; tarsi I–IV 4 asl (on left leg) and
<emphasis id="B95EEAB6CC08FFA7FD52C72817A5B823" bold="true" box="[668,680,359,383]" italics="true" pageId="14" pageNumber="2061">/</emphasis>
or 3asl (on right), 1 famulus pit, 1 dtsl, 2 tsl, 22 sts–1 bsl, 1 dtsl, 1 tsl, 19 sts–1 tsl, 17 sts –17 sts.
</paragraph>
<paragraph id="8B9536A4CC08FFA7FF1DC7881498B882" blockId="14.[211,405,454,479]" box="[211,405,454,479]" pageId="14" pageNumber="2061">
<heading id="D0DD81C8CC08FFA7FF1DC7881498B882" box="[211,405,454,479]" fontSize="10" level="3" pageId="14" pageNumber="2061" reason="5">
<emphasis id="B95EEAB6CC08FFA7FF1DC7881400B883" box="[211,269,455,479]" italics="true" pageId="14" pageNumber="2061">Male</emphasis>
. Unknown.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330652FCC08FFA7FF1DC46C1120BB3F" box="[211,1069,547,611]" pageId="14" pageNumber="2061" type="etymology">
<paragraph id="8B9536A4CC08FFA7FF1DC46C1446BB67" blockId="14.[211,1069,547,611]" box="[211,331,547,571]" pageId="14" pageNumber="2061">
<heading id="D0DD81C8CC08FFA7FF1DC46C1446BB67" box="[211,331,547,571]" fontSize="10" level="3" pageId="14" pageNumber="2061" reason="5">
<emphasis id="B95EEAB6CC08FFA7FF1DC46C1446BB67" box="[211,331,547,571]" italics="true" pageId="14" pageNumber="2061">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9536A4CC08FFA7FF1DC4041120BB3F" blockId="14.[211,1069,547,611]" box="[211,1069,587,611]" pageId="14" pageNumber="2061">This species is named after the town Razan, where this species was collected.</paragraph>
</subSubSection>
</treatment>
</document>
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