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<document id="25556240FD652ACA69CB2EC8A0DE58E4" ID-DOI="10.11646/zootaxa.3985.4.3" ID-GBIF-Dataset="48cc8fed-5ec0-4ece-a82b-c2fa1ddd8893" ID-ISSN="1175-5326" ID-Zenodo-Dep="235350" ID-ZooBank="ECE53B6E-DBAF-4E98-BF8A-96FD9106EF35" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461259944332" checkinUser="plazi" docAuthor="Antić, Dragan Ž., Mock, Andrej &amp; Enghoff, Henrik" docDate="2015" docId="682F87EBAD16074F84F4FC5BFBD85CB9" docLanguage="en" docName="zt03985p540.pdf" docOrigin="Zootaxa 3985 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cibiniulus slovacus Antic, Mock &amp; Enghoff, sp. n." docType="treatment" docVersion="9" lastPageNumber="529" masterDocId="9416FF93AD1707498463FFDEFF865D31" masterDocTitle="Two new species of the millipede family Blaniulidae (Diplopoda, Julida) from caves in central and southeastern Europe" masterLastPageNumber="540" masterPageNumber="523" pageNumber="524" updateTime="1698609266564" updateUser="plazi">
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<mods:title id="6DF18FF25D974A1B632DA44DDF33A6BA">Two new species of the millipede family Blaniulidae (Diplopoda, Julida) from caves in central and southeastern Europe</mods:title>
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<mods:name id="D6056C0B635686EF9F411BAE01363CF7" type="personal">
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<mods:namePart id="C08A22BC64EA465C761F8E9351CA6960">Antić, Dragan Ž.</mods:namePart>
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<mods:name id="96307D079F10952FDC6848EDC74B9C89" type="personal">
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<mods:namePart id="BB59256D3F87DE718A73A5FFAB09BF6E">Mock, Andrej</mods:namePart>
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<mods:role id="2D360ECFE5DAFB734B7BE4E2594BB98B">
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<mods:namePart id="CB6BDB89C61250C695236844C143D7EF">Enghoff, Henrik</mods:namePart>
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<mods:title id="9C80F86D1D86F5B1E37CBF4171F98678">Zootaxa</mods:title>
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<mods:date id="57D91C13F6B36FAC72562FC0B09E54B5">2015</mods:date>
<mods:detail id="534230E9A6289D2FCAC20096253747B1" type="volume">
<mods:number id="BFB851BDB6F33DAD426A3783A32FB273">3985</mods:number>
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<mods:classification id="791F94AF8CDECD64CE0FFF26D25A34B6">journal article</mods:classification>
<mods:identifier id="59D3E5EA7F606732DF157AC4DC95C747" type="DOI">10.11646/zootaxa.3985.4.3</mods:identifier>
<mods:identifier id="6EE458B531BF6D7D4A8C9189C4E71692" type="GBIF-Dataset">48cc8fed-5ec0-4ece-a82b-c2fa1ddd8893</mods:identifier>
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<treatment id="682F87EBAD16074F84F4FC5BFBD85CB9" ID-DOI="http://doi.org/10.5281/zenodo.5620896" ID-GBIF-Taxon="119636792" ID-Zenodo-Dep="5620896" LSID="urn:lsid:plazi:treatment:682F87EBAD16074F84F4FC5BFBD85CB9" httpUri="http://treatment.plazi.org/id/682F87EBAD16074F84F4FC5BFBD85CB9" lastPageId="6" lastPageNumber="529" pageId="1" pageNumber="524">
<subSubSection id="A89C6576AD16074884F4FC5BFE805EF1" pageId="1" pageNumber="524" type="nomenclature">
<paragraph id="E03936FDAD16074884F4FC5BFC855EAE" blockId="1.[151,771,901,960]" box="[151,771,901,927]" pageId="1" pageNumber="524">
<heading id="BB718191AD16074884F4FC5BFC855EAE" bold="true" box="[151,771,901,927]" fontSize="11" level="1" pageId="1" pageNumber="524" reason="1">
<emphasis id="D2F2EAEFAD16074884F4FC5BFC855EAE" bold="true" box="[151,771,901,927]" pageId="1" pageNumber="524">
<taxonomicName id="27864D7EAD16074884F4FC5BFD325EAE" ID-CoL="847K6" authority="Antic, Mock &amp; Enghoff" authorityName="Antic, Mock &amp; Enghoff" box="[151,692,901,927]" class="Diplopoda" family="Blaniulidae" genus="Cibiniulus" kingdom="Animalia" order="Julida" pageId="1" pageNumber="524" phylum="Arthropoda" rank="species" species="slovacus" status="sp. nov.">
<emphasis id="D2F2EAEFAD16074884F4FC5BFE025EAE" bold="true" box="[151,388,901,927]" italics="true" pageId="1" pageNumber="524">Cibiniulus slovacus</emphasis>
Antić, Mock &amp; Enghoff
</taxonomicName>
,
<taxonomicNameLabel id="C9C15794AD16074886A3FC5BFC855EAE" box="[704,771,901,927]" pageId="1" pageNumber="524" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E03936FDAD16074884F4FC79FE805EF1" blockId="1.[151,771,901,960]" box="[151,262,935,960]" pageId="1" pageNumber="524">
<figureCitation id="78BD2A78AD16074884F4FC79FE805EF1" box="[151,262,935,960]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 9" captionStart-2="FIGURES 10 – 12" captionStartId-0="2.[151,264,1933,1955]" captionStartId-1="3.[151,264,1868,1890]" captionStartId-2="4.[151,264,1931,1953]" captionTargetBox-0="[204,1372,195,1912]" captionTargetBox-1="[223,1340,193,1835]" captionTargetBox-2="[207,1356,193,1891]" captionTargetId-0="figure@2.[204,1372,195,1913]" captionTargetId-1="figure@3.[223,1340,193,1837]" captionTargetId-2="figure@4.[207,1357,193,1892]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1 – 4. Cibiniulus slovacus sp. n., paratype male. 1. anterior gonopods, posterior view; 2. anterior gonopods, posterolateral view; 3. anterior gonopods, lateral view; 4. second pair of legs with penis, posterior view. – f: postero-mesal flange, p: penis, t: telopodite, cp: coxal processes, ls: lanceolate seta. Scale lines: 0.1 mm." captionText-1="FIGURES 5 – 9. Cibiniulus slovacus sp. n., paratype male. 5. posterior gonopod, lateral view; 6. apical half of posterior gonopod, lateral view; 7. posterior gonopod, mesal view; 8. left first leg, posterior view; 9. apical part of right first leg, anterior view. a: apophyses / modified setae, s: sternum, cl: claw, dl: dorsal lobe, fs: femoral striations, ll: leaf-shaped lamella, sh: setiferous hump, st: striations on leaf-shaped lamella, ta: tarsus, ti: tibia, tr: triangular area, vl: ventral lobe. Scale lines: 0.1 mm." captionText-2="FIGURES 10 – 12. Cibiniulus slovacus sp. n., 10. holotype male posterior gonopods, lateral and mesal view; 11. holotype male posterior gonopods apex, lateral and mesal view; 12. female vulva, posterior view. s: sternum, dl: dorsal lobe, ll: leaf-shaped lamella, rs: receptaculum seminis, st: striations on leaf-shaped lamella, vl: ventral lobe." httpUri-0="https://zenodo.org/record/235351/files/figure.png" httpUri-1="https://zenodo.org/record/235352/files/figure.png" httpUri-2="https://zenodo.org/record/235353/files/figure.png" pageId="1" pageNumber="524">Figs 1–12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A89C6576AD16074884F4FC31FDA8598C" pageId="1" pageNumber="524" type="materials_examined">
<paragraph id="E03936FDAD16074884F4FC31FDA8598C" blockId="1.[151,1437,1007,1968]" pageId="1" pageNumber="524">
<emphasis id="D2F2EAEFAD16074884F4FC31FEEF5939" bold="true" box="[151,361,1007,1032]" pageId="1" pageNumber="524">Material studied.</emphasis>
<typeStatus id="3F3D885FAD1607488517FC31FE5A5939" box="[372,476,1007,1032]" pageId="1" pageNumber="524" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="8697AE38AD1607488651FC2EFD0C5939" box="[562,650,1008,1032]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="524">ZMUC</collectionCode>
): Jazvínska Cave, Tribeč Mountains,
<collectingCountry id="9891766DAD1607488020FC31FB2E5939" box="[1091,1192,1007,1032]" name="Slovakia" pageId="1" pageNumber="524">Slovakia</collectingCountry>
, pitfall traps, 14.04–
<date id="9438103DAD16074884F4FBCAFF5E591D" box="[151,216,1044,1068]" pageId="1" pageNumber="524" value="2000-10-17">17.10</date>
.2000, leg. Roman Mlejnek.
<typeStatus id="3F3D885FAD160748867FFBCBFD0F591C" box="[540,649,1045,1069]" pageId="1" pageNumber="524" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="8697AE38AD16074886FAFBCAFD77591D" box="[665,753,1044,1068]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="524">ZMUC</collectionCode>
): four females (one fragmented) same data as
<typeStatus id="3F3D885FAD1607488163FBCAFAE0591C" box="[1280,1382,1044,1069]" pageId="1" pageNumber="524" type="holotype">holotype</typeStatus>
; two females: Dobrotínska Cave, Tribeč Mountains,
<collectingCountry id="9891766DAD16074886ADFBE9FCB55961" box="[718,819,1079,1104]" name="Slovakia" pageId="1" pageNumber="524">Slovakia</collectingCountry>
, baited pitfall traps, 14.04–
<date id="9438103DAD16074880E2FBE6FA845961" box="[1153,1282,1080,1104]" pageId="1" pageNumber="524" value="2000-10-17">17.10.2000</date>
, leg. Roman Mlejnek; one female and one male: Gemersko-teplická Cave, Slovak Karst,
<collectingCountry id="9891766DAD1607488067FB82FBEF5944" box="[1028,1129,1116,1141]" name="Slovakia" pageId="1" pageNumber="524">Slovakia</collectingCountry>
,
<date id="9438103DAD1607488015FB82FB715945" box="[1142,1271,1116,1140]" pageId="1" pageNumber="524" value="2003-07-27">27.07.2003</date>
, leg. Vladimír Papáč. Additional material: three females and one juvenile male: Kolónia II Cave, Volovské vrchy Mountain,
<collectingCountry id="9891766DAD16074884F4FB7AFF7A598C" box="[151,252,1188,1213]" name="Slovakia" pageId="1" pageNumber="524">Slovakia</collectingCountry>
, 0 6.03.2010, leg. A. Mock.
</paragraph>
</subSubSection>
<subSubSection id="A89C6576AD16074884A4FB19FD8659D1" box="[199,512,1223,1248]" pageId="1" pageNumber="524" type="etymology">
<paragraph id="E03936FDAD16074884A4FB19FD8659D1" blockId="1.[151,1437,1007,1968]" box="[199,512,1223,1248]" pageId="1" pageNumber="524">
<emphasis id="D2F2EAEFAD16074884A4FB19FEC859D1" bold="true" box="[199,334,1223,1248]" pageId="1" pageNumber="524">Etymology.</emphasis>
After
<collectingCountry id="9891766DAD16074885F5FB19FE7A59D1" box="[406,508,1223,1248]" name="Slovakia" pageId="1" pageNumber="524">Slovakia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A89C6576AD16074884A4FB32FBDD5841" pageId="1" pageNumber="524" type="diagnosis">
<paragraph id="E03936FDAD16074884A4FB32FBDD5841" blockId="1.[151,1437,1007,1968]" pageId="1" pageNumber="524">
<emphasis id="D2F2EAEFAD16074884A4FB32FEC75834" bold="true" box="[199,321,1260,1285]" pageId="1" pageNumber="524">Diagnosis.</emphasis>
Differs from
<taxonomicName id="27864D7EAD16074885BDFB32FD355834" box="[478,691,1260,1285]" class="Diplopoda" family="Blaniulidae" genus="Cibiniulus" kingdom="Animalia" order="Julida" pageId="1" pageNumber="524" phylum="Arthropoda" rank="species" species="phlepsii">
<emphasis id="D2F2EAEFAD16074885BDFB32FD355834" box="[478,691,1260,1285]" italics="true" pageId="1" pageNumber="524">Cibiniulus phlepsii</emphasis>
</taxonomicName>
by numerous aspects in both habitus and posterior gonopods (see
<tableCitation id="AD040346AD16074884F4FAD1FF6E5819" box="[151,232,1295,1320]" captionStart="TABLE 1" captionStartId="5.[151,239,155,178]" captionTargetBox="[159,1406,206,1101]" captionTargetPageId="5" captionText="TABLE 1. Differences between Cibiniulus phlepsii (Verhoeff, 1897) and C. slovacus sp. n." httpUri="http://table.plazi.org/id/B4F96675AD12074C84F4FF45FBE35D83" pageId="1" pageNumber="524" tableUuid="B4F96675AD12074C84F4FF45FBE35D83">Table 1</tableCitation>
), e.g., absence of ocelli and absence of a lateral lamella on the posterior gonopods. Differs from other blind blaniulids occurring in Central Europe by the habitus and setation of vulva: without setation in
<taxonomicName id="27864D7EAD1607488087FAEBFAE1587C" box="[1252,1383,1332,1357]" class="Diplopoda" family="Blaniulidae" genus="Cibiniulus" kingdom="Animalia" order="Julida" pageId="1" pageNumber="524" phylum="Arthropoda" rank="species" species="slovacus">
<emphasis id="D2F2EAEFAD1607488087FAEBFAE1587C" box="[1252,1383,1332,1357]" italics="true" pageId="1" pageNumber="524">C. slovacus</emphasis>
</taxonomicName>
, but setose in
<emphasis id="D2F2EAEFAD160748849CFA87FD935841" box="[255,533,1367,1392]" italics="true" pageId="1" pageNumber="524">
<taxonomicName id="27864D7EAD160748849CFA87FE175841" box="[255,401,1367,1392]" class="Diplopoda" family="Blaniulidae" genus="Blaniulus" kingdom="Animalia" order="Julida" pageId="1" pageNumber="524" phylum="Arthropoda" rank="species" species="guttulatus">B. guttulatus</taxonomicName>
,
<taxonomicName id="27864D7EAD16074885FFFA86FD935841" box="[412,533,1367,1392]" class="Diplopoda" family="Blaniulidae" genus="Archiboreoiulus" kingdom="Animalia" order="Julida" pageId="1" pageNumber="524" phylum="Arthropoda" rank="species" species="pallidus">A. pallidus</taxonomicName>
</emphasis>
and
<taxonomicName id="27864D7EAD160748862FFA87FD285841" box="[588,686,1369,1392]" class="Diplopoda" family="Blaniulidae" genus="Boreoiulus" kingdom="Animalia" order="Julida" pageId="1" pageNumber="524" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="D2F2EAEFAD160748862FFA87FD285841" box="[588,686,1369,1392]" italics="true" pageId="1" pageNumber="524">B. tenuis</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="84174B0CAD1607488681FA89FC2B5841" author="Broelemann" box="[738,941,1367,1392]" pageId="1" pageNumber="524" refString="Broelemann, H. W. (1923) Biospeologica XLVIII. Blaniulidae, Myriapodes (Premiere Serie). Archives de Zoologie Experimentale et generale, 61, 99 - 453, pls. I - XVI." type="journal article" year="1923">Broelemann 1923</bibRefCitation>
;
<bibRefCitation id="84174B0CAD16074887DAFA89FBC85841" author="Blower" box="[953,1102,1367,1392]" pageId="1" pageNumber="524" refString="Blower, J. G. (1985) Millipedes. Synopses of the British Fauna, New Series, 35, 1 - 242." type="journal article" year="1985">Blower 1985</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A89C6576AD16074F84A4FAA2FB7C5C54" lastPageId="6" lastPageNumber="529" pageId="1" pageNumber="524" type="description">
<paragraph id="E03936FDAD16074884A4FAA2FADE58EC" blockId="1.[151,1437,1007,1968]" pageId="1" pageNumber="524">
<emphasis id="D2F2EAEFAD16074884A4FAA2FEDF58A4" bold="true" box="[199,345,1404,1429]" pageId="1" pageNumber="524">Description.</emphasis>
<typeStatus id="3F3D885FAD1607488503FAA2FE4E58A4" box="[352,456,1404,1429]" pageId="1" pageNumber="524" type="holotype">Holotype</typeStatus>
adult male: L
<quantity id="277E9B18AD160748860CFAA2FD6358A5" box="[623,741,1404,1428]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="1" pageNumber="524" unit="mm" value="12.3">12.30 mm</quantity>
, H
<quantity id="277E9B18AD160748876EFAA2FCF258A5" box="[781,884,1404,1428]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="524" unit="mm" value="0.48">0.48 mm</quantity>
, 48 podous + 2 apodous rings + telson.
<typeStatus id="3F3D885FAD1607488159FAA3FA1A58A4" box="[1338,1436,1405,1429]" pageId="1" pageNumber="524" type="paratype">Paratype</typeStatus>
adult male: L
<quantity id="277E9B18AD1607488555FA7EFE1B5886" box="[310,413,1440,1464]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="1" pageNumber="524" unit="mm" value="9.2">9.20 mm</quantity>
, H
<quantity id="277E9B18AD16074885A6FA7EFDAA5886" box="[453,556,1440,1464]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="524" unit="mm" value="0.48">0.48 mm</quantity>
, 41 podous + 2 apodous rings + telson.
<typeStatus id="3F3D885FAD1607488792FA7EFBD55889" box="[1009,1107,1440,1464]" pageId="1" pageNumber="524" type="paratype">Paratype</typeStatus>
females up to L
<quantity id="277E9B18AD1607488176FA41FAFD5886" box="[1301,1403,1439,1464]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="1" pageNumber="524" unit="mm" value="18.5">18.5 mm</quantity>
, H
<quantity id="277E9B18AD16074884F4FA1AFF7B58ED" box="[151,253,1476,1501]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="524" unit="mm" value="0.58">0.58 mm</quantity>
, 38–56 podous + 2 apodous rings + telson. Juvenile male with 33 podous +3 apodous ring + telson.
</paragraph>
<paragraph id="E03936FDAD16074884A4FA39FD425B31" blockId="1.[151,1437,1007,1968]" box="[199,708,1511,1536]" pageId="1" pageNumber="524">
<emphasis id="D2F2EAEFAD16074884A4FA39FE9D5B31" box="[199,283,1511,1536]" italics="true" pageId="1" pageNumber="524">Colour.</emphasis>
Depigmented, whitish to pale yellow.
</paragraph>
<paragraph id="E03936FDAD16074884A4F9D2FA985B5C" blockId="1.[151,1437,1007,1968]" pageId="1" pageNumber="524">
<emphasis id="D2F2EAEFAD16074884A4F9D2FE8D5B14" box="[199,267,1548,1573]" italics="true" pageId="1" pageNumber="524">Head.</emphasis>
Without ocelli. Antennae quite long, length 203 % of H in
<typeStatus id="3F3D885FAD16074887BBF9D3FBBE5B14" box="[984,1080,1549,1573]" pageId="1" pageNumber="524" type="paratype">paratype</typeStatus>
male, 188 % of H in female. Relative lengths of antennomeres 1–8 (8 = apical sensilla):
<date id="9438103DAD1607488758F9F1FC165B79" box="[827,912,1583,1608]" pageId="1" pageNumber="524" value="1918-07-20">7/20/18</date>
/14/
<date id="9438103DAD16074887DEF9F1FB965B79" box="[957,1040,1583,1608]" pageId="1" pageNumber="524" value="1916-07-14">16/14/7</date>
/4 % in
<typeStatus id="3F3D885FAD160748800EF9EEFB485B79" box="[1133,1230,1584,1608]" pageId="1" pageNumber="524" type="paratype">paratype</typeStatus>
male,
<date id="9438103DAD1607488178F9F1FAF75B79" box="[1307,1393,1583,1608]" pageId="1" pageNumber="524" value="1918-07-20">7/20/18</date>
/15/
<date id="9438103DAD16074884F4F98AFF6B5B5D" box="[151,237,1620,1644]" pageId="1" pageNumber="524" value="1917-07-13">17/13/7</date>
/3 % in female. In males, mandibles with usual parrot-bill-like modification of cardo and stipes.
</paragraph>
<paragraph id="E03936FDAD16074884A4F9A9FBDD5B84" blockId="1.[151,1437,1007,1968]" pageId="1" pageNumber="524">
<emphasis id="D2F2EAEFAD16074884A4F9A9FEE95BA1" box="[199,367,1655,1680]" italics="true" pageId="1" pageNumber="524">Midbody rings</emphasis>
with ca. 12–14 setae on posterior margin, length of setae ca. 15–17% of H in both sexes. Latero-ventral sides of rings with longitudinal striations which extend into prozonites.
</paragraph>
<paragraph id="E03936FDAD16074884A4F91FFB5C5A11" blockId="1.[151,1437,1007,1968]" pageId="1" pageNumber="524">
<emphasis id="D2F2EAEFAD16074884A4F91FFF7A5BE9" box="[199,252,1729,1752]" italics="true" pageId="1" pageNumber="524">Legs</emphasis>
: length 95 % of H in
<typeStatus id="3F3D885FAD1607488598F91EFDDA5BE9" box="[507,604,1728,1752]" pageId="1" pageNumber="524" type="paratype">paratype</typeStatus>
male, 85 % of H in female. Relative lengths of podomeres from coxa to claw
<date id="9438103DAD16074884B1F93AFEB35BCD" box="[210,309,1764,1788]" pageId="1" pageNumber="524" value="1917-12-14">17/14/12</date>
/
<date id="9438103DAD160748855CF93AFE155BCD" box="[319,403,1764,1788]" pageId="1" pageNumber="524" value="1923-08-13">8/13/23</date>
/13 % in
<typeStatus id="3F3D885FAD1607488594F93BFDDE5BCC" box="[503,600,1765,1789]" pageId="1" pageNumber="524" type="paratype">paratype</typeStatus>
male, 20/
<date id="9438103DAD16074886ABF93AFCAE5BCD" box="[712,808,1764,1788]" pageId="1" pageNumber="524" value="2014-12-14">14/14/12</date>
/
<date id="9438103DAD1607488751F93AFC005BCD" box="[818,902,1764,1788]" pageId="1" pageNumber="524" value="2011-09-20">9/20/11</date>
% in female; claw length/height
<quantity id="277E9B18AD1607488095F93AFAB05BCC" box="[1270,1334,1764,1789]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" pageId="1" pageNumber="524" unit="in" value="4.5">4.5 in</quantity>
<typeStatus id="3F3D885FAD160748815FF93BFA1A5BCC" box="[1340,1436,1765,1789]" pageId="1" pageNumber="524" type="paratype">paratype</typeStatus>
male and
<quantity id="277E9B18AD1607488567F8D6FEA95A11" box="[260,303,1799,1824]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="1" pageNumber="524" unit="in" value="4.0">4 in</quantity>
female. Lanceolate setae present on femur (1), postfemur (1) and tibia (2) in males.
</paragraph>
<paragraph id="E03936FDAD16074884A4F8F3FDCE5A81" blockId="1.[151,1437,1007,1968]" pageId="1" pageNumber="524">
<emphasis id="D2F2EAEFAD16074884A4F8F3FE555A74" box="[199,467,1836,1861]" italics="true" pageId="1" pageNumber="524">First pair of male legs</emphasis>
(
<figureCitation id="78BD2A78AD1607488585F8F2FDF95A75" box="[486,639,1836,1861]" captionStart="FIGURES 5 – 9" captionStartId="3.[151,264,1868,1890]" captionTargetBox="[223,1340,193,1835]" captionTargetId="figure@3.[223,1340,193,1837]" captionTargetPageId="3" captionText="FIGURES 5 – 9. Cibiniulus slovacus sp. n., paratype male. 5. posterior gonopod, lateral view; 6. apical half of posterior gonopod, lateral view; 7. posterior gonopod, mesal view; 8. left first leg, posterior view; 9. apical part of right first leg, anterior view. a: apophyses / modified setae, s: sternum, cl: claw, dl: dorsal lobe, fs: femoral striations, ll: leaf-shaped lamella, sh: setiferous hump, st: striations on leaf-shaped lamella, ta: tarsus, ti: tibia, tr: triangular area, vl: ventral lobe. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/235352/files/figure.png" pageId="1" pageNumber="524">Figs 8 and 9</figureCitation>
) highly modified, incrassate. A narrow triangular area (
<emphasis id="D2F2EAEFAD1607488177F8F2FAAC5A74" bold="true" box="[1300,1322,1836,1861]" pageId="1" pageNumber="524">tr</emphasis>
) between prefemur and femur. Femur with characteristic longitudinal striations (
<emphasis id="D2F2EAEFAD16074887DDF891FC545A59" bold="true" box="[958,978,1871,1896]" pageId="1" pageNumber="524">fs</emphasis>
). Tibia (
<emphasis id="D2F2EAEFAD160748805BF891FBCF5A59" bold="true" box="[1080,1097,1871,1896]" pageId="1" pageNumber="524">ti</emphasis>
) very robust, with two mesal apophyses/modified setae (
<emphasis id="D2F2EAEFAD16074885A4F8AAFE535ABC" bold="true" box="[455,469,1908,1933]" pageId="1" pageNumber="524">a</emphasis>
), laterally with anterior setiferous hump (
<emphasis id="D2F2EAEFAD16074887C5F8AAFC465ABC" bold="true" box="[934,960,1908,1933]" pageId="1" pageNumber="524">sh</emphasis>
). Tarsus (
<emphasis id="D2F2EAEFAD1607488053F8AAFBC15ABC" bold="true" box="[1072,1095,1908,1933]" pageId="1" pageNumber="524">ta</emphasis>
) in the form of a cap on top of tibia, carrying a rudimentary claw (
<emphasis id="D2F2EAEFAD1607488640F849FDB15A81" bold="true" box="[547,567,1943,1968]" pageId="1" pageNumber="524">cl</emphasis>
).
</paragraph>
<caption id="B4F96675AD15074B84F4F853FC465AD0" httpUri="https://zenodo.org/record/235351/files/figure.png" pageId="2" pageNumber="525" targetBox="[204,1372,195,1912]" targetPageId="2">
<paragraph id="E03936FDAD15074B84F4F853FC465AD0" blockId="2.[151,1436,1933,2017]" pageId="2" pageNumber="525">
<emphasis id="D2F2EAEFAD15074B84F4F853FEBC5A92" bold="true" box="[151,314,1933,1955]" pageId="2" pageNumber="525">FIGURES 1–4.</emphasis>
<taxonomicName id="27864D7EAD15074B8523F853FD815A92" box="[320,519,1933,1955]" class="Diplopoda" family="Blaniulidae" genus="Cibiniulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="525" phylum="Arthropoda" rank="species" species="slovacus" status="sp. nov.">
<emphasis id="D2F2EAEFAD15074B8523F853FD815A92" box="[320,519,1933,1955]" italics="true" pageId="2" pageNumber="525">Cibiniulus slovacus</emphasis>
</taxonomicName>
<emphasis id="D2F2EAEFAD15074B866DF853FDCB5A92" bold="true" box="[526,589,1933,1955]" pageId="2" pageNumber="525">
<taxonomicNameLabel id="C9C15794AD15074B866DF853FDC15A92" box="[526,583,1933,1955]" pageId="2" pageNumber="525" rank="species">sp. n.</taxonomicNameLabel>
,
</emphasis>
paratype male. 1. anterior gonopods, posterior view; 2. anterior gonopods, posterolateral view; 3. anterior gonopods, lateral view; 4. second pair of legs with penis, posterior view. –
<emphasis id="D2F2EAEFAD15074B80EDF872FB105AF3" bold="true" box="[1166,1174,1964,1986]" pageId="2" pageNumber="525">f</emphasis>
: postero-mesal flange,
<emphasis id="D2F2EAEFAD15074B81E4F872FA135AF3" bold="true" box="[1415,1429,1964,1986]" pageId="2" pageNumber="525">p</emphasis>
: penis,
<emphasis id="D2F2EAEFAD15074B84BAF815FF675AD0" bold="true" box="[217,225,1995,2017]" pageId="2" pageNumber="525">t</emphasis>
: telopodite,
<emphasis id="D2F2EAEFAD15074B8502F815FEFC5AD0" bold="true" box="[353,378,1995,2017]" pageId="2" pageNumber="525">cp</emphasis>
: coxal processes,
<emphasis id="D2F2EAEFAD15074B8651F815FDC55AD0" bold="true" box="[562,579,1995,2017]" pageId="2" pageNumber="525">ls</emphasis>
: lanceolate seta. Scale lines: 0.1 mm.
</paragraph>
</caption>
<caption id="B4F96675AD14074A84F4F892FF405AED" httpUri="https://zenodo.org/record/235352/files/figure.png" pageId="3" pageNumber="526" targetBox="[223,1340,193,1835]" targetPageId="3">
<paragraph id="E03936FDAD14074A84F4F892FF405AED" blockId="3.[151,1436,1868,2012]" pageId="3" pageNumber="526">
<emphasis id="D2F2EAEFAD14074A84F4F892FEB95A53" bold="true" box="[151,319,1868,1890]" pageId="3" pageNumber="526">FIGURES 5–9.</emphasis>
<taxonomicName id="27864D7EAD14074A852AF892FD925A53" box="[329,532,1868,1890]" class="Diplopoda" family="Blaniulidae" genus="Cibiniulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="slovacus" status="sp. nov.">
<emphasis id="D2F2EAEFAD14074A852AF892FD925A53" box="[329,532,1868,1890]" italics="true" pageId="3" pageNumber="526">Cibiniulus slovacus</emphasis>
</taxonomicName>
<emphasis id="D2F2EAEFAD14074A867CF892FDDB5A53" bold="true" box="[543,605,1868,1890]" pageId="3" pageNumber="526">
<taxonomicNameLabel id="C9C15794AD14074A867CF892FDDB5A53" box="[543,605,1868,1890]" pageId="3" pageNumber="526" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, paratype male. 5. posterior gonopod, lateral view; 6. apical half of posterior gonopod, lateral view; 7. posterior gonopod, mesal view; 8. left first leg, posterior view; 9. apical part of right first leg, anterior view.
<emphasis id="D2F2EAEFAD14074A84B4F857FF655AAE" bold="true" box="[215,227,1929,1951]" pageId="3" pageNumber="526">a</emphasis>
: apophyses/modified setae,
<emphasis id="D2F2EAEFAD14074A866DF857FD9E5AAE" bold="true" box="[526,536,1929,1951]" pageId="3" pageNumber="526">s</emphasis>
: sternum,
<emphasis id="D2F2EAEFAD14074A86EFF857FD185AAE" bold="true" box="[652,670,1929,1951]" pageId="3" pageNumber="526">cl</emphasis>
: claw,
<emphasis id="D2F2EAEFAD14074A868DF857FC855AAE" bold="true" box="[750,771,1929,1951]" pageId="3" pageNumber="526">dl</emphasis>
: dorsal lobe,
<emphasis id="D2F2EAEFAD14074A87FAF857FC2D5AAE" bold="true" box="[921,939,1929,1951]" pageId="3" pageNumber="526">fs</emphasis>
: femoral striations,
<emphasis id="D2F2EAEFAD14074A80E0F857FB175AAE" bold="true" box="[1155,1169,1929,1951]" pageId="3" pageNumber="526">ll</emphasis>
: leaf-shaped lamella,
<emphasis id="D2F2EAEFAD14074A811EF857FA135AAE" bold="true" box="[1405,1429,1929,1951]" pageId="3" pageNumber="526">sh</emphasis>
: setiferous hump,
<emphasis id="D2F2EAEFAD14074A852AF876FEDD5A8F" bold="true" box="[329,347,1960,1982]" pageId="3" pageNumber="526">st</emphasis>
: striations on leaf-shaped lamella,
<emphasis id="D2F2EAEFAD14074A86A5F876FD5D5A8F" bold="true" box="[710,731,1960,1982]" pageId="3" pageNumber="526">ta</emphasis>
: tarsus,
<emphasis id="D2F2EAEFAD14074A8757F876FCC55A8F" bold="true" box="[820,835,1960,1982]" pageId="3" pageNumber="526">ti</emphasis>
: tibia,
<emphasis id="D2F2EAEFAD14074A87EDF876FC245A8F" bold="true" box="[910,930,1960,1982]" pageId="3" pageNumber="526">tr</emphasis>
: triangular area,
<emphasis id="D2F2EAEFAD14074A8037F876FBEE5A8F" bold="true" box="[1108,1128,1960,1982]" pageId="3" pageNumber="526">vl</emphasis>
: ventral lobe. Scale lines: 0.1 mm.
</paragraph>
</caption>
<caption id="B4F96675AD13074D84F4F855FB885AEF" httpUri="https://zenodo.org/record/235353/files/figure.png" pageId="4" pageNumber="527" targetBox="[207,1356,193,1891]" targetPageId="4">
<paragraph id="E03936FDAD13074D84F4F855FB885AEF" blockId="4.[151,1436,1931,2014]" pageId="4" pageNumber="527">
<emphasis id="D2F2EAEFAD13074D84F4F855FED55A90" bold="true" box="[151,339,1931,1953]" pageId="4" pageNumber="527">FIGURES 10–12.</emphasis>
<taxonomicName id="27864D7EAD13074D853AF855FDA65A90" box="[345,544,1931,1953]" class="Diplopoda" family="Blaniulidae" genus="Cibiniulus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="slovacus" status="sp. nov.">
<emphasis id="D2F2EAEFAD13074D853AF855FDA65A90" box="[345,544,1931,1953]" italics="true" pageId="4" pageNumber="527">Cibiniulus slovacus</emphasis>
</taxonomicName>
<emphasis id="D2F2EAEFAD13074D8645F855FDD95A90" bold="true" box="[550,607,1931,1953]" pageId="4" pageNumber="527">
<taxonomicNameLabel id="C9C15794AD13074D8645F855FDD95A90" box="[550,607,1931,1953]" pageId="4" pageNumber="527" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, 10. holotype male posterior gonopods, lateral and mesal view; 11. holotype male posterior gonopods apex, lateral and mesal view; 12. female vulva, posterior view.
<emphasis id="D2F2EAEFAD13074D878CF877FC7F5A8E" bold="true" box="[1007,1017,1961,1983]" pageId="4" pageNumber="527">s</emphasis>
: sternum,
<emphasis id="D2F2EAEFAD13074D8004F877FBFA5A8E" bold="true" box="[1127,1148,1961,1983]" pageId="4" pageNumber="527">dl</emphasis>
: dorsal lobe,
<emphasis id="D2F2EAEFAD13074D8169F877FA9E5A8E" bold="true" box="[1290,1304,1961,1983]" pageId="4" pageNumber="527">ll</emphasis>
: leaf-shaped lamella,
<emphasis id="D2F2EAEFAD13074D848DF816FE855AEF" bold="true" box="[238,259,1992,2014]" pageId="4" pageNumber="527">rs</emphasis>
: receptaculum seminis,
<emphasis id="D2F2EAEFAD13074D859AF816FD8D5AEF" bold="true" box="[505,523,1992,2014]" pageId="4" pageNumber="527">st</emphasis>
: striations on leaf-shaped lamella,
<emphasis id="D2F2EAEFAD13074D870EF816FC075AEF" bold="true" box="[877,897,1992,2014]" pageId="4" pageNumber="527">vl</emphasis>
: ventral lobe.
</paragraph>
</caption>
<caption id="B4F96675AD12074C84F4FF45FBE35D83" ID-Table-UUID="B4F96675AD12074C84F4FF45FBE35D83" box="[151,1125,155,178]" httpUri="http://table.plazi.org/id/B4F96675AD12074C84F4FF45FBE35D83" pageId="5" pageNumber="528" targetBox="[159,1406,206,1101]" targetIsTable="true" targetPageId="5">
<paragraph id="E03936FDAD12074C84F4FF45FBE35D83" blockId="5.[151,1125,155,178]" box="[151,1125,155,178]" pageId="5" pageNumber="528">
<emphasis id="D2F2EAEFAD12074C84F4FF45FE8C5D83" bold="true" box="[151,266,155,178]" pageId="5" pageNumber="528">TABLE 1.</emphasis>
Differences between
<taxonomicName id="27864D7EAD12074C8591FF45FCF35D83" authority="Verhoeff, 1897" authorityName="Verhoeff" authorityYear="1897" box="[498,885,155,178]" class="Diplopoda" family="Blaniulidae" genus="Cibiniulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="528" phylum="Arthropoda" rank="species" species="phlepsii">
<emphasis id="D2F2EAEFAD12074C8591FF45FD3A5D83" box="[498,700,155,178]" italics="true" pageId="5" pageNumber="528">Cibiniulus phlepsii</emphasis>
(Verhoeff, 1897)
</taxonomicName>
and
<taxonomicName id="27864D7EAD12074C87CBFF45FBA45D83" box="[936,1058,155,178]" class="Diplopoda" family="Blaniulidae" genus="Cibiniulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="528" phylum="Arthropoda" rank="species" species="slovacus" status="sp. nov.">
<emphasis id="D2F2EAEFAD12074C87CBFF45FBA45D83" box="[936,1058,155,178]" italics="true" pageId="5" pageNumber="528">C. slovacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C9C15794AD12074C8049FF45FBE35D83" box="[1066,1125,155,178]" pageId="5" pageNumber="528" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="E03936FDAD12074C84FCFF10FB11591E" pageId="5" pageNumber="528">
<table id="9286C45DAD12F8B684FCFF10FAF8597C" box="[159,1406,206,1101]" gridcols="3" gridrows="11" pageId="5" pageNumber="528">
<tr id="5EB634BFAD12F8B684FCFF10FAF85DD5" box="[159,1406,206,228]" gridrow="0" pageId="5" pageNumber="528">
<th id="1D675DC3AD12F8B684FCFF10FDB05DD5" box="[159,566,206,228]" gridcol="0" gridrow="0" pageId="5" pageNumber="528">Characters</th>
<th id="1D675DC3AD12F8B68628FF10FC655DD5" box="[587,995,206,228]" gridcol="1" gridrow="0" pageId="5" pageNumber="528">
<taxonomicName id="27864D7EAD12074C862FFF10FC3C5DD5" authority="Verhoeff, 1897" authorityName="Verhoeff" authorityYear="1897" box="[588,954,206,228]" class="Diplopoda" family="Blaniulidae" genus="Cibiniulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="528" phylum="Arthropoda" rank="species" species="phlepsii">
<emphasis id="D2F2EAEFAD12074C862FFF10FC8A5DD5" box="[588,780,206,228]" italics="true" pageId="5" pageNumber="528">Cibiniulus phlepsii</emphasis>
(Verhoeff, 1897)
</taxonomicName>
</th>
<th id="1D675DC3AD12F8B6879BFF10FAF85DD5" box="[1016,1406,206,228]" gridcol="2" gridrow="0" pageId="5" pageNumber="528">
<taxonomicName id="27864D7EAD12074C879BFF10FB395DD5" box="[1016,1215,206,228]" class="Diplopoda" family="Blaniulidae" genus="Cibiniulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="528" phylum="Arthropoda" rank="species" species="slovacus" status="sp. nov.">
<emphasis id="D2F2EAEFAD12074C879BFF10FB395DD5" box="[1016,1215,206,228]" italics="true" pageId="5" pageNumber="528">Cibiniulus slovacus</emphasis>
</taxonomicName>
<emphasis id="D2F2EAEFAD12074C80A5FF10FB795DD5" bold="true" box="[1222,1279,206,228]" pageId="5" pageNumber="528">
<taxonomicNameLabel id="C9C15794AD12074C80A5FF10FB795DD5" box="[1222,1279,206,228]" pageId="5" pageNumber="528" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="5EB634BFAD12F8B684FCFEC8FAF85C1D" box="[159,1406,278,300]" gridrow="1" pageId="5" pageNumber="528">
<th id="1D675DC3AD12F8B684FCFEC8FDB05C1D" box="[159,566,278,300]" gridcol="0" gridrow="1" pageId="5" pageNumber="528">Ocelli</th>
<td id="1D675DC3AD12F8B68628FEC8FC655C1D" box="[587,995,278,300]" gridcol="1" gridrow="1" pageId="5" pageNumber="528">Present</td>
<td id="1D675DC3AD12F8B6879BFEC8FAF85C1D" box="[1016,1406,278,300]" gridcol="2" gridrow="1" pageId="5" pageNumber="528">Absent</td>
</tr>
<tr id="5EB634BFAD12F8B684FCFE80FAF85C45" box="[159,1406,350,372]" gridrow="2" pageId="5" pageNumber="528">
<th id="1D675DC3AD12F8B684FCFE80FDB05C45" box="[159,566,350,372]" gridcol="0" gridrow="2" pageId="5" pageNumber="528">Antennae</th>
<td id="1D675DC3AD12F8B68628FE80FC655C45" box="[587,995,350,372]" gridcol="1" gridrow="2" pageId="5" pageNumber="528">Shorter</td>
<td id="1D675DC3AD12F8B6879BFE80FAF85C45" box="[1016,1406,350,372]" gridcol="2" gridrow="2" pageId="5" pageNumber="528">Longer</td>
</tr>
<tr id="5EB634BFAD12F8B684FCFE78FAF85CEB" box="[159,1406,422,474]" gridrow="3" pageId="5" pageNumber="528">
<th id="1D675DC3AD12F8B684FCFE78FDB05CEB" box="[159,566,422,474]" gridcol="0" gridrow="3" pageId="5" pageNumber="528">Tibia of first pair of legs in ♂♂</th>
<td id="1D675DC3AD12F8B68628FE78FC655CEB" box="[587,995,422,474]" gridcol="1" gridrow="3" pageId="5" pageNumber="528">Without apophyses and without strongly developed antero-lateral hump</td>
<td id="1D675DC3AD12F8B6879BFE78FAF85CEB" box="[1016,1406,422,474]" gridcol="2" gridrow="3" pageId="5" pageNumber="528">With two apophyses and with strongly developed antero-lateral hump</td>
</tr>
<tr id="5EB634BFAD12F8B684FCFDD2FAF85F13" box="[159,1406,524,546]" gridrow="4" pageId="5" pageNumber="528">
<th id="1D675DC3AD12F8B684FCFDD2FDB05F13" box="[159,566,524,546]" gridcol="0" gridrow="4" pageId="5" pageNumber="528">Tarsi of first pair of legs in ♂♂</th>
<td id="1D675DC3AD12F8B68628FDD2FC655F13" box="[587,995,524,546]" gridcol="1" gridrow="4" pageId="5" pageNumber="528">Very low</td>
<td id="1D675DC3AD12F8B6879BFDD2FAF85F13" box="[1016,1406,524,546]" gridcol="2" gridrow="4" pageId="5" pageNumber="528">In the form of a cap on top of tibia</td>
</tr>
<tr id="5EB634BFAD12F8B684FCFD8AFAF85FB8" box="[159,1406,596,649]" gridrow="5" pageId="5" pageNumber="528">
<th id="1D675DC3AD12F8B684FCFD8AFDB05FB8" box="[159,566,596,649]" gridcol="0" gridrow="5" pageId="5" pageNumber="528">Anterior gonopods with narrowing at border between:</th>
<td id="1D675DC3AD12F8B68628FD8AFC655FB8" box="[587,995,596,649]" gridcol="1" gridrow="5" pageId="5" pageNumber="528">second and third ⅓ of length of coxal processes</td>
<td id="1D675DC3AD12F8B6879BFD8AFAF85FB8" box="[1016,1406,596,649]" gridcol="2" gridrow="5" pageId="5" pageNumber="528">first and second ⅓ of length of coxal processes</td>
</tr>
<tr id="5EB634BFAD12F8B684FCFD65FAF85FC1" box="[159,1406,699,752]" gridrow="6" pageId="5" pageNumber="528">
<th id="1D675DC3AD12F8B684FCFD65FDB05FC1" box="[159,566,699,752]" gridcol="0" gridrow="6" pageId="5" pageNumber="528">Postero-mesal flange of anterior gonopods in lateral view</th>
<td id="1D675DC3AD12F8B68628FD65FC655FC1" box="[587,995,699,752]" gridcol="1" gridrow="6" pageId="5" pageNumber="528">Bilobed</td>
<td id="1D675DC3AD12F8B6879BFD65FAF85FC1" box="[1016,1406,699,752]" gridcol="2" gridrow="6" pageId="5" pageNumber="528">Almost rectilinear</td>
</tr>
<tr id="5EB634BFAD12F8B684FCFCFCFAF85E09" box="[159,1406,802,824]" gridrow="7" pageId="5" pageNumber="528">
<th id="1D675DC3AD12F8B684FCFCFCFDB05E09" box="[159,566,802,824]" gridcol="0" gridrow="7" pageId="5" pageNumber="528">Telopodites of anterior gonopods</th>
<td id="1D675DC3AD12F8B68628FCFCFC655E09" box="[587,995,802,824]" gridcol="1" gridrow="7" pageId="5" pageNumber="528">Without setae</td>
<td id="1D675DC3AD12F8B6879BFCFCFAF85E09" box="[1016,1406,802,824]" gridcol="2" gridrow="7" pageId="5" pageNumber="528">With or without setae</td>
</tr>
<tr id="5EB634BFAD12F8B684FCFCB4FAF85EB1" box="[159,1406,874,896]" gridrow="8" pageId="5" pageNumber="528">
<th id="1D675DC3AD12F8B684FCFCB4FDB05EB1" box="[159,566,874,896]" gridcol="0" gridrow="8" pageId="5" pageNumber="528">Posterior gonopods</th>
<td id="1D675DC3AD12F8B68628FCB4FC655EB1" box="[587,995,874,896]" gridcol="1" gridrow="8" pageId="5" pageNumber="528">With lateral lamella</td>
<td id="1D675DC3AD12F8B6879BFCB4FAF85EB1" box="[1016,1406,874,896]" gridcol="2" gridrow="8" pageId="5" pageNumber="528">Without lateral lamella</td>
</tr>
<tr id="5EB634BFAD12F8B684FCFC6CFAF85ED6" box="[159,1406,946,999]" gridrow="9" pageId="5" pageNumber="528">
<th id="1D675DC3AD12F8B684FCFC6CFDB05ED6" box="[159,566,946,999]" gridcol="0" gridrow="9" pageId="5" pageNumber="528">Dorsal lobe of mesal lamella of posterior gonopods</th>
<td id="1D675DC3AD12F8B68628FC6CFC655ED6" box="[587,995,946,999]" gridcol="1" gridrow="9" pageId="5" pageNumber="528">Acuminate</td>
<td id="1D675DC3AD12F8B6879BFC6CFAF85ED6" box="[1016,1406,946,999]" gridcol="2" gridrow="9" pageId="5" pageNumber="528">Almost rounded</td>
</tr>
<tr id="5EB634BFAD12F8B684FCFBC7FAF8597C" box="[159,1406,1049,1101]" gridrow="10" pageId="5" pageNumber="528">
<th id="1D675DC3AD12F8B684FCFBC7FDB0597C" box="[159,566,1049,1101]" gridcol="0" gridrow="10" pageId="5" pageNumber="528">Ventral lobe of mesal lamella of posterior gonopods</th>
<td id="1D675DC3AD12F8B68628FBC7FC65597C" box="[587,995,1049,1101]" gridcol="1" gridrow="10" pageId="5" pageNumber="528">With fringes</td>
<td id="1D675DC3AD12F8B6879BFBC7FAF8597C" box="[1016,1406,1049,1101]" gridcol="2" gridrow="10" pageId="5" pageNumber="528">Without fringes</td>
</tr>
</table>
</paragraph>
<paragraph id="E03936FDAD12074C84A4FB4FFEB859FC" blockId="5.[151,1437,1169,2022]" pageId="5" pageNumber="528">
<emphasis id="D2F2EAEFAD12074C84A4FB4FFE6A599B" box="[199,492,1169,1194]" italics="true" pageId="5" pageNumber="528">Second pair of male legs</emphasis>
(
<figureCitation id="78BD2A78AD12074C8663FB4FFDCE5998" box="[512,584,1169,1194]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1933,1955]" captionTargetBox="[204,1372,195,1912]" captionTargetId="figure@2.[204,1372,195,1913]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Cibiniulus slovacus sp. n., paratype male. 1. anterior gonopods, posterior view; 2. anterior gonopods, posterolateral view; 3. anterior gonopods, lateral view; 4. second pair of legs with penis, posterior view. – f: postero-mesal flange, p: penis, t: telopodite, cp: coxal processes, ls: lanceolate seta. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/235351/files/figure.png" pageId="5" pageNumber="528">Fig. 4</figureCitation>
) with two ventral, lanceolate setae (
<emphasis id="D2F2EAEFAD12074C8795FB4FFB8F599B" bold="true" box="[1014,1033,1169,1194]" pageId="5" pageNumber="528">ls</emphasis>
) on femur, postfemur and tibia in
<typeStatus id="3F3D885FAD12074C84F4FB6AFF7F59FC" box="[151,249,1204,1229]" pageId="5" pageNumber="528" type="holotype">holotype</typeStatus>
male.
</paragraph>
<paragraph id="E03936FDAD12074C84A4FB04FD3359C0" blockId="5.[151,1437,1169,2022]" box="[199,693,1241,1266]" pageId="5" pageNumber="528">
<emphasis id="D2F2EAEFAD12074C84A4FB04FE8059C0" box="[199,262,1242,1265]" italics="true" pageId="5" pageNumber="528">Penis</emphasis>
(
<figureCitation id="78BD2A78AD12074C8577FB07FEDE59C0" box="[276,344,1241,1266]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1933,1955]" captionTargetBox="[204,1372,195,1912]" captionTargetId="figure@2.[204,1372,195,1913]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Cibiniulus slovacus sp. n., paratype male. 1. anterior gonopods, posterior view; 2. anterior gonopods, posterolateral view; 3. anterior gonopods, lateral view; 4. second pair of legs with penis, posterior view. – f: postero-mesal flange, p: penis, t: telopodite, cp: coxal processes, ls: lanceolate seta. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/235351/files/figure.png" pageId="5" pageNumber="528">Fig. 4</figureCitation>
,
<emphasis id="D2F2EAEFAD12074C8507FB07FEF559C3" bold="true" box="[356,371,1241,1266]" pageId="5" pageNumber="528">p</emphasis>
) simple, with rounded apex.
</paragraph>
<paragraph id="E03936FDAD12074C84A4FB23FC5C5827" blockId="5.[151,1437,1169,2022]" box="[199,986,1277,1302]" pageId="5" pageNumber="528">
<emphasis id="D2F2EAEFAD12074C84A4FB23FD1A5827" box="[199,668,1277,1302]" italics="true" pageId="5" pageNumber="528">
Ventral margin of pleurotergite
<quantity id="277E9B18AD12074C864AFB20FDD25824" box="[553,596,1278,1301]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="5" pageNumber="528" unit="in" value="7.0">7 in</quantity>
males
</emphasis>
without a horizontal flange.
</paragraph>
<paragraph id="E03936FDAD12074C84A4FAFCFB3D58FB" blockId="5.[151,1437,1169,2022]" pageId="5" pageNumber="528">
<emphasis id="D2F2EAEFAD12074C84A4FAFCFE1A580B" box="[199,412,1313,1338]" italics="true" pageId="5" pageNumber="528">Anterior gonopods</emphasis>
(
<figureCitation id="78BD2A78AD12074C85CEFAFFFD945808" box="[429,530,1313,1338]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1933,1955]" captionTargetBox="[204,1372,195,1912]" captionTargetId="figure@2.[204,1372,195,1913]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Cibiniulus slovacus sp. n., paratype male. 1. anterior gonopods, posterior view; 2. anterior gonopods, posterolateral view; 3. anterior gonopods, lateral view; 4. second pair of legs with penis, posterior view. – f: postero-mesal flange, p: penis, t: telopodite, cp: coxal processes, ls: lanceolate seta. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/235351/files/figure.png" pageId="5" pageNumber="528">Figs 1–3</figureCitation>
) with separated slender coxal processes (
<emphasis id="D2F2EAEFAD12074C8792FAFFFB8B580B" bold="true" box="[1009,1037,1313,1338]" pageId="5" pageNumber="528">cp</emphasis>
) which are somewhat wider at the base, narrowing at the border between ⅓ and ⅔ of the length, and then gradually widening towards a rounded apex. A postero-mesal flange (
<emphasis id="D2F2EAEFAD12074C85D5FAB7FE3958B3" bold="true" box="[438,447,1385,1410]" pageId="5" pageNumber="528">f</emphasis>
) extending almost the entire length of coxal process, with a thickening at the border between ⅓ and ⅔ of the length. Flange almost rectilinear in lateral view. Telopodites (
<emphasis id="D2F2EAEFAD12074C8008FA53FBF25897" bold="true" box="[1131,1140,1421,1446]" pageId="5" pageNumber="528">t</emphasis>
) rudimentary, less than ¼ the length of coxal processes, with 1+2 setae in
<typeStatus id="3F3D885FAD12074C86D3FA6FFC9458FB" box="[688,786,1457,1482]" pageId="5" pageNumber="528" type="holotype">holotype</typeStatus>
male, without setae in
<typeStatus id="3F3D885FAD12074C8076FA6CFBF058FB" box="[1045,1142,1458,1482]" pageId="5" pageNumber="528" type="paratype">paratype</typeStatus>
male.
</paragraph>
<paragraph id="E03936FDAD12074C84A4FA08FEEE5B4F" blockId="5.[151,1437,1169,2022]" pageId="5" pageNumber="528">
<emphasis id="D2F2EAEFAD12074C84A4FA08FE2158DF" box="[199,423,1493,1518]" italics="true" pageId="5" pageNumber="528">Posterior gonopods</emphasis>
(
<figureCitation id="78BD2A78AD12074C85DBFA0BFDA658DF" box="[440,544,1493,1518]" captionStart="FIGURES 5 – 9" captionStartId="3.[151,264,1868,1890]" captionTargetBox="[223,1340,193,1835]" captionTargetId="figure@3.[223,1340,193,1837]" captionTargetPageId="3" captionText="FIGURES 5 – 9. Cibiniulus slovacus sp. n., paratype male. 5. posterior gonopod, lateral view; 6. apical half of posterior gonopod, lateral view; 7. posterior gonopod, mesal view; 8. left first leg, posterior view; 9. apical part of right first leg, anterior view. a: apophyses / modified setae, s: sternum, cl: claw, dl: dorsal lobe, fs: femoral striations, ll: leaf-shaped lamella, sh: setiferous hump, st: striations on leaf-shaped lamella, ta: tarsus, ti: tibia, tr: triangular area, vl: ventral lobe. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/235352/files/figure.png" pageId="5" pageNumber="528">Figs 5–7</figureCitation>
,
<figureCitation id="78BD2A78AD12074C864DFA0BFD2758DC" box="[558,673,1493,1518]" captionStart="FIGURES 10 – 12" captionStartId="4.[151,264,1931,1953]" captionTargetBox="[207,1356,193,1891]" captionTargetId="figure@4.[207,1357,193,1892]" captionTargetPageId="4" captionText="FIGURES 10 – 12. Cibiniulus slovacus sp. n., 10. holotype male posterior gonopods, lateral and mesal view; 11. holotype male posterior gonopods apex, lateral and mesal view; 12. female vulva, posterior view. s: sternum, dl: dorsal lobe, ll: leaf-shaped lamella, rs: receptaculum seminis, st: striations on leaf-shaped lamella, vl: ventral lobe." httpUri="https://zenodo.org/record/235353/files/figure.png" pageId="5" pageNumber="528">10 and 11</figureCitation>
) with prolonged sternum (
<emphasis id="D2F2EAEFAD12074C87BBFA0BFC6558DF" bold="true" box="[984,995,1493,1518]" pageId="5" pageNumber="528">s</emphasis>
). Apical half with wide and thin leafshaped lamella (
<emphasis id="D2F2EAEFAD12074C8537FA27FEE55B23" bold="true" box="[340,355,1529,1554]" pageId="5" pageNumber="528">ll</emphasis>
) divided into a dorsal, somewhat rounded lobe (
<emphasis id="D2F2EAEFAD12074C87FBFA27FC295B23" bold="true" box="[920,943,1529,1554]" pageId="5" pageNumber="528">dl</emphasis>
), and a ventral, acuminate lobe (
<emphasis id="D2F2EAEFAD12074C814CFA27FAC35B23" bold="true" box="[1327,1349,1529,1554]" pageId="5" pageNumber="528">vl</emphasis>
). Leafshaped lamella with numerous striations (
<emphasis id="D2F2EAEFAD12074C86E0F9C3FD115B07" bold="true" box="[643,663,1565,1590]" pageId="5" pageNumber="528">st</emphasis>
) which at first glance resemble setae. Lateral lamella completely absent. On the SEM images, the apical half of the posterior gonopods is distorted, probably due to the air-drying method employed.
</paragraph>
<paragraph id="E03936FDAD12074C84A4F957FC955B93" blockId="5.[151,1437,1169,2022]" box="[199,787,1673,1698]" pageId="5" pageNumber="528">
<emphasis id="D2F2EAEFAD12074C84A4F957FE765B93" box="[199,496,1673,1698]" italics="true" pageId="5" pageNumber="528">Second pair of female legs</emphasis>
without any peculiarities.
</paragraph>
<paragraph id="E03936FDAD12074C84A4F973FC7C5BDB" blockId="5.[151,1437,1169,2022]" pageId="5" pageNumber="528">
<emphasis id="D2F2EAEFAD12074C84A4F973FE975BF7" box="[199,273,1709,1734]" italics="true" pageId="5" pageNumber="528">Vulvae</emphasis>
(
<figureCitation id="78BD2A78AD12074C8547F973FEF15BF4" box="[292,375,1709,1734]" captionStart="FIGURES 10 – 12" captionStartId="4.[151,264,1931,1953]" captionTargetBox="[207,1356,193,1891]" captionTargetId="figure@4.[207,1357,193,1892]" captionTargetPageId="4" captionText="FIGURES 10 – 12. Cibiniulus slovacus sp. n., 10. holotype male posterior gonopods, lateral and mesal view; 11. holotype male posterior gonopods apex, lateral and mesal view; 12. female vulva, posterior view. s: sternum, dl: dorsal lobe, ll: leaf-shaped lamella, rs: receptaculum seminis, st: striations on leaf-shaped lamella, vl: ventral lobe." httpUri="https://zenodo.org/record/235353/files/figure.png" pageId="5" pageNumber="528">Fig. 12</figureCitation>
) subspherical, operculum and bursa of about equal length, both without setae; receptaculum seminis (
<emphasis id="D2F2EAEFAD12074C849FF90FFE955BDB" bold="true" box="[252,275,1745,1770]" pageId="5" pageNumber="528">rs</emphasis>
) single, ellipsoid. Vulval invaginations reaching back to ring five.
</paragraph>
<paragraph id="E03936FDAD12074F84A4F92BFC695DC6" blockId="5.[151,1437,1169,2022]" lastBlockId="6.[151,1437,151,392]" lastPageId="6" lastPageNumber="529" pageId="5" pageNumber="528">
<emphasis id="D2F2EAEFAD12074C84A4F92BFD255A3F" bold="true" box="[199,675,1781,1806]" pageId="5" pageNumber="528">Note on habitat and coexisting species.</emphasis>
The new
<taxonomicName id="27864D7EAD12074C877FF92BFC165A3F" box="[796,912,1781,1806]" class="Diplopoda" family="Blaniulidae" genus="Cibiniulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="528" phylum="Arthropoda" rank="genus">
<emphasis id="D2F2EAEFAD12074C877FF92BFC165A3F" box="[796,912,1781,1806]" italics="true" pageId="5" pageNumber="528">Cibiniulus</emphasis>
</taxonomicName>
species was found in karstic caves, with the elevation of the entrances ranging from
<quantity id="277E9B18AD12074C8634F8C7FD6A5A00" box="[599,748,1817,1842]" metricMagnitude="2" metricUnit="m" metricValue="5.275" metricValueMax="8.0" metricValueMin="2.55" pageId="5" pageNumber="528" unit="m" value="527.5" valueMax="800.0" valueMin="255.0">255 to 800 m</quantity>
a.s.l. Some of the caves (Jazvínska, Dobrotínska, Kolónia II) are quite short (
<quantity id="277E9B18AD12074C8531F8E3FE3E5A64" box="[338,440,1853,1878]" metricMagnitude="1" metricUnit="m" metricValue="3.45" metricValueMax="4.4" metricValueMin="2.5" pageId="5" pageNumber="528" unit="m" value="34.5" valueMax="44.0" valueMin="25.0">25-44 m</quantity>
), the last one (Gemersko-teplická Cave) has long a narrow corridor (
<quantity id="277E9B18AD12074C8087F8E3FAB45A64" box="[1252,1330,1853,1878]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="5" pageNumber="528" unit="m" value="510.0">510 m</quantity>
) with an underground stream. The cave floors are horizontal except for the cave Kolónia II (-
<quantity id="277E9B18AD12074C80E7F8BFFB4E5A48" box="[1156,1224,1889,1914]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="528" unit="m" value="25.0">25 m</quantity>
). Specimens were collected on decaying wood or in pitfall traps (100 ml plastic bottles with a mixture of beer and ethyleneglycole with cheese or salami under the opening; the traps were placed deep in the debris). All records are from the inner part of the caves (deeper than
<quantity id="277E9B18AD12074C8593F813FDA95AD4" box="[496,559,1997,2021]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="528" unit="m" value="20.0">20 m</quantity>
). All of the caves have a well preserved environment and are not open to the public. There were no other obligate cavernicolous millipedes co-occurring with
<taxonomicName id="27864D7EAD11074F8031FF46FB5E5D81" box="[1106,1240,151,176]" class="Diplopoda" family="Blaniulidae" genus="Cibiniulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="slovacus" status="sp. nov.">
<emphasis id="D2F2EAEFAD11074F8031FF46FB5E5D81" box="[1106,1240,151,176]" italics="true" pageId="6" pageNumber="529">C. slovacus</emphasis>
</taxonomicName>
<emphasis id="D2F2EAEFAD11074F8087FF49FAAE5D81" bold="true" box="[1252,1320,151,176]" pageId="6" pageNumber="529">
<taxonomicNameLabel id="C9C15794AD11074F8087FF49FAAE5D81" box="[1252,1320,151,176]" pageId="6" pageNumber="529" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, but only common species like
<taxonomicName id="27864D7EAD11074F85CBFF62FC3A5DE5" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" box="[424,956,188,213]" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="D2F2EAEFAD11074F85CBFF62FD4E5DE4" box="[424,712,188,213]" italics="true" pageId="6" pageNumber="529">Polydesmus denticulatus</emphasis>
C. L. Koch, 1847
</taxonomicName>
(Jazvínska, Dobrotínska caves) or the eutroglophilous
<taxonomicName id="27864D7EAD11074F852EFF01FD6A5DC9" authority="Waga, 1857" authorityName="Waga" authorityYear="1857" box="[333,748,223,248]" class="Diplopoda" family="Glomeridae" genus="Trachysphaera" kingdom="Animalia" order="Glomerida" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="D2F2EAEFAD11074F852EFF01FDC85DC9" box="[333,590,223,248]" italics="true" pageId="6" pageNumber="529">Trachysphaera costata</emphasis>
(Waga, 1857)
</taxonomicName>
in the cave Kolónia II.
</paragraph>
<paragraph id="E03936FDAD11074F84A4FEDAFB7C5C54" blockId="6.[151,1437,151,392]" pageId="6" pageNumber="529">
<emphasis id="D2F2EAEFAD11074F84A4FEDAFE7E5C2C" bold="true" box="[199,504,260,285]" pageId="6" pageNumber="529">Note on cave adaptations.</emphasis>
The depigmented body, the absence of ocelli and the somewhat elongated antennae of the new species are characters all shared with several other soil-dwelling blaniulids and cannot be regarded as troglomorphic traits. Thus,
<taxonomicName id="27864D7EAD11074F85A9FE93FDCD5C54" box="[458,587,332,357]" class="Diplopoda" family="Blaniulidae" genus="Cibiniulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="slovacus" status="sp. nov.">
<emphasis id="D2F2EAEFAD11074F85A9FE93FDCD5C54" box="[458,587,332,357]" italics="true" pageId="6" pageNumber="529">C. slovacus</emphasis>
</taxonomicName>
<emphasis id="D2F2EAEFAD11074F8631FE92FD175C54" bold="true" box="[594,657,332,357]" pageId="6" pageNumber="529">
<taxonomicNameLabel id="C9C15794AD11074F8631FE92FD175C54" box="[594,657,332,357]" pageId="6" pageNumber="529" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
should probably be classified as a troglophilic species.
</paragraph>
</subSubSection>
<subSubSection id="A89C6576AD11074F84A4FEB1FBD85CB9" box="[199,1118,367,392]" pageId="6" pageNumber="529" type="distribution">
<paragraph id="E03936FDAD11074F84A4FEB1FBD85CB9" blockId="6.[151,1437,151,392]" box="[199,1118,367,392]" pageId="6" pageNumber="529">
<emphasis id="D2F2EAEFAD11074F84A4FEB1FEDF5CB9" bold="true" box="[199,345,367,392]" pageId="6" pageNumber="529">Distribution</emphasis>
. Known only from several caves in
<collectingCountry id="9891766DAD11074F868CFEB1FCD75CB9" box="[751,849,367,392]" name="Slovakia" pageId="6" pageNumber="529">Slovakia</collectingCountry>
(
<figureCitation id="78BD2A78AD11074F8703FEB1FC355CB9" box="[864,947,367,392]" captionStart="FIGURE 43" captionStartId="16.[151,250,1117,1139]" captionTargetBox="[151,1436,193,1095]" captionTargetId="figure@16.[151,1436,193,1095]" captionTargetPageId="16" captionText="FIGURE 43. Distribution of the genera Cibiniulus Verhoeff, 1897 and Thassoblaniulus Mauriès, 1985." httpUri="https://zenodo.org/record/235362/files/figure.png" pageId="6" pageNumber="529">Fig. 43</figureCitation>
, green circles).
</paragraph>
</subSubSection>
</treatment>
</document>
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