244 lines
24 KiB
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244 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.879.37684" ID-GBIF-Dataset="b0dc35b9-f482-44c7-86ee-9cace2d308f5" ID-PMC="PMC6795606" ID-Pensoft-Pub="1313-2970-879-33" ID-Pensoft-UUID="19C55ED885CC5DD797A08E86976CB190" ID-PubMed="31636498" ID-ZooBank="668885D2C2184402B4308672EC98E81E" ModsDocID="1313-2970-879-33" checkinTime="1570746154165" checkinUser="pensoft" docAuthor="Sokolov, Igor M." docDate="2019" docId="C1B6AF79563C50C8A5E1049E0FD7DBAF" docLanguage="en" docName="ZooKeys 879: 33-56" docOrigin="ZooKeys 879" docSource="http://dx.doi.org/10.3897/zookeys.879.37684" docTitle="Pelodiaetus sulcatipennis Jeannel" docType="treatment" docVersion="5" id="19C55ED885CC5DD797A08E86976CB190" lastPageNumber="48" masterDocId="19C55ED885CC5DD797A08E86976CB190" masterDocTitle="A review of the genus Pelodiaetus Jeannel (Coleoptera, Carabidae, Anillini) of New Zealand, with re-description of the genus, description of a new species, and notes on the evolutionary history" masterLastPageNumber="56" masterPageNumber="33" pageNumber="44" updateTime="1668167867040" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the genus Pelodiaetus Jeannel (Coleoptera, Carabidae, Anillini) of New Zealand, with re-description of the genus, description of a new species, and notes on the evolutionary history</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sokolov, Igor M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>879</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>56</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.879.37684</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.879.37684</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-879-33</mods:identifier>
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<mods:identifier type="ZooBank">668885D2C2184402B4308672EC98E81E</mods:identifier>
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<mods:identifier type="Pensoft-UUID">19C55ED885CC5DD797A08E86976CB190</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159690802" LSID="urn:lsid:plazi:treatment:C1B6AF79563C50C8A5E1049E0FD7DBAF" httpUri="http://treatment.plazi.org/id/C1B6AF79563C50C8A5E1049E0FD7DBAF" lastPageId="15" lastPageNumber="48" pageId="11" pageNumber="44">
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<subSubSection pageId="11" pageNumber="44" type="nomenclature">
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<paragraph pageId="11" pageNumber="44">
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<taxonomicName LSID="C1B6AF79-563C-50C8-A5E1-049E0FD7DBAF" authority="Jeannel" class="Insecta" family="Carabidae" genus="Pelodiaetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelodiaetus sulcatipennis" order="Coleoptera" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="sulcatipennis">
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<pageBreakToken pageId="11" pageNumber="44" start="start">Pelodiaetus</pageBreakToken>
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sulcatipennis Jeannel
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="45" type="reference_group">
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<paragraph pageId="12" pageNumber="45">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pelodiaetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelodiaetus sulcatipennis" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="sulcatipennis">
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<pageBreakToken pageId="12" pageNumber="45" start="start">Pelodiaetus</pageBreakToken>
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sulcatipennis
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</taxonomicName>
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Jeannel, 1937: 277 (original description);
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<bibRefCitation author="Jeannel, R" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="21" pageNumber="54" pagination="33 - 204" refId="B19" refString="Jeannel, R, 1963. Monographie des "Anillini", Bembidiides endoges [ColeopteraTrechidae]. Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie 28: 33 - 204" title="Monographie des " Anillini ", Bembidiides endoges [ColeopteraTrechidae]." volume="28" year="1963">Jeannel 1963</bibRefCitation>
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(figures, key, distribution);
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<bibRefCitation DOI="https://doi.org/10.1080/03014223.1980.10423793" author="Moore, BP" journalOrPublisher="New Zealand Journal of Zoology" pageId="22" pageNumber="55" pagination="399 - 406" refId="B25" refString="Moore, BP, 1980. A synopsis of the New Zealand Anillini (Coleoptera: Carabidae: Bembidiinae), with descriptions of new genera and species. New Zealand Journal of Zoology 7: 399 - 406" title="A synopsis of the New Zealand Anillini (Coleoptera: Carabidae: Bembidiinae), with descriptions of new genera and species." url="https://doi.org/10.1080/03014223.1980.10423793" volume="7" year="1980">Moore 1980</bibRefCitation>
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(key, distribution).
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</paragraph>
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<paragraph pageId="12" pageNumber="45">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pelodiaetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelodiaetus lewisi" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="lewisi">Pelodiaetus lewisi</taxonomicName>
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Jeannel, 1937: 277 (original description),
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<emphasis bold="true" pageId="12" pageNumber="45">syn. nov.</emphasis>
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;
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<bibRefCitation author="Jeannel, R" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="21" pageNumber="54" pagination="33 - 204" refId="B19" refString="Jeannel, R, 1963. Monographie des "Anillini", Bembidiides endoges [ColeopteraTrechidae]. Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie 28: 33 - 204" title="Monographie des " Anillini ", Bembidiides endoges [ColeopteraTrechidae]." volume="28" year="1963">Jeannel 1963</bibRefCitation>
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(key, distribution);
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<bibRefCitation DOI="https://doi.org/10.1080/03014223.1980.10423793" author="Moore, BP" journalOrPublisher="New Zealand Journal of Zoology" pageId="22" pageNumber="55" pagination="399 - 406" refId="B25" refString="Moore, BP, 1980. A synopsis of the New Zealand Anillini (Coleoptera: Carabidae: Bembidiinae), with descriptions of new genera and species. New Zealand Journal of Zoology 7: 399 - 406" title="A synopsis of the New Zealand Anillini (Coleoptera: Carabidae: Bembidiinae), with descriptions of new genera and species." url="https://doi.org/10.1080/03014223.1980.10423793" volume="7" year="1980">Moore 1980</bibRefCitation>
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(key, distribution).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="14" lastPageNumber="47" pageId="12" pageNumber="45" type="materials_examined">
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<paragraph pageId="12" pageNumber="45">Investigated material.</paragraph>
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<paragraph pageId="12" pageNumber="45">Stewart Island: Ulva Island Nov-Dec 2003 /coastal forest litter (5 specimens).</paragraph>
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<paragraph pageId="13" pageNumber="46">
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<pageBreakToken pageId="13" pageNumber="46" start="start">Southland</pageBreakToken>
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: Tussock Creek Forest Hill Res 1 Sep 07 / Washed soil sample. Wet Podocarp/broadleaf forest (4 specimens); New Zealand SL Forest Hill Res nr Edendale 1 Sep 07 / Washed soil sample. Kahikitea forest (3 specimens); New Zealand SL Alton Burn Tuatapere SR 16 Nov 08 / Washed soil sample, totara/beech forest / NZMS 260 D45: 992419 38m (4 specimens); New Zealand SL Tuatapere Domain Tuatapere SR 16 Nov 08 / Washed soil sample, beech forest / NZMS 260 D45: 993403 30m (1 specimen); New Zealand SL Tuatapere Scenic Reserve at Tuatapere Domain, 30m J.Nunn 16 Nov 08 / Molecular voucher # 104 Sokolov I.M 2009 (1 specimen); New Zealand SL Tuatapere Scenic Reserve at Tuatapere Domain, 30m J.Nunn 16 Nov 08 / Molecular voucher # 107 Sokolov I.M 2009 (1 specimen); New Zealand SL Bog Burn Taringatura Forest 2 Jan 09 / Washed soil sample, beech forest / NZMS 260 E45: 397596 250m (1 specimen); New Zealand SL Dunsdale Stream, Hokonui Hills, 110m, washed soil sample J.Nunn 26 April 08 / Molecular voucher # 100 Sokolov I.M 2009 (1 specimen); New Zealand MC Pudding Hill Reserve Mt. Hutt 700m J.Nunn 25 Oct 08 / Molecular voucher # 105 Sokolov I.M 2009 (1 specimen); SL Chloris Pass, Catlins 17 Jun 06 (7 specimens); Rakahouka 2 Sep 07 /Neaopanax forest (2 specimens).
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</paragraph>
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<paragraph pageId="13" pageNumber="46">Dunedin: Woodside Glen, Outram 18 Nov 06 (3 specimens); Sutton, Salt Lake 21 Jun 09 / tussock and improved pasture (2 specimens); Picnic Gulley, Taieri Mouth 18 May 06 / under stone after rain (29 specimens); New Zealand DN Start of Government Tck Waipori Valley / Washed soil sample 17 Dec 06 (1 specimen).</paragraph>
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<paragraph pageId="14" pageNumber="47">
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<pageBreakToken pageId="14" pageNumber="47" start="start">Central</pageBreakToken>
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Otago: Logan Burn 900m 13 Dec 1982 / ex.
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<taxonomicName class="Liliopsida" family="Cyperaceae" genus="Oreobolus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Oreobolus pectinatus" order="Poales" pageId="14" pageNumber="47" phylum="Tracheophyta" rank="species" species="pectinatus">
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<emphasis italics="true" pageId="14" pageNumber="47">Oreobolus pectinatus</emphasis>
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</taxonomicName>
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(3 specimens); Cromwell, Kawarau Gorge, Roaring Meg 500m 12 Mar 1959 (2 specimens); Garden V., Raggedy Range 10 Sept 68 / Raoulia (1 specimen); Alexandra Hills, the Knobbies 10 Sept 68 / Raoulia (1 specimen).
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</paragraph>
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<paragraph pageId="14" pageNumber="47">South Canterbury: Guns Bush Waimate 23 Dec 06 / Washed soil sample, broadleaf forest (1 female).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="15" lastPageNumber="48" pageId="14" pageNumber="47" type="discussion">
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<paragraph pageId="14" pageNumber="47">Discussion.</paragraph>
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<paragraph pageId="14" pageNumber="47">
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As mentioned above, the genus
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<taxonomicName class="Insecta" family="Carabidae" genus="Pelodiaetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelodiaetus" order="Coleoptera" pageId="14" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="14" pageNumber="47">Pelodiaetus</emphasis>
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</taxonomicName>
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was established by
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<bibRefCitation author="Jeannel, R" journalOrPublisher="Revue francaise d'Entomologie" pageId="21" pageNumber="54" pagination="241 - 339" refId="B18" refString="Jeannel, R, 1937. Les Bembidiides endoges (Col. Carabidae). Revue francaise d'Entomologie 3: 241 - 339" title="Les Bembidiides endoges (Col. Carabidae)." volume="3" year="1937">Jeannel (1937)</bibRefCitation>
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for two species:
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<taxonomicName lsidName="P. sulcatipennis" pageId="14" pageNumber="47" rank="species" species="sulcatipennis">
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<emphasis italics="true" pageId="14" pageNumber="47">P. sulcatipennis</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="P. lewisi" pageId="14" pageNumber="47" rank="species" species="lewisi">
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<emphasis italics="true" pageId="14" pageNumber="47">P. lewisi</emphasis>
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</taxonomicName>
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. Both species were collected together in Dunedin, Otago, by G Lewis, and, according to
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<bibRefCitation author="Jeannel, R" journalOrPublisher="Revue francaise d'Entomologie" pageId="21" pageNumber="54" pagination="241 - 339" refId="B18" refString="Jeannel, R, 1937. Les Bembidiides endoges (Col. Carabidae). Revue francaise d'Entomologie 3: 241 - 339" title="Les Bembidiides endoges (Col. Carabidae)." volume="3" year="1937">Jeannel (1937</bibRefCitation>
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,
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<bibRefCitation author="Jeannel, R" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="21" pageNumber="54" pagination="33 - 204" refId="B19" refString="Jeannel, R, 1963. Monographie des "Anillini", Bembidiides endoges [ColeopteraTrechidae]. Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie 28: 33 - 204" title="Monographie des " Anillini ", Bembidiides endoges [ColeopteraTrechidae]." volume="28" year="1963">1963</bibRefCitation>
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), they can be distinguished from each other by the shape of the pronotum and elytra. In their descriptions, Jeannel did not provide comparison of the genitalia of the two species, and obviously did not make the necessary measurements for comparison of above-mentioned body parts. He simply restricted description to two couplets of a key with very general diagnoses: a more transverse and rounded pronotum with shorter and more convex elytra for
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<taxonomicName lsidName="P. sulcatipennis" pageId="14" pageNumber="47" rank="species" species="sulcatipennis">
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<emphasis italics="true" pageId="14" pageNumber="47">P. sulcatipennis</emphasis>
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</taxonomicName>
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, and a correspondingly less transverse and rounded pronotum, and narrower and more elongated elytra, for
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<taxonomicName lsidName="P. lewisi" pageId="14" pageNumber="47" rank="species" species="lewisi">
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<emphasis italics="true" pageId="14" pageNumber="47">P. lewisi</emphasis>
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</taxonomicName>
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. This short and very general comparison, together with the same type locality, made the validity of two forms questionable. Later,
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<bibRefCitation DOI="https://doi.org/10.1080/03014223.1980.10423793" author="Moore, BP" journalOrPublisher="New Zealand Journal of Zoology" pageId="22" pageNumber="55" pagination="399 - 406" refId="B25" refString="Moore, BP, 1980. A synopsis of the New Zealand Anillini (Coleoptera: Carabidae: Bembidiinae), with descriptions of new genera and species. New Zealand Journal of Zoology 7: 399 - 406" title="A synopsis of the New Zealand Anillini (Coleoptera: Carabidae: Bembidiinae), with descriptions of new genera and species." url="https://doi.org/10.1080/03014223.1980.10423793" volume="7" year="1980">Moore (1980)</bibRefCitation>
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, reviewing the New Zealand
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<taxonomicName lsidName="" pageId="14" pageNumber="47" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
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, paid attention to the presence of intermediates in the body shape between the two species and stated that both names may belong to extremes of the same species, but did not formally synonymize these taxa because of the scarcity of the available material.
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</paragraph>
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<paragraph lastPageId="15" lastPageNumber="48" pageId="14" pageNumber="47">
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Preparing this taxonomic review, I had an opportunity to investigate many more representatives of the genus. For analysis of the variation in body part proportions, measurements of 57 members of the genus were completed, including seven specimens of
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<taxonomicName lsidName="P. nunni" pageId="14" pageNumber="47" rank="species" species="nunni">
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<emphasis italics="true" pageId="14" pageNumber="47">P. nunni</emphasis>
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</taxonomicName>
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and 50 specimens of
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<taxonomicName lsidName="P. sulcatipennis" pageId="14" pageNumber="47" rank="species" species="sulcatipennis">
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<emphasis italics="true" pageId="14" pageNumber="47">P. sulcatipennis</emphasis>
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</taxonomicName>
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. Because of the wide range of
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<taxonomicName lsidName="P. sulcatipennis" pageId="14" pageNumber="47" rank="species" species="sulcatipennis">
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<emphasis italics="true" pageId="14" pageNumber="47">P. sulcatipennis</emphasis>
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</taxonomicName>
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, the aim of this investigation was to compare main body ratios between the northeastern and southwestern parts of its range. All investigated ratios showed no difference between the two populations of
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<taxonomicName lsidName="P. sulcatipennis" pageId="14" pageNumber="47" rank="species" species="sulcatipennis">
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<emphasis italics="true" pageId="14" pageNumber="47">P. sulcatipennis</emphasis>
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</taxonomicName>
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, as well as between the
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<normalizedToken originalValue="latter’s">latter's</normalizedToken>
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populations and the representatives of
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<taxonomicName lsidName="P. nunni." pageId="14" pageNumber="47" rank="species" species="nunni.">
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<emphasis italics="true" pageId="14" pageNumber="47">P. nunni.</emphasis>
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</taxonomicName>
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At the same time, many ratios showed rather high interspecies variability; this was especially true for W/L - ratio of maximal width to length along the midline of the pronotum, one of the characters used by Jeannel for distinguishing his species. According to the obtained data, the variation in the proportions of the pronotum (
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Variation in proportions of pronotum among different taxa and populations of Pelodiaetus. A - P. nunni; B - P. sulcatipennis, mid-eastern populations; C - P. sulcatipennis, south-western populations. Map of southern half of South Island, New Zealand, shows localities from where specimens were measured. Each species location dot (yellow or white) is identical to Fig. 8 but they are additionally marked with the appropriate bar graph color: pink, orange, and blue. X-axis values represent W / L - ratio of maximal width to length along the midline of pronotum. Red curve - expected normal distribution with sample mean and standard deviation. Elevation scale bars are given in meters." figureDoi="10.3897/zookeys.879.37684.figure9" httpUri="https://binary.pensoft.net/fig/345434" pageId="14" pageNumber="47" tableDoi="10.3897/zookeys.879.37684.figure9">Fig. 9</figureCitation>
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) is similar across different species of
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<taxonomicName class="Insecta" family="Carabidae" genus="Pelodiaetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelodiaetus" order="Coleoptera" pageId="14" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="14" pageNumber="47">Pelodiaetus</emphasis>
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</taxonomicName>
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(T-test for independent groups, p = 0.693, n = 7 for
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<taxonomicName lsidName="P. nunni" pageId="14" pageNumber="47" rank="species" species="nunni">
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<emphasis italics="true" pageId="14" pageNumber="47">P. nunni</emphasis>
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</taxonomicName>
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, and n = 50 for
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<taxonomicName lsidName="P. sulcatipennis" pageId="14" pageNumber="47" rank="species" species="sulcatipennis">
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<emphasis italics="true" pageId="14" pageNumber="47">P. sulcatipennis</emphasis>
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</taxonomicName>
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) and between different populations of
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<taxonomicName lsidName="P. sulcatipennis" pageId="14" pageNumber="47" rank="species" species="sulcatipennis">
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<emphasis italics="true" pageId="14" pageNumber="47">P. sulcatipennis</emphasis>
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</taxonomicName>
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(T-test for independent groups, p = 0.095, n = 23 for mid-eastern, and n = 27 for south-western populations). The only difference between populations of
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<taxonomicName lsidName="P. sulcatipennis" pageId="14" pageNumber="47" rank="species" species="sulcatipennis">
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<emphasis italics="true" pageId="14" pageNumber="47">P. sulcatipennis</emphasis>
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</taxonomicName>
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can be seen in the light shift towards the prevalence of specimens with a slightly more transverse pronotum in the southwestern part of range. However, this deviation lies within the error range and is not statistically significant (see above). The type locality of
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<taxonomicName lsidName="P. sulcatipennis" pageId="14" pageNumber="47" rank="species" species="sulcatipennis">
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<emphasis italics="true" pageId="14" pageNumber="47">P. sulcatipennis</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="P. lewisi" pageId="14" pageNumber="47" rank="species" species="lewisi">
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<emphasis italics="true" pageId="14" pageNumber="47">P. lewisi</emphasis>
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</taxonomicName>
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(Dunedin, Otago) is located exactly within the range of the mid-eastern population, thus supporting the point of view of Moore that both
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<normalizedToken originalValue="Jeannel’s">Jeannel's</normalizedToken>
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species may belong to extremes of a single species. In addition, in investigating male genitalia, I was unable to find any constant difference in the shape of the median lobe and in the armature of the internal sac. On the basis of this evidence,
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<taxonomicName lsidName="P. sulcatipennis" pageId="15" pageNumber="48" rank="species" species="sulcatipennis">
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<emphasis italics="true" pageId="15" pageNumber="48">
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<pageBreakToken pageId="15" pageNumber="48" start="start">P</pageBreakToken>
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. sulcatipennis
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</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="P. lewisi" pageId="15" pageNumber="48" rank="species" species="lewisi">
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<emphasis italics="true" pageId="15" pageNumber="48">P. lewisi</emphasis>
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</taxonomicName>
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should be considered as variations of the same species and be synonymized under the name of
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<taxonomicName class="Insecta" family="Carabidae" genus="Pelodiaetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelodiaetus sulcatipennis" order="Coleoptera" pageId="15" pageNumber="48" phylum="Arthropoda" rank="species" species="sulcatipennis">
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<emphasis italics="true" pageId="15" pageNumber="48">Pelodiaetus sulcatipennis</emphasis>
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</taxonomicName>
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Jeannel.
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</paragraph>
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<caption doi="10.3897/zookeys.879.37684.figure8" httpUri="https://binary.pensoft.net/fig/345433" pageId="15" pageNumber="48" start="Figure 8" startId="F8">
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<paragraph pageId="15" pageNumber="48">
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<emphasis bold="true" pageId="15" pageNumber="48">Figure 8.</emphasis>
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Map of southern half of South Island, New Zealand, showing positions of locality records for the species of
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<taxonomicName class="Insecta" family="Carabidae" genus="Pelodiaetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelodiaetus" order="Coleoptera" pageId="15" pageNumber="48" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="15" pageNumber="48">Pelodiaetus</emphasis>
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</taxonomicName>
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:
|
||
<taxonomicName lsidName="P. nunni" pageId="15" pageNumber="48" rank="species" species="nunni">
|
||
<emphasis italics="true" pageId="15" pageNumber="48">P. nunni</emphasis>
|
||
</taxonomicName>
|
||
, white circles;
|
||
<taxonomicName lsidName="P. sulcatipennis" pageId="15" pageNumber="48" rank="species" species="sulcatipennis">
|
||
<emphasis italics="true" pageId="15" pageNumber="48">P. sulcatipennis</emphasis>
|
||
</taxonomicName>
|
||
, yellow circles. Elevation scale bars are given in meters.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.879.37684.figure9" httpUri="https://binary.pensoft.net/fig/345434" pageId="15" pageNumber="48" start="Figure 9" startId="F9">
|
||
<paragraph pageId="15" pageNumber="48">
|
||
<emphasis bold="true" pageId="15" pageNumber="48">Figure 9.</emphasis>
|
||
Variation in proportions of pronotum among different taxa and populations of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Pelodiaetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelodiaetus" order="Coleoptera" pageId="15" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="48">Pelodiaetus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="15" pageNumber="48">A</emphasis>
|
||
-
|
||
<taxonomicName lsidName="P. nunni" pageId="15" pageNumber="48" rank="species" species="nunni">
|
||
<emphasis italics="true" pageId="15" pageNumber="48">P. nunni</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis bold="true" pageId="15" pageNumber="48">B</emphasis>
|
||
-
|
||
<taxonomicName lsidName="P. sulcatipennis" pageId="15" pageNumber="48" rank="species" species="sulcatipennis">
|
||
<emphasis italics="true" pageId="15" pageNumber="48">P. sulcatipennis</emphasis>
|
||
</taxonomicName>
|
||
, mid-eastern populations;
|
||
<emphasis bold="true" pageId="15" pageNumber="48">C</emphasis>
|
||
-
|
||
<taxonomicName lsidName="P. sulcatipennis" pageId="15" pageNumber="48" rank="species" species="sulcatipennis">
|
||
<emphasis italics="true" pageId="15" pageNumber="48">P. sulcatipennis</emphasis>
|
||
</taxonomicName>
|
||
, south-western populations. Map of southern half of South Island, New Zealand, shows localities from where specimens were measured. Each species location dot (yellow or white) is identical to
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Map of southern half of South Island, New Zealand, showing positions of locality records for the species of Pelodiaetus: P. nunni, white circles; P. sulcatipennis, yellow circles. Elevation scale bars are given in meters." figureDoi="10.3897/zookeys.879.37684.figure8" httpUri="https://binary.pensoft.net/fig/345433" pageId="15" pageNumber="48" tableDoi="10.3897/zookeys.879.37684.figure8">Fig. 8</figureCitation>
|
||
but they are additionally marked with the appropriate bar graph color: pink, orange, and blue. X-axis values represent W/L - ratio of maximal width to length along the midline of pronotum. Red curve - expected normal distribution with sample mean and standard deviation. Elevation scale bars are given in meters.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |