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<document id="DBFD28444A8AA8BD6CE1812D1B7084B0" ID-CLB-Dataset="46124" ID-DOI="10.11646/zootaxa.3784.2.1" ID-GBIF-Dataset="f4530593-bcbe-4092-a140-30de91a6f699" ID-ISSN="1175-5326" ID-Zenodo-Dep="225955" ID-ZooBank="0075CB1A-38B8-4E3E-ABCC-699E3113AAF9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460402627306" checkinUser="plazi" docAuthor="Kaiser, Stefanie" docDate="2014" docId="830310144130FFD4FF183EDAFA5AFEC9" docLanguage="en" docName="zt03784p119.pdf" docOrigin="Zootaxa 3784 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hebefustis vecino Kaiser, 2014, sp. n." docType="treatment" docVersion="10" lastPageNumber="118" masterDocId="7F3A686C413DFFC5FF8F3E4DFFC6FF89" masterDocTitle="New species of Hebefustis Siebenaller &amp; Hessler 1977 (Isopoda, Asellota, Nannoniscidae) from the Clarion Clipperton Fracture Zone (equatorial NE Pacific)" masterLastPageNumber="119" masterPageNumber="101" pageNumber="114" updateTime="1698299397550" updateUser="ExternalLinkService">
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<mods:title id="27FE6DA8A73B8404130C57EC1DD1EFAD">New species of Hebefustis Siebenaller &amp; Hessler 1977 (Isopoda, Asellota, Nannoniscidae) from the Clarion Clipperton Fracture Zone (equatorial NE Pacific)</mods:title>
</mods:titleInfo>
<mods:name id="C10D200583882549E6C3DB4F54D4DCCC" type="personal">
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<mods:roleTerm id="D18BDFEFC1F6E7AA0D559E81FE3C54FB">Author</mods:roleTerm>
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<mods:namePart id="4919494A86E1EE2252FB85D2EA78140B">Kaiser, Stefanie</mods:namePart>
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<mods:title id="9880224584AEDCA2E323F7EDCE12C1FD">Zootaxa</mods:title>
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<mods:part id="6499177482FDE5284429C58FB85E665D">
<mods:date id="E454190047BCD5F27EC4CC9B9EF68A48">2014</mods:date>
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<mods:number id="24E6783F079FDC20A68A94BD0A06D8CF">3784</mods:number>
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<mods:identifier id="FAA7A1043743DCE46070438F0BC864D8" type="DOI">10.11646/zootaxa.3784.2.1</mods:identifier>
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<treatment id="830310144130FFD4FF183EDAFA5AFEC9" ID-DOI="http://doi.org/10.5281/zenodo.5698411" ID-GBIF-Taxon="119420214" ID-Zenodo-Dep="5698411" LSID="urn:lsid:plazi:treatment:830310144130FFD4FF183EDAFA5AFEC9" httpUri="http://treatment.plazi.org/id/830310144130FFD4FF183EDAFA5AFEC9" lastPageId="17" lastPageNumber="118" pageId="13" pageNumber="114">
<subSubSection id="43B0F2894130FFC8FF183EDAFECCFF5A" pageId="13" pageNumber="114" type="nomenclature">
<paragraph id="0B15A1024130FFC8FF183EDAFE70FF3B" blockId="13.[151,438,151,211]" box="[151,438,151,178]" pageId="13" pageNumber="114">
<heading id="505D166E4130FFC8FF183EDAFE70FF3B" bold="true" box="[151,438,151,178]" fontSize="11" level="1" pageId="13" pageNumber="114" reason="1">
<emphasis id="39DE7D104130FFC8FF183EDAFE70FF3B" bold="true" box="[151,438,151,178]" pageId="13" pageNumber="114">
<taxonomicName id="CCAADA814130FFC8FF183EDAFEABFF38" ID-CoL="3JXHD" box="[151,365,151,177]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="vecino" status="sp. nov.">
<emphasis id="39DE7D104130FFC8FF183EDAFEABFF38" bold="true" box="[151,365,151,177]" italics="true" pageId="13" pageNumber="114">Hebefustis vecino</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B4130FFC8FEFB3ED5FE70FF3B" box="[372,438,152,178]" pageId="13" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0B15A1024130FFC8FF183EF7FECCFF5A" blockId="13.[151,438,151,211]" box="[151,266,186,211]" pageId="13" pageNumber="114">
(
<figureCitation id="9391BD874130FFC8FF103EF7FEC4FF5A" box="[159,258,186,211]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="14.[151,250,1974,1996]" captionStartId-1="15.[151,250,1976,1998]" captionTargetBox-0="[245,1335,213,1943]" captionTargetBox-1="[184,1391,211,1943]" captionTargetId-0="figure@14.[234,1355,193,1954]" captionTargetId-1="figure@15.[162,1424,193,1954]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURE 8. Hebefustis vecino sp. n.; A E, holotype female (ZMH-K- 43954); A, habitus, dorsal view. B, antennula, antenna podomere articles 1 4; C, operculum; D, pleotelson, ventral view. E, uropod. Scale bar: A, D = 500 Μm, B C, E = 250 µm." captionText-1="FIGURE 9. Hebefustis vecino sp. n.; A D, holotype female (ZMH-K- 43954); A, habitus, dorsal view. B, maxilliped. C E, pereopods I III, detail: dorsal and ventral claw. Scale bar: A = 500 Μm, B E = 250 µm." httpUri-0="https://zenodo.org/record/225963/files/figure.png" httpUri-1="https://zenodo.org/record/225964/files/figure.png" pageId="13" pageNumber="114">Figs 89</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="43B0F2894130FFC8FF183F4FFCE6FE23" pageId="13" pageNumber="114" type="materials_examined">
<paragraph id="0B15A1024130FFC8FF183F4FFCB4FE92" blockId="13.[151,1437,258,1975]" box="[151,882,258,283]" pageId="13" pageNumber="114">
<emphasis id="39DE7D104130FFC8FF183F4FFEFAFE92" bold="true" box="[151,316,258,283]" pageId="13" pageNumber="114">
<typeStatus id="D4111FA04130FFC8FF183F4FFF14FE92" box="[151,210,258,283]" pageId="13" pageNumber="114">Type</typeStatus>
fixation.
</emphasis>
<typeStatus id="D4111FA04130FFC8FECD3F4FFE6BFE92" box="[322,429,258,283]" pageId="13" pageNumber="114" type="holotype">Holotype</typeStatus>
, non-ovigerous female, here designated.
</paragraph>
<paragraph id="0B15A1024130FFC8FF483F6BFE08FE0E" blockId="13.[151,1437,258,1975]" pageId="13" pageNumber="114">
<emphasis id="39DE7D104130FFC8FF483F6BFE76FEB6" bold="true" box="[199,432,294,319]" pageId="13" pageNumber="114">Material examined.</emphasis>
<emphasis id="39DE7D104130FFC8FE383F6BFDE0FEB7" box="[439,550,294,318]" italics="true" pageId="13" pageNumber="114">
<typeStatus id="D4111FA04130FFC8FE383F6BFDE6FEB7" box="[439,544,294,318]" pageId="13" pageNumber="114" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="1DAC6A8B4130FFC8FDA13F6AFD49FEB6" box="[558,655,294,319]" pageId="13" pageNumber="114" type="female">1 female</specimenCount>
(non-ovigerous),
<collectionCode id="6DBB39C74130FFC8FCD73F6AFC67FEB6" box="[856,929,295,319]" pageId="13" pageNumber="114">CCFZ</collectionCode>
, equatorial NE Pacific, BIONOD expedition, RV LAtalante,
<collectionCode id="6DBB39C74130FFC8FEC73F06FEB8FEEA" box="[328,382,331,355]" pageId="13" pageNumber="114">EBS</collectionCode>
,
<collectionCode id="6DBB39C74130FFC8FE063F07FE7AFEEA" box="[393,444,330,355]" httpUri="http://grbio.org/cool/d4va-b13i" name="University of the Basque Country" pageId="13" pageNumber="114">BIO</collectionCode>
12-51 (start:
<geoCoordinate id="6E9EC7C54130FFC8FDC43F07FD17FEEA" box="[587,721,330,355]" direction="north" orientation="latitude" pageId="13" pageNumber="114" precision="9" value="12.491667">12°29.50N</geoCoordinate>
, 118°13.47,
<quantity id="CC520CE74130FFC8FCE83F07FC79FEEB" box="[871,959,330,355]" metricMagnitude="3" metricUnit="m" metricValue="4.226" pageId="13" pageNumber="114" unit="m" value="4226.0">4226 m</quantity>
;
<geoCoordinate id="6E9EC7C54130FFC8FC443F06FB94FEEA" box="[971,1106,331,355]" direction="north" orientation="latitude" pageId="13" pageNumber="114" precision="9" value="12.523833">12°31.43N</geoCoordinate>
,
<geoCoordinate id="6E9EC7C54130FFC8FBD13F06FB33FEEA" box="[1118,1269,331,355]" direction="west" orientation="longitude" pageId="13" pageNumber="114" precision="9" value="-118.2235">118°13.41W</geoCoordinate>
,
<quantity id="CC520CE74130FFC8FA8F3F06FA90FEEA" box="[1280,1366,331,355]" metricMagnitude="3" metricUnit="m" metricValue="4.234" pageId="13" pageNumber="114" unit="m" value="4234.0">4234 m</quantity>
) date:
<date id="7F1487C24130FFC8FF183F22FEDEFE0E" box="[151,280,367,391]" pageId="13" pageNumber="114" value="2012-04-11">11/04/2012</date>
, ZMH-K-43954
</paragraph>
<paragraph id="0B15A1024130FFC8FF483FDFFCE6FE23" blockId="13.[151,1437,258,1975]" box="[199,800,402,427]" pageId="13" pageNumber="114">
<emphasis id="39DE7D104130FFC8FF483FDFFEA3FE22" bold="true" box="[199,357,402,427]" pageId="13" pageNumber="114">
<typeStatus id="D4111FA04130FFC8FF483FDFFEC4FE22" box="[199,258,402,427]" pageId="13" pageNumber="114">Type</typeStatus>
locality.
</emphasis>
<collectionCode id="6DBB39C74130FFC8FEE33FDEFE73FE22" box="[364,437,403,427]" pageId="13" pageNumber="114">CCFZ</collectionCode>
, Equatorial NE Pacific,
<quantity id="CC520CE74130FFC8FD4C3FDFFCDAFE23" box="[707,796,402,427]" metricMagnitude="3" metricUnit="m" metricValue="4.226" pageId="13" pageNumber="114" unit="m" value="4226.0">4226 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="43B0F2894130FFC8FF483FFBFD0EFE46" box="[199,712,438,463]" pageId="13" pageNumber="114" type="distribution">
<paragraph id="0B15A1024130FFC8FF483FFBFD0EFE46" blockId="13.[151,1437,258,1975]" box="[199,712,438,463]" pageId="13" pageNumber="114">
<emphasis id="39DE7D104130FFC8FF483FFBFEA6FE46" bold="true" box="[199,352,438,463]" pageId="13" pageNumber="114">Distribution.</emphasis>
Only known from
<typeStatus id="D4111FA04130FFC8FDB83FFAFDA1FE46" box="[567,615,439,463]" pageId="13" pageNumber="114">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="43B0F2894130FFC8FF483F97FABAFD9E" pageId="13" pageNumber="114" type="etymology">
<paragraph id="0B15A1024130FFC8FF483F97FABAFD9E" blockId="13.[151,1437,258,1975]" pageId="13" pageNumber="114">
<emphasis id="39DE7D104130FFC8FF483F97FE8AFE7A" bold="true" box="[199,332,474,499]" pageId="13" pageNumber="114">Etymology.</emphasis>
The name (nom., masculine) comes from the Spanish word for neighbouring and emphasizes both the new species morphological resemblance as well as the close vicinity of its
<typeStatus id="D4111FA04130FFC8FB8C3FB2FBF5FD9E" box="[1027,1075,511,535]" pageId="13" pageNumber="114">type</typeStatus>
locality to
<taxonomicName id="CCAADA814130FFC8FB203FB2FAF0FD9F" box="[1199,1334,511,534]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="juansenii" status="sp. nov.">
<emphasis id="39DE7D104130FFC8FB203FB2FAF0FD9F" box="[1199,1334,511,534]" italics="true" pageId="13" pageNumber="114">H. juansenii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B4130FFC8FAB23FB3FABAFD9E" box="[1341,1404,510,535]" pageId="13" pageNumber="114" rank="species">
<emphasis id="39DE7D104130FFC8FAB23FB3FABAFD9E" bold="true" box="[1341,1404,510,535]" pageId="13" pageNumber="114">sp. n.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="43B0F2894130FFC8FF483C6FFB78FD2E" pageId="13" pageNumber="114" type="diagnosis">
<paragraph id="0B15A1024130FFC8FF483C6FFB78FD2E" blockId="13.[151,1437,258,1975]" pageId="13" pageNumber="114">
<emphasis id="39DE7D104130FFC8FF483C6FFE87FDB2" bold="true" box="[199,321,546,571]" pageId="13" pageNumber="114">Diagnosis.</emphasis>
Maxilliped epipodite reaching mid of palpal article 3. Pereonites anterior margins 1
<emphasis id="39DE7D104130FFC8FA9A3C6EFAE5FDB3" box="[1301,1315,547,570]" italics="true" pageId="13" pageNumber="114"></emphasis>
4 frontally directed, rounded; pereonite 2 with few simple setae laterally; pereonite 4 anterior margin slightly concave; pereonites 6 and 7 lateral margins rounded. Pleotelson with two pairs of posterolateral spines; proximal pair inserting 0.7 from anterior margin. Uropods not extending beyond pleotelson posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="43B0F2894130FFC8FF483CFFFA8AFD42" box="[199,1356,690,715]" pageId="13" pageNumber="114" type="discussion">
<paragraph id="0B15A1024130FFC8FF483CFFFA8AFD42" blockId="13.[151,1437,258,1975]" box="[199,1356,690,715]" pageId="13" pageNumber="114">
<emphasis id="39DE7D104130FFC8FF483CFFFEFDFD42" bold="true" box="[199,315,690,715]" pageId="13" pageNumber="114">Remarks.</emphasis>
All appendages of
<typeStatus id="D4111FA04130FFC8FD9B3CFFFDB0FD42" box="[532,630,690,715]" pageId="13" pageNumber="114" type="holotype">holotype</typeStatus>
female were drawn
<emphasis id="39DE7D104130FFC8FCD43CFEFC67FD43" box="[859,929,691,714]" italics="true" pageId="13" pageNumber="114">in situ</emphasis>
to minimize damage of the specimen.
</paragraph>
</subSubSection>
<subSubSection id="43B0F2894130FFD5FF483C9BFED6FC09" lastPageId="16" lastPageNumber="117" pageId="13" pageNumber="114" type="description">
<paragraph id="0B15A1024130FFC8FF483C9BFBD7FBDE" blockId="13.[151,1437,258,1975]" pageId="13" pageNumber="114">
<emphasis id="39DE7D104130FFC8FF483C9BFDF9FD66" bold="true" box="[199,575,726,751]" pageId="13" pageNumber="114">
Description of
<typeStatus id="D4111FA04130FFC8FEF53C9BFE27FD66" box="[378,481,726,751]" pageId="13" pageNumber="114" type="holotype">holotype</typeStatus>
female.
</emphasis>
<emphasis id="39DE7D104130FFC8FDC73C9BFD67FD67" box="[584,673,726,750]" italics="true" pageId="13" pageNumber="114">Habitus</emphasis>
(
<figureCitation id="9391BD874130FFC8FD3E3C9BFD3EFD66" box="[689,760,726,751]" captionStart="FIGURE 8" captionStartId="14.[151,250,1974,1996]" captionTargetBox="[245,1335,213,1943]" captionTargetId="figure@14.[234,1355,193,1954]" captionTargetPageId="14" captionText="FIGURE 8. Hebefustis vecino sp. n.; A E, holotype female (ZMH-K- 43954); A, habitus, dorsal view. B, antennula, antenna podomere articles 1 4; C, operculum; D, pleotelson, ventral view. E, uropod. Scale bar: A, D = 500 Μm, B C, E = 250 µm." httpUri="https://zenodo.org/record/225963/files/figure.png" pageId="13" pageNumber="114">Figs 8</figureCitation>
A, D, 9A). Body length 3.7 pereonite 3.0 width. Surface of dorsal cuticle with numerous setules. Pereonites 13 increasing in length posteriorly; pereonite 1 width 0.9 pereonite 2 width; pereonites 23 with frontally directed rounded antero-lateral margins. Pereonites 2 and 3 of equal width, widest; pereonite 2 width 3.3 length; pereonite 3 longest, width 2.5 length. Pereonites 4 width 0.9 pereonite 2 width, width 3.2 length, with slightly convex anterior margin. Pereonites 57 of similar width; pereonites 6 and 7 medially fused, with rounded lateral margins; pereonite 6 with convex anterior margin. Pleotelson 0.23 body length, length 0.9 width; width 0.9 pereonite 2 width, with two pairs of well-developed posterolateral spines; proximal pair inserting 0.7 from anterior margin, distal pair inserting 0.9 from anterior margin; posterior margin strongly convex; anterior margin straight. Anus (
<figureCitation id="9391BD874130FFC8FB9E3DBBFB9CFB86" box="[1041,1114,1014,1039]" captionStart="FIGURE 2" captionStartId="5.[151,250,1946,1968]" captionTargetBox="[184,1406,217,1880]" captionTargetId="figure@5.[166,1429,193,1897]" captionTargetPageId="5" captionText="FIGURE 2. Hebefustis juansenii sp. n.; A D, holotype female (ZMH K 43951); A, habitus, dorsal view. B, pleotelson, ventral view. C, uropod. D, operculum. Scale bar: A B = 500 Μm, C D = 250 µm." httpUri="https://zenodo.org/record/225957/files/figure.png" pageId="13" pageNumber="114">Fig. 2</figureCitation>
D) covered by anus valves laterally. Uropods inserting closely to the anus valves, short in relation to pleotelson, not projecting beyond posterior margin. Operculum length 0.7 pleotelson length (ventrally measured).
</paragraph>
<paragraph id="0B15A1024130FFC8FF483A2EFCBBFB16" blockId="13.[151,1437,258,1975]" pageId="13" pageNumber="114">
<emphasis id="39DE7D104130FFC8FF483A2EFEA9FBF2" box="[199,367,1123,1147]" italics="true" pageId="13" pageNumber="114">Cephalothorax</emphasis>
(
<figureCitation id="9391BD874130FFC8FE0C3A2FFE09FBF2" box="[387,463,1122,1147]" captionStart="FIGURE 8" captionStartId="14.[151,250,1974,1996]" captionTargetBox="[245,1335,213,1943]" captionTargetId="figure@14.[234,1355,193,1954]" captionTargetPageId="14" captionText="FIGURE 8. Hebefustis vecino sp. n.; A E, holotype female (ZMH-K- 43954); A, habitus, dorsal view. B, antennula, antenna podomere articles 1 4; C, operculum; D, pleotelson, ventral view. E, uropod. Scale bar: A, D = 500 Μm, B C, E = 250 µm." httpUri="https://zenodo.org/record/225963/files/figure.png" pageId="13" pageNumber="114">Figs 8</figureCitation>
A, 9A). Length 0.9 width. Anterior margin slightly rounded, posterior and lateral margins straight. Antennae inserting frontolaterally in a deep fold.
</paragraph>
<paragraph id="0B15A1024130FFC8FF483AE6FED6FADA" blockId="13.[151,1437,258,1975]" pageId="13" pageNumber="114">
<taxonomicName id="CCAADA814130FFC8FF483AE6FEFCFB4A" box="[199,314,1195,1219]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="13" pageNumber="114" phylum="Ascomycota" rank="genus">
<emphasis id="39DE7D104130FFC8FF483AE6FEFCFB4A" box="[199,314,1195,1219]" italics="true" pageId="13" pageNumber="114">Antennula</emphasis>
</taxonomicName>
(
<figureCitation id="9391BD874130FFC8FEC63AE7FE4DFB4A" box="[329,395,1194,1219]" captionStart="FIGURE 8" captionStartId="14.[151,250,1974,1996]" captionTargetBox="[245,1335,213,1943]" captionTargetId="figure@14.[234,1355,193,1954]" captionTargetPageId="14" captionText="FIGURE 8. Hebefustis vecino sp. n.; A E, holotype female (ZMH-K- 43954); A, habitus, dorsal view. B, antennula, antenna podomere articles 1 4; C, operculum; D, pleotelson, ventral view. E, uropod. Scale bar: A, D = 500 Μm, B C, E = 250 µm." httpUri="https://zenodo.org/record/225963/files/figure.png" pageId="13" pageNumber="114">Fig. 8</figureCitation>
B). Length 0.11 body length; with 5 articles. First article circular and broadest; with 1 simple seta distolaterally; second article length 1.5 article 1 length, width 0.4 length; with 2 broom setae distally. Article 3 and 4 equal in length, length 0.2 article 2 length; article 3 with 2 simple setae distally, article 4 with 1 small broom seta distally. Article 5 length 4.8 article 4 length, length 8.0 width; with 1 long aesthetasc and 2 simple setae terminally.
</paragraph>
<paragraph id="0B15A1024130FFC8FF483B12FC48FA12" blockId="13.[151,1437,258,1975]" pageId="13" pageNumber="114">
<emphasis id="39DE7D104130FFC8FF483B12FEE2FAFF" box="[199,292,1375,1398]" italics="true" pageId="13" pageNumber="114">Antenna</emphasis>
(
<figureCitation id="9391BD874130FFC8FEBB3B13FEB0FAFE" box="[308,374,1374,1399]" captionStart="FIGURE 8" captionStartId="14.[151,250,1974,1996]" captionTargetBox="[245,1335,213,1943]" captionTargetId="figure@14.[234,1355,193,1954]" captionTargetPageId="14" captionText="FIGURE 8. Hebefustis vecino sp. n.; A E, holotype female (ZMH-K- 43954); A, habitus, dorsal view. B, antennula, antenna podomere articles 1 4; C, operculum; D, pleotelson, ventral view. E, uropod. Scale bar: A, D = 500 Μm, B C, E = 250 µm." httpUri="https://zenodo.org/record/225963/files/figure.png" pageId="13" pageNumber="114">Fig. 8</figureCitation>
B). Broken off, only podomere articles 14 present. Podomere articles 14 short; article 3 with 1 robust spine tipped with a small setule and 2 simple setae distally.
</paragraph>
<paragraph id="0B15A1024130FFC8FF483BEBFB2DF91E" blockId="13.[151,1437,258,1975]" pageId="13" pageNumber="114">
<emphasis id="39DE7D104130FFC8FF483BEBFE86FA37" box="[199,320,1446,1470]" italics="true" pageId="13" pageNumber="114">Maxilliped</emphasis>
(
<figureCitation id="9391BD874130FFC8FEDF3BEBFE52FA36" box="[336,404,1446,1471]" captionStart="FIGURE 9" captionStartId="15.[151,250,1976,1998]" captionTargetBox="[184,1391,211,1943]" captionTargetId="figure@15.[162,1424,193,1954]" captionTargetPageId="15" captionText="FIGURE 9. Hebefustis vecino sp. n.; A D, holotype female (ZMH-K- 43954); A, habitus, dorsal view. B, maxilliped. C E, pereopods I III, detail: dorsal and ventral claw. Scale bar: A = 500 Μm, B E = 250 µm." httpUri="https://zenodo.org/record/225964/files/figure.png" pageId="13" pageNumber="114">Fig. 9</figureCitation>
B). Left and right maxilliped connected by 3 coupling hooks. Epipodite smooth, triangular, slender, length 3.1 width, reaching mid of palpal article 3. Palpal article 1 short, width 2.5 length; with 1 robust sensory seta laterally. Article 2 longest, length 1.8 article 1 length, width 1.5 length; with 3 simple setae laterally, with 5 simple setae distally. Article 3 length 0.8 article 2 length, width 1.5 length, distal margin saw-toothed; with 5 robust sensillae. Article 4 length 1.5 article 3 length, width 0.5 length; with 3 slender setae distally. Article 5 length 0.6 article 1 length, twice as long as wide; with 3 slender setae terminally. Endite distal margin with some robust, dentate setae and several fine setae laterally. Protopod coxa quadrangular; length 3.2 width.
</paragraph>
<paragraph id="0B15A1024130FFC8FF5F38EEFEE0F83E" blockId="13.[151,1437,258,1975]" pageId="13" pageNumber="114">
<emphasis id="39DE7D104130FFC8FF5F38EEFE89F933" box="[208,335,1699,1723]" italics="true" pageId="13" pageNumber="114">Pereopod I</emphasis>
(
<figureCitation id="9391BD874130FFC8FEEE38EFFE60F932" box="[353,422,1698,1723]" captionStart="FIGURE 9" captionStartId="15.[151,250,1976,1998]" captionTargetBox="[184,1391,211,1943]" captionTargetId="figure@15.[162,1424,193,1954]" captionTargetPageId="15" captionText="FIGURE 9. Hebefustis vecino sp. n.; A D, holotype female (ZMH-K- 43954); A, habitus, dorsal view. B, maxilliped. C E, pereopods I III, detail: dorsal and ventral claw. Scale bar: A = 500 Μm, B E = 250 µm." httpUri="https://zenodo.org/record/225964/files/figure.png" pageId="13" pageNumber="114">Fig. 9</figureCitation>
C) more robust than pereopod II. Basis length 5 width. Ischium about half basis length, length 1.9 width. Merus length 0.6 ischium length, length 1.5 width. Carpus length 1.9 merus length, length 2.7 width; with 2 simple setae dorsally, ventral margin with numerous small setae, membranously embedded, and 4 robust unequally bifid setae in between (1 broken off), with 1 long simple seta distoventrally. Propodus length 0.8 carpus length, 2.5 width; with 3 simple setae dorsally, with numerous small setae, membranously embedded and 2 robust unequally bifid setae in between ventrally. Dactylus length 0.4 propodus length, twice as long as wide; with 4 slender setae medially. Dorsal claw about as long as dactylus; with 2 long, slender setae between dorsal and ventral claw.
</paragraph>
<caption id="5FD5F18A4133FFCBFF1839FBFAB3F862" httpUri="https://zenodo.org/record/225963/files/figure.png" pageId="14" pageNumber="115" targetBox="[245,1335,213,1943]" targetPageId="14">
<paragraph id="0B15A1024133FFCBFF1839FBFAB3F862" blockId="14.[151,1436,1974,2027]" pageId="14" pageNumber="115">
<emphasis id="39DE7D104133FFCBFF1839FBFED2F845" bold="true" box="[151,276,1974,1996]" pageId="14" pageNumber="115">FIGURE 8.</emphasis>
<taxonomicName id="CCAADA814133FFCBFE9339FBFE16F844" box="[284,464,1974,1997]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="vecino" status="sp. nov.">
<emphasis id="39DE7D104133FFCBFE9339FBFE16F844" box="[284,464,1974,1997]" italics="true" pageId="14" pageNumber="115">Hebefustis vecino</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B4133FFCBFE5739FAFDD4F845" box="[472,530,1975,1996]" pageId="14" pageNumber="115" rank="species">
<emphasis id="39DE7D104133FFCBFE5739FAFDD4F845" bold="true" box="[472,530,1975,1996]" pageId="14" pageNumber="115">sp. n.</emphasis>
</taxonomicNameLabel>
; AE, holotype female (ZMH-K-43954); A, habitus, dorsal view. B,
<taxonomicName id="CCAADA814133FFCBFB5239FBFA84F845" box="[1245,1346,1974,1996]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="14" pageNumber="114" phylum="Ascomycota" rank="genus">antennula</taxonomicName>
, antenna podomere articles 14; C, operculum; D, pleotelson, ventral view. E, uropod. Scale bar: A, D = 500 Μm, BC, E = 250 µm.
</paragraph>
</caption>
<caption id="5FD5F18A4132FFCAFF1839F5FBCDF865" httpUri="https://zenodo.org/record/225964/files/figure.png" pageId="15" pageNumber="116" targetBox="[184,1391,211,1943]" targetPageId="15">
<paragraph id="0B15A1024132FFCAFF1839F5FBCDF865" blockId="15.[151,1436,1976,2029]" pageId="15" pageNumber="116">
<emphasis id="39DE7D104132FFCAFF1839F5FED3F844" bold="true" box="[151,277,1976,1998]" pageId="15" pageNumber="116">FIGURE 9.</emphasis>
<taxonomicName id="CCAADA814132FFCAFE9239F5FE15F847" box="[285,467,1976,1998]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="vecino" status="sp. nov.">
<emphasis id="39DE7D104132FFCAFE9239F5FE15F847" box="[285,467,1976,1998]" italics="true" pageId="15" pageNumber="116">Hebefustis vecino</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B4132FFCAFE5439F5FDD0F844" box="[475,534,1976,1997]" pageId="15" pageNumber="116" rank="species">
<emphasis id="39DE7D104132FFCAFE5439F5FDD0F844" bold="true" box="[475,534,1976,1997]" pageId="15" pageNumber="116">sp. n.</emphasis>
</taxonomicNameLabel>
; AD, holotype female (ZMH-K-43954); A, habitus, dorsal view. B, maxilliped. CE, pereopods IIII, detail: dorsal and ventral claw. Scale bar: A = 500 Μm, B
<emphasis id="39DE7D104132FFCAFCF03995FC4DF864" box="[895,907,2008,2029]" italics="true" pageId="15" pageNumber="116"></emphasis>
E = 250 µm.
</paragraph>
</caption>
<paragraph id="0B15A102412DFFD5FF483ED5FC65FE24" blockId="16.[151,1437,151,1220]" pageId="16" pageNumber="117">
<emphasis id="39DE7D10412DFFD5FF483ED5FE88FF39" box="[199,334,152,176]" italics="true" pageId="16" pageNumber="117">Pereopod II</emphasis>
(
<figureCitation id="9391BD87412DFFD5FED03EDAFE62FF39" box="[351,420,151,176]" captionStart="FIGURE 9" captionStartId="15.[151,250,1976,1998]" captionTargetBox="[184,1391,211,1943]" captionTargetId="figure@15.[162,1424,193,1954]" captionTargetPageId="15" captionText="FIGURE 9. Hebefustis vecino sp. n.; A D, holotype female (ZMH-K- 43954); A, habitus, dorsal view. B, maxilliped. C E, pereopods I III, detail: dorsal and ventral claw. Scale bar: A = 500 Μm, B E = 250 µm." httpUri="https://zenodo.org/record/225964/files/figure.png" pageId="16" pageNumber="117">Fig. 9</figureCitation>
D). Basis length 5.6 width; with 1 simple seta ventrally. Ischium half basis length, length twice width; with 1 simple seta dorsally. Merus length 0.75 ischium length, length 1.8 width; with 2 robust setae distodorsally, with 2 more slender setae ventrally. Carpus length 2.3 merus length, length 4.6 width; with 1 simple seta dorsally, with 1 broom seta distodorsally, with numerous small setae, membranously embedded and 6 stout unequally bifid setae in between ventrally. Propodus length 0.8 carpus length, length 5.5 width; with 4 simple setae dorsally, with numerous small setae, membranously embedded, and with 4 stout unequally bifid setae ventrally. Dactylus length one-third propodus length, length 3.1 width; with 3 simple setae medially. Dorsal claw length 0.8 dactylus length; with 2 slender setae between dorsal and ventral claw.
</paragraph>
<paragraph id="0B15A102412DFFD5FF483FF5FB8AFD21" blockId="16.[151,1437,151,1220]" pageId="16" pageNumber="117">
<emphasis id="39DE7D10412DFFD5FF483FF5FE93FE59" box="[199,341,440,464]" italics="true" pageId="16" pageNumber="117">Pereopod III</emphasis>
(
<figureCitation id="9391BD87412DFFD5FEEB3FFAFE61FE59" box="[356,423,439,464]" captionStart="FIGURE 9" captionStartId="15.[151,250,1976,1998]" captionTargetBox="[184,1391,211,1943]" captionTargetId="figure@15.[162,1424,193,1954]" captionTargetPageId="15" captionText="FIGURE 9. Hebefustis vecino sp. n.; A D, holotype female (ZMH-K- 43954); A, habitus, dorsal view. B, maxilliped. C E, pereopods I III, detail: dorsal and ventral claw. Scale bar: A = 500 Μm, B E = 250 µm." httpUri="https://zenodo.org/record/225964/files/figure.png" pageId="16" pageNumber="117">Fig. 9</figureCitation>
E). Basis length 5.4 width. Ischium length 0.6 basis length, length 2.4 width; with 1 simple seta dorsally. Merus length 0.6 ischium length, length 1.8 width. Carpus length 2.4 merus length, length 5.2 width; with 2 simple setae dorsally, with numerous small setae, membranously embedded and 7 stout unequally bifid setae in between ventrally. Propodus length 0.9 carpus length, length 5.8 width; with 3 simple setae dorsally, with 1 broom seta distodorsally, with numerous small setae, membranously embedded, and with 4 stout unequally bifid setae ventrally. Dactylus length one-third of propodus length, length 3 width; with 3 simple setae medially. Dorsal claw about as long as dactylus; with 2 slender setae between dorsal and ventral claw.
</paragraph>
<paragraph id="0B15A102412DFFD5FF483CF9FE2CFD66" blockId="16.[151,1437,151,1220]" pageId="16" pageNumber="117">
<emphasis id="39DE7D10412DFFD5FF483CF9FE82FD45" box="[199,324,692,716]" italics="true" pageId="16" pageNumber="117">Operculum</emphasis>
(female pleopod 2;
<figureCitation id="9391BD87412DFFD5FDAA3CF9FDA1FD45" box="[549,615,692,717]" captionStart="FIGURE 8" captionStartId="14.[151,250,1974,1996]" captionTargetBox="[245,1335,213,1943]" captionTargetId="figure@14.[234,1355,193,1954]" captionTargetPageId="14" captionText="FIGURE 8. Hebefustis vecino sp. n.; A E, holotype female (ZMH-K- 43954); A, habitus, dorsal view. B, antennula, antenna podomere articles 1 4; C, operculum; D, pleotelson, ventral view. E, uropod. Scale bar: A, D = 500 Μm, B C, E = 250 µm." httpUri="https://zenodo.org/record/225963/files/figure.png" pageId="16" pageNumber="117">Fig. 8</figureCitation>
C, D). About as long as wide. Lateral and posterior margin rounded; with numerous (&gt; 50) simple setae.
</paragraph>
<paragraph id="0B15A102412DFFD5FF483CB1FED6FC09" blockId="16.[151,1437,151,1220]" pageId="16" pageNumber="117">
<emphasis id="39DE7D10412DFFD5FF483CB1FEEEFC9D" box="[199,296,764,788]" italics="true" pageId="16" pageNumber="117">Uropods</emphasis>
(
<figureCitation id="9391BD87412DFFD5FEB43CB1FE46FC9D" box="[315,384,764,789]" captionStart="FIGURE 8" captionStartId="14.[151,250,1974,1996]" captionTargetBox="[245,1335,213,1943]" captionTargetId="figure@14.[234,1355,193,1954]" captionTargetPageId="14" captionText="FIGURE 8. Hebefustis vecino sp. n.; A E, holotype female (ZMH-K- 43954); A, habitus, dorsal view. B, antennula, antenna podomere articles 1 4; C, operculum; D, pleotelson, ventral view. E, uropod. Scale bar: A, D = 500 Μm, B C, E = 250 µm." httpUri="https://zenodo.org/record/225963/files/figure.png" pageId="16" pageNumber="117">Fig. 8</figureCitation>
D, E). Biramous. Protopod trapezoid, nearly as long as wide; with 2 simple setae distally. Exopodite about as long as protopod, length 3.2 width; with 1 simple seta terminally. Endopodite length 1.3 exopodite length; with 7 simple setae laterally, with 3 broom setae (one broken off) and 1 long, slender simple seta terminally.
</paragraph>
</subSubSection>
<subSubSection id="43B0F289412DFFD5FF483DC1FE4DFB4D" pageId="16" pageNumber="117" type="discussion">
<paragraph id="0B15A102412DFFD5FF483DC1FB99FB99" blockId="16.[151,1437,151,1220]" pageId="16" pageNumber="117">
<emphasis id="39DE7D10412DFFD5FF483DC1FEFDFC2C" bold="true" box="[199,315,908,933]" pageId="16" pageNumber="117">Remarks.</emphasis>
<taxonomicName id="CCAADA81412DFFD5FECE3DC1FDC1FC2D" box="[321,519,908,932]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="vecino" status="sp. nov.">
<emphasis id="39DE7D10412DFFD5FECE3DC1FDC1FC2D" box="[321,519,908,932]" italics="true" pageId="16" pageNumber="117">Hebefustis vecino</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B412DFFD5FD823DC1FD8BFC2C" box="[525,589,908,933]" pageId="16" pageNumber="117" rank="species">
<emphasis id="39DE7D10412DFFD5FD823DC1FD8BFC2C" bold="true" box="[525,589,908,933]" pageId="16" pageNumber="117">sp. n.</emphasis>
</taxonomicNameLabel>
is only represented by a single female specimen, which highlights a general problem in deep-sea taxonomy. Many deep-sea species are just recorded from a single locality with one or two individuals (e.g.,
<bibRefCitation id="6F3BDCF3412DFFD5FED33D99FDEAFC64" author="Brandt" box="[348,556,980,1005]" pageId="16" pageNumber="117" refString="Brandt, A., Brenke, N., Andres, H. G., Brix, S., Guerrero-Kommritz, J., Muhlenhardt-Siegel, U. &amp; Wagele, J. W. (2005). Diversity of peracarid crustaceans (Malacostraca) from the abyssal plain of the Angola Basin. Organisms Diversity &amp; Evolution, 5, 105 - 112." type="journal article" year="2005">
Brandt
<emphasis id="39DE7D10412DFFD5FE3F3D98FE2FFC65" box="[432,489,980,1004]" italics="true" pageId="16" pageNumber="117">et al.</emphasis>
2005
</bibRefCitation>
;
<bibRefCitation id="6F3BDCF3412DFFD5FDB73D99FCC1FC65" author="Brandt" box="[568,775,980,1005]" pageId="16" pageNumber="117" refString="Brandt, A., Gooday, A. J., Brandao, S. N., Brix, S., Brokeland, W., Cedhagen, T., Choudhury, M., Cornelius, N., Danis, B., De Mesel, I., Diaz, R. J., Gillan, D. C., Hilbig, B., Howe, J., Janussen, D., Kaiser, S., Linse, K., Malyutina, M., Pawlowski, J., Raupach, M. &amp; Vanreusel, A. (2007) First insights into the biodiversity and biogeography of the Southern Ocean deep sea. Nature, 447, 307 - 311. http: // dx. doi. org / 10.1038 / nature 05827" type="journal article" year="2007">
Brandt
<emphasis id="39DE7D10412DFFD5FD033D98FD03FC65" box="[652,709,980,1004]" italics="true" pageId="16" pageNumber="117">et al.</emphasis>
2007
</bibRefCitation>
) representing a major impediment for both taxonomic and molecular examinations (e.g., with regard to the assessment of intraspecific variation).
</paragraph>
<paragraph id="0B15A102412DFFD5FF483A51FE4DFB4D" blockId="16.[151,1437,151,1220]" pageId="16" pageNumber="117">
Although
<taxonomicName id="CCAADA81412DFFD5FEB73A51FE39FBBD" box="[312,511,1052,1076]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="vecino" status="sp. nov.">
<emphasis id="39DE7D10412DFFD5FEB73A51FE39FBBD" box="[312,511,1052,1076]" italics="true" pageId="16" pageNumber="117">Hebefustis vecino</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B412DFFD5FD883A51FD81FBBC" box="[519,583,1052,1077]" pageId="16" pageNumber="117" rank="species">
<emphasis id="39DE7D10412DFFD5FD883A51FD81FBBC" bold="true" box="[519,583,1052,1077]" pageId="16" pageNumber="117">sp. n.</emphasis>
</taxonomicNameLabel>
shares many features with
<taxonomicName id="CCAADA81412DFFD5FCF33A50FBC0FBBD" box="[892,1030,1053,1076]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="juansenii" status="sp. nov.">
<emphasis id="39DE7D10412DFFD5FCF33A50FBC0FBBD" box="[892,1030,1053,1076]" italics="true" pageId="16" pageNumber="117">H. juansenii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B412DFFD5FB823A51FB8BFBBC" box="[1037,1101,1052,1077]" pageId="16" pageNumber="117" rank="species">
<emphasis id="39DE7D10412DFFD5FB823A51FB8BFBBC" bold="true" box="[1037,1101,1052,1077]" pageId="16" pageNumber="117">sp. n.</emphasis>
</taxonomicNameLabel>
, it, nevertheless, shows some significant differences: maxilliped epipodite reaching mid of palpal article 3 (vs. upper third of palpal article
<quantity id="CC520CE7412DFFD5FAFF3A0DFA5AFBD1" box="[1392,1436,1087,1112]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="16" pageNumber="117" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="CCAADA81412DFFD5FF183A28FEE7FBF5" box="[151,289,1125,1148]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="juansenii">
<emphasis id="39DE7D10412DFFD5FF183A28FEE7FBF5" box="[151,289,1125,1148]" italics="true" pageId="16" pageNumber="117">H. juansenii</emphasis>
</taxonomicName>
), maxilliped palpal article 2 with a strong sensory seta distally (vs. no such seta present); pereonite 2 lacking a robust spine on anterolateral margin (vs. robust spine present); pereonite 4 anterior margin slightly concave (vs. convex).
</paragraph>
</subSubSection>
<subSubSection id="43B0F289412DFFD4FF183B5AFA5AFEC9" lastPageId="17" lastPageNumber="118" pageId="16" pageNumber="117" type="key">
<paragraph id="0B15A102412DFFD5FF183B5AFC91FAB9" blockId="16.[151,855,1303,1328]" box="[151,855,1303,1328]" pageId="16" pageNumber="117">
<emphasis id="39DE7D10412DFFD5FF183B5AFDCDFAB9" bold="true" box="[151,523,1303,1328]" pageId="16" pageNumber="117">
Key to the species of
<taxonomicName id="CCAADA81412DFFD5FE1F3B5AFDCDFAB9" ID-CoL="7P5JG" box="[400,523,1303,1328]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="39DE7D10412DFFD5FE1F3B5AFDCDFAB9" bold="true" box="[400,523,1303,1328]" italics="true" pageId="16" pageNumber="117">Hebefustis</emphasis>
</taxonomicName>
</emphasis>
(modified from
<bibRefCitation id="6F3BDCF3412DFFD5FD4A3B5AFC89FAB9" author="Kaiser" box="[709,847,1303,1328]" pageId="16" pageNumber="117" refString="Kaiser, S. (2008) First record and new species of the genus Hebefustis Siebenaller &amp; Hessler, 1977 (Isopoda: Asellota: Nannoniscidae) from the deep Weddell Sea Basin. Zootaxa, 1866, 467 - 481." type="journal article" year="2008">Kaiser 2008</bibRefCitation>
)
</paragraph>
<key id="7F3BF633412DFFD4FF183B12FA5AFEC9" lastPageId="17" lastPageNumber="118" pageId="16" pageNumber="117">
<keyStep id="B05EB9A7412DFFD5FF183B12FA5DFA1B" pageId="16" pageNumber="117">
<paragraph id="0B15A102412DFFD5FF183B12FA5DFAFC" blockId="16.[151,1436,1375,2013]" box="[151,1435,1375,1397]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF183B12FA5DFAFC" box="[151,1435,1375,1397]" pageId="16" pageNumber="117">1a. Pereonite 6 and 7 with acute, distally directed posterolateral margins........................................... 2</keyLead>
</paragraph>
<paragraph id="0B15A102412DFFD5FF183B31FA5DFA1B" blockId="16.[151,1436,1375,2013]" box="[151,1435,1404,1426]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF183B31FA5DFA1B" box="[151,1435,1404,1426]" pageId="16" pageNumber="117">1b. Pereonite 6 and 7 with rounded posterolateral margins....................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="B05EB9A7412DFFD5FF183BD4FA5AFA44" pageId="16" pageNumber="117">
<paragraph id="0B15A102412DFFD5FF183BD4FA5DFA26" blockId="16.[151,1436,1375,2013]" box="[151,1435,1433,1455]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF183BD4FA5DFA26" box="[151,1435,1433,1455]" pageId="16" pageNumber="117">2a. Male pleotelson with rounded posterior margin............................................................. 3</keyLead>
</paragraph>
<paragraph id="0B15A102412DFFD5FF183BFAFA5AFA44" blockId="16.[151,1436,1375,2013]" box="[151,1436,1463,1485]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF183BFAFA5AFA44" box="[151,1436,1463,1485]" pageId="16" pageNumber="117">
2b. Male pleotelson angularly shaped...............................................................
<taxonomicName id="CCAADA81412DFFD5FAA43BF5FA5AFA44" ID-CoL="3JXH3" box="[1323,1436,1464,1485]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="39DE7D10412DFFD5FAA43BF5FA5AFA44" box="[1323,1436,1464,1485]" italics="true" pageId="16" pageNumber="117">H. cornutus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B05EB9A7412DFFD5FF183B99FA5DF98E" pageId="16" pageNumber="117">
<paragraph id="0B15A102412DFFD5FF183B99FA5DFA63" blockId="16.[151,1436,1375,2013]" box="[151,1435,1492,1514]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF183B99FA5DFA63" box="[151,1435,1492,1514]" pageId="16" pageNumber="117">
3a. Pereonite
<quantity id="CC520CE7412DFFD5FECE3B99FEA0FA63" box="[321,358,1492,1514]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="16" pageNumber="117" unit="in" value="2.0">2 in</quantity>
female and/or male with a robust seta or spine on the anterolateral corner............................. 4
</keyLead>
</paragraph>
<paragraph id="0B15A102412DFFD5FF183BBCFA5DF98E" blockId="16.[151,1436,1375,2013]" box="[151,1435,1521,1543]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF183BBCFA5DF98E" box="[151,1435,1521,1543]" pageId="16" pageNumber="117">
3b. Pereonite
<quantity id="CC520CE7412DFFD5FECE3BBCFEA0F98E" box="[321,358,1521,1543]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="16" pageNumber="117" unit="in" value="2.0">2 in</quantity>
female and/or male without a seta or spine on the anterolateral corner....................
<taxonomicName id="CCAADA81412DFFD5FA8C3BBFFA5DF98E" ID-CoL="3JXH5" box="[1283,1435,1521,1543]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="hexadentium">
<emphasis id="39DE7D10412DFFD5FA8C3BBFFA5DF98E" box="[1283,1435,1521,1543]" italics="true" pageId="16" pageNumber="117">H. hexadentium</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B05EB9A7412DFFD5FF183842FA5AF9CB" pageId="16" pageNumber="117">
<paragraph id="0B15A102412DFFD5FF183842FA5DF9AC" blockId="16.[151,1436,1375,2013]" box="[151,1435,1551,1573]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF183842FA5DF9AC" box="[151,1435,1551,1573]" pageId="16" pageNumber="117">4a. Operculum length ≤ 0.2 body length...................................................................... 5</keyLead>
</paragraph>
<paragraph id="0B15A102412DFFD5FF183861FA5AF9CB" blockId="16.[151,1436,1375,2013]" box="[151,1436,1580,1602]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF183861FA5AF9CB" box="[151,1436,1580,1602]" pageId="16" pageNumber="117">
4b. Operculum length&gt; 0.3 body length............................................................... ..
<taxonomicName id="CCAADA81412DFFD5FACB3860FA5AF9CB" ID-CoL="6LGYH" box="[1348,1436,1581,1602]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="vitjazi">
<emphasis id="39DE7D10412DFFD5FACB3860FA5AF9CB" box="[1348,1436,1581,1602]" italics="true" pageId="16" pageNumber="117">H. vitjazi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B05EB9A7412DFFD5FF183804FA5DF9F4" pageId="16" pageNumber="117">
<paragraph id="0B15A102412DFFD5FF183804FA5DF9D6" blockId="16.[151,1436,1375,2013]" box="[151,1435,1609,1631]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF183804FA5DF9D6" box="[151,1435,1609,1631]" pageId="16" pageNumber="117">
5b.
<taxonomicName id="CCAADA81412DFFD5FF6E3804FE83F9D6" box="[225,325,1609,1631]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="16" pageNumber="114" phylum="Ascomycota" rank="genus">Antennula</taxonomicName>
article 5 proximally and distally of equal width.....................................................
<emphasis id="39DE7D10412DFFD5FA003804FA5DF9D6" box="[1423,1435,1609,1631]" italics="true" pageId="16" pageNumber="117">6</emphasis>
</keyLead>
</paragraph>
<paragraph id="0B15A102412DFFD5FF18382AFA5DF9F4" blockId="16.[151,1436,1375,2013]" box="[151,1435,1639,1661]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF18382AFA5DF9F4" box="[151,1435,1639,1661]" pageId="16" pageNumber="117">
5a.
<taxonomicName id="CCAADA81412DFFD5FF6E382AFE83F9F4" box="[225,325,1639,1661]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="16" pageNumber="114" phylum="Ascomycota" rank="genus">Antennula</taxonomicName>
article 5 increasing in width distally.......................................................
<taxonomicName id="CCAADA81412DFFD5FACE3825FA5DF9F4" ID-CoL="3JXH4" box="[1345,1435,1639,1661]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="39DE7D10412DFFD5FACE3825FA5DF9F4" box="[1345,1435,1639,1661]" italics="true" pageId="16" pageNumber="117">H. dispar</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B05EB9A7412DFFD5FF1838C9FA5DF93E" pageId="16" pageNumber="117">
<paragraph id="0B15A102412DFFD5FF1838C9FA5DF913" blockId="16.[151,1436,1375,2013]" box="[151,1435,1668,1690]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF1838C9FA5DF913" box="[151,1435,1668,1690]" pageId="16" pageNumber="117">
6a. Male pleopod 1 distal margin with ≤ 3 setae, male pleopod 2 outer margin with ≥ 11 setae....................
<taxonomicName id="CCAADA81412DFFD5FABE38C8FA5DF913" ID-CoL="3JXH6" box="[1329,1435,1668,1690]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="39DE7D10412DFFD5FABE38C8FA5DF913" box="[1329,1435,1668,1690]" italics="true" pageId="16" pageNumber="117">H. hirsutus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="0B15A102412DFFD5FF1838ECFA5DF93E" blockId="16.[151,1436,1375,2013]" box="[151,1435,1697,1719]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF1838ECFA5DF93E" box="[151,1435,1697,1719]" pageId="16" pageNumber="117">
6b. Male pleopod 1 distal margin with ≥ 4 setae, male pleopod 2 outer margin with ≤ 5 setae.................
<taxonomicName id="CCAADA81412DFFD5FA9D38EFFA5DF93E" ID-CoL="3JXH2" box="[1298,1435,1697,1719]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="clareolithis">
<emphasis id="39DE7D10412DFFD5FA9D38EFFA5DF93E" box="[1298,1435,1697,1719]" italics="true" pageId="16" pageNumber="117">H. clareolithis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B05EB9A7412DFFD5FF1838F2FA5DF97B" pageId="16" pageNumber="117">
<paragraph id="0B15A102412DFFD5FF1838F2FA5DF95C" blockId="16.[151,1436,1375,2013]" box="[151,1435,1727,1749]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF1838F2FA5DF95C" box="[151,1435,1727,1749]" pageId="16" pageNumber="117">7a. Pleotelson with one pair of posterolateral spines............................................................ 8</keyLead>
</paragraph>
<paragraph id="0B15A102412DFFD5FF183891FA5DF97B" blockId="16.[151,1436,1375,2013]" box="[151,1435,1756,1778]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF183891FA5DF97B" box="[151,1435,1756,1778]" pageId="16" pageNumber="117">7b. Pleotelson with two pairs of posterolateral spines.......................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="B05EB9A7412DFFD5FF1838B4FA5DF8A4" pageId="16" pageNumber="117">
<paragraph id="0B15A102412DFFD5FF1838B4FA5DF886" blockId="16.[151,1436,1375,2013]" box="[151,1435,1785,1807]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF1838B4FA5DF886" box="[151,1435,1785,1807]" pageId="16" pageNumber="117">8a. Pleotelson posterolateral spines ≥ 0.8 from anterior margin, body length &lt;3.8 width................................ 9</keyLead>
</paragraph>
<paragraph id="0B15A102412DFFD5FF18395AFA5DF8A4" blockId="16.[151,1436,1375,2013]" box="[151,1435,1815,1837]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF18395AFA5DF8A4" box="[151,1435,1815,1837]" pageId="16" pageNumber="117">8b. Pleotelson posterolateral spines ≤ 0.6 from anterior margin.................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="B05EB9A7412DFFD5FF183979FA5DF8EE" pageId="16" pageNumber="117">
<paragraph id="0B15A102412DFFD5FF183979FA5DF8C3" blockId="16.[151,1436,1375,2013]" box="[151,1435,1844,1866]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF183979FA5DF8C3" box="[151,1435,1844,1866]" pageId="16" pageNumber="117">
9a.
<taxonomicName id="CCAADA81412DFFD5FF6E3979FE83F8C3" box="[225,325,1844,1866]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="16" pageNumber="114" phylum="Ascomycota" rank="genus">Antennula</taxonomicName>
article 5 length approximately article 2 length.................................................
<taxonomicName id="CCAADA81412DFFD5FAD23978FA5DF8C3" ID-CoL="3JXH9" box="[1373,1435,1845,1866]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="par">
<emphasis id="39DE7D10412DFFD5FAD23978FA5DF8C3" box="[1373,1435,1845,1866]" italics="true" pageId="16" pageNumber="117">H. par</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="0B15A102412DFFD5FF18391CFA5DF8EE" blockId="16.[151,1436,1375,2013]" box="[151,1435,1873,1895]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF18391CFA5DF8EE" box="[151,1435,1873,1895]" pageId="16" pageNumber="117">
9b.
<taxonomicName id="CCAADA81412DFFD5FF6E391CFE83F8EE" box="[225,325,1873,1895]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="16" pageNumber="114" phylum="Ascomycota" rank="genus">Antennula</taxonomicName>
article 5 length 0.7 article 2 length.........................................................
<taxonomicName id="CCAADA81412DFFD5FAC8391FFA5DF8EE" ID-CoL="3JXGZ" box="[1351,1435,1873,1895]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="alleni">
<emphasis id="39DE7D10412DFFD5FAC8391FFA5DF8EE" box="[1351,1435,1873,1895]" italics="true" pageId="16" pageNumber="117">H. alleni</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B05EB9A7412DFFD5FF183922FA5DF82B" pageId="16" pageNumber="117">
<paragraph id="0B15A102412DFFD5FF183922FA5DF80C" blockId="16.[151,1436,1375,2013]" box="[151,1435,1903,1925]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF183922FA5DF80C" box="[151,1435,1903,1925]" pageId="16" pageNumber="117">10a. Operculum distal margin softly rounded.................................................................. 11</keyLead>
</paragraph>
<paragraph id="0B15A102412DFFD5FF1839C1FA5DF82B" blockId="16.[151,1436,1375,2013]" box="[151,1435,1932,1954]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF1839C1FA5DF82B" box="[151,1435,1932,1954]" pageId="16" pageNumber="117">
10b. Operculum distal margin straight.................................................................
<taxonomicName id="CCAADA81412DFFD5FAA639C0FA5DF82B" ID-CoL="3JXHB" box="[1321,1435,1932,1954]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="39DE7D10412DFFD5FAA639C0FA5DF82B" box="[1321,1435,1932,1954]" italics="true" pageId="16" pageNumber="117">H. robustus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B05EB9A7412DFFD5FF1839E7FA5DF854" pageId="16" pageNumber="117">
<paragraph id="0B15A102412DFFD5FF1839E7FA5DF849" blockId="16.[151,1436,1375,2013]" box="[151,1435,1962,1984]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF1839E7FA5DF849" box="[151,1435,1962,1984]" pageId="16" pageNumber="117">
11a.
<taxonomicName id="CCAADA81412DFFD5FF6E39E7FE83F849" box="[225,325,1962,1984]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="16" pageNumber="114" phylum="Ascomycota" rank="genus">Antennula</taxonomicName>
article 5 length 4.1 width; pleotelson posterior margin rounded..................................
<taxonomicName id="CCAADA81412DFFD5FAC339E6FA5DF849" ID-CoL="3JXHC" box="[1356,1435,1962,1984]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="vafer">
<emphasis id="39DE7D10412DFFD5FAC339E6FA5DF849" box="[1356,1435,1962,1984]" italics="true" pageId="16" pageNumber="117">H. vafer</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="0B15A102412DFFD5FF18398AFA5DF854" blockId="16.[151,1436,1375,2013]" box="[151,1435,1991,2013]" pageId="16" pageNumber="117">
<keyLead id="B05B0237412DFFD5FF18398AFA5DF854" box="[151,1435,1991,2013]" pageId="16" pageNumber="117">
11b.
<taxonomicName id="CCAADA81412DFFD5FF6E398AFE83F854" box="[225,325,1991,2013]" class="Dothideomycetes" family="Euantennariaceae" genus="Antennula" higherTaxonomySource="GBIF" kingdom="Fungi" order="Capnodiales" pageId="16" pageNumber="114" phylum="Ascomycota" rank="genus">Antennula</taxonomicName>
article 5 length 3.2 width; pleotelson posterior margin tapering distally........................
<taxonomicName id="CCAADA81412DFFD5FA9E3985FA5DF854" ID-CoL="3JXH8" box="[1297,1435,1991,2013]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="117" phylum="Arthropoda" rank="species" species="mollicellus">
<emphasis id="39DE7D10412DFFD5FA9E3985FA5DF854" box="[1297,1435,1991,2013]" italics="true" pageId="16" pageNumber="117">H. mollicellus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B05EB9A7412CFFD4FF183EDAFA5AFF43" pageId="17" pageNumber="118">
<paragraph id="0B15A102412CFFD4FF183EDAFA5DFF24" blockId="17.[151,1436,151,320]" box="[151,1435,151,173]" pageId="17" pageNumber="118">
<keyLead id="B05B0237412CFFD4FF183EDAFA5DFF24" box="[151,1435,151,173]" pageId="17" pageNumber="118">12a. Pereonites 14 anterolateral margins rounded; uropods not visible from dorsal view............................... 13</keyLead>
</paragraph>
<paragraph id="0B15A102412CFFD4FF183EF9FA5AFF43" blockId="17.[151,1436,151,320]" box="[151,1436,180,202]" pageId="17" pageNumber="118">
<keyLead id="B05B0237412CFFD4FF183EF9FA5AFF43" box="[151,1436,180,202]" pageId="17" pageNumber="118">
12b. Pereonites 14 anterolateral margins acute; uropods clearly projecting beyond pleotelson posterior margin.....
<taxonomicName id="CCAADA81412CFFD4FA923EF8FA5AFF43" ID-CoL="6LGYN" box="[1309,1436,181,202]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="primitivus">
<emphasis id="39DE7D10412CFFD4FA923EF8FA5AFF43" box="[1309,1436,181,202]" italics="true" pageId="17" pageNumber="118">H. primitivus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B05EB9A7412CFFD4FF183E9CFA5AFEC9" pageId="17" pageNumber="118">
<paragraph id="0B15A102412CFFD4FF183E9CFA5AFE8C" blockId="17.[151,1436,151,320]" pageId="17" pageNumber="118">
<keyLead id="B05B0237412CFFD4FF183E9CFA5AFE8C" pageId="17" pageNumber="118">
13a. Pereonite 2 antero-lateral margin with a strong spine in both male and female, maxilliped epipodite reaching upper third of pal- pal article 2............................................................................
<taxonomicName id="CCAADA81412CFFD4FB643EBDFAA6FE8C" ID-CoL="3JXH7" box="[1259,1376,240,261]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="juansenii" status="sp. nov.">
<emphasis id="39DE7D10412CFFD4FB643EBDFAA6FE8C" box="[1259,1376,240,261]" italics="true" pageId="17" pageNumber="118">H. juansenii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B412CFFD4FAE93EA2FA5AFE8C" box="[1382,1436,239,261]" pageId="17" pageNumber="118" rank="species">
<emphasis id="39DE7D10412CFFD4FAE93EA2FA5AFE8C" bold="true" box="[1382,1436,239,261]" pageId="17" pageNumber="118">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="0B15A102412CFFD4FF183F41FA5AFEC9" blockId="17.[151,1436,151,320]" pageId="17" pageNumber="118">
<keyLead id="B05B0237412CFFD4FF183F41FA5AFEC9" pageId="17" pageNumber="118">
13b. Pereonite 2 antero-lateral margin without a strong spine, maxilliped epipodite reaching mid of palpal article 3.....................................................................................................
<taxonomicName id="CCAADA81412CFFD4FA8B3F67FA99FEB6" ID-CoL="3JXHD" box="[1284,1375,298,319]" class="Malacostraca" family="Nannoniscidae" genus="Hebefustis" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="vecino" status="sp. nov.">
<emphasis id="39DE7D10412CFFD4FA8B3F67FA99FEB6" box="[1284,1375,298,319]" italics="true" pageId="17" pageNumber="118">H. vecino</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22EDC06B412CFFD4FAE93F67FA5AFEC9" box="[1382,1436,298,320]" pageId="17" pageNumber="118" rank="species">
<emphasis id="39DE7D10412CFFD4FAE93F67FA5AFEC9" bold="true" box="[1382,1436,298,320]" pageId="17" pageNumber="118">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>