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<document id="312F1B45C8BD53967A5F6278131797D9" ID-DOI="10.11646/zootaxa.3985.4.3" ID-GBIF-Dataset="48cc8fed-5ec0-4ece-a82b-c2fa1ddd8893" ID-ISSN="1175-5326" ID-Zenodo-Dep="235350" ID-ZooBank="ECE53B6E-DBAF-4E98-BF8A-96FD9106EF35" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461259944332" checkinUser="plazi" docAuthor="Antić, Dragan Ž., Mock, Andrej &amp; Enghoff, Henrik" docDate="2015" docId="682F87EBAD11074484F4FDC8FACA5C71" docLanguage="en" docName="zt03985p540.pdf" docOrigin="Zootaxa 3985 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thassoblaniulus radjai Antic &amp; Enghoff, sp. n." docType="treatment" docVersion="9" lastPageNumber="536" masterDocId="9416FF93AD1707498463FFDEFF865D31" masterDocTitle="Two new species of the millipede family Blaniulidae (Diplopoda, Julida) from caves in central and southeastern Europe" masterLastPageNumber="540" masterPageNumber="523" pageNumber="529" updateTime="1698609266564" updateUser="plazi">
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<mods:title id="DA30B27B1895A3F885BF6CD480832886">Two new species of the millipede family Blaniulidae (Diplopoda, Julida) from caves in central and southeastern Europe</mods:title>
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<mods:namePart id="0F7988D6CD5D3776C2676B702A1C4425">Antić, Dragan Ž.</mods:namePart>
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<mods:namePart id="1FCECA79A04E6DEBF880A073607999C4">Mock, Andrej</mods:namePart>
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<mods:namePart id="851DCBE241B28CFDC039EC47B1C425B5">Enghoff, Henrik</mods:namePart>
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<mods:title id="11251D73E3D0AFDD6D27B61CEFB170C3">Zootaxa</mods:title>
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<mods:date id="E10C746387D3A8D10A469AE04316268B">2015</mods:date>
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<mods:number id="75D7DC8C52686237734CED2B0493A84A">3985</mods:number>
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<mods:identifier id="F0438099E5D177ED49CFDCA0F8AD5199" type="DOI">10.11646/zootaxa.3985.4.3</mods:identifier>
<mods:identifier id="0F97CFC0B3369F32F6A50E956946F058" type="GBIF-Dataset">48cc8fed-5ec0-4ece-a82b-c2fa1ddd8893</mods:identifier>
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<treatment id="682F87EBAD11074484F4FDC8FACA5C71" ID-DOI="http://doi.org/10.5281/zenodo.5620898" ID-GBIF-Taxon="119636795" ID-Zenodo-Dep="5620898" LSID="urn:lsid:plazi:treatment:682F87EBAD11074484F4FDC8FACA5C71" httpUri="http://treatment.plazi.org/id/682F87EBAD11074484F4FDC8FACA5C71" lastPageId="13" lastPageNumber="536" pageId="6" pageNumber="529">
<subSubSection id="A89C6576AD11074F84F4FDC8FE925F60" pageId="6" pageNumber="529" type="nomenclature">
<paragraph id="E03936FDAD11074F84F4FDC8FD515F01" blockId="6.[151,727,534,594]" box="[151,727,534,560]" pageId="6" pageNumber="529">
<heading id="BB718191AD11074F84F4FDC8FD515F01" bold="true" box="[151,727,534,560]" fontSize="11" level="1" pageId="6" pageNumber="529" reason="1">
<emphasis id="D2F2EAEFAD11074F84F4FDC8FD515F01" bold="true" box="[151,727,534,560]" pageId="6" pageNumber="529">
<taxonomicName id="27864D7EAD11074F84F4FDC8FD0E5F01" ID-CoL="84DSX" authority="Antic &amp; Enghoff" authorityName="Antic &amp; Enghoff" box="[151,648,534,560]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="radjai" status="sp. nov.">
<emphasis id="D2F2EAEFAD11074F84F4FDC8FE2B5F01" bold="true" box="[151,429,534,560]" italics="true" pageId="6" pageNumber="529">Thassoblaniulus radjai</emphasis>
Antić &amp; Enghoff
</taxonomicName>
,
<taxonomicNameLabel id="C9C15794AD11074F86F6FDC8FD515F01" box="[661,727,534,560]" pageId="6" pageNumber="529" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E03936FDAD11074F84F4FDE7FE925F60" blockId="6.[151,727,534,594]" box="[151,276,569,594]" pageId="6" pageNumber="529">
<figureCitation id="78BD2A78AD11074F84F4FDE7FE925F60" box="[151,276,569,594]" captionStart-0="FIGURES 13 – 16" captionStart-1="FIGURES 17 – 20" captionStart-2="FIGURES 21 and 22" captionStart-3="FIGURES 23 – 26" captionStart-4="FIGURES 27 – 32" captionStart-5="FIGURE 33" captionStartId-0="7.[151,264,1951,1973]" captionStartId-1="8.[151,264,1836,1858]" captionStartId-2="9.[151,264,1293,1315]" captionStartId-3="10.[151,264,1826,1848]" captionStartId-4="11.[151,264,1942,1964]" captionStartId-5="12.[151,250,1178,1200]" captionTargetBox-0="[151,1436,193,1929]" captionTargetBox-1="[151,1436,193,1814]" captionTargetBox-2="[151,1436,193,1271]" captionTargetBox-3="[151,1436,193,1804]" captionTargetBox-4="[210,1361,193,1917]" captionTargetBox-5="[151,1436,193,1156]" captionTargetId-0="figure@7.[151,1436,193,1930]" captionTargetId-1="figure@8.[151,1436,193,1815]" captionTargetId-2="figure@9.[151,1436,193,1271]" captionTargetId-3="figure@10.[151,1436,193,1805]" captionTargetId-4="figure@11.[210,1361,193,1917]" captionTargetId-5="figure@12.[151,1436,193,1157]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="12" captionText-0="FIGURES 13 – 16. Thassoblaniulus radjai sp. n., paratype male anterior gonopods. 13. anterior view; 14. lateral view; 15. posterior view; 16. distal view. f: postero-mesal flange, t: telopodite, cp: coxal processes, lsh: lateral shoulder. Scale lines: 0.1 mm." captionText-1="FIGURES 17 – 20. Thassoblaniulus radjai sp. n., paratype male posterior gonopod tip. 17. lateral view; 18. anterior view; 19. mesal view; 20. posterior view. – ll: leaf-shaped lamella, so: solenomere. Scale lines: 0.1 for 17, 18 and 19; 0.05 mm for 20." captionText-2="FIGURES 21 and 22. Thassoblaniulus radjai sp. n., paratype male posterior gonopod. 21. solenomere; 22. detail of lamella. Scale lines: 0.01 mm." captionText-3="FIGURES 23 – 26. Thassoblaniulus radjai sp. n., paratype male. 23. first pair of legs, antero-lateral view; 24. first pair of legs, anterior view; 25. second pair of legs with penis, posterior view; 26. body ring 7, ventral view. – p: penis, cl: claw, ls: lanceolate seta, ta: tarsus, ti: tibia. Scale lines: 0.1 mm." captionText-4="FIGURES 27 – 32. Thassoblaniulus radjai sp. n., holotype male. 27. anterior gonopods, posterior view; 28. anterior gonopods, lateral view; 29. penis, posterior view; 30. posterior gonopod, lateral view; 31. first pair of legs, posterior view; 32. second leg. Scale line: 0.4 mm." captionText-5="FIGURE 33. Thassoblaniulus radjai sp. n., female vulva, lateral view. b: bursa, o: operculum, rs: receptaculum seminis. Scale line: 0.05 mm." httpUri-0="https://zenodo.org/record/235354/files/figure.png" httpUri-1="https://zenodo.org/record/235355/files/figure.png" httpUri-2="https://zenodo.org/record/235356/files/figure.png" httpUri-3="https://zenodo.org/record/235357/files/figure.png" httpUri-4="https://zenodo.org/record/235358/files/figure.png" httpUri-5="https://zenodo.org/record/235359/files/figure.png" pageId="6" pageNumber="529">Figs 13–33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A89C6576AD11074F84F4FD5FFC445FD3" pageId="6" pageNumber="529" type="materials_examined">
<paragraph id="E03936FDAD11074F84F4FD5FFC445FD3" blockId="6.[151,1436,641,918]" pageId="6" pageNumber="529">
<emphasis id="D2F2EAEFAD11074F84F4FD5FFEED5FAB" bold="true" box="[151,363,641,666]" pageId="6" pageNumber="529">Material studied.</emphasis>
<typeStatus id="3F3D885FAD11074F851BFD5FFE665FAB" box="[376,480,641,666]" pageId="6" pageNumber="529" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="8697AE38AD11074F865AFD5CFD175FAB" box="[569,657,642,666]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="6" pageNumber="529">ZMUC</collectionCode>
): Shpella e Zezë (Shpella e Pëllumbasit, Black Cave), village of Pëllumbas, near Tirana,
<collectingCountry id="9891766DAD11074F85C6FD7BFD865F8F" box="[421,512,677,702]" name="Albania" pageId="6" pageNumber="529">Albania</collectingCountry>
, 0 3.08.2003, leg. T. Rađa.
<typeStatus id="3F3D885FAD11074F8748FD78FC1E5F8F" box="[811,920,678,702]" pageId="6" pageNumber="529" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="8697AE38AD11074F87C5FD78FC7D5F8F" box="[934,1019,678,702]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="6" pageNumber="529">ZMUC</collectionCode>
,
<collectionCode id="8697AE38AD11074F8065FD78FBBE5F8F" box="[1030,1080,678,702]" pageId="6" pageNumber="529">IZB</collectionCode>
): two adult males (one with lost gonopods), seven juvenile males and
<specimenCount id="F680FD74AD11074F8658FD14FD3F5FD3" box="[571,697,713,738]" pageId="6" pageNumber="529" type="female">13 females</specimenCount>
: same data as
<typeStatus id="3F3D885FAD11074F873BFD17FC385FD3" box="[856,958,713,738]" pageId="6" pageNumber="529" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="A89C6576AD11074F84A4FD33FE5D5E1B" pageId="6" pageNumber="529" type="etymology">
<paragraph id="E03936FDAD11074F84A4FD33FE5D5E1B" blockId="6.[151,1436,641,918]" pageId="6" pageNumber="529">
<emphasis id="D2F2EAEFAD11074F84A4FD33FEC85E37" bold="true" box="[199,334,749,774]" pageId="6" pageNumber="529">Etymology.</emphasis>
The new species is named after Tonći Rađa, a well-known Croatian speleologist who discovered numerous new cave animals.
</paragraph>
</subSubSection>
<subSubSection id="A89C6576AD11074F84A4FCEBFDA25EA7" pageId="6" pageNumber="529" type="diagnosis">
<paragraph id="E03936FDAD11074F84A4FCEBFDA25EA7" blockId="6.[151,1436,641,918]" pageId="6" pageNumber="529">
<emphasis id="D2F2EAEFAD11074F84A4FCEBFEC75E7F" bold="true" box="[199,321,821,846]" pageId="6" pageNumber="529">Diagnosis.</emphasis>
Differs from
<taxonomicName id="27864D7EAD11074F85BEFCEBFC955E7F" box="[477,787,821,846]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="simplarius">
<emphasis id="D2F2EAEFAD11074F85BEFCEBFC955E7F" box="[477,787,821,846]" italics="true" pageId="6" pageNumber="529">Thassoblaniulus simplarius</emphasis>
</taxonomicName>
(see
<tableCitation id="AD040346AD11074F8733FCEBFC225E7F" box="[848,932,821,846]" captionStart="TABLE 2" captionStartId="6.[151,239,961,984]" captionTargetBox="[159,1427,1012,1804]" captionTargetPageId="6" captionText="TABLE 2. Differences between Thassoblaniulus simplarius Mauriès, 1985 and T. radjai sp. n." httpUri="http://table.plazi.org/id/B4F96675AD11074F84F4FC1FFB0C5EE9" pageId="6" pageNumber="529" tableUuid="B4F96675AD11074F84F4FC1FFB0C5EE9">Table 2</tableCitation>
) and all other blaniulid species by the shape of the distal part of the anterior gonopod coxal processes, which are characterised by very broad lateral shoulders and axe-like postero-mesal flanges.
</paragraph>
</subSubSection>
<caption id="B4F96675AD11074F84F4FC1FFB0C5EE9" ID-Table-UUID="B4F96675AD11074F84F4FC1FFB0C5EE9" box="[151,1162,961,984]" httpUri="http://table.plazi.org/id/B4F96675AD11074F84F4FC1FFB0C5EE9" pageId="6" pageNumber="529" targetBox="[159,1427,1012,1804]" targetIsTable="true" targetPageId="6">
<paragraph id="E03936FDAD11074F84F4FC1FFB0C5EE9" blockId="6.[151,1162,961,984]" box="[151,1162,961,984]" pageId="6" pageNumber="529">
<emphasis id="D2F2EAEFAD11074F84F4FC1FFE8F5EE9" bold="true" box="[151,265,961,984]" pageId="6" pageNumber="529">TABLE 2.</emphasis>
Differences between
<taxonomicName id="27864D7EAD11074F8591FC1FFC3F5EE9" authority="Mauries, 1985" authorityName="Mauries" authorityYear="1985" box="[498,953,961,984]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="simplarius">
<emphasis id="D2F2EAEFAD11074F8591FC1FFC9E5EE9" box="[498,792,961,984]" italics="true" pageId="6" pageNumber="529">Thassoblaniulus simplarius</emphasis>
Mauriès, 1985
</taxonomicName>
and
<taxonomicName id="27864D7EAD11074F878EFC1FFBCE5EE9" box="[1005,1096,961,984]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="radjai" status="sp. nov.">
<emphasis id="D2F2EAEFAD11074F878EFC1FFBCE5EE9" box="[1005,1096,961,984]" italics="true" pageId="6" pageNumber="529">T. radjai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C9C15794AD11074F802CFC1FFB0C5EE9" box="[1103,1162,961,984]" pageId="6" pageNumber="529" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="E03936FDAD11074F84C6FC2AFB245A3D" pageId="6" pageNumber="529">
<table id="9286C45DAD11F8B684FCFC2AFA155A3D" box="[159,1427,1012,1804]" gridcols="3" gridrows="8" pageId="6" pageNumber="529">
<tr id="5EB634BFAD11F8B684FCFC2AFA15593B" box="[159,1427,1012,1034]" gridrow="0" pageId="6" pageNumber="529">
<th id="1D675DC3AD11F8B684FCFC2AFE7C593B" box="[159,506,1012,1034]" gridcol="0" gridrow="0" pageId="6" pageNumber="529">Characters</th>
<th id="1D675DC3AD11F8B6865BFC2AFC6E593B" box="[568,1000,1012,1034]" gridcol="1" gridrow="0" pageId="6" pageNumber="529">
<taxonomicName id="27864D7EAD11074F865BFC2AFC6E593B" authority="Mauries, 1985" authorityName="Mauries" authorityYear="1985" box="[568,1000,1012,1034]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="simplarius">
<emphasis id="D2F2EAEFAD11074F865BFC2AFCC9593B" box="[568,847,1012,1034]" italics="true" pageId="6" pageNumber="529">Thassoblaniulus simplarius</emphasis>
Mauriès, 1985
</taxonomicName>
</th>
<th id="1D675DC3AD11F8B68068FC2AFA15593B" box="[1035,1427,1012,1034]" gridcol="2" gridrow="0" pageId="6" pageNumber="529">
<taxonomicName id="27864D7EAD11074F8068FC2AFB73593B" box="[1035,1269,1012,1034]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="529" phylum="Arthropoda" rank="species" species="radjai" status="sp. nov.">
<emphasis id="D2F2EAEFAD11074F8068FC2AFB73593B" box="[1035,1269,1012,1034]" italics="true" pageId="6" pageNumber="529">Thassoblaniulus radjai</emphasis>
</taxonomicName>
<emphasis id="D2F2EAEFAD11074F809FFC2AFAB2593B" bold="true" box="[1276,1332,1012,1034]" pageId="6" pageNumber="529">
<taxonomicNameLabel id="C9C15794AD11074F809FFC2AFAB2593B" box="[1276,1332,1012,1034]" pageId="6" pageNumber="529" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="5EB634BFAD11F8B684FCFBE2FA155963" box="[159,1427,1084,1106]" gridrow="1" pageId="6" pageNumber="529">
<th id="1D675DC3AD11F8B684FCFBE2FE7C5963" box="[159,506,1084,1106]" gridcol="0" gridrow="1" pageId="6" pageNumber="529">Ocelli</th>
<td id="1D675DC3AD11F8B6865BFBE2FC6E5963" box="[568,1000,1084,1106]" gridcol="1" gridrow="1" pageId="6" pageNumber="529">♂♂ 3–6, ♀♀ 4–7; one or two rows</td>
<td id="1D675DC3AD11F8B68068FBE2FA155963" box="[1035,1427,1084,1106]" gridcol="2" gridrow="1" pageId="6" pageNumber="529">2–4 in both sexes; one row</td>
</tr>
<tr id="5EB634BFAD11F8B684FCFB5AFA155831" box="[159,1427,1156,1280]" gridrow="2" pageId="6" pageNumber="529">
<th id="1D675DC3AD11F8B684FCFB5AFE7C5831" box="[159,506,1156,1280]" gridcol="0" gridrow="2" pageId="6" pageNumber="529">L and H of body (mm) Length/height ratio of claws of midbody legs</th>
<td id="1D675DC3AD11F8B6865BFB5AFC6E5831" box="[568,1000,1156,1280]" gridcol="1" gridrow="2" pageId="6" pageNumber="529">♂♂ 6–10 L, 0.35–0.6 H ♀♀ up to 13 L, 0.7 H 4</td>
<td id="1D675DC3AD11F8B68068FB5AFA155831" box="[1035,1427,1156,1280]" gridcol="2" gridrow="2" pageId="6" pageNumber="529">♂♂ 10–13 L, 0.6–0.7 H ♀♀ up to 15 L, 0.78 H 6</td>
</tr>
<tr id="5EB634BFAD11F8B684FCFAECFA155879" box="[159,1427,1330,1352]" gridrow="3" pageId="6" pageNumber="529">
<th id="1D675DC3AD11F8B684FCFAECFE7C5879" box="[159,506,1330,1352]" gridcol="0" gridrow="3" pageId="6" pageNumber="529">First pair of legs in ♂♂</th>
<td id="1D675DC3AD11F8B6865BFAECFC6E5879" box="[568,1000,1330,1352]" gridcol="1" gridrow="3" pageId="6" pageNumber="529">Without claws</td>
<td id="1D675DC3AD11F8B68068FAECFA155879" box="[1035,1427,1330,1352]" gridcol="2" gridrow="3" pageId="6" pageNumber="529">With duble claws</td>
</tr>
<tr id="5EB634BFAD11F8B684FCFAA4FA1558A1" box="[159,1427,1402,1424]" gridrow="4" pageId="6" pageNumber="529">
<th id="1D675DC3AD11F8B684FCFAA4FE7C58A1" box="[159,506,1402,1424]" gridcol="0" gridrow="4" pageId="6" pageNumber="529">Telopodites of anterior gonopods</th>
<td id="1D675DC3AD11F8B6865BFAA4FC6E58A1" box="[568,1000,1402,1424]" gridcol="1" gridrow="4" pageId="6" pageNumber="529">Less than ¼ the length of coxal processes</td>
<td id="1D675DC3AD11F8B68068FAA4FA1558A1" box="[1035,1427,1402,1424]" gridcol="2" gridrow="4" pageId="6" pageNumber="529">ca. ¼ to ⅓ of length of coxal processes</td>
</tr>
<tr id="5EB634BFAD11F8B684FCFA1CFA155B0E" box="[159,1427,1474,1599]" gridrow="5" pageId="6" pageNumber="529">
<th id="1D675DC3AD11F8B684FCFA1CFE7C5B0E" box="[159,506,1474,1599]" gridcol="0" gridrow="5" pageId="6" pageNumber="529">Lateral margins of distal parts of anterior gonopods Postero-mesal flange of anterior gonopods</th>
<td id="1D675DC3AD11F8B6865BFA1CFC6E5B0E" box="[568,1000,1474,1599]" gridcol="1" gridrow="5" pageId="6" pageNumber="529">Gradually tapering to the top Bilobed</td>
<td id="1D675DC3AD11F8B68068FA1CFA155B0E" box="[1035,1427,1474,1599]" gridcol="2" gridrow="5" pageId="6" pageNumber="529">Suddenly getting narrow, forming lateral shoulder Axe-like</td>
</tr>
<tr id="5EB634BFAD11F8B684FCF9AFFA155BB6" box="[159,1427,1649,1671]" gridrow="6" pageId="6" pageNumber="529">
<th id="1D675DC3AD11F8B684FCF9AFFE7C5BB6" box="[159,506,1649,1671]" gridcol="0" gridrow="6" pageId="6" pageNumber="529">Basal part of anterior gonopods</th>
<td id="1D675DC3AD11F8B6865BF9AFFC6E5BB6" box="[568,1000,1649,1671]" gridcol="1" gridrow="6" pageId="6" pageNumber="529">Low</td>
<td id="1D675DC3AD11F8B68068F9AFFA155BB6" box="[1035,1427,1649,1671]" gridcol="2" gridrow="6" pageId="6" pageNumber="529">High</td>
</tr>
<tr id="5EB634BFAD11F8B684FCF967FA155A3D" box="[159,1427,1721,1804]" gridrow="7" pageId="6" pageNumber="529">
<th id="1D675DC3AD11F8B684FCF967FE7C5A3D" box="[159,506,1721,1804]" gridcol="0" gridrow="7" pageId="6" pageNumber="529">Posterior gonopods in lateral view</th>
<td id="1D675DC3AD11F8B6865BF967FC6E5A3D" box="[568,1000,1721,1804]" gridcol="1" gridrow="7" pageId="6" pageNumber="529">With narrow basal part which gradually widens up to leaf-shaped lamella</td>
<td id="1D675DC3AD11F8B68068F967FA155A3D" box="[1035,1427,1721,1804]" gridcol="2" gridrow="7" pageId="6" pageNumber="529">With somewhat broader basal part which gradually tapers up to the leafshaped lamella</td>
</tr>
</table>
</paragraph>
<subSubSection id="A89C6576AD11074484A4F893FE0B5C2C" lastPageId="13" lastPageNumber="536" pageId="6" pageNumber="529" type="description">
<paragraph id="E03936FDAD11074F84A4F893FB595AC7" blockId="6.[151,1437,1869,2038]" pageId="6" pageNumber="529">
<emphasis id="D2F2EAEFAD11074F84A4F893FEDF5A57" bold="true" box="[199,345,1869,1894]" pageId="6" pageNumber="529">Description.</emphasis>
<typeStatus id="3F3D885FAD11074F8503F893FE4E5A57" box="[352,456,1869,1894]" pageId="6" pageNumber="529" type="holotype">Holotype</typeStatus>
male: L
<quantity id="277E9B18AD11074F864DF890FDF95A54" box="[558,639,1870,1894]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="529" unit="mm" value="10.0">10 mm</quantity>
, H
<quantity id="277E9B18AD11074F86C4F893FD795A54" box="[679,767,1869,1894]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="529" unit="mm" value="0.6">0.6 mm</quantity>
,
<typeStatus id="3F3D885FAD11074F8768F890FCED5A57" box="[779,875,1870,1894]" pageId="6" pageNumber="529" type="paratype">paratype</typeStatus>
adult male: L
<quantity id="277E9B18AD11074F8072F890FBE75A54" box="[1041,1121,1870,1894]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="529" unit="mm" value="11.0">11 mm</quantity>
, H
<quantity id="277E9B18AD11074F80EAF893FB775A54" box="[1161,1265,1869,1894]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="529" unit="mm" value="0.65">0.65 mm</quantity>
;
<typeStatus id="3F3D885FAD11074F809EF890FADB5A57" box="[1277,1373,1870,1894]" pageId="6" pageNumber="529" type="paratype">paratype</typeStatus>
adult male (with lost gonopods): L
<quantity id="277E9B18AD11074F8592F8ACFDC35AB8" box="[497,581,1906,1930]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="529" unit="mm" value="13.0">13 mm</quantity>
, H
<quantity id="277E9B18AD11074F8612F8ACFD4A5AB8" box="[625,716,1906,1930]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="529" unit="mm" value="0.7">0.7 mm</quantity>
, females up to L
<quantity id="277E9B18AD11074F87FFF8AFFC765AB8" box="[924,1008,1905,1930]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="529" unit="mm" value="15.0">15 mm</quantity>
, H
<quantity id="277E9B18AD11074F807FF8ACFB005AB8" box="[1052,1158,1906,1930]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="6" pageNumber="529" unit="mm" value="0.78">0.78 mm</quantity>
. Number of body rings:
<typeStatus id="3F3D885FAD11074F84F4F84BFF7F5A9F" box="[151,249,1941,1966]" pageId="6" pageNumber="529" type="holotype">holotype</typeStatus>
male: 31 podous + 2 apodous + telson;
<typeStatus id="3F3D885FAD11074F86A9F848FCAC5A9F" box="[714,810,1942,1966]" pageId="6" pageNumber="529" type="paratype">paratype</typeStatus>
adult male: 34 podous + 2 apodous + telson,
<typeStatus id="3F3D885FAD11074F815FF848FA1A5A9F" box="[1340,1436,1942,1966]" pageId="6" pageNumber="529" type="paratype">paratype</typeStatus>
adult male (with lost gonopods): 36 podous + 2 apodous + telson, juvenile males: 26–37 body rings + telson (including 1–4 apodous rings), females: 28–47 body rings + telson (including 1–5 apodous rings).
</paragraph>
<caption id="B4F96675AD10074E84F4F841FF405AC3" httpUri="https://zenodo.org/record/235354/files/figure.png" pageId="7" pageNumber="530" targetBox="[151,1436,193,1929]" targetPageId="7">
<paragraph id="E03936FDAD10074E84F4F841FF405AC3" blockId="7.[151,1436,1951,2034]" pageId="7" pageNumber="530">
<emphasis id="D2F2EAEFAD10074E84F4F841FED05A84" bold="true" box="[151,342,1951,1973]" pageId="7" pageNumber="530">FIGURES 13–16.</emphasis>
<taxonomicName id="27864D7EAD10074E853CF841FDCB5A84" box="[351,589,1951,1973]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="530" phylum="Arthropoda" rank="species" species="radjai" status="sp. nov.">
<emphasis id="D2F2EAEFAD10074E853CF841FDCB5A84" box="[351,589,1951,1973]" italics="true" pageId="7" pageNumber="530">Thassoblaniulus radjai</emphasis>
</taxonomicName>
<emphasis id="D2F2EAEFAD10074E8635F841FD145A84" bold="true" box="[598,658,1951,1973]" pageId="7" pageNumber="530">
<taxonomicNameLabel id="C9C15794AD10074E8635F841FD145A84" box="[598,658,1951,1973]" pageId="7" pageNumber="530" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, paratype male anterior gonopods. 13. anterior view; 14. lateral view; 15. posterior view; 16. distal view.
<emphasis id="D2F2EAEFAD10074E85BBF860FE665AE5" bold="true" box="[472,480,1982,2004]" pageId="7" pageNumber="530">f</emphasis>
: postero-mesal flange,
<emphasis id="D2F2EAEFAD10074E86ACF860FD515AE5" bold="true" box="[719,727,1982,2004]" pageId="7" pageNumber="530">t</emphasis>
: telopodite,
<emphasis id="D2F2EAEFAD10074E873BF860FCF75AE5" bold="true" box="[856,881,1982,2004]" pageId="7" pageNumber="530">cp</emphasis>
: coxal processes,
<emphasis id="D2F2EAEFAD10074E8048F860FBCC5AE5" bold="true" box="[1067,1098,1982,2004]" pageId="7" pageNumber="530">lsh</emphasis>
: lateral shoulder. Scale lines: 0.1 mm.
</paragraph>
</caption>
<caption id="B4F96675AD1F074184F4F8F2FAF95A50" httpUri="https://zenodo.org/record/235355/files/figure.png" pageId="8" pageNumber="531" targetBox="[151,1436,193,1814]" targetPageId="8">
<paragraph id="E03936FDAD1F074184F4F8F2FAF95A50" blockId="8.[151,1436,1836,1889]" pageId="8" pageNumber="531">
<emphasis id="D2F2EAEFAD1F074184F4F8F2FED35A73" bold="true" box="[151,341,1836,1858]" pageId="8" pageNumber="531">FIGURES 17–20.</emphasis>
<taxonomicName id="27864D7EAD1F07418538F8F2FDC05A73" box="[347,582,1836,1858]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="531" phylum="Arthropoda" rank="species" species="radjai" status="sp. nov.">
<emphasis id="D2F2EAEFAD1F07418538F8F2FDC05A73" box="[347,582,1836,1858]" italics="true" pageId="8" pageNumber="531">Thassoblaniulus radjai</emphasis>
</taxonomicName>
<emphasis id="D2F2EAEFAD1F0741862EF8F2FD015A73" bold="true" box="[589,647,1836,1858]" pageId="8" pageNumber="531">
<taxonomicNameLabel id="C9C15794AD1F0741862EF8F2FD015A73" box="[589,647,1836,1858]" pageId="8" pageNumber="531" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, paratype male posterior gonopod tip. 17. lateral view; 18. anterior view; 19. mesal view; 20. posterior view. –
<emphasis id="D2F2EAEFAD1F0741858FF895FE7C5A50" bold="true" box="[492,506,1867,1889]" pageId="8" pageNumber="531">ll</emphasis>
: leaf-shaped lamella,
<emphasis id="D2F2EAEFAD1F074186B9F895FD765A50" bold="true" box="[730,752,1867,1889]" pageId="8" pageNumber="531">so</emphasis>
: solenomere. Scale lines: 0.1 for 17, 18 and 19; 0.05 mm for 20.
</paragraph>
</caption>
<caption id="B4F96675AD1E074084F4FAD3FEF45873" httpUri="https://zenodo.org/record/235356/files/figure.png" pageId="9" pageNumber="532" targetBox="[151,1436,193,1271]" targetPageId="9">
<paragraph id="E03936FDAD1E074084F4FAD3FEF45873" blockId="9.[151,1437,1293,1346]" pageId="9" pageNumber="532">
<emphasis id="D2F2EAEFAD1E074084F4FAD3FEF95812" bold="true" box="[151,383,1293,1315]" pageId="9" pageNumber="532">FIGURES 21 and 22.</emphasis>
<taxonomicName id="27864D7EAD1E074085E5FAD3FDF75812" box="[390,625,1293,1315]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="532" phylum="Arthropoda" rank="species" species="radjai" status="sp. nov.">
<emphasis id="D2F2EAEFAD1E074085E5FAD3FDF75812" box="[390,625,1293,1315]" italics="true" pageId="9" pageNumber="532">Thassoblaniulus radjai</emphasis>
</taxonomicName>
<emphasis id="D2F2EAEFAD1E0740861BFAD3FD345812" bold="true" box="[632,690,1293,1315]" pageId="9" pageNumber="532">
<taxonomicNameLabel id="C9C15794AD1E0740861BFAD3FD345812" box="[632,690,1293,1315]" pageId="9" pageNumber="532" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, paratype male posterior gonopod. 21. solenomere; 22. detail of lamella. Scale lines: 0.01 mm.
</paragraph>
</caption>
<paragraph id="E03936FDAD1E074084A4FAB0FAD158B6" blockId="9.[151,1436,1390,1991]" box="[199,1367,1390,1415]" pageId="9" pageNumber="532">
<emphasis id="D2F2EAEFAD1E074084A4FAB0FE9D58B6" box="[199,283,1390,1415]" italics="true" pageId="9" pageNumber="532">Colour.</emphasis>
Yellowish white to pale brown. Some specimens with clearly visible, large and darker ozadenes.
</paragraph>
<paragraph id="E03936FDAD1E074084A4FA4CFAC15B0A" blockId="9.[151,1436,1390,1991]" pageId="9" pageNumber="532">
<emphasis id="D2F2EAEFAD1E074084A4FA4CFE8D589A" box="[199,267,1426,1451]" italics="true" pageId="9" pageNumber="532">Head.</emphasis>
With one row of a very small number of ocelli on each side.
<typeStatus id="3F3D885FAD1E074087CDFA4CFB90589A" box="[942,1046,1426,1451]" pageId="9" pageNumber="532" type="holotype">Holotype</typeStatus>
and
<typeStatus id="3F3D885FAD1E0740802FFA4DFB2A589A" box="[1100,1196,1427,1451]" pageId="9" pageNumber="532" type="paratype">paratype</typeStatus>
adult males with 2+2 ocelli,
<typeStatus id="3F3D885FAD1E07408485FA69FEC058FE" box="[230,326,1463,1487]" pageId="9" pageNumber="532" type="paratype">paratype</typeStatus>
adult male (with lost gonopods) with 4+4 ocelli, juvenile males and females with 2–4 ocelli on each side (2+2, 2+3, 3+2, 3+3, 3+4, 4+3, 4+4). Length of antennae 169 % of H in
<typeStatus id="3F3D885FAD1E0740802AFA05FB2F58C2" box="[1097,1193,1499,1523]" pageId="9" pageNumber="532" type="paratype">paratype</typeStatus>
male, 130 % of H in female. Relative lengths of antennomeres 1–8 (8 = apical sensilla):
<date id="9438103DAD1E074087EAFA20FC5B5B26" box="[905,989,1534,1559]" pageId="9" pageNumber="532" value="1918-07-19">7/19/18</date>
/15/
<date id="9438103DAD1E0740806BFA20FBD95B26" box="[1032,1119,1534,1559]" pageId="9" pageNumber="532" value="1920-05-14">20/14/5</date>
/2 % in
<typeStatus id="3F3D885FAD1E074080D4FA21FA915B26" box="[1207,1303,1535,1559]" pageId="9" pageNumber="532" type="paratype">paratype</typeStatus>
male,
<date id="9438103DAD1E07408100FA21FF365B0A" pageId="9" pageNumber="532" value="1915-08-17">8/17/ 15</date>
/17/
<date id="9438103DAD1E074084BFF9FCFEB55B0A" box="[220,307,1570,1595]" pageId="9" pageNumber="532" value="1920-05-14">20/14/5</date>
/
<quantity id="277E9B18AD1E07408558F9FCFEE15B0A" box="[315,359,1570,1595]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="9" pageNumber="532" unit="in" value="4.0">4 in</quantity>
female. In males, mandibles with usual parrot-bill-like modification of cardo and stipes.
</paragraph>
<paragraph id="E03936FDAD1E074084A4F998FAB35BB2" blockId="9.[151,1436,1390,1991]" pageId="9" pageNumber="532">
<emphasis id="D2F2EAEFAD1E074084A4F998FEEA5B6F" box="[199,364,1606,1631]" italics="true" pageId="9" pageNumber="532">Midbody rings</emphasis>
with ca. 14 short setae on posterior margin, length of setae ca. 6 % of H in
<typeStatus id="3F3D885FAD1E074080A1F999FAA45B6E" box="[1218,1314,1607,1631]" pageId="9" pageNumber="532" type="paratype">paratype</typeStatus>
male, 5 % of H in female. Latero-ventral sides of the rings with longitudinal striations which extend into prozonites.
</paragraph>
<paragraph id="E03936FDAD1E074084A4F951FF535A22" blockId="9.[151,1436,1390,1991]" pageId="9" pageNumber="532">
<emphasis id="D2F2EAEFAD1E074084A4F951FF7A5B97" box="[199,252,1679,1702]" italics="true" pageId="9" pageNumber="532">Legs</emphasis>
: length 92 % of H in
<typeStatus id="3F3D885FAD1E07408598F951FDDA5B96" box="[507,604,1679,1703]" pageId="9" pageNumber="532" type="paratype">paratype</typeStatus>
male, 77 % of H in female. Relative lengths of podomeres from coxa to claw 16/
<date id="9438103DAD1E07408494F96CFEDC5BFA" box="[247,346,1714,1739]" pageId="9" pageNumber="532" value="2014-10-17">14/17/10</date>
/
<date id="9438103DAD1E07408500F96CFE405BFA" box="[355,454,1714,1739]" pageId="9" pageNumber="532" value="2010-12-21">10/21/12</date>
% in
<typeStatus id="3F3D885FAD1E07408668F96DFDED5BFA" box="[523,619,1715,1739]" pageId="9" pageNumber="532" type="paratype">paratype</typeStatus>
male, 15/
<date id="9438103DAD1E074086B8F96CFCA95BFA" box="[731,815,1714,1739]" pageId="9" pageNumber="532" value="1915-09-17">15/17/9</date>
/
<date id="9438103DAD1E0740875BF96CFC165BFA" box="[824,912,1714,1739]" pageId="9" pageNumber="532" value="2013-09-22">9/22/13</date>
% in female; claw length/height
<quantity id="277E9B18AD1E0740816BF96CFAB25BFA" box="[1288,1332,1714,1739]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="9" pageNumber="532" unit="in" value="6.0">6 in</quantity>
<typeStatus id="3F3D885FAD1E0740815FF96DFA1A5BFA" box="[1340,1436,1715,1739]" pageId="9" pageNumber="532" type="paratype">paratype</typeStatus>
male and female. No accessory claw. Lanceolate setae present on femur (1), postfemur (2) and tibia (2) in
<typeStatus id="3F3D885FAD1E0740815FF909FA1A5BDE" box="[1340,1436,1751,1775]" pageId="9" pageNumber="532" type="paratype">paratype</typeStatus>
male.
</paragraph>
<paragraph id="E03936FDAD1E074084A4F8C1FE3C5A4E" blockId="9.[151,1436,1390,1991]" pageId="9" pageNumber="532">
<emphasis id="D2F2EAEFAD1E074084A4F8C1FE435A06" box="[199,453,1822,1847]" italics="true" pageId="9" pageNumber="532">First pair of male legs</emphasis>
(
<figureCitation id="78BD2A78AD1E074085B6F8C0FDD35A06" box="[469,597,1822,1847]" captionStart="FIGURES 23 – 26" captionStartId="10.[151,264,1826,1848]" captionTargetBox="[151,1436,193,1804]" captionTargetId="figure@10.[151,1436,193,1805]" captionTargetPageId="10" captionText="FIGURES 23 – 26. Thassoblaniulus radjai sp. n., paratype male. 23. first pair of legs, antero-lateral view; 24. first pair of legs, anterior view; 25. second pair of legs with penis, posterior view; 26. body ring 7, ventral view. – p: penis, cl: claw, ls: lanceolate seta, ta: tarsus, ti: tibia. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/235357/files/figure.png" pageId="9" pageNumber="532">Figs 23, 24</figureCitation>
and
<figureCitation id="78BD2A78AD1E074086EEF8C1FD2C5A06" box="[653,682,1823,1847]" captionStart="FIGURES 27 – 32" captionStartId="11.[151,264,1942,1964]" captionTargetBox="[210,1361,193,1917]" captionTargetId="figure@11.[210,1361,193,1917]" captionTargetPageId="11" captionText="FIGURES 27 – 32. Thassoblaniulus radjai sp. n., holotype male. 27. anterior gonopods, posterior view; 28. anterior gonopods, lateral view; 29. penis, posterior view; 30. posterior gonopod, lateral view; 31. first pair of legs, posterior view; 32. second leg. Scale line: 0.4 mm." httpUri="https://zenodo.org/record/235358/files/figure.png" pageId="9" pageNumber="532">31</figureCitation>
) simple, with six short podomeres. Division between tibia (
<emphasis id="D2F2EAEFAD1E07408132F8C0FAE45A06" bold="true" box="[1361,1378,1822,1847]" pageId="9" pageNumber="532">ti</emphasis>
) and tarsus (
<emphasis id="D2F2EAEFAD1E07408489F89CFE875A6A" bold="true" box="[234,257,1858,1883]" pageId="9" pageNumber="532">ta</emphasis>
) incomplete and poorly visible. Prefemur with one long ventral seta. Tarsus with a rudimentary double claw (
<emphasis id="D2F2EAEFAD1E074084BFF8B8FF765A4E" bold="true" box="[220,240,1894,1919]" pageId="9" pageNumber="532">cl</emphasis>
) on a small knob.
</paragraph>
<paragraph id="E03936FDAD1E074084A4F854FF575AF6" blockId="9.[151,1436,1390,1991]" pageId="9" pageNumber="532">
<emphasis id="D2F2EAEFAD1E074084A4F854FE635A92" box="[199,485,1930,1955]" italics="true" pageId="9" pageNumber="532">Second pair of male legs</emphasis>
(
<figureCitation id="78BD2A78AD1E07408594F854FDC85A92" box="[503,590,1930,1955]" captionStart="FIGURES 23 – 26" captionStartId="10.[151,264,1826,1848]" captionTargetBox="[151,1436,193,1804]" captionTargetId="figure@10.[151,1436,193,1805]" captionTargetPageId="10" captionText="FIGURES 23 – 26. Thassoblaniulus radjai sp. n., paratype male. 23. first pair of legs, antero-lateral view; 24. first pair of legs, anterior view; 25. second pair of legs with penis, posterior view; 26. body ring 7, ventral view. – p: penis, cl: claw, ls: lanceolate seta, ta: tarsus, ti: tibia. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/235357/files/figure.png" pageId="9" pageNumber="532">Figs 25</figureCitation>
and
<figureCitation id="78BD2A78AD1E074086EAF855FD205A92" box="[649,678,1931,1955]" captionStart="FIGURES 27 – 32" captionStartId="11.[151,264,1942,1964]" captionTargetBox="[210,1361,193,1917]" captionTargetId="figure@11.[210,1361,193,1917]" captionTargetPageId="11" captionText="FIGURES 27 – 32. Thassoblaniulus radjai sp. n., holotype male. 27. anterior gonopods, posterior view; 28. anterior gonopods, lateral view; 29. penis, posterior view; 30. posterior gonopod, lateral view; 31. first pair of legs, posterior view; 32. second leg. Scale line: 0.4 mm." httpUri="https://zenodo.org/record/235358/files/figure.png" pageId="9" pageNumber="532">32</figureCitation>
) with one and two ventral, lanceolate setae (
<emphasis id="D2F2EAEFAD1E074080CDF854FB475A92" bold="true" box="[1198,1217,1930,1955]" pageId="9" pageNumber="532">ls</emphasis>
) on postfemur and tibia.
</paragraph>
<caption id="B4F96675AD1D074384F4F8FCFDDC5A44" httpUri="https://zenodo.org/record/235357/files/figure.png" pageId="10" pageNumber="533" targetBox="[151,1436,193,1804]" targetPageId="10">
<paragraph id="E03936FDAD1D074384F4F8FCFDDC5A44" blockId="10.[151,1436,1826,1909]" pageId="10" pageNumber="533">
<emphasis id="D2F2EAEFAD1D074384F4F8FCFED25A09" bold="true" box="[151,340,1826,1848]" pageId="10" pageNumber="533">FIGURES 23–26.</emphasis>
<taxonomicName id="27864D7EAD1D07438539F8FCFDC25A09" box="[346,580,1826,1848]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="533" phylum="Arthropoda" rank="species" species="radjai" status="sp. nov.">
<emphasis id="D2F2EAEFAD1D07438539F8FCFDC25A09" box="[346,580,1826,1848]" italics="true" pageId="10" pageNumber="533">Thassoblaniulus radjai</emphasis>
</taxonomicName>
<emphasis id="D2F2EAEFAD1D07438628F8FCFD025A09" bold="true" box="[587,644,1826,1848]" pageId="10" pageNumber="533">
<taxonomicNameLabel id="C9C15794AD1D07438628F8FCFD025A09" box="[587,644,1826,1848]" pageId="10" pageNumber="533" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, paratype male. 23. first pair of legs, antero-lateral view; 24. first pair of legs, anterior view; 25. second pair of legs with penis, posterior view; 26. body ring 7, ventral view. –
<emphasis id="D2F2EAEFAD1D07438001F89FFBF65A66" bold="true" box="[1122,1136,1857,1879]" pageId="10" pageNumber="533">p</emphasis>
: penis,
<emphasis id="D2F2EAEFAD1D074380DDF89FFB565A66" bold="true" box="[1214,1232,1857,1879]" pageId="10" pageNumber="533">cl</emphasis>
: claw,
<emphasis id="D2F2EAEFAD1D0743817BF89FFAAF5A66" bold="true" box="[1304,1321,1857,1879]" pageId="10" pageNumber="533">ls</emphasis>
: lanceolate seta,
<emphasis id="D2F2EAEFAD1D074384A8F881FF665A44" bold="true" box="[203,224,1887,1909]" pageId="10" pageNumber="533">ta</emphasis>
: tarsus,
<emphasis id="D2F2EAEFAD1D07438556F881FEC35A44" bold="true" box="[309,325,1887,1909]" pageId="10" pageNumber="533">ti</emphasis>
: tibia. Scale lines: 0.1 mm.
</paragraph>
</caption>
<caption id="B4F96675AD1C074284F4F848FEDA5ADB" httpUri="https://zenodo.org/record/235358/files/figure.png" pageId="11" pageNumber="534" targetBox="[210,1361,193,1917]" targetPageId="11">
<paragraph id="E03936FDAD1C074284F4F848FEDA5ADB" blockId="11.[151,1437,1942,2026]" pageId="11" pageNumber="534">
<emphasis id="D2F2EAEFAD1C074284F4F848FED25A9D" bold="true" box="[151,340,1942,1964]" pageId="11" pageNumber="534">FIGURES 27–32.</emphasis>
<taxonomicName id="27864D7EAD1C07428539F848FDC25A9D" box="[346,580,1942,1964]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="11" pageNumber="534" phylum="Arthropoda" rank="species" species="radjai" status="sp. nov.">
<emphasis id="D2F2EAEFAD1C07428539F848FDC25A9D" box="[346,580,1942,1964]" italics="true" pageId="11" pageNumber="534">Thassoblaniulus radjai</emphasis>
</taxonomicName>
<emphasis id="D2F2EAEFAD1C07428628F848FD055A9D" bold="true" box="[587,643,1942,1964]" pageId="11" pageNumber="534">
<taxonomicNameLabel id="C9C15794AD1C07428628F848FD055A9D" box="[587,643,1942,1964]" pageId="11" pageNumber="534" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype male. 27. anterior gonopods, posterior view; 28. anterior gonopods, lateral view; 29. penis, posterior view; 30. posterior gonopod, lateral view; 31. first pair of legs, posterior view; 32. second leg. Scale line: 0.4 mm.
</paragraph>
</caption>
<caption id="B4F96675AD1B074584F4FB44FEAD59FE" httpUri="https://zenodo.org/record/235359/files/figure.png" pageId="12" pageNumber="535" targetBox="[151,1436,193,1156]" targetPageId="12">
<paragraph id="E03936FDAD1B074584F4FB44FEAD59FE" blockId="12.[151,1436,1178,1231]" pageId="12" pageNumber="535">
<emphasis id="D2F2EAEFAD1B074584F4FB44FE995981" bold="true" box="[151,287,1178,1200]" pageId="12" pageNumber="535">FIGURE 33.</emphasis>
<taxonomicName id="27864D7EAD1B07458546FB44FD895981" box="[293,527,1178,1200]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="12" pageNumber="535" phylum="Arthropoda" rank="species" species="radjai" status="sp. nov.">
<emphasis id="D2F2EAEFAD1B07458546FB44FD895981" box="[293,527,1178,1200]" italics="true" pageId="12" pageNumber="535">Thassoblaniulus radjai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C9C15794AD1B07458676FB44FDCB5981" box="[533,589,1178,1200]" pageId="12" pageNumber="535" rank="species">
<emphasis id="D2F2EAEFAD1B07458676FB44FDCB5981" bold="true" box="[533,589,1178,1200]" pageId="12" pageNumber="535">sp. n.</emphasis>
</taxonomicNameLabel>
, female vulva, lateral view.
<emphasis id="D2F2EAEFAD1B07458709FB44FCFE5981" bold="true" box="[874,888,1178,1200]" pageId="12" pageNumber="535">b</emphasis>
: bursa,
<emphasis id="D2F2EAEFAD1B074587AAFB44FC535981" bold="true" box="[969,981,1178,1200]" pageId="12" pageNumber="535">o</emphasis>
: operculum,
<emphasis id="D2F2EAEFAD1B07458038FB44FBF65981" bold="true" box="[1115,1136,1178,1200]" pageId="12" pageNumber="535">rs</emphasis>
: receptaculum seminis. Scale line: 0.05 mm.
</paragraph>
</caption>
<paragraph id="E03936FDAD1B074584A4FB23FD205809" blockId="12.[151,1437,1275,2021]" pageId="12" pageNumber="535">
<emphasis id="D2F2EAEFAD1B074584A4FB23FE805825" box="[199,262,1277,1300]" italics="true" pageId="12" pageNumber="535">Penis</emphasis>
(
<figureCitation id="78BD2A78AD1B07458570FB25FEEF5825" box="[275,361,1275,1300]" captionStart="FIGURES 23 – 26" captionStartId="10.[151,264,1826,1848]" captionTargetBox="[151,1436,193,1804]" captionTargetId="figure@10.[151,1436,193,1805]" captionTargetPageId="10" captionText="FIGURES 23 – 26. Thassoblaniulus radjai sp. n., paratype male. 23. first pair of legs, antero-lateral view; 24. first pair of legs, anterior view; 25. second pair of legs with penis, posterior view; 26. body ring 7, ventral view. – p: penis, cl: claw, ls: lanceolate seta, ta: tarsus, ti: tibia. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/235357/files/figure.png" pageId="12" pageNumber="535">Figs 25</figureCitation>
,
<emphasis id="D2F2EAEFAD1B07458516FB25FE025825" bold="true" box="[373,388,1275,1300]" pageId="12" pageNumber="535">p</emphasis>
and 29) in the form of a high isosceles subtriangle or triangle, gradually tapering from base to top, apically truncate or completely acuminate.
</paragraph>
<paragraph id="E03936FDAD1B074584A4FA9DFADF586D" blockId="12.[151,1437,1275,2021]" box="[199,1369,1347,1372]" pageId="12" pageNumber="535">
<emphasis id="D2F2EAEFAD1B074584A4FA9DFDEE586D" box="[199,616,1347,1372]" italics="true" pageId="12" pageNumber="535">
Ventral margin of segment
<quantity id="277E9B18AD1B07458596FA9AFDA6586D" box="[501,544,1348,1372]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="12" pageNumber="535" unit="in" value="7.0">7 in</quantity>
males
</emphasis>
(
<figureCitation id="78BD2A78AD1B07458614FA9DFD4F586D" box="[631,713,1347,1372]" captionStart="FIGURES 23 – 26" captionStartId="10.[151,264,1826,1848]" captionTargetBox="[151,1436,193,1804]" captionTargetId="figure@10.[151,1436,193,1805]" captionTargetPageId="10" captionText="FIGURES 23 – 26. Thassoblaniulus radjai sp. n., paratype male. 23. first pair of legs, antero-lateral view; 24. first pair of legs, anterior view; 25. second pair of legs with penis, posterior view; 26. body ring 7, ventral view. – p: penis, cl: claw, ls: lanceolate seta, ta: tarsus, ti: tibia. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/235357/files/figure.png" pageId="12" pageNumber="535">Fig. 26</figureCitation>
) with a stout ridge, but without a horizontal mesad flange.
</paragraph>
<paragraph id="E03936FDAD1B074584A4FAB6FE565B69" blockId="12.[151,1437,1275,2021]" pageId="12" pageNumber="535">
<emphasis id="D2F2EAEFAD1B074584A4FAB6FE1A58B1" box="[199,412,1383,1408]" italics="true" pageId="12" pageNumber="535">Anterior gonopods</emphasis>
(
<figureCitation id="78BD2A78AD1B074585CDFAB9FDB758B1" box="[430,561,1383,1408]" captionStart="FIGURES 13 – 16" captionStartId="7.[151,264,1951,1973]" captionTargetBox="[151,1436,193,1929]" captionTargetId="figure@7.[151,1436,193,1930]" captionTargetPageId="7" captionText="FIGURES 13 – 16. Thassoblaniulus radjai sp. n., paratype male anterior gonopods. 13. anterior view; 14. lateral view; 15. posterior view; 16. distal view. f: postero-mesal flange, t: telopodite, cp: coxal processes, lsh: lateral shoulder. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/235354/files/figure.png" pageId="12" pageNumber="535">Figs 13–16</figureCitation>
,
<figureCitation id="78BD2A78AD1B07458623FAB6FD3358B1" box="[576,693,1383,1408]" captionStart="FIGURES 27 – 32" captionStartId="11.[151,264,1942,1964]" captionTargetBox="[210,1361,193,1917]" captionTargetId="figure@11.[210,1361,193,1917]" captionTargetPageId="11" captionText="FIGURES 27 – 32. Thassoblaniulus radjai sp. n., holotype male. 27. anterior gonopods, posterior view; 28. anterior gonopods, lateral view; 29. penis, posterior view; 30. posterior gonopod, lateral view; 31. first pair of legs, posterior view; 32. second leg. Scale line: 0.4 mm." httpUri="https://zenodo.org/record/235358/files/figure.png" pageId="12" pageNumber="535">27 and 28</figureCitation>
) with very distinctive, separated coxal processes (
<emphasis id="D2F2EAEFAD1B0745809FFAB9FA9158B1" bold="true" box="[1276,1303,1383,1408]" pageId="12" pageNumber="535">cp</emphasis>
) which are widest at base. From base to top, in the middle part, lateral margins first slightly converging and then slightly diverging. In the apicalmost ⅓, lateral margins suddenly narrowing, forming lateral shoulders (
<emphasis id="D2F2EAEFAD1B0745809DFA71FAA658F9" bold="true" box="[1278,1312,1455,1480]" pageId="12" pageNumber="535">lsh</emphasis>
) and very narrow tips when seen in oral or caudal view. In lateral view, the apical part with very characteristic axe-like postero-mesal flanges (
<emphasis id="D2F2EAEFAD1B074585FCFA29FE2E5B21" bold="true" box="[415,424,1527,1552]" pageId="12" pageNumber="535">f</emphasis>
). Telopodites (
<emphasis id="D2F2EAEFAD1B07458637FA29FDDB5B21" bold="true" box="[596,605,1527,1552]" pageId="12" pageNumber="535">t</emphasis>
) short, ca. ¼ - ⅓ the length of coxal processes; without setae; somewhat curved orally; almost of the same width over the entire length and with a rounded tip or with lower mesal side of tip, from the posterior view.
</paragraph>
<paragraph id="E03936FDAD1B074584A4F9BBFC6F5A01" blockId="12.[151,1437,1275,2021]" pageId="12" pageNumber="535">
<emphasis id="D2F2EAEFAD1B074584A4F9BBFE205B4D" box="[199,422,1635,1660]" italics="true" pageId="12" pageNumber="535">Posterior gonopods</emphasis>
(
<figureCitation id="78BD2A78AD1B074585D6F9BDFD8F5B4D" box="[437,521,1635,1660]" captionStart="FIGURES 17 – 20" captionStartId="8.[151,264,1836,1858]" captionTargetBox="[151,1436,193,1814]" captionTargetId="figure@8.[151,1436,193,1815]" captionTargetPageId="8" captionText="FIGURES 17 – 20. Thassoblaniulus radjai sp. n., paratype male posterior gonopod tip. 17. lateral view; 18. anterior view; 19. mesal view; 20. posterior view. – ll: leaf-shaped lamella, so: solenomere. Scale lines: 0.1 for 17, 18 and 19; 0.05 mm for 20." httpUri="https://zenodo.org/record/235355/files/figure.png" pageId="12" pageNumber="535">Figs 17</figureCitation>
–22 and 30) simple and elongated. Lateral view: arched (or S-shaped); first ⅔ of almost constant width, with somewhat narrower second ⅓. Border between second and third ⅓ is the narrower part. The last third is the widest part, in the form of a wide leaf-shaped lamella (
<emphasis id="D2F2EAEFAD1B07458796F975FB825BF5" bold="true" box="[1013,1028,1707,1732]" pageId="12" pageNumber="535">ll</emphasis>
) with numerous minute setules and microspines (Fig. 22). Lamella laterally with a tubular solenomere (
<emphasis id="D2F2EAEFAD1B074587EAF911FC245BD9" bold="true" box="[905,930,1743,1768]" pageId="12" pageNumber="535">so</emphasis>
) (
<figureCitation id="78BD2A78AD1B074587D9F911FB8B5BD9" box="[954,1037,1743,1768]" captionStart="FIGURES 21 and 22" captionStartId="9.[151,264,1293,1315]" captionTargetBox="[151,1436,193,1271]" captionTargetId="figure@9.[151,1436,193,1271]" captionTargetPageId="9" captionText="FIGURES 21 and 22. Thassoblaniulus radjai sp. n., paratype male posterior gonopod. 21. solenomere; 22. detail of lamella. Scale lines: 0.01 mm." httpUri="https://zenodo.org/record/235356/files/figure.png" pageId="12" pageNumber="535">Fig. 21</figureCitation>
).
<emphasis id="D2F2EAEFAD1B07458042F90FFBEE5BD9" box="[1057,1128,1745,1768]" italics="true" pageId="12" pageNumber="535">In situ</emphasis>
the leaf-shaped lamellae fit nicely with the narrow distal parts of anterior gonopods on the both sides. Antero-posterior view: second ⅓ is the widest, gradually tapering to very narrow part, the beginning of the lamella.
</paragraph>
<paragraph id="E03936FDAD1B074584A4F8E5FC955A65" blockId="12.[151,1437,1275,2021]" box="[199,787,1851,1876]" pageId="12" pageNumber="535">
<emphasis id="D2F2EAEFAD1B074584A4F8E5FE765A65" box="[199,496,1851,1876]" italics="true" pageId="12" pageNumber="535">Second pair of female legs</emphasis>
without any peculiarities.
</paragraph>
<paragraph id="E03936FDAD1B074584A4F881FB255AAD" blockId="12.[151,1437,1275,2021]" pageId="12" pageNumber="535">
<emphasis id="D2F2EAEFAD1B074584A4F881FE975A49" box="[199,273,1887,1912]" italics="true" pageId="12" pageNumber="535">Vulvae</emphasis>
(
<figureCitation id="78BD2A78AD1B07458541F881FEF25A49" box="[290,372,1887,1912]" captionStart="FIGURE 33" captionStartId="12.[151,250,1178,1200]" captionTargetBox="[151,1436,193,1156]" captionTargetId="figure@12.[151,1436,193,1157]" captionTargetPageId="12" captionText="FIGURE 33. Thassoblaniulus radjai sp. n., female vulva, lateral view. b: bursa, o: operculum, rs: receptaculum seminis. Scale line: 0.05 mm." httpUri="https://zenodo.org/record/235359/files/figure.png" pageId="12" pageNumber="535">Fig. 33</figureCitation>
) ellipsoid, with somewhat longer operculum (
<emphasis id="D2F2EAEFAD1B0745871DF881FC0A5A49" bold="true" box="[894,908,1887,1912]" pageId="12" pageNumber="535">o</emphasis>
). Both operculum and bursa (
<emphasis id="D2F2EAEFAD1B074580BCF881FB685A49" bold="true" box="[1247,1262,1887,1912]" pageId="12" pageNumber="535">b</emphasis>
) without setae. Receptaculum seminis (
<emphasis id="D2F2EAEFAD1B074585C7F85DFE3D5AAD" bold="true" box="[420,443,1923,1948]" pageId="12" pageNumber="535">rs</emphasis>
) single, ellipsoid. Vulval invaginations reaching back to ring four.
</paragraph>
<paragraph id="E03936FDAD1B074484A4F879FE0B5C2C" blockId="12.[151,1437,1275,2021]" lastBlockId="13.[151,1436,151,320]" lastPageId="13" lastPageNumber="536" pageId="12" pageNumber="535">
<emphasis id="D2F2EAEFAD1B074584A4F879FE7E5AF1" bold="true" box="[199,504,1959,1984]" pageId="12" pageNumber="535">Note on cave adaptations.</emphasis>
Although the new species has so far been found only in a cave, it is not a real cavedweller. Evidence for this claim includes the pigmented, mainly pale brown body (a few specimens are with brighter coloration which may indicate that they were caught shortly after molting), short setae on metazonites, “normal” length of walking legs and antennae as well as presence of ocelli. The number of ocelli is reduced to 2–4 on each side of the head, which may be interpreted as one step towards troglomorphism. Probably
<taxonomicName id="27864D7EAD1A074480BDFF3FFABB5DC9" box="[1246,1341,223,248]" class="Diplopoda" family="Blaniulidae" genus="Thassoblaniulus" kingdom="Animalia" order="Julida" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="radjai" status="sp. nov.">
<emphasis id="D2F2EAEFAD1A074480BDFF3FFABB5DC9" box="[1246,1341,223,248]" italics="true" pageId="13" pageNumber="536">T. radjai</emphasis>
</taxonomicName>
<emphasis id="D2F2EAEFAD1A07448127FF01FA055DC9" bold="true" box="[1348,1411,223,248]" pageId="13" pageNumber="536">
<taxonomicNameLabel id="C9C15794AD1A07448127FF01FA055DC9" box="[1348,1411,223,248]" pageId="13" pageNumber="536" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is a troglophilic species.
</paragraph>
</subSubSection>
<subSubSection id="A89C6576AD1A074484A4FEF9FACA5C71" box="[199,1356,295,320]" pageId="13" pageNumber="536" type="distribution">
<paragraph id="E03936FDAD1A074484A4FEF9FACA5C71" blockId="13.[151,1436,151,320]" box="[199,1356,295,320]" pageId="13" pageNumber="536">
<emphasis id="D2F2EAEFAD1A074484A4FEF9FEE65C71" bold="true" box="[199,352,295,320]" pageId="13" pageNumber="536">Distribution.</emphasis>
Known only from the
<typeStatus id="3F3D885FAD1A07448603FEF6FD165C71" box="[608,656,296,320]" pageId="13" pageNumber="536">type</typeStatus>
locality (cave) near Tirana,
<collectingCountry id="9891766DAD1A074487AFFEF9FBA35C71" box="[972,1061,295,320]" name="Albania" pageId="13" pageNumber="536">Albania</collectingCountry>
(
<figureCitation id="78BD2A78AD1A07448057FEF9FB015C71" box="[1076,1159,295,320]" captionStart="FIGURE 43" captionStartId="16.[151,250,1117,1139]" captionTargetBox="[151,1436,193,1095]" captionTargetId="figure@16.[151,1436,193,1095]" captionTargetPageId="16" captionText="FIGURE 43. Distribution of the genera Cibiniulus Verhoeff, 1897 and Thassoblaniulus Mauriès, 1985." httpUri="https://zenodo.org/record/235362/files/figure.png" pageId="13" pageNumber="536">Fig. 43</figureCitation>
, yellow triangle).
</paragraph>
</subSubSection>
</treatment>
</document>
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