251 lines
26 KiB
XML
251 lines
26 KiB
XML
<document id="074F901791DE092CBFDDFE7156B381BB" ID-DOI="10.11646/zootaxa.4061.1.7" ID-GBIF-Dataset="267ffc8e-f126-43f3-badb-adb07b0984d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="257921" ID-ZooBank="6E5A0C08-460B-4646-9C5B-E3EC1EAC9668" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458391255366" checkinUser="plazi" docAuthor="Kataev, Boris M." docDate="2016" docId="847687C3FFCFFFE4FF4AFDEAFDB7FD74" docLanguage="en" docName="zootaxa.4061.1.7.pdf" docOrigin="Zootaxa 4061 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bottchrus Jedlicka 1935" docType="treatment" docVersion="8" masterDocId="784FFFBBFFCEFFE6FFDDFFF4FFB3FFF1" masterDocTitle="On taxonomic status of Bottchrus, with a description of a new brachypterous species of the genus Trichotichnus (Coleoptera: Carabidae) from the Philippines" updateTime="1698185509331" updateUser="plazi">
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<mods:mods id="CD2EEF4AA1A71DE01D405E9ACFEE7AB4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="42614EEDC2132F556C43C269B7E248E0">
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<mods:title id="050BF7EDD19C799E19E59FD111CD21DE">On taxonomic status of Bottchrus, with a description of a new brachypterous species of the genus Trichotichnus (Coleoptera: Carabidae) from the Philippines</mods:title>
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</mods:titleInfo>
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<mods:name id="8A8685C8C23F41F09C48DBC3A546C9F3" type="personal">
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<mods:role id="FD55ED8F65C380375812D97C3BB19FA9">
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<mods:roleTerm id="378FC8FFE36747D768F3A02C72C52D5C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0C69CD3E9C0E9EA5CF85A5A2D1D9BBBE">Kataev, Boris M.</mods:namePart>
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<mods:typeOfResource id="0CB2F514FD4EAB1DD40630F2FF0BFE7D">text</mods:typeOfResource>
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<mods:titleInfo id="B0548D3B531F3FE2435971613595F484">
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<mods:title id="39A977001F2D91C49486F83CF9DBE8D4">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E32208A43990E184DCD7942C757AD3E2">
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<mods:date id="3F16AED46E7C97D6DE1A074AE2AD8EBA">2016</mods:date>
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<mods:detail id="F37F478BC36F073C100B08D071E3D965" type="volume">
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<mods:number id="BFEDB4921F4B29EECF585712DBE5FF28">4061</mods:number>
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</mods:detail>
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<mods:detail id="2A48A908F9B3F117DA3CD1FF61345FF7" type="issue">
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<mods:number id="896A04042F5382CAE98FBF5544CE70B6">1</mods:number>
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</mods:detail>
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</mods:part>
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<mods:classification id="2DED7AA863EC4E5992228E148FF2B1FC">journal volume</mods:classification>
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<mods:identifier id="5AD58C2ECE6770E35FBFEAC053B39356" type="DOI">10.11646/zootaxa.4061.1.7</mods:identifier>
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<mods:identifier id="98D7BAC8E63C7E44482FBA83AC50375D" type="GBIF-Dataset">267ffc8e-f126-43f3-badb-adb07b0984d8</mods:identifier>
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<mods:identifier id="B074CC672CFA64C82A0F9F4781B1930E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="402966B039B5CC3E086AFB515F7F7131" type="Zenodo-Dep">257921</mods:identifier>
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</mods:mods>
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<treatment id="847687C3FFCFFFE4FF4AFDEAFDB7FD74" ID-DOI="http://doi.org/10.5281/zenodo.6084997" ID-GBIF-Taxon="119208562" ID-Zenodo-Dep="6084997" LSID="urn:lsid:plazi:treatment:847687C3FFCFFFE4FF4AFDEAFDB7FD74" httpUri="http://treatment.plazi.org/id/847687C3FFCFFFE4FF4AFDEAFDB7FD74" lastPageId="2" pageId="1">
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<subSubSection id="44C5655EFFCFFFE7FF4AFDEAFD6FFDC9" box="[151,732,542,568]" pageId="1" type="nomenclature">
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<paragraph id="0C6036D5FFCFFFE7FF4AFDEAFD6FFDC9" blockId="1.[151,732,542,568]" box="[151,732,542,568]" pageId="1">
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||
<heading id="572881B9FFCFFFE7FF4AFDEAFD6FFDC9" bold="true" box="[151,732,542,568]" fontSize="11" level="1" pageId="1" reason="1">
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<emphasis id="3EABEAC7FFCFFFE7FF4AFDEAFD6FFDC9" bold="true" box="[151,732,542,568]" pageId="1">
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The subgenus
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<taxonomicName id="CBDF4D56FFCFFFE7FE97FDEAFDC8FDC9" authority="Jedlicka, 1935" authorityName="Jedlicka" authorityYear="1935" box="[330,635,542,568]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="3EABEAC7FFCFFFE7FE97FDEAFE72FDC9" bold="true" box="[330,449,542,568]" italics="true" pageId="1">Bottchrus</emphasis>
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Jedlička, 1935
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</taxonomicName>
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, stat. n.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="44C5655EFFCFFFE7FF4AFD91FE2BFD07" pageId="1" type="description">
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<paragraph id="0C6036D5FFCFFFE7FF4AFD91FCF0FD6B" blockId="1.[151,1436,613,758]" pageId="1">
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<taxonomicName id="CBDF4D56FFCFFFE7FF4AFD91FE1FFD8A" authority="Jedlicka, 1935: 8" authorityName="Jedlicka" authorityPageNumber="8" authorityYear="1935" box="[151,428,613,635]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="3EABEAC7FFCFFFE7FF4AFD91FF49FD8A" box="[151,250,613,635]" italics="true" pageId="1">Bottchrus</emphasis>
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Jedlička, 1935: 8
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</taxonomicName>
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[as genus].
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<typeStatus id="D3648877FFCFFFE7FDFBFD92FDEBFD8A" box="[550,600,614,635]" pageId="1">Type</typeStatus>
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species:
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<taxonomicName id="CBDF4D56FFCFFFE7FD68FD91FB98FD8A" authority="Jedlicka, 1935" authorityName="Jedlicka" authorityYear="1935" box="[693,1067,613,635]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="philippinus">
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<emphasis id="3EABEAC7FFCFFFE7FD68FD91FC23FD8A" box="[693,912,613,635]" italics="true" pageId="1">Bottchrus philippinus</emphasis>
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Jedlička, 1935
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</taxonomicName>
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, by monotypy.
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<taxonomicName id="CBDF4D56FFCFFFE7FF4AFD70FEC4FD6B" box="[151,375,644,666]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="javanus">
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<emphasis id="3EABEAC7FFCFFFE7FF4AFD70FEC4FD6B" box="[151,375,644,666]" italics="true" pageId="1">Trichotichnus javanus</emphasis>
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</taxonomicName>
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species group sensu Schauberger, 1935: 108.
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</paragraph>
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<paragraph id="0C6036D5FFCFFFE7FF4AFD56FE2BFD07" blockId="1.[151,1436,613,758]" pageId="1">
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<emphasis id="3EABEAC7FFCFFFE7FF4AFD56FEBBFD49" box="[151,264,674,696]" italics="true" pageId="1">Bellogenus</emphasis>
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Clarke, 1971: 264 [as genus],
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<emphasis id="3EABEAC7FFCFFFE7FDE3FD57FD30FD49" bold="true" box="[574,643,675,696]" pageId="1">syn. n.</emphasis>
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<typeStatus id="D3648877FFCFFFE7FD57FD57FD0FFD49" box="[650,700,675,696]" pageId="1">Type</typeStatus>
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species:
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<emphasis id="3EABEAC7FFCFFFE7FCC4FD56FC3AFD49" box="[793,905,674,696]" italics="true" pageId="1">Bellogenus</emphasis>
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<taxonomicName id="CBDF4D56FFCFFFE7FC4DFD57FBC1FD49" authority="Clarke, 1971" authorityName="Clarke" authorityYear="1971" box="[912,1138,674,696]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="amazeus">
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<emphasis id="3EABEAC7FFCFFFE7FC4DFD57FC54FD49" box="[912,999,675,696]" italics="true" pageId="1">amazeus</emphasis>
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Clarke, 1971
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</taxonomicName>
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, by original designation.
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<taxonomicName id="CBDF4D56FFCFFFE7FF4AFD35FDACFD26" authority="Habu, 1973: 225" authorityName="Habu" authorityPageNumber="225" authorityYear="1973" box="[151,543,705,727]" class="Insecta" family="Carabidae" genus="Pseudotrichotichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="3EABEAC7FFCFFFE7FF4AFD35FED4FD26" box="[151,359,705,727]" italics="true" pageId="1">Pseudotrichotichnus</emphasis>
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Habu, 1973: 225
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</taxonomicName>
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[as a subgenus of
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<taxonomicName id="CBDF4D56FFCFFFE7FD34FD35FCC7FD26" box="[745,884,705,727]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="3EABEAC7FFCFFFE7FD34FD35FCC7FD26" box="[745,884,705,727]" italics="true" pageId="1">Trichotichnus</emphasis>
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</taxonomicName>
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].
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<typeStatus id="D3648877FFCFFFE7FC51FD36FC0DFD26" box="[908,958,706,727]" pageId="1">Type</typeStatus>
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species:
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<taxonomicName id="CBDF4D56FFCFFFE7FBFDFD35FAC6FD26" authority="Habu, 1954" authorityName="Habu" authorityYear="1954" box="[1056,1397,705,727]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="nanus">
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<emphasis id="3EABEAC7FFCFFFE7FBFDFD35FB42FD26" box="[1056,1265,705,727]" italics="true" pageId="1">Trichotichnus nanus</emphasis>
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Habu, 1954
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</taxonomicName>
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, by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="44C5655EFFCFFFE7FF4AFCD6FF6AFA32" pageId="1" type="diagnosis">
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<paragraph id="0C6036D5FFCFFFE7FF4AFCD6FCCDFAC3" blockId="1.[151,1436,802,2015]" pageId="1">
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<emphasis id="3EABEAC7FFCFFFE7FF4AFCD6FEB9FCCA" bold="true" box="[151,266,802,827]" pageId="1">Diagnosis</emphasis>
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. The most distinctive feature of
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<taxonomicName id="CBDF4D56FFCFFFE7FD58FCD6FD41FCCA" box="[645,754,802,827]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="3EABEAC7FFCFFFE7FD58FCD6FD41FCCA" box="[645,754,802,827]" italics="true" pageId="1">Bottchrus</emphasis>
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</taxonomicName>
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is the clypeo-ocular furrows deep throughout, reaching the deep furrows around the inner margin of eyes. By this character this subgenus is rather easily distinguished from other subgenera of
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<taxonomicName id="CBDF4D56FFCFFFE7FEAEFC9EFDBEFC72" box="[371,525,874,899]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="3EABEAC7FFCFFFE7FEAEFC9EFDBEFC72" box="[371,525,874,899]" italics="true" pageId="1">Trichotichnus</emphasis>
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</taxonomicName>
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. Members of
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<taxonomicName id="CBDF4D56FFCFFFE7FD73FC9EFCA8FC72" box="[686,795,874,899]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="3EABEAC7FFCFFFE7FD73FC9EFCA8FC72" box="[686,795,874,899]" italics="true" pageId="1">Bottchrus</emphasis>
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</taxonomicName>
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are also characterized by the following character states: mentum with acute median tooth, either separated from submentum by complete transverse suture or fused laterally and separated medially; epilobes narrow, with inner margin more or less straight; ligular sclerite narrow at apex, with two ventral setae near apical margin; paraglossae moderately wide or somewhat narrow, rounded at apex, separated apically from ligular sclerite by narrow notch; genae wide; elytral intervals impunctate and glabrous; abdominal sternites with very short and fine pubescence or glabrous; last visible abdominal sternite in male and female with two pairs of marginal setigerous pores; metafemur with two (rarely three) setigerous pores along posterior margin; male pro- and mesotarsi with ventral biseriate vestiture; median lobe of aedeagus with apical orifice shifted leftwards; and apical stylomere with one or two peg-like spines on both ventral and dorsal edges of outer margin. In most species the males have a well developed apical capitulum of the median lobe of aedeagus, and mesotarsomere 1 not widened, slightly shorter than 2+3, without adhesive vestiture ventrally. Setae on genae, mentioned by Habu (1973) as one of the distinctive characters of
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<taxonomicName id="CBDF4D56FFCFFFE7FC56FB02FBC2FAFE" box="[907,1137,1270,1295]" class="Insecta" family="Carabidae" genus="Pseudotrichotichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="3EABEAC7FFCFFFE7FC56FB02FBC2FAFE" box="[907,1137,1270,1295]" italics="true" pageId="1">Pseudotrichotichnus</emphasis>
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</taxonomicName>
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, are present only in some members and should be excluded from the diagnosis of this taxon.
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</paragraph>
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<paragraph id="0C6036D5FFCFFFE7FF1AFACAFF6AFA32" blockId="1.[151,1436,802,2015]" pageId="1">
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<emphasis id="3EABEAC7FFCFFFE7FF1AFACAFD98FAA6" bold="true" box="[199,555,1342,1367]" pageId="1">Composition and distribution</emphasis>
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. This subgenus includes about 50 described species, with most distributed in the Southeastern Asia from
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<collectingCountry id="74C87645FFCFFFE7FE10FA96FD89FA8A" box="[461,570,1378,1403]" name="Sri Lanka" pageId="1">Sri Lanka</collectingCountry>
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and
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<collectingCountry id="74C87645FFCFFFE7FDB2FA96FD51FA8A" box="[623,738,1378,1403]" name="India" pageId="1">Hindustan</collectingCountry>
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to
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<collectingCountry id="74C87645FFCFFFE7FCD8FA96FCDCFA8A" box="[773,879,1378,1403]" name="Australia" pageId="1">Australia</collectingCountry>
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. One species,
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<taxonomicName id="CBDF4D56FFCFFFE7FBD1FA90FA9FFA8A" authority="Clarke 1971" authorityName="Clarke" authorityYear="1971" box="[1036,1324,1378,1403]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="amazeus">
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<emphasis id="3EABEAC7FFCFFFE7FBD1FA90FB3BFA8A" box="[1036,1160,1380,1403]" italics="true" pageId="1">T. amazeus</emphasis>
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(Clarke 1971)
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</taxonomicName>
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, is known from
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<collectingCountry id="74C87645FFCFFFE7FF0BFA72FE8AFA6E" box="[214,313,1414,1439]" name="Ethiopia" pageId="1">Ethiopia</collectingCountry>
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. The taxonomic position of some species described from New
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<collectingCountry id="74C87645FFCFFFE7FBD1FA72FBEEFA6E" box="[1036,1117,1414,1439]" name="Guinea-Bissau" pageId="1">Guinea</collectingCountry>
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and
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<collectingCountry id="74C87645FFCFFFE7FB45FA72FB4CFA6E" box="[1176,1279,1414,1439]" name="Australia" pageId="1">Australia</collectingCountry>
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needs further study.
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</paragraph>
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</subSubSection>
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<subSubSection id="44C5655EFFCFFFE4FF1AFA3AFDB7FD74" lastPageId="2" pageId="1" type="discussion">
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<paragraph id="0C6036D5FFCFFFE4FF1AFA3AFEA3FE04" blockId="1.[151,1436,802,2015]" lastBlockId="2.[151,1437,151,645]" lastPageId="2" pageId="1">
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<emphasis id="3EABEAC7FFCFFFE7FF1AFA3AFE88FA16" bold="true" box="[199,315,1486,1511]" pageId="1">Remarks.</emphasis>
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Schauberger (1935) was the first to recognize this taxon with 11 Oriental species placed in the
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<taxonomicName id="CBDF4D56FFCFFFE7FF4AFA00FF5DF9FA" box="[151,238,1524,1547]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="javanus">
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<emphasis id="3EABEAC7FFCFFFE7FF4AFA00FF5DF9FA" box="[151,238,1524,1547]" italics="true" pageId="1">javanus</emphasis>
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</taxonomicName>
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species group of the genus
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<taxonomicName id="CBDF4D56FFCFFFE7FDEBFA06FD63F9FA" box="[566,720,1522,1547]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="3EABEAC7FFCFFFE7FDEBFA06FD63F9FA" box="[566,720,1522,1547]" italics="true" pageId="1">Trichotichnus</emphasis>
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</taxonomicName>
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. More recently, Habu (1973) described the same taxon as the subgenus
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<taxonomicName id="CBDF4D56FFCFFFE7FED7F9E2FE43F9DE" box="[266,496,1558,1583]" class="Insecta" family="Carabidae" genus="Pseudotrichotichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
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<emphasis id="3EABEAC7FFCFFFE7FED7F9E2FE43F9DE" box="[266,496,1558,1583]" italics="true" pageId="1">Pseudotrichotichnus</emphasis>
|
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</taxonomicName>
|
||
originally for three species of
|
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<taxonomicName id="CBDF4D56FFCFFFE7FC82F9E2FC4AF9DE" box="[863,1017,1558,1583]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
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<emphasis id="3EABEAC7FFCFFFE7FC82F9E2FC4AF9DE" box="[863,1017,1558,1583]" italics="true" pageId="1">Trichotichnus</emphasis>
|
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</taxonomicName>
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from
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<collectingCountry id="74C87645FFCFFFE7FB98F9E3FB36F9DF" box="[1093,1157,1559,1582]" name="Japan" pageId="1">Japan</collectingCountry>
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and later (Habu 1975a, 1975b, 1980) described several new species from
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<collectingCountry id="74C87645FFCFFFE7FD38F9CEFC84F9A2" box="[741,823,1594,1619]" name="Taiwan" pageId="1">Taiwan</collectingCountry>
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and Ryukyus. Noonan (1985) in his revision of the Selenophori group synonymized
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<taxonomicName id="CBDF4D56FFCFFFE7FDD7F9AAFD43F986" box="[522,752,1630,1655]" class="Insecta" family="Carabidae" genus="Pseudotrichotichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="3EABEAC7FFCFFFE7FDD7F9AAFD43F986" box="[522,752,1630,1655]" italics="true" pageId="1">Pseudotrichotichnus</emphasis>
|
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</taxonomicName>
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with
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<emphasis id="3EABEAC7FFCFFFE7FCEDF9AAFC1FF986" box="[816,940,1630,1655]" italics="true" pageId="1">Bellogenus</emphasis>
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, which was described by Clarke (1971) as a monotypical genus from
|
||
<collectingCountry id="74C87645FFCFFFE7FE6AF976FDA5F96A" box="[439,534,1666,1691]" name="Ethiopia" pageId="1">Ethiopia</collectingCountry>
|
||
without comparison with any Asian species. This synonymy was accepted by Kirschenhofer (1992), Lorenz (1998, 2005) and Bousquet (2012). For unknown reasons Ito (1991, 1996, 1997a, 1997b, 1997c, 1998a, 1998b, 1999, 2009, 2014) continued to describe new Oriental species as members of the subgenus
|
||
<taxonomicName id="CBDF4D56FFCFFFE7FEDBF91AFE5FF8F6" box="[262,492,1774,1799]" class="Insecta" family="Carabidae" genus="Pseudotrichotichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3EABEAC7FFCFFFE7FEDBF91AFE5FF8F6" box="[262,492,1774,1799]" italics="true" pageId="1">Pseudotrichotichnus</emphasis>
|
||
</taxonomicName>
|
||
and cited this name as valid in the Catalogue of the Palaearctic
|
||
<taxonomicName id="CBDF4D56FFCFFFE7FB63F91AFEFDF8DA" authority="Lobl & Smetana, 2003" authorityName="Lobl & Smetana" authorityYear="2003" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="order">Coleoptera (Löbl & Smetana, 2003)</taxonomicName>
|
||
. My examination of the male and two females of the
|
||
<typeStatus id="D3648877FFCFFFE7FC11F8E7FC4FF8DA" box="[972,1020,1811,1835]" pageId="1">type</typeStatus>
|
||
series of
|
||
<emphasis id="3EABEAC7FFCFFFE7FBAEF8E6FAE9F8DA" box="[1139,1370,1810,1835]" italics="true" pageId="1">
|
||
Bellogenus
|
||
<taxonomicName id="CBDF4D56FFCFFFE7FB27F8E0FAE9F8DA" box="[1274,1370,1812,1835]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="amazeus">amazeus</taxonomicName>
|
||
</emphasis>
|
||
from Badabuna Forest and Belleta Forest,
|
||
<collectingCountry id="74C87645FFCFFFE7FDEFF8C2FD22F8BE" box="[562,657,1846,1871]" name="Ethiopia" pageId="1">Ethiopia</collectingCountry>
|
||
(MRACT, MNHN) and comparison them with the East Asian and the Oriental species of the subgenus
|
||
<taxonomicName id="CBDF4D56FFCFFFE7FDB8F8AEFC4AF882" authority="Habu" authorityName="Habu" box="[613,1017,1882,1907]" class="Insecta" family="Carabidae" genus="Pseudotrichotichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3EABEAC7FFCFFFE7FDB8F8AEFCF8F882" box="[613,843,1882,1907]" italics="true" pageId="1">Pseudotrichotichnus</emphasis>
|
||
sensu Habu
|
||
</taxonomicName>
|
||
and sensu Ito confirmed that
|
||
<taxonomicName id="CBDF4D56FFCFFFE7FF4AF88AFECEF866" box="[151,381,1918,1943]" class="Insecta" family="Carabidae" genus="Pseudotrichotichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3EABEAC7FFCFFFE7FF4AF88AFECEF866" box="[151,381,1918,1943]" italics="true" pageId="1">Pseudotrichotichnus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="3EABEAC7FFCFFFE7FE69F88AFD83F866" box="[436,560,1918,1943]" italics="true" pageId="1">Bellogenus</emphasis>
|
||
are synonyms as stated by Noonan (1985). The Ethiopian species shares with the East Asian and the Oriental species all the distinctive characters of the latter including the deep clypeo-ocular furrows reaching deep furrows around inner margin of eyes, the acute median tooth of mentum, the ligular sclerite narrowed to apex, the apical stylomere with one peg-like spine on both ventral and dorsal edges of the outer margin, and the median lobe of aedeagus with the apical orifice shifted to the left (labium, and male and female genitalia of
|
||
<emphasis id="3EABEAC7FFCCFFE4FEF8FF2BFDBEFF09" box="[293,525,223,248]" italics="true" pageId="2">
|
||
Bellogenus
|
||
<taxonomicName id="CBDF4D56FFCCFFE4FE71FF15FDBEFF09" box="[428,525,225,248]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="amazeus">amazeus</taxonomicName>
|
||
</emphasis>
|
||
are illustrated in
|
||
<figureCitation id="94E42A50FFCCFFE4FD3FFF2BFCF9FF09" box="[738,842,223,248]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,263,1857,1880]" captionTargetBox="[213,1372,706,1820]" captionTargetId="figure@2.[205,1381,700,1834]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Trichotichnus (Bottchrus) amazeus (paratypes). 1, labium; 2, left laterotergite and stylus; 3, stylus; 4 – 5, median lobe of aedeagus. 1 – 2, ventral view; 3 – 4, lateral view, 5, dorsal view. Scales = 0.5 mm (A: Figs 2 – 3; B: Figs 1, 4 – 5)." httpUri="https://zenodo.org/record/257922/files/figure.png" pageId="2">Figs 1–5</figureCitation>
|
||
). Additional characteristics of
|
||
<emphasis id="3EABEAC7FFCCFFE4FB69FF2BFA2FFF09" box="[1204,1436,223,248]" italics="true" pageId="2">
|
||
Bellogenus
|
||
<taxonomicName id="CBDF4D56FFCCFFE4FAE1FF15FA2FFF09" box="[1340,1436,225,248]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="amazeus">amazeus</taxonomicName>
|
||
</emphasis>
|
||
include: left mandible truncate at apex; mentum and submentum separated by complete transverse suture; pronotal apical and basal margins bordered throughout; elytra without a discal pore on interval 3, with rudimentary parascutellar striole and with parascutellar pope; prosternum and abdominal sternites glabrous; metepisternum wider than long; and tarsi with sparse short setae dorsally.
|
||
<taxonomicName id="CBDF4D56FFCCFFE4FCF9FE9BFBA5FE79" box="[804,1046,367,392]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="philippinus">
|
||
<emphasis id="3EABEAC7FFCCFFE4FCF9FE9BFBA5FE79" box="[804,1046,367,392]" italics="true" pageId="2">Bottchrus philippinus</emphasis>
|
||
</taxonomicName>
|
||
described by Jedlička (1935) from the
|
||
<collectingCountry id="74C87645FFCCFFE4FF1DFE60FE8DFE5C" box="[192,318,404,429]" name="Philippines" pageId="2">Philippines</collectingCountry>
|
||
also possesses all the distinctive characters of
|
||
<emphasis id="3EABEAC7FFCCFFE4FC94FE60FC75FE5C" box="[841,966,404,429]" italics="true" pageId="2">Bellogenus</emphasis>
|
||
and
|
||
<taxonomicName id="CBDF4D56FFCCFFE4FC21FE60FB51FE5C" box="[1020,1250,404,429]" class="Insecta" family="Carabidae" genus="Pseudotrichotichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3EABEAC7FFCCFFE4FC21FE60FB51FE5C" box="[1020,1250,404,429]" italics="true" pageId="2">Pseudotrichotichnus</emphasis>
|
||
</taxonomicName>
|
||
. Thus
|
||
<taxonomicName id="CBDF4D56FFCCFFE4FAF2FE60FA2FFE5C" box="[1327,1436,404,429]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3EABEAC7FFCCFFE4FAF2FE60FA2FFE5C" box="[1327,1436,404,429]" italics="true" pageId="2">Bottchrus</emphasis>
|
||
</taxonomicName>
|
||
becomes the valid name of the subgenus discussed and
|
||
<emphasis id="3EABEAC7FFCCFFE4FC95FE43FC76FE21" box="[840,965,439,464]" italics="true" pageId="2">Bellogenus</emphasis>
|
||
and
|
||
<taxonomicName id="CBDF4D56FFCCFFE4FBD7FE43FB43FE21" box="[1034,1264,439,464]" class="Insecta" family="Carabidae" genus="Pseudotrichotichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3EABEAC7FFCCFFE4FBD7FE43FB43FE21" box="[1034,1264,439,464]" italics="true" pageId="2">Pseudotrichotichnus</emphasis>
|
||
</taxonomicName>
|
||
are its junior synonyms.
|
||
</paragraph>
|
||
<paragraph id="0C6036D5FFCCFFE4FF1AFE0BFDB7FD74" blockId="2.[151,1437,151,645]" pageId="2">
|
||
The subgenus
|
||
<taxonomicName id="CBDF4D56FFCCFFE4FEA9FE0BFE52FDE9" box="[372,481,511,536]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3EABEAC7FFCCFFE4FEA9FE0BFE52FDE9" box="[372,481,511,536]" italics="true" pageId="2">Bottchrus</emphasis>
|
||
</taxonomicName>
|
||
, having the deep clypeo-ocular furrows reaching the deep furrows around inner margin of eyes, is rather isolated from other members of
|
||
<taxonomicName id="CBDF4D56FFCCFFE4FCC8FDD0FC1CFDCC" box="[789,943,548,573]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3EABEAC7FFCCFFE4FCC8FDD0FC1CFDCC" box="[789,943,548,573]" italics="true" pageId="2">Trichotichnus</emphasis>
|
||
</taxonomicName>
|
||
and further investigations may demonstrate that it warrants a generic status. In the present paper, its subgeneric status is conserved because many species of
|
||
<taxonomicName id="CBDF4D56FFCCFFE4FF4AFD98FE82FD74" box="[151,305,620,645]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3EABEAC7FFCCFFE4FF4AFD98FE82FD74" box="[151,305,620,645]" italics="true" pageId="2">Trichotichnus</emphasis>
|
||
</taxonomicName>
|
||
should be revised.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |