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<document id="F6031027A297BABFEB1BB5966035EFC0" ID-CLB-Dataset="20458" ID-DOI="10.11646/palaeoentomology.5.1.11" ID-GBIF-Dataset="a8c2895d-e4a0-4793-b9f9-cda04a191af6" ID-ISSN="2624-2834" ID-Zenodo-Dep="6280600" ID-ZooBank="C21D37B9-E67D-455C-B04D-0CB130BEE0CD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1645778115009" checkinUser="plazi" docAuthor="Fedotova, Zoya A., Perkovsky, Evgeny E., Ross, Andrew J. &amp; Zhang, Qingqing" docDate="2022" docId="D34F46249735FFDEFF29F9E0FA91FEDF" docLanguage="en" docName="palaeoentomology.5.1.11.pdf" docOrigin="Palaeoentomology 5 (1)" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Fushuniola Fedotova &amp; Perkovsky &amp; Ross &amp; Zhang 2022, gen. nov." docType="treatment" docVersion="7" lastPageNumber="95" masterDocId="2F763E5C9731FFDBFFA1FF9CFFA5FFFD" masterDocTitle="A new genus and species of gall midges the tribe Winnertziini (Diptera, Cecidomyiidae, Porricondylinae) from lower Eocene Fushun amber from China" masterLastPageNumber="98" masterPageNumber="90" pageNumber="94" updateTime="1699317416577" updateUser="ExternalLinkService">
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<mods:title id="930A388B5703054FD821906072ECB31C">A new genus and species of gall midges the tribe Winnertziini (Diptera, Cecidomyiidae, Porricondylinae) from lower Eocene Fushun amber from China</mods:title>
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<mods:name id="349859435254DE6CD37D459559CAD5E9" type="personal">
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<mods:namePart id="38FB8651242384BF670EB55692BC2E09">Fedotova, Zoya A.</mods:namePart>
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<mods:namePart id="090BF837F68FD5FC6390E6BC6DB4D149">Perkovsky, Evgeny E.</mods:namePart>
<mods:affiliation id="C5AB8A2C86BBE1BE03C7B9BCEDB38E6E">A. A. Borissiak Paleontological Institute of the Russian Academy of Sciences, Profsoyuznaya Str. 123, Moscow, 117647, Russia Department of Natural Sciences, National Museum of Scotland, Chambers St., Edinburgh EH 1 1 JF, UK</mods:affiliation>
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<mods:namePart id="97A62FA90825101D351970B0FC5888F6">Ross, Andrew J.</mods:namePart>
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<mods:namePart id="257620B437D920885ED17520D915287C">Zhang, Qingqing</mods:namePart>
<mods:affiliation id="2C8468498FF401D435121E9EF44F7B3A">Institute of Geosciences, University of Bonn, Bonn 53115, Germany &amp; State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
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<mods:title id="8E8F922D2EA262602F3ACCDB978E1804">Palaeoentomology</mods:title>
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<mods:date id="E84E24FB1BEB06A64156625D145717EF">2022</mods:date>
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<mods:number id="19DFD97D61ECD4D5BF9AEAF25EF7C331">2022-02-25</mods:number>
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<mods:number id="B1D158AABF1CCCB0742C8E00B6340B08">5</mods:number>
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<mods:number id="38CC36ACB7EAC24038156B1449932FB3">1</mods:number>
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<treatment id="D34F46249735FFDEFF29F9E0FA91FEDF" ID-DOI="http://doi.org/10.5281/zenodo.6317489" ID-GBIF-Taxon="193838230" ID-Zenodo-Dep="6317489" LSID="urn:lsid:plazi:treatment:D34F46249735FFDEFF29F9E0FA91FEDF" httpUri="http://treatment.plazi.org/id/D34F46249735FFDEFF29F9E0FA91FEDF" lastPageId="5" lastPageNumber="95" pageId="4" pageNumber="94">
<subSubSection id="13FCA4B99735FFDFFF29F9E0FD54F96A" box="[136,753,1660,1687]" pageId="4" pageNumber="94" type="nomenclature">
<paragraph id="5B59F7329735FFDFFF29F9E0FD54F96A" blockId="4.[136,753,1660,1687]" box="[136,753,1660,1687]" pageId="4" pageNumber="94">
<heading id="0011405E9735FFDFFF29F9E0FD54F96A" box="[136,753,1660,1687]" fontSize="10" level="2" pageId="4" pageNumber="94" reason="2">
<emphasis id="69922B209735FFDFFF29F9E0FD54F96A" bold="true" box="[136,753,1660,1687]" pageId="4" pageNumber="94">
Genus
<taxonomicName id="9CE68CB19735FFDFFF7AF9E0FDDEF96A" ID-CoL="9HZYV" authority="Fedotova &amp; Perkovsky" authorityName="Fedotova &amp; Perkovsky &amp; Ross &amp; Zhang" authorityYear="2022" box="[219,635,1660,1687]" class="Insecta" family="Cecidomyiidae" genus="Fushuniola" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="69922B209735FFDFFF7AF9E0FEC7F96A" bold="true" box="[219,354,1660,1687]" italics="true" pageId="4" pageNumber="94">Fushuniola</emphasis>
Fedotova &amp; Perkovsky
</taxonomicName>
,
<taxonomicNameLabel id="72A1965B9735FFDFFD29F9E1FD54F96A" box="[648,753,1661,1687]" pageId="4" pageNumber="94" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="13FCA4B99735FFDFFF29F95BFE6EF8F8" pageId="4" pageNumber="94" type="type_taxon">
<paragraph id="5B59F7329735FFDFFF29F95BFE6EF8F8" blockId="4.[136,777,1734,2092]" pageId="4" pageNumber="94">
<emphasis id="69922B209735FFDFFF29F95BFE87F91D" bold="true" box="[136,290,1734,1761]" pageId="4" pageNumber="94">
<typeStatus id="845D49909735FFDFFF29F95BFF66F91C" box="[136,195,1735,1761]" pageId="4" pageNumber="94">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="9CE68CB19735FFDFFE89F95AFD7BF91D" ID-CoL="9HZYT" authority="Fedotova &amp; Perkovsky" authorityName="Fedotova &amp; Perkovsky" authorityYear="2022" box="[296,734,1734,1761]" class="Insecta" family="Cecidomyiidae" genus="Fushuniola" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="mai" status="sp. nov.">
<emphasis id="69922B209735FFDFFE89F95AFE72F91D" box="[296,471,1734,1760]" italics="true" pageId="4" pageNumber="94">Fushuniola mai</emphasis>
Fedotova &amp; Perkovsky
</taxonomicName>
,
<emphasis id="69922B209735FFDFFD49F95BFF1CF8FB" bold="true" pageId="4" pageNumber="94">
<taxonomicNameLabel id="72A1965B9735FFDFFD49F95BFF1CF8FB" pageId="4" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="13FCA4B99735FFDFFF1CF88CFF71F8B2" pageId="4" pageNumber="94" type="etymology">
<paragraph id="5B59F7329735FFDFFF1CF88CFF71F8B2" blockId="4.[136,777,1734,2092]" pageId="4" pageNumber="94">
<emphasis id="69922B209735FFDFFF1CF88CFEE1F8D7" bold="true" box="[189,324,1808,1834]" pageId="4" pageNumber="94">Etymology.</emphasis>
The genus is named after Fushun amber.
</paragraph>
</subSubSection>
<subSubSection id="13FCA4B99735FFDFFF1CF8C6FBCDFA47" pageId="4" pageNumber="94" type="diagnosis">
<paragraph id="5B59F7329735FFDFFF1CF8C6FBCDFA47" blockId="4.[136,777,1734,2092]" lastBlockId="4.[810,1452,1292,2093]" pageId="4" pageNumber="94">
<emphasis id="69922B209735FFDFFF1CF8C6FE9DF889" bold="true" box="[189,312,1882,1908]" pageId="4" pageNumber="94">Diagnosis.</emphasis>
Head of female is broad, enlarged frontally, palpi much longer than height of head, long lingua developed, antennae 2+10-segmented, all flagellomeres with distinct very short and narrow necks, 4-8
<superScript id="AC935A7A9735FFDFFF0CF871FF1FF806" attach="left" box="[173,186,2029,2043]" fontSize="6" pageId="4" pageNumber="94">th</superScript>
with short hair-shaped sensoria, vein R5 strongly curved apically, cell between veins C and R1+2 slightly sclerotized,
<emphasis id="69922B209735FFDFFC17FA91FC59FADB" box="[950,1020,1293,1318]" italics="true" pageId="4" pageNumber="94">rm+m</emphasis>
slightly curved medially, prolonged projection R1+2 from Rs to point of fusion with C, vein Sc absence, hind femora longer than their tibia, tarsal claws simple, apical lobe of ovipositor is wide, covered by strongly prolonged setae.
</paragraph>
</subSubSection>
<subSubSection id="13FCA4B99735FFDEFCFEFA58FEE5FC3F" lastPageId="5" lastPageNumber="95" pageId="4" pageNumber="94" type="description">
<paragraph id="5B59F7329735FFDEFCFEFA58FEE5FC3F" blockId="4.[810,1452,1292,2093]" lastBlockId="5.[136,778,158,2093]" lastPageId="5" lastPageNumber="95" pageId="4" pageNumber="94">
<emphasis id="69922B209735FFDFFCFEFA58FC54FA23" bold="true" box="[863,1009,1476,1502]" pageId="4" pageNumber="94">Description.</emphasis>
Female (
<figureCitation id="C3DDEBB79735FFDFFBFFFA58FB7DFA22" box="[1118,1240,1476,1503]" captionStart="FIGURE 1" captionStartId="3.[136,235,1680,1704]" captionTargetBox="[151,1435,189,1657]" captionTargetId="figure-143@3.[151,1435,189,1657]" captionTargetPageId="3" captionText="FIGURE 1. Fushuniola mai Fedotova &amp; Perkovsky, gen. et sp. nov. in lower Eocene Fushun amber, female. A, General appearance with two separated fore tarsi and one hind tarsus. B, Antenna. C, Tarsomeres 35 of hind leg and tarsomere 5 of fore leg. D, Head. E, Flagellomeres 35. F, Apex of abdomen. G, Ovipositor. H, Wing. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6280602" httpUri="https://zenodo.org/record/6280602/files/figure.png" pageId="4" pageNumber="94">Fig. 1AH</figureCitation>
,
<figureCitation id="C3DDEBB79735FFDFFB45FA58FAF2FA22" box="[1252,1367,1476,1503]" captionStart="FIGURE 2" captionStartId="4.[136,235,1139,1163]" captionTargetBox="[170,1413,210,1115]" captionTargetId="figure-334@4.[151,1436,189,1115]" captionTargetPageId="4" captionText="FIGURE 2. Fushuniola mai Fedotova &amp; Perkovsky, gen. et sp. nov. in lower Eocene Fushun amber, female.A, General appearance. B, C, tarsomeres 25 of fore leg. D, Tarsomeres 25 of hind leg. E, Tarsomeres 45 of hind leg. F, Flagellomeres 410. G, 5th flagellomere. H, Scape, pedicel and flagellomeres 13. I, Mouthparts and palpi. J, Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6280604" httpUri="https://zenodo.org/record/6280604/files/figure.png" pageId="4" pageNumber="94">Fig. 2AJ</figureCitation>
). Body slightly longer than wings. Head is broad, enlarged frontally; eyes fused, positioned around the head, with wide bridge, occiput wide rounded. Numerous occipital setae very long. Mouthparts much elongate and wide. Mouthparts longer than half part of the face length. Long lingua is developed. Palpi much longer than height of head, 4-segmented, the 3
<superScript id="AC935A7A9735FFDFFB6AF95AFB7CF929" attach="left" box="[1227,1241,1734,1748]" fontSize="6" pageId="4" pageNumber="94">rd</superScript>
and 4
<superScript id="AC935A7A9735FFDFFA88F95AFA93F929" attach="left" box="[1321,1334,1734,1748]" fontSize="6" pageId="4" pageNumber="94">th</superScript>
segments are long. Antennae 2+10-segmented, scape enlarged distally, pedicel slightly broad. Basal enlargement of 1
<superScript id="AC935A7A9735FFDFFC99F8A9FCE6F8BE" attach="left" box="[824,835,1845,1859]" fontSize="6" pageId="4" pageNumber="94">st</superScript>
and 2
<superScript id="AC935A7A9735FFDFFC32F8A9FC06F8BE" attach="left" box="[915,931,1845,1859]" fontSize="6" pageId="4" pageNumber="94">nd</superScript>
flagellomeres slightly more prolonged than others. All flagellomeres with distinct very short and narrow necks, but basal enlargements slightly shorted from base to apex of flagellum. Basal enlargement on proximal flagellomeres with narrowing near middle and enlarged basally. Medial flagellomeres with very long setae in basal whorls and basally curved medial whorls of rare setae. Flagellomeres 48
<superScript id="AC935A7A9734FFDEFE57FF02FDA6FF51" attach="left" box="[502,515,158,172]" fontSize="6" pageId="5" pageNumber="95">th</superScript>
with short hair-shaped sensoria. Necks of flagellomeres always distinct but very short, especially of the middle flagellomeres. The head is located on a very long neck, which passes into a similarly narrowed part of the apical part of the prothorax. Wing evenly and very strongly widened medially. Costal vein with break is beyond joining one and R5. Cell between veins C and R1+2 slightly sclerotized. Vein
<emphasis id="69922B209734FFDEFDCBFE0AFD14FE52" box="[618,689,406,431]" italics="true" pageId="5" pageNumber="95">rm+m</emphasis>
slightly curved medially, R5 strongly curved apically, joining margin of wing little before the wing tip, Rs very strong; slightly curved CuA and M3+4 simple, positioned far from margin of wing, not reaching of wing edge, forms large anal lobe. Vein Rs more than 4 times longer than projection R1+2 from Rs to point of fusion with C. Legs completely densely covered by short setae and rare long setae. Fore femora shorter than tibia. Hind femora longer than their tibia. Tarsal claws simple, slightly curved. Abdomen strongly swollen basally (1
<superScript id="AC935A7A9734FFDEFD8EFD6BFD9FFCF8" attach="none" box="[559,570,759,773]" fontSize="6" pageId="5" pageNumber="95">st</superScript>
3
<superScript id="AC935A7A9734FFDEFDF7FD6BFDC1FCF8" attach="left" box="[598,612,759,773]" fontSize="6" pageId="5" pageNumber="95">rd</superScript>
segments) and narrowed apically. Abdominal segments covered by very long setae. Ovipositor very long and thin, curved dorsally with 3-segmented apical plates (lateral view,
<figureCitation id="C3DDEBB79734FFDEFDDAFCFDFD77FC86" box="[635,722,865,891]" captionStart="FIGURE 1" captionStartId="3.[136,235,1680,1704]" captionTargetBox="[151,1435,189,1657]" captionTargetId="figure-143@3.[151,1435,189,1657]" captionTargetPageId="3" captionText="FIGURE 1. Fushuniola mai Fedotova &amp; Perkovsky, gen. et sp. nov. in lower Eocene Fushun amber, female. A, General appearance with two separated fore tarsi and one hind tarsus. B, Antenna. C, Tarsomeres 35 of hind leg and tarsomere 5 of fore leg. D, Head. E, Flagellomeres 35. F, Apex of abdomen. G, Ovipositor. H, Wing. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6280602" httpUri="https://zenodo.org/record/6280602/files/figure.png" pageId="5" pageNumber="95">Fig. 1G</figureCitation>
;
<figureCitation id="C3DDEBB79734FFDEFD7CFCFDFF00FC62" captionStart="FIGURE 2" captionStartId="4.[136,235,1139,1163]" captionTargetBox="[170,1413,210,1115]" captionTargetId="figure-334@4.[151,1436,189,1115]" captionTargetPageId="4" captionText="FIGURE 2. Fushuniola mai Fedotova &amp; Perkovsky, gen. et sp. nov. in lower Eocene Fushun amber, female.A, General appearance. B, C, tarsomeres 25 of fore leg. D, Tarsomeres 25 of hind leg. E, Tarsomeres 45 of hind leg. F, Flagellomeres 410. G, 5th flagellomere. H, Scape, pedicel and flagellomeres 13. I, Mouthparts and palpi. J, Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6280604" httpUri="https://zenodo.org/record/6280604/files/figure.png" pageId="5" pageNumber="95">Fig. 2J</figureCitation>
). Apical lobe of ovipositor is wide, covered by strongly prolonged setae.
</paragraph>
</subSubSection>
<subSubSection id="13FCA4B99734FFDEFF1CFC57FA91FEDF" pageId="5" pageNumber="95" type="discussion">
<paragraph id="5B59F7329734FFDEFF1CFC57FE27F7F4" blockId="5.[136,778,158,2093]" pageId="5" pageNumber="95">
<emphasis id="69922B209734FFDEFF1CFC57FE94FC18" bold="true" box="[189,305,971,997]" pageId="5" pageNumber="95">Remarks.</emphasis>
The shape of the body, 2+10-segmented female antennae, reduced wing venation with slightly curved veins CuA and M3+4, long legs and ovipositor the new genus is closely similar to modern genus
<taxonomicName id="9CE68CB19734FFDEFF29FBC4FE5BFB8F" authority="Mamaev, 1964" authorityName="Mamaev" authorityYear="1964" box="[136,510,1112,1139]" class="Insecta" family="Cecidomyiidae" genus="Rhipidoxylomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="69922B209734FFDEFF29FBC4FEE3FB8F" box="[136,326,1112,1138]" italics="true" pageId="5" pageNumber="95">Rhipidoxylomyia</emphasis>
<bibRefCitation id="3F778AC39734FFDEFEF0FBC5FE5BFB8F" author="Mamaev, B. M." box="[337,510,1112,1139]" pageId="5" pageNumber="95" pagination="894 - 913" refId="ref6840" refString="Mamaev, B. M. (1964) [Gall midges of the USSR. 6. New species of the tribe Porricondylini (Diptera, Cecidomyiidae).] Entomologicheskoe Obozrenie, 43 (4), 894 - 913. [In Russian]" type="journal article" year="1964">Mamaev, 1964</bibRefCitation>
</taxonomicName>
, which is found in the Palaearctic and Oriental regions, as well as in Rovno amber, comprising 18 species, 15 of which are described from the Palaearctic (
<bibRefCitation id="3F778AC39734FFDEFE38FB5EFCA0FB20" author="Fedotova, Z. A. &amp; Sidorenko, V. S." box="[409,773,1218,1245]" pageId="5" pageNumber="95" pagination="69 - 103" refId="ref5977" refString="Fedotova, Z. A. &amp; Sidorenko, V. S. (2007) New taxa of gall midges from tribes Porricondylini, Bryocryptini, Asynaptini and Winnertziini (Diptera, Cecidomyiidae, Porricondylinae) from the Russian Far East. An International Journal of Dipterological Research, 18 (2), 69 - 103." type="journal article" year="2007">Fedotova &amp; Sidorenko, 2007</bibRefCitation>
;
<bibRefCitation id="3F778AC39734FFDEFF29FB7AFE7DFAFD" author="Fedotova, Z. A. &amp; Perkovsky, E. E." box="[136,472,1254,1280]" pageId="5" pageNumber="95" pagination="27 - 40" refId="ref5529" refString="Fedotova, Z. A. &amp; Perkovsky, E. E. (2008) New taxa of gall midges (Diptera, Cecidomyiidae) from Dubrovitsa (Rovno amber). Vestnik Zoologii, 42 (1), 27 - 40. https: // doi. org / 10.1007 / s 11492 - 008 - 2007 - 6" type="journal article" year="2008">Fedotova &amp; Perkovsky, 2008</bibRefCitation>
;
<bibRefCitation id="3F778AC39734FFDEFE47FB7AFCA0FAFD" author="Gagne, R. J. &amp; Jaschhof, M." box="[486,773,1254,1280]" pageId="5" pageNumber="95" refId="ref6067" refString="Gagne, R. J. &amp; Jaschhof, M. (2021) A catalog of the Cecidomyiidae (Diptera) of the world. 5 th Edition. Digital. 813 pp. https: // www. ars. usda. gov / ARSUserFiles / 80420580 / Gagne _" type="book" year="2021">Gagné &amp; Jaschhof, 2021</bibRefCitation>
;
<bibRefCitation id="3F778AC39734FFDEFF29FA95FF5BFADE" author="Nel, A." box="[136,254,1289,1315]" pageId="5" pageNumber="95" pagination="620 - 628" refId="ref7014" refString="Nel, A. (2021) New Cecidomyiidae from the Lowermost Eocene French amber (Diptera). Palaeoentomology, 4 (6), 620 - 628. https: // doi. org / 10.11646 / palaeoentomology. 4.6.11" type="journal article" year="2021">Nel, 2021</bibRefCitation>
). Previously, 6 extant species were described from
<collectingCountry id="23F1B7A29734FFDEFF6BFAB1FEA8FABA" box="[202,269,1325,1351]" name="China" pageId="5" pageNumber="95">China</collectingCountry>
(
<bibRefCitation id="3F778AC39734FFDEFEBEFAB1FE50FABA" author="Jiang, Y. X. &amp; Bu, W. J." box="[287,501,1325,1351]" pageId="5" pageNumber="95" pagination="357 - 364" refId="ref6605" refString="Jiang, Y. X. &amp; Bu, W. J. (2004) Six new species of the genus Rhipidoxylomyia from China (Diptera, Cecidomyiidae). Acta Zootaxonomica Sinica, 29, 357 - 364." type="journal article" year="2004">Jiang &amp; Bu, 2004</bibRefCitation>
) trapped in Malaise. In the forest zone, the larvae of known
<taxonomicName id="9CE68CB19734FFDEFDEDFACCFF7CFA70" class="Insecta" family="Cecidomyiidae" genus="Rhipidoxylomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="95" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="69922B209734FFDEFDEDFACCFCAFFA97" box="[588,778,1360,1386]" italics="true" pageId="5" pageNumber="95">Rhipidoxylomyia</emphasis>
species
</taxonomicName>
live exclusively in decaying wood (
<bibRefCitation id="3F778AC39734FFDEFD24FAEFFEFCFA4C" author="Mamaev, B. M. &amp; Krivosheina, N. P." pageId="5" pageNumber="95" refId="ref6887" refString="Mamaev, B. M. &amp; Krivosheina, N. P. (1965) [Larvae of gall midges (Diptera, Cecidomyiidae). Comparative morphology, biology and keys.] Nauka, Moskow-Leningrad, 278 pp. [In Russian]" type="book" year="1965">Mamaev &amp; Krivosheina, 1965</bibRefCitation>
). Representatives of the genus have not been found in Baltic amber. Only two species are fossil. The single species from Rovno amber,
<taxonomicName id="9CE68CB19734FFDEFDEDFA41FDBEF9E6" authority="Fedotova &amp; Perkovsky, 2008" authorityName="Fedotova &amp; Perkovsky" authorityYear="2008" class="Insecta" family="Cecidomyiidae" genus="Rhipidoxylomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="95" phylum="Arthropoda" rank="species" species="vaga">
<emphasis id="69922B209734FFDEFDEDFA41FF1BF9E7" italics="true" pageId="5" pageNumber="95">Rhipidoxylomyia vaga</emphasis>
<bibRefCitation id="3F778AC39734FFDEFF69F99DFDBEF9E6" author="Fedotova, Z. A. &amp; Perkovsky, E. E." box="[200,539,1537,1563]" pageId="5" pageNumber="95" pagination="27 - 40" refId="ref5529" refString="Fedotova, Z. A. &amp; Perkovsky, E. E. (2008) New taxa of gall midges (Diptera, Cecidomyiidae) from Dubrovitsa (Rovno amber). Vestnik Zoologii, 42 (1), 27 - 40. https: // doi. org / 10.1007 / s 11492 - 008 - 2007 - 6" type="journal article" year="2008">Fedotova &amp; Perkovsky, 2008</bibRefCitation>
</taxonomicName>
, was described from the Dubrovitsa deposit (
<bibRefCitation id="3F778AC39734FFDEFE02F9B8FD5BF9C3" author="Fedotova, Z. A. &amp; Perkovsky, E. E." box="[419,766,1572,1598]" pageId="5" pageNumber="95" pagination="27 - 40" refId="ref5529" refString="Fedotova, Z. A. &amp; Perkovsky, E. E. (2008) New taxa of gall midges (Diptera, Cecidomyiidae) from Dubrovitsa (Rovno amber). Vestnik Zoologii, 42 (1), 27 - 40. https: // doi. org / 10.1007 / s 11492 - 008 - 2007 - 6" type="journal article" year="2008">Fedotova &amp; Perkovsky, 2008</bibRefCitation>
). The second fossil species
<taxonomicName id="9CE68CB19734FFDEFE0EF9D4FD10F99F" authority="Nel, 2021" authorityName="Nel" authorityYear="2021" box="[431,693,1607,1634]" class="Insecta" family="Cecidomyiidae" genus="Rhipidoxylomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="95" phylum="Arthropoda" rank="species" species="rasnitsyni">
<emphasis id="69922B209734FFDEFE0EF9D4FE65F99C" box="[431,448,1608,1633]" italics="true" pageId="5" pageNumber="95">R</emphasis>
.
<emphasis id="69922B209734FFDEFE6EF9D4FD98F99C" box="[463,573,1608,1633]" italics="true" pageId="5" pageNumber="95">rasnitsyni</emphasis>
<bibRefCitation id="3F778AC39734FFDEFDE7F9DBFD10F99F" author="Nel, A." box="[582,693,1607,1634]" pageId="5" pageNumber="95" pagination="620 - 628" refId="ref7014" refString="Nel, A. (2021) New Cecidomyiidae from the Lowermost Eocene French amber (Diptera). Palaeoentomology, 4 (6), 620 - 628. https: // doi. org / 10.11646 / palaeoentomology. 4.6.11" type="journal article" year="2021">Nel, 2021</bibRefCitation>
</taxonomicName>
known from Oise amber (
<bibRefCitation id="3F778AC39734FFDEFEF3F9F7FE60F978" author="Nel, A." box="[338,453,1643,1669]" pageId="5" pageNumber="95" pagination="620 - 628" refId="ref7014" refString="Nel, A. (2021) New Cecidomyiidae from the Lowermost Eocene French amber (Diptera). Palaeoentomology, 4 (6), 620 - 628. https: // doi. org / 10.11646 / palaeoentomology. 4.6.11" type="journal article" year="2021">Nel, 2021</bibRefCitation>
). The new genus differs from
<taxonomicName id="9CE68CB19734FFDEFF29F912FEE3F955" authorityName="Mamaev" authorityYear="1964" box="[136,326,1678,1704]" class="Insecta" family="Cecidomyiidae" genus="Rhipidoxylomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="69922B209734FFDEFF29F912FEE3F955" box="[136,326,1678,1704]" italics="true" pageId="5" pageNumber="95">Rhipidoxylomyia</emphasis>
</taxonomicName>
in the form of head, 2+10-segmented antennae (instead 2+12-segmented as fossil species
<taxonomicName id="9CE68CB19734FFDEFD53F92EFF1BF912" authorityName="Fedotova &amp; Perkovsky" authorityYear="2008" class="Insecta" family="Cecidomyiidae" genus="Rhipidoxylomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="95" phylum="Arthropoda" rank="species" species="vaga">
<emphasis id="69922B209734FFDEFD53F92EFCA6F936" box="[754,771,1714,1739]" italics="true" pageId="5" pageNumber="95">R</emphasis>
.
<emphasis id="69922B209734FFDEFF29F94AFF1BF912" box="[136,190,1750,1775]" italics="true" pageId="5" pageNumber="95">vaga</emphasis>
</taxonomicName>
and
<taxonomicName id="9CE68CB19734FFDEFF50F94AFED9F912" authorityName="Nel" authorityYear="2021" box="[241,380,1749,1775]" class="Insecta" family="Cecidomyiidae" genus="Rhipidoxylomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="95" phylum="Arthropoda" rank="species" species="rasnitsyni">
<emphasis id="69922B209734FFDEFF50F94AFEA7F912" box="[241,258,1750,1775]" italics="true" pageId="5" pageNumber="95">R</emphasis>
.
<emphasis id="69922B209734FFDEFEAFF94AFED9F912" box="[270,380,1750,1775]" italics="true" pageId="5" pageNumber="95">rasnitsyni</emphasis>
</taxonomicName>
) elongated segments of mouth parts with long lingua and palpi; very large eyes; absence of narrow eye bridge; different shape of flagellomeres; very short neck of flagellomeres; presence of very rare long setae of medial whorl on basal enlargement; presence of hair-shaped sensoria of middle flagellomeres; wider and shorter wing, vein R5 terminating near of tip of the wing and smaller body size; very long ovipositor with strongly enlarged apical plates.
</paragraph>
<paragraph id="5B59F7329734FFDEFF1CF78EFA91FEDF" blockId="5.[136,778,158,2093]" lastBlockId="5.[810,1452,158,290]" pageId="5" pageNumber="95">
The similar genus
<taxonomicName id="9CE68CB19734FFDEFE08F78EFDC2F7D1" authorityName="Mamaev" authorityYear="1964" box="[425,615,2066,2092]" class="Insecta" family="Cecidomyiidae" genus="Rhipidoxylomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="69922B209734FFDEFE08F78EFDC2F7D1" box="[425,615,2066,2092]" italics="true" pageId="5" pageNumber="95">Rhipidoxylomyia</emphasis>
</taxonomicName>
known from Holarctic and Oriental Region includes 16 extant species and 2 fossils (
<bibRefCitation id="3F778AC39734FFDEFC7AFF5EFAAEFF21" author="Gagne, R. J. &amp; Jaschhof, M." box="[987,1291,193,220]" pageId="5" pageNumber="95" refId="ref6067" refString="Gagne, R. J. &amp; Jaschhof, M. (2021) A catalog of the Cecidomyiidae (Diptera) of the world. 5 th Edition. Digital. 813 pp. https: // www. ars. usda. gov / ARSUserFiles / 80420580 / Gagne _" type="book" year="2021">Gagné &amp; Jaschhof, 2021</bibRefCitation>
;
<bibRefCitation id="3F778AC39734FFDEFABEFF5DFC4DFF02" author="Fedotova, Z. A. &amp; Perkovsky, E. E." pageId="5" pageNumber="95" pagination="27 - 40" refId="ref5529" refString="Fedotova, Z. A. &amp; Perkovsky, E. E. (2008) New taxa of gall midges (Diptera, Cecidomyiidae) from Dubrovitsa (Rovno amber). Vestnik Zoologii, 42 (1), 27 - 40. https: // doi. org / 10.1007 / s 11492 - 008 - 2007 - 6" type="journal article" year="2008">Fedotova &amp; Perkovsky, 2008</bibRefCitation>
;
<bibRefCitation id="3F778AC39734FFDEFC57FF79FBCFFF02" author="Nel, A." box="[1014,1130,229,255]" pageId="5" pageNumber="95" pagination="620 - 628" refId="ref7014" refString="Nel, A. (2021) New Cecidomyiidae from the Lowermost Eocene French amber (Diptera). Palaeoentomology, 4 (6), 620 - 628. https: // doi. org / 10.11646 / palaeoentomology. 4.6.11" type="journal article" year="2021">Nel, 2021</bibRefCitation>
). Larvae of the three known European species are found in decaying wood.
</paragraph>
</subSubSection>
</treatment>
</document>