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<mods:title id="2C79D253CBFA22864F87264F495CCD88">A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species</mods:title>
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<heading id="357B81C5FFBFFFDCFF7E47259175672A" box="[151,785,798,825]" fontSize="11" level="2" pageId="33" pageNumber="34" reason="5">
<taxonomicName id="A98C4D2AFFBFFFDCFF7E472590F4672A" authority="(Smits van Burgst, 1914)" authorityName="Galsworthy &amp; Shaw &amp; Haraldseide" authorityYear="2023" baseAuthorityName="Smits van Burgst" baseAuthorityYear="1914" box="[151,656,798,825]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alpicola" status="comb. nov.">
<emphasis id="5CF8EABBFFBFFFDCFF7E472593E4672B" bold="true" box="[151,384,798,825]" pageId="33" pageNumber="34">
<emphasis id="5CF8EABBFFBFFFDCFF7E47259314672A" bold="true" box="[151,368,798,825]" italics="true" pageId="33" pageNumber="34">Hyposoter alpicola</emphasis>
(
</emphasis>
<bibRefCitation id="0A1D4B58FFBFFFDCFE69472590EC672A" author="Smits van Burgst, C. A. L." box="[384,648,799,825]" pageId="33" pageNumber="34" pagination="325 - 333" refId="ref55373" refString="Smits van Burgst, C. A. L. (1914) Ichneumonidae captured in the environs of Bozen (Tyrol) in June 1913. Deutsche Entomologische Zeitschrift, 1914, 325 - 333." type="journal article" year="1914">Smits van Burgst, 1914</bibRefCitation>
)
</taxonomicName>
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<taxonomicNameLabel id="47CB57C0FFBFFFDCFD7E47259175672A" box="[663,785,799,825]" pageId="33" pageNumber="34" rank="species">comb.nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="6E3336A9FFBFFFDCFF7E476496746766" blockId="33.[151,1436,862,1039]" box="[151,1040,862,888]" pageId="33" pageNumber="34">
<taxonomicName id="A98C4D2AFFBFFFDCFF7E4764904B6765" authority="Smits van Burgst, 1914" authorityName="Smits van Burgst" authorityYear="1914" box="[151,559,862,886]" class="Insecta" family="Noctuidae" genus="Angitia" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis id="5CF8EABBFFBFFFDCFF7E4764935F6765" box="[151,315,862,886]" italics="true" pageId="33" pageNumber="34">Angitia alpicola</emphasis>
<materialsCitation id="DEE43CF4FFBFFFDCFEA84764904B6765" ID-GBIF-Occurrence="4116948316" box="[321,559,862,886]" collectionCode="LEW" pageId="33" pageNumber="34" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="0A1D4B58FFBFFFDCFEA84764904B6765" author="Smits van Burgst, C. A. L." box="[321,559,862,886]" pageId="33" pageNumber="34" pagination="325 - 333" refId="ref55373" refString="Smits van Burgst, C. A. L. (1914) Ichneumonidae captured in the environs of Bozen (Tyrol) in June 1913. Deutsche Entomologische Zeitschrift, 1914, 325 - 333." type="journal article" year="1914">Smits van Burgst, 1914</bibRefCitation>
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</taxonomicName>
;
<typeStatus id="B137880BFFBFFFDCFDD5476490F16765" box="[572,661,862,886]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
♁ in LEW, examined by Horstmann.
</paragraph>
<paragraph id="6E3336A9FFBFFFDCFF7E474790B6601C" blockId="33.[151,1436,862,1039]" pageId="33" pageNumber="34">
syn.
<taxonomicName id="A98C4D2AFFBFFFDCFF20474790D46787" ID-CoL="3NYZ3" authority="Aubert, 1964" authorityName="Aubert" authorityYear="1964" box="[201,688,892,916]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="caedator" subSpecies="postcaedator">
<emphasis id="5CF8EABBFFBFFFDCFF20474790446787" box="[201,544,892,916]" italics="true" pageId="33" pageNumber="34">Hyposoter caedator postcaedator</emphasis>
<bibRefCitation id="0A1D4B58FFBFFFDCFDC1474690D46787" author="Aubert, J. - F." box="[552,688,892,916]" pageId="33" pageNumber="34" pagination="144 - 164" refId="ref51849" refString="Aubert, J. - F. (1964) Les Ichneumonides du rivage mediterraneen francais (Hym.). 7 e serie: Ichneumoninae, Cryptinae, Ophioninae et Mesochorinae de l'Herault et des Bouches-du-Rhone. Bulletin de la Societe Entomologique de France, 69, 144 - 164. https: // doi. org / 10.3406 / bsef. 1964.20764" type="journal article" year="1964">Aubert, 1964</bibRefCitation>
</taxonomicName>
<emphasis id="5CF8EABBFFBFFFDCFD5E474791726786" bold="true" box="[695,790,893,917]" pageId="33" pageNumber="34">
<taxonomicNameLabel id="47CB57C0FFBFFFDCFD5E474791726786" box="[695,790,893,917]" pageId="33" pageNumber="34">syn. nov.</taxonomicNameLabel>
</emphasis>
:
<typeStatus id="B137880BFFBFFFDCFCCC474691E16787" box="[805,901,892,916]" pageId="33" pageNumber="34" type="lectotype">lectotype</typeStatus>
, here designated. Label data: TYPE;
<taxonomicName id="A98C4D2AFFBFFFDCFAEE474790EE67A0" ID-CoL="3NYVS" authority="Grav. POSTCAEDATOR Aub" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="caedator">HYPOSOTER CAEDATOR Grav. POSTCAEDATOR Aub</taxonomicName>
; J F Aubert 1962 BdR VAUVERT; Si ce nest pas une forme de CAEDATOR ne correspond aucune autre espèce decrite [in Auberts handwriting];
<typeStatus id="B137880BFFBFFFDCFC6A478091B867C1" box="[899,988,954,978]" pageId="33" pageNumber="34" type="syntype">syntypes</typeStatus>
1 &amp; 2/?22
<taxonomicName id="A98C4D2AFFBFFFDCFBA34781933167E3" ID-CoL="3NYZ3" authority="Aubert 1964 c" authorityName="Aubert" authorityYear="1964" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="caedator" subSpecies="postcaedator">
<emphasis id="5CF8EABBFFBFFFDCFBA3478197F867C1" box="[1098,1436,954,978]" italics="true" pageId="33" pageNumber="34">Hyposoter caedator postcaedator</emphasis>
<bibRefCitation id="0A1D4B58FFBFFFDCFF2E47E2932E67E3" author="Aubert, J. - F." box="[199,330,984,1008]" pageId="33" pageNumber="34" pagination="144 - 164" refId="ref51849" refString="Aubert, J. - F. (1964) Les Ichneumonides du rivage mediterraneen francais (Hym.). 7 e serie: Ichneumoninae, Cryptinae, Ophioninae et Mesochorinae de l'Herault et des Bouches-du-Rhone. Bulletin de la Societe Entomologique de France, 69, 144 - 164. https: // doi. org / 10.3406 / bsef. 1964.20764" type="journal article" year="1964">Aubert 1964</bibRefCitation>
c
</taxonomicName>
, labelled by S. Klopfstein 2009. GBIFCH 00832761 [with barcode]; Note: there are
<specimenCount id="788AFD20FFBFFFDCFB5747E3973167E3" box="[1214,1365,984,1008]" pageId="33" pageNumber="34" type="generic">two specimens</specimenCount>
on the pin; the top one is chosen as the
<typeStatus id="B137880BFFBFFFDCFDF947CD9014601C" box="[528,624,1015,1039]" pageId="33" pageNumber="34" type="lectotype">lectotype</typeStatus>
. (
<figureCitation id="F6B72A2CFFBFFFDCFD6B47CD90AF601C" box="[642,715,1015,1039]" captionStart="FIGURES 3943" captionStartId="65.[152,270,1262,1286]" captionTargetBox="[151,1436,181,1237]" captionTargetId="figure-19@65.[151,1436,181,1237]" captionTargetPageId="65" captionText="FIGURES 3943. 3940: Hyposoter rufonigrator sp. n. Habitus: 39 lateral, 40 dorsal. 41: Hyposoter insulator lectotype. Habitus. 42: Hyposoter praecaedator lectotype. Habitus. 43: Hyposoter postcaedator lectotype. Habitus." figureDoi="http://doi.org/10.5281/zenodo.7959276" httpUri="https://zenodo.org/record/7959276/files/figure.png" pageId="33" pageNumber="34">Fig. 43</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="26966522FFBFFFDCFF7E4003934561A9" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="6E3336A9FFBFFFDCFF7E4003938860AD" blockId="33.[151,1437,1080,1466]" pageId="33" pageNumber="34">
<bibRefCitation id="0A1D4B58FFBFFFDCFF7E4003930D6041" author="Horstmann, K." box="[151,361,1080,1107]" pageId="33" pageNumber="34" pagination="413 - 472" refId="ref53346" refString="Horstmann, K. (1969) Typenrevision der europaischen Arten der Gattung Diadegma F ˆ rster (syn. Angitia Holmgren). Beitrage zur Entomologie, 19, 413 - 472." type="journal article" year="1969">Horstmann (1969)</bibRefCitation>
, after examining the
<typeStatus id="B137880BFFBFFFDCFDB7400290A46041" box="[606,704,1080,1106]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A98C4D2AFFBFFFDCFD00400291216041" authorityName="Smits van Burgst" authorityYear="1914" box="[745,837,1080,1106]" class="Insecta" family="Noctuidae" genus="Angitia" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis id="5CF8EABBFFBFFFDCFD00400291216041" box="[745,837,1080,1106]" italics="true" pageId="33" pageNumber="34">alpicola</emphasis>
</taxonomicName>
, speculated that this species belonged in
<taxonomicName id="A98C4D2AFFBFFFDCFAC8400397F26041" authorityName="Forster" authorityYear="1869" box="[1313,1430,1081,1106]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="5CF8EABBFFBFFFDCFAC8400397F26041" box="[1313,1430,1081,1106]" italics="true" pageId="33" pageNumber="34">Hyposoter</emphasis>
</taxonomicName>
. His notes, and his draft key, confirm that he had intended to transfer it to
<taxonomicName id="A98C4D2AFFBFFFDCFC57406796576065" authorityName="Forster" authorityYear="1869" box="[958,1075,1117,1142]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="5CF8EABBFFBFFFDCFC57406796576065" box="[958,1075,1117,1142]" italics="true" pageId="33" pageNumber="34">Hyposoter</emphasis>
</taxonomicName>
, and that he regarded
<taxonomicName id="A98C4D2AFFBFFFDCFAC1406793A16089" authority="Aubert, 1964" authorityName="Aubert" authorityYear="1964" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="postcaedator">
<emphasis id="5CF8EABBFFBFFFDCFAC14067934D6089" italics="true" pageId="33" pageNumber="34">Hyposoter postcaedator</emphasis>
<bibRefCitation id="0A1D4B58FFBFFFDCFEC640BA93A16089" author="Aubert, J. - F." box="[303,453,1152,1178]" pageId="33" pageNumber="34" pagination="144 - 164" refId="ref51849" refString="Aubert, J. - F. (1964) Les Ichneumonides du rivage mediterraneen francais (Hym.). 7 e serie: Ichneumoninae, Cryptinae, Ophioninae et Mesochorinae de l'Herault et des Bouches-du-Rhone. Bulletin de la Societe Entomologique de France, 69, 144 - 164. https: // doi. org / 10.3406 / bsef. 1964.20764" type="journal article" year="1964">Aubert, 1964</bibRefCitation>
</taxonomicName>
, the type material of which he had also examined, as a junior synonym of it. These intentions are now formalised here.
</paragraph>
<paragraph id="6E3336A9FFBFFFDCFF2E40F396B7615C" blockId="33.[151,1437,1080,1466]" pageId="33" pageNumber="34">
We have not ourselves seen the
<typeStatus id="B137880BFFBFFFDCFDCE40F290ED60F1" box="[551,649,1224,1250]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A98C4D2AFFBFFFDCFD4440F2916D60F1" authorityName="Smits van Burgst" authorityYear="1914" box="[685,777,1224,1250]" class="Insecta" family="Noctuidae" genus="Angitia" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis id="5CF8EABBFFBFFFDCFD4440F2916D60F1" box="[685,777,1224,1250]" italics="true" pageId="33" pageNumber="34">alpicola</emphasis>
</taxonomicName>
, and rely on Horstmanns notes. We have however seen the
<typeStatus id="B137880BFFBFFFDCFF7E40D6929A6115" box="[151,254,1260,1286]" pageId="33" pageNumber="34" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="A98C4D2AFFBFFFDCFECB40D693D06115" authorityName="Aubert" authorityYear="1964" box="[290,436,1260,1286]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="postcaedator">
<emphasis id="5CF8EABBFFBFFFDCFECB40D693D06115" box="[290,436,1260,1286]" italics="true" pageId="33" pageNumber="34">postcaedator</emphasis>
</taxonomicName>
designated above. This is distinct from
<taxonomicName id="A98C4D2AFFBFFFDCFC8740D796426115" authorityName="Aubert" authorityYear="1963" box="[878,1062,1260,1286]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="praecaedator">
<emphasis id="5CF8EABBFFBFFFDCFC8740D796426115" box="[878,1062,1260,1286]" italics="true" pageId="33" pageNumber="34">H. praecaedator</emphasis>
</taxonomicName>
in having the last 2 tergites of the metasoma darkened, and a narrow dark apex to tergite 2. The mesopleuron is fully granulate. The species resembles
<taxonomicName id="A98C4D2AFFBFFFDCFF7E410F9379615D" box="[151,285,1332,1358]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="caedator">
<emphasis id="5CF8EABBFFBFFFDCFF7E410F9379615D" box="[151,285,1332,1358]" italics="true" pageId="33" pageNumber="34">H. caedator</emphasis>
</taxonomicName>
more than
<taxonomicName id="A98C4D2AFFBFFFDCFE75410E9057615D" authorityName="Aubert" authorityYear="1963" box="[412,563,1332,1358]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="praecaedator">
<emphasis id="5CF8EABBFFBFFFDCFE75410E9057615D" box="[412,563,1332,1358]" italics="true" pageId="33" pageNumber="34">praecaedator</emphasis>
</taxonomicName>
but, unlike
<taxonomicName id="A98C4D2AFFBFFFDCFD51410E9178615D" box="[696,796,1332,1358]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="caedator">
<emphasis id="5CF8EABBFFBFFFDCFD51410E9178615D" box="[696,796,1332,1358]" italics="true" pageId="33" pageNumber="34">caedator</emphasis>
</taxonomicName>
, it has a yellow underside to the scape.
</paragraph>
<paragraph id="6E3336A9FFBFFFDCFF2E4162934561A9" blockId="33.[151,1437,1080,1466]" pageId="33" pageNumber="34">
The
<typeStatus id="B137880BFFBFFFDCFF13416293386161" box="[250,348,1368,1394]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A98C4D2AFFBFFFDCFE68416390646161" authorityName="Smits van Burgst" authorityYear="1914" box="[385,512,1368,1394]" class="Insecta" family="Noctuidae" genus="Angitia" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis id="5CF8EABBFFBFFFDCFE68416390646161" box="[385,512,1368,1394]" italics="true" pageId="33" pageNumber="34">H. alpicola</emphasis>
</taxonomicName>
was from near Bolzano in the Italian Alps.
<taxonomicName id="A98C4D2AFFBFFFDCFC00416396936161" authorityName="Aubert" authorityYear="1964" box="[1001,1271,1368,1394]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="postcaedator">
<emphasis id="5CF8EABBFFBFFFDCFC00416396936161" box="[1001,1271,1368,1394]" italics="true" pageId="33" pageNumber="34">Hyposoter postcaedator</emphasis>
</taxonomicName>
was described from the south of
<collectingCountry id="169B7639FFBFFFDCFEB4414793C96184" box="[349,429,1405,1431]" name="France" pageId="33" pageNumber="34">France</collectingCountry>
. We have seen
<specimenCount id="788AFD20FFBFFFDCFDBC4147909E6185" box="[597,762,1404,1431]" pageId="33" pageNumber="34" type="generic">two specimens</specimenCount>
of
<taxonomicName id="A98C4D2AFFBFFFDCFCF7414791FF6185" authorityName="Smits van Burgst" authorityYear="1914" box="[798,923,1404,1430]" class="Insecta" family="Noctuidae" genus="Angitia" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis id="5CF8EABBFFBFFFDCFCF7414791FF6185" box="[798,923,1404,1430]" italics="true" pageId="33" pageNumber="34">H. alpicola</emphasis>
</taxonomicName>
from
<collectingCountry id="169B7639FFBFFFDCFC374147965D6184" box="[990,1081,1405,1431]" name="Norway" pageId="33" pageNumber="34">Norway</collectingCountry>
, both taken in July. The biology is unknown.
</paragraph>
</subSubSection>
</treatment>
</document>