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<document id="847A583E69C8F2E5CDA39DC97F5C261C" ID-CLB-Dataset="29197" ID-DOI="10.11646/zootaxa.4479.1.1" ID-GBIF-Dataset="73cc6f7d-b3ec-40c5-adc7-52b9e0cbf236" ID-ISSN="1175-5326" ID-Zenodo-Dep="1452913" ID-ZooBank="93C84828-6EEF-4758-BEA1-97EEEF115245" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539087676732" checkinUser="plazi" docAuthor="Eiseman, Charles S. &amp; Lonsdale, Owen" docDate="2018" docId="03D287EFFFF9E410A8E551BB41EFFBF1" docLanguage="en" docName="zootaxa.4479.1.1.pdf" docOrigin="Zootaxa 4479 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Phytomyza verbenae Eiseman &amp; Lonsdale 2018, spec. nov." docType="treatment" docVersion="8" lastPageNumber="91" masterDocId="FFEBFF97FFA0E44AA872526F4103FFE1" masterDocTitle="New state and host records for Agromyzidae (Diptera) in the United States, with the description of thirty new species" masterLastPageNumber="156" masterPageNumber="1" pageNumber="90" updateTime="1698570476436" updateUser="ExternalLinkService">
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<mods:title id="D58EAFA9F5C4C227C38235BDDD65162E">New state and host records for Agromyzidae (Diptera) in the United States, with the description of thirty new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="1B4335A3F2C74F170F8AE82DDE1F2773">Eiseman, Charles S.</mods:namePart>
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<mods:namePart id="BC1B7B01B25B7B3B4D689ACC9406897F">Lonsdale, Owen</mods:namePart>
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<mods:title id="5F7055C01F1C66CB82C2648860B98783">Zootaxa</mods:title>
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<mods:part id="47711D57030732ACEB4CCC4066094ABE">
<mods:date id="BB66723FEA8122957FD35338BC3C2841">2018</mods:date>
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<mods:number id="516B13F13FD5DAE2BD281C2FA25B97AE">2018-09-14</mods:number>
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<mods:number id="F2F408994CE585A6C47B68ADBCA01BC2">4479</mods:number>
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<mods:classification id="96F9BABB70BBEBC5FFBB880EF0A5F315">journal article</mods:classification>
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<treatment id="03D287EFFFF9E410A8E551BB41EFFBF1" ID-DOI="http://doi.org/10.5281/zenodo.5997962" ID-GBIF-Taxon="148663604" ID-Zenodo-Dep="5997962" LSID="urn:lsid:plazi:treatment:03D287EFFFF9E410A8E551BB41EFFBF1" httpUri="http://treatment.plazi.org/id/03D287EFFFF9E410A8E551BB41EFFBF1" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90">
<subSubSection id="C3616572FFF9E413A8E551BB40D5FBEE" pageId="89" pageNumber="90" type="nomenclature">
<paragraph id="8BC436F9FFF9E413A8E551BB430DFC0F" blockId="89.[151,526,980,1039]" box="[151,526,980,1006]" pageId="89" pageNumber="90">
<heading id="D08C8195FFF9E413A8E551BB430DFC0F" bold="true" box="[151,526,980,1006]" fontSize="11" level="1" pageId="89" pageNumber="90" reason="1">
<emphasis id="B90FEAEBFFF9E413A8E551BB430DFC0F" bold="true" box="[151,526,980,1006]" pageId="89" pageNumber="90">
<taxonomicName id="4C7B4D7AFFF9E413A8E551BB408CFC0F" ID-CoL="8DCQL" authority="Eiseman &amp; Lonsdale, 2018" authorityName="Eiseman &amp; Lonsdale" authorityYear="2018" box="[151,399,980,1006]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="verbenae" status="sp. nov.">
<emphasis id="B90FEAEBFFF9E413A8E551BB408CFC0F" bold="true" box="[151,399,980,1006]" italics="true" pageId="89" pageNumber="90">Phytomyza verbenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C5790FFF9E413A9E451BB430DFC0F" box="[406,526,980,1006]" pageId="89" pageNumber="90" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC436F9FFF9E413A8E5519940D5FBEE" blockId="89.[151,526,980,1039]" box="[151,470,1014,1039]" pageId="89" pageNumber="90">
(
<figureCitation id="13402A7CFFF9E413A8ED51994024FBEE" box="[159,295,1014,1039]" captionStart="FIGURES 5862" captionStartId="107.[151,264,1660,1683]" captionTargetBox="[151,1437,193,1639]" captionTargetId="figure@107.[151,1437,193,1639]" captionTargetPageId="107" captionText="FIGURES 5862. Figures 5859. Phytomyza verbenae spec. nov.; 58: adult male; 59: empty puparium. Figures 6061: P. ziziae spec. nov.; 60: adult female; 61: puparium. Figure 62: Phytomyza sp. 3, puparium." httpUri="https://zenodo.org/record/1452931/files/figure.png" pageId="89" pageNumber="90">Figs. 5859</figureCitation>
,
<figureCitation id="13402A7CFFF9E413A9405199405CFBEE" box="[306,351,1014,1039]" captionStart="FIGURES 208216" captionStartId="123.[151,264,1660,1683]" captionTargetBox="[151,1437,193,1639]" captionTargetId="figure@123.[151,1437,193,1639]" captionTargetPageId="123" captionText="FIGURES 208216. leaf mines; 208: Phytomyza syngenesiae (Hardy) in Tanacetum parthenium; 209: P. tarnwoodensis spec. nov. in Diervilla lonicera; 210: P. thalictrivora Spencer in Thalictrum pubescens; 211: P. tigris spec. nov. in Tiarella cordifolia; 212: P. triangularidis spec. nov. in Senecio triangularis; 213: same; 214: P. vancouveriella spec. nov. in Ƒancouveria hexandra; 215: P. verbenae spec. nov. in Ƒerbena lasiostachys; 216: P. ziziae spec. nov. in Zizia aurea (dark brown puparium on leaf surface, near margin)." httpUri="https://zenodo.org/record/1452963/files/figure.png" pageId="89" pageNumber="90">215</figureCitation>
,
<figureCitation id="13402A7CFFF9E413A918519840CDFBEE" box="[362,462,1015,1039]" captionStart="FIGURES 389393" captionStartId="154.[151,264,1820,1843]" captionTargetBox="[197,1420,214,1771]" captionTargetId="figure@154.[151,1436,193,1800]" captionTargetPageId="154" captionText="FIGURES 389393. Phytomyza verbenae spec. nov., male genitalia; 389: external components, posterior; 390: postgonite, left lateral; 391: hypandrium, ventral; 392: phallus, ventral; 393: phallus, left lateral." httpUri="https://zenodo.org/record/1453027/files/figure.png" pageId="89" pageNumber="90">389393</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFF9E413A8E5565142F7FB7E" pageId="89" pageNumber="90" type="materials_examined">
<paragraph id="8BC436F9FFF9E413A8E556514374FB9A" blockId="89.[151,1437,1086,2011]" pageId="89" pageNumber="90">
<emphasis id="B90FEAEBFFF9E413A8E5565140CDFBB6" bold="true" box="[151,462,1086,1111]" pageId="89" pageNumber="90">
<typeStatus id="54C0885BFFF9E413A8E556514005FBB6" box="[151,262,1086,1111]" pageId="89" pageNumber="90" type="holotype">Holotype</typeStatus>
.
<collectingRegion id="49BFF81BFFF9E413A961565140C9FBB6" box="[275,458,1086,1111]" country="United States of America" name="California" pageId="89" pageNumber="90">CALIFORNIA</collectingRegion>
.
</emphasis>
Mariposa Co., near El Portal,
<date id="FFC51039FFF9E413AB5B5650429FFBB6" box="[809,924,1087,1111]" pageId="89" pageNumber="90" value="2012-10-30">30.x.2012</date>
, em. by
<date id="FFC51039FFF9E413AB8C56514569FBB6" box="[1022,1130,1086,1111]" pageId="89" pageNumber="90" value="2012-11-05">5.xi.2012</date>
, C.S. Eiseman, ex
<taxonomicName id="4C7B4D7AFFF9E413AD3256514023FB9A" class="Magnoliopsida" family="Verbenaceae" genus="Verbena" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="89" pageNumber="90" phylum="Tracheophyta" rank="species" species="lasiostachys">
<emphasis id="B90FEAEBFFF9E413AD3256514023FB9A" italics="true" pageId="89" pageNumber="90">Verbena lasiostachys</emphasis>
</taxonomicName>
, #CSE118,
<collectionCode id="ED6AAE3CFFF9E413A9D4560D40DCFB9A" box="[422,479,1122,1147]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="89" pageNumber="90">CNC</collectionCode>
751734 (
<specimenCount id="9D7DFD70FFF9E413AA37560C436AFB9A" box="[581,617,1123,1147]" pageId="89" pageNumber="90" type="male">1♂</specimenCount>
).
</paragraph>
<paragraph id="8BC436F9FFF9E413A8E556E942F7FB7E" blockId="89.[151,1437,1086,2011]" box="[151,1012,1158,1183]" pageId="89" pageNumber="90">
<emphasis id="B90FEAEBFFF9E413A8E556E940DBFB7E" bold="true" box="[151,472,1158,1183]" pageId="89" pageNumber="90">
<typeStatus id="54C0885BFFF9E413A8E556E94012FB7E" box="[151,273,1158,1183]" pageId="89" pageNumber="90" type="paratype">Paratypes</typeStatus>
.
<collectingRegion id="49BFF81BFFF9E413A96E56E940D0FB7E" box="[284,467,1158,1183]" country="United States of America" name="California" pageId="89" pageNumber="90">CALIFORNIA</collectingRegion>
.
</emphasis>
same collection as
<typeStatus id="54C0885BFFF9E413AAC156E9421BFB7E" box="[691,792,1158,1183]" pageId="89" pageNumber="90" type="holotype">holotype</typeStatus>
,
<collectionCode id="ED6AAE3CFFF9E413AB5156E9425EFB7E" box="[803,861,1158,1183]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="89" pageNumber="90">CNC</collectionCode>
751735 (
<specimenCount id="9D7DFD70FFF9E413ABB156E842E7FB7E" box="[963,996,1159,1183]" pageId="89" pageNumber="90" type="female">1♀</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFF9E413A8B556C542BFFB23" box="[199,956,1194,1219]" pageId="89" pageNumber="90" type="etymology">
<paragraph id="8BC436F9FFF9E413A8B556C542BFFB23" blockId="89.[151,1437,1086,2011]" box="[199,956,1194,1219]" pageId="89" pageNumber="90">
<emphasis id="B90FEAEBFFF9E413A8B556C5404DFB22" bold="true" box="[199,334,1194,1219]" pageId="89" pageNumber="90">Etymology.</emphasis>
The specific epithet refers to the host plant,
<taxonomicName id="4C7B4D7AFFF9E413AB3256C542BFFB23" authority="L." authorityName="L." box="[832,956,1194,1219]" class="Magnoliopsida" family="Verbenaceae" genus="Verbena" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="89" pageNumber="90" phylum="Tracheophyta" rank="genus">
<emphasis id="B90FEAEBFFF9E413AB3256C5429FFB22" box="[832,924,1194,1219]" italics="true" pageId="89" pageNumber="90">Verbena</emphasis>
L.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFF9E413A8B556A14565FAB2" pageId="89" pageNumber="90" type="biology_ecology">
<paragraph id="8BC436F9FFF9E413A8B556A143DAFB06" blockId="89.[151,1437,1086,2011]" box="[199,729,1230,1255]" pageId="89" pageNumber="90">
<emphasis id="B90FEAEBFFF9E413A8B556A14005FB06" bold="true" box="[199,262,1230,1255]" pageId="89" pageNumber="90">Host.</emphasis>
<taxonomicName id="4C7B4D7AFFF9E413A97F56A1409CFB06" authorityName="" authorityYear="1805" box="[269,415,1230,1255]" family="Verbenaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="89" pageNumber="90" phylum="Magnoliopsida" rank="family">Verbenaceae</taxonomicName>
:
<taxonomicName id="4C7B4D7AFFF9E413A9DB56A143DAFB06" authority="Link." authorityName="Link." box="[425,729,1230,1255]" class="Magnoliopsida" family="Verbenaceae" genus="Verbena" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="89" pageNumber="90" phylum="Tracheophyta" rank="species" species="lasiostachys">
<emphasis id="B90FEAEBFFF9E413A9DB56A14396FB06" box="[425,661,1230,1255]" italics="true" pageId="89" pageNumber="90">Verbena lasiostachys</emphasis>
Link.
</taxonomicName>
</paragraph>
<paragraph id="8BC436F9FFF9E413A8B5569D4565FAB2" blockId="89.[151,1437,1086,2011]" pageId="89" pageNumber="90">
<emphasis id="B90FEAEBFFF9E413A8B5569D4051FAEA" bold="true" box="[199,338,1266,1291]" pageId="89" pageNumber="90">Leaf mine.</emphasis>
(
<figureCitation id="13402A7CFFF9E413A91F569D40DAFAEA" box="[365,473,1266,1291]" captionStart="FIGURES 208216" captionStartId="123.[151,264,1660,1683]" captionTargetBox="[151,1437,193,1639]" captionTargetId="figure@123.[151,1437,193,1639]" captionTargetPageId="123" captionText="FIGURES 208216. leaf mines; 208: Phytomyza syngenesiae (Hardy) in Tanacetum parthenium; 209: P. tarnwoodensis spec. nov. in Diervilla lonicera; 210: P. thalictrivora Spencer in Thalictrum pubescens; 211: P. tigris spec. nov. in Tiarella cordifolia; 212: P. triangularidis spec. nov. in Senecio triangularis; 213: same; 214: P. vancouveriella spec. nov. in Ƒancouveria hexandra; 215: P. verbenae spec. nov. in Ƒerbena lasiostachys; 216: P. ziziae spec. nov. in Zizia aurea (dark brown puparium on leaf surface, near margin)." httpUri="https://zenodo.org/record/1452963/files/figure.png" pageId="89" pageNumber="90">Fig. 215</figureCitation>
) Linear, yellowish-green to whitish, on the upper surface, sometimes partly interparenchymal; typically following up and down most of the length of the midrib, in addition to meandering elsewhere in the leaf. The dark green to black frass is in discrete grains along the sides.
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFF9E413A8B5573140D8FA7A" pageId="89" pageNumber="90" type="description">
<paragraph id="8BC436F9FFF9E413A8B5573140D8FA7A" blockId="89.[151,1437,1086,2011]" pageId="89" pageNumber="90">
<emphasis id="B90FEAEBFFF9E413A8B55731404BFA96" bold="true" box="[199,328,1374,1399]" pageId="89" pageNumber="90">Puparium.</emphasis>
(
<figureCitation id="13402A7CFFF9E413A924573140ABFA96" box="[342,424,1374,1399]" captionStart="FIGURES 5862" captionStartId="107.[151,264,1660,1683]" captionTargetBox="[151,1437,193,1639]" captionTargetId="figure@107.[151,1437,193,1639]" captionTargetPageId="107" captionText="FIGURES 5862. Figures 5859. Phytomyza verbenae spec. nov.; 58: adult male; 59: empty puparium. Figures 6061: P. ziziae spec. nov.; 60: adult female; 61: puparium. Figure 62: Phytomyza sp. 3, puparium." httpUri="https://zenodo.org/record/1452931/files/figure.png" pageId="89" pageNumber="90">Fig. 59</figureCitation>
) White; formed within the leaf, on the lower surface, with the anterior spiracles projecting through the lower epidermis.
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFF9E413A8B557C94331F9AE" pageId="89" pageNumber="90" type="distribution">
<paragraph id="8BC436F9FFF9E413A8B557C940DFFA5E" blockId="89.[151,1437,1086,2011]" box="[199,476,1446,1471]" pageId="89" pageNumber="90">
<emphasis id="B90FEAEBFFF9E413A8B557C94063FA5E" bold="true" box="[199,352,1446,1471]" pageId="89" pageNumber="90">Distribution.</emphasis>
<collectingCountry id="F36C7669FFF9E413A91557C840A0FA5E" box="[359,419,1447,1471]" name="United States of America" pageId="89" pageNumber="90">USA</collectingCountry>
: CA.
</paragraph>
<paragraph id="8BC436F9FFF9E413A8B557A54331F9AE" blockId="89.[151,1437,1086,2011]" pageId="89" pageNumber="90">
<emphasis id="B90FEAEBFFF9E413A8B557A540A1FA02" bold="true" box="[199,418,1482,1507]" pageId="89" pageNumber="90">Adult description.</emphasis>
Wing length
<quantity id="4C839B1CFFF9E413AA3357A44393FA02" box="[577,656,1483,1507]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="89" pageNumber="90" unit="mm" value="2.3">2.3mm</quantity>
(
<specimenCount id="9D7DFD70FFF9E413AAD057A443BAFA02" box="[674,697,1483,1507]" pageId="89" pageNumber="90" type="male"></specimenCount>
),
<quantity id="4C839B1CFFF9E413AAA357A44223FA02" box="[721,800,1483,1507]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="89" pageNumber="90" unit="mm" value="2.7">2.7mm</quantity>
(
<specimenCount id="9D7DFD70FFF9E413AB4657A44244FA02" box="[820,839,1483,1507]" pageId="89" pageNumber="90" type="female"></specimenCount>
). Eye height divided by gena height: 3.03.1. Male first flagellomere rounded, small, slightly higher than long; covered with long hairs approximately as long as width of arista base, except at base; female first flagellomere enlarged, slightly more than 1/3 eye height. Cheek distinct. Notum pruinose. Vein dm-cu absent.
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFF9E410A8B554354008FD64" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90" type="description">
<paragraph id="8BC436F9FFF9E413A8B5543543EDF8E2" blockId="89.[151,1437,1086,2011]" pageId="89" pageNumber="90">
<emphasis id="B90FEAEBFFF9E413A8B554354044F992" box="[199,327,1626,1651]" italics="true" pageId="89" pageNumber="90">Chaetotaxy</emphasis>
: Two ors, two ori; slightly decreasing in length anteriorly, male with anterior ori thin, half length of posterior ori. Eye sparsely haired. Postvertical seta slightly shorter and thinner than ors, ocellar seta slightly smaller. Five strong dorsocentral setae, decreasing in length anteriorly, with fourth seta approximately 1/52/5 length first seta, and fifth dorsocentral nearly setula-like. Acrostichal setulae in four rows, reduced to two or three behind suture, ending at level of second dorsocentral.
</paragraph>
<paragraph id="8BC436F9FFF9E410A8B5556143B2FF19" blockId="89.[151,1437,1086,2011]" lastBlockId="90.[151,1437,151,1040]" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90">
<emphasis id="B90FEAEBFFF9E413A8B555614041F8C6" box="[199,322,1806,1831]" italics="true" pageId="89" pageNumber="90">Coloration</emphasis>
: (
<figureCitation id="13402A7CFFF9E413A929556140ADF8C6" box="[347,430,1806,1831]" captionStart="FIGURES 5862" captionStartId="107.[151,264,1660,1683]" captionTargetBox="[151,1437,193,1639]" captionTargetId="figure@107.[151,1437,193,1639]" captionTargetPageId="107" captionText="FIGURES 5862. Figures 5859. Phytomyza verbenae spec. nov.; 58: adult male; 59: empty puparium. Figures 6061: P. ziziae spec. nov.; 60: adult female; 61: puparium. Figure 62: Phytomyza sp. 3, puparium." httpUri="https://zenodo.org/record/1452931/files/figure.png" pageId="89" pageNumber="90">Fig. 58</figureCitation>
) Setae dark brown. Head mostly light yellow; antenna, lunule and adjacent orbital plate and parafacial, posterior margin of vertex, posterolateral corner of frons to base of inner vertical seta, small rounded subtriangular spot around tubercle, dorsal half of face, clypeus, palpus, ventral line on gena and back of head dark brown with blackish tint. Body with light pruinosity that is denser on thorax, becoming grayish and almost matt on notum. Thorax dark brown to blackish with postpronotum (excluding large spot) and notopleuron light yellow, and dorsal margin of anepisternum light yellow with stripe widening to level of seta posteriorly. Calypter yellow to white. Legs dark brown to blackish, apex of fore femur and base of fore tibia narrowly light yellow, tarsi slightly paler. Abdomen dark brown to blackish, lateral margin of tergites with light yellow stripe that is widest on tergite 2 and narrowest on tergite 6.
</paragraph>
<paragraph id="8BC436F9FFFAE410A8B5536B4008FD64" blockId="90.[151,1437,151,1040]" pageId="90" pageNumber="91">
<emphasis id="B90FEAEBFFFAE410A8B5536B4032FEFC" box="[199,305,260,285]" italics="true" pageId="90" pageNumber="91">Genitalia</emphasis>
: (
<figureCitation id="13402A7CFFFAE410A938536B40F3FEFD" box="[330,496,260,285]" captionStart="FIGURES 389393" captionStartId="154.[151,264,1820,1843]" captionTargetBox="[197,1420,214,1771]" captionTargetId="figure@154.[151,1436,193,1800]" captionTargetPageId="154" captionText="FIGURES 389393. Phytomyza verbenae spec. nov., male genitalia; 389: external components, posterior; 390: postgonite, left lateral; 391: hypandrium, ventral; 392: phallus, ventral; 393: phallus, left lateral." httpUri="https://zenodo.org/record/1453027/files/figure.png" pageId="90" pageNumber="91">Figs. 389393</figureCitation>
) Surstylus small, rounded, fused to epandrium, with small setae on inner surface. Posteromedial margin of epandrium with cluster of 910 tubercle-like setae in front of small thin pointed plate. Cercus long, tapering apically. Hypandrium relatively long and narrow, subtriangular with apex widely rounded; inner lobe small with two short setae. Postgonite simple with thicker apex and two empty dorsoapical sockets. Phallus relatively large and pale, apex mostly membranous with sclerites ill-defined. Phallophorus small, lateral bands not found. Basiphallus composed of two outwardly bowed sclerites with apices nearly separated into separate sclerites about as long as high; right sclerite fused to phallophorus. Hypophallus lightly sclerotized with thickened medial fold confluent with somewhat paddle-shaped lateral sclerited region. Paraphallus mostly membranous, weakly and irregularly sclerotized basally, apically fused to broad membranous hood covering distiphallus. Mesophallus not evident. Distiphallus very small, linear, strongly curved at base. Ejaculatory apodeme not found.
</paragraph>
</subSubSection>
<subSubSection id="C3616572FFFAE410A8B550E041EFFBF1" pageId="90" pageNumber="91" type="discussion">
<paragraph id="8BC436F9FFFAE410A8B550E04501FCD9" blockId="90.[151,1437,151,1040]" pageId="90" pageNumber="91">
<emphasis id="B90FEAEBFFFAE410A8B550E0404CFD49" bold="true" box="[199,335,655,680]" pageId="90" pageNumber="91">Comments.</emphasis>
Extreme reduction and weakening of the distal sclerites of the phallus, including the narrow, unified meso/distiphallus, would suggest placement of
<taxonomicName id="4C7B4D7AFFFAE410AB6150DB42FCFD2C" authorityName="Eiseman &amp; Lonsdale" authorityYear="2018" box="[787,1023,692,717]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="verbenae">
<emphasis id="B90FEAEBFFFAE410AB6150DB42FCFD2C" box="[787,1023,692,717]" italics="true" pageId="90" pageNumber="91">Phytomyza verbenae</emphasis>
</taxonomicName>
in the
<taxonomicName id="4C7B4D7AFFFAE410AC2750DA45F8FD2D" baseAuthorityName="Hardy" baseAuthorityYear="1849" box="[1109,1275,693,716]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="syngenesiae">
<emphasis id="B90FEAEBFFFAE410AC2750DA45F8FD2D" box="[1109,1275,693,716]" italics="true" pageId="90" pageNumber="91">P. syngenesiae</emphasis>
</taxonomicName>
group. While these reductive characters are prone to homoplasy, the new species also shares with this group a mostly dark body covered with a gray pruinosity, a contrastingly pale head (excluding the black antenna), somewhat reduced acrostichal setulae and an enlarged female first flagellomere (as in
<taxonomicName id="4C7B4D7AFFFAE410ABF6514E42F2FCD9" authorityName="Frost" authorityYear="1924" box="[900,1009,799,824]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="lactuca">
<emphasis id="B90FEAEBFFFAE410ABF6514E42F2FCD9" box="[900,1009,799,824]" italics="true" pageId="90" pageNumber="91">P. lactuca</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BC436F9FFFAE410A8B5512B41EFFBF1" blockId="90.[151,1437,151,1040]" pageId="90" pageNumber="91">
Aside from these potentially superficial similarities,
<taxonomicName id="4C7B4D7AFFFAE410AB51512B450DFCBC" authorityName="Eiseman &amp; Lonsdale" authorityYear="2018" box="[803,1038,836,861]" class="Insecta" family="Agromyzidae" genus="Phytomyza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="verbenae">
<emphasis id="B90FEAEBFFFAE410AB51512B450DFCBC" box="[803,1038,836,861]" italics="true" pageId="90" pageNumber="91">Phytomyza verbenae</emphasis>
</taxonomicName>
is otherwise distinct. The calypter hairs are white (not brown), the notopleuron and postpronotum are light yellow, the basiphallus is long, pale, bowed medially and broken subapically, the epandrium has tubercle-like setae (a very uncommon and highly diagnostic character in the genus group), the hypophallus has a pale oar-shaped lateral sclerite and a weak, folded medial sclerite, and the mesophallus+distiphallus is very small, C-shaped and produced past the duct insertion basally.
</paragraph>
</subSubSection>
</treatment>
</document>