290 lines
21 KiB
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290 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.116.27301" ID-GBIF-Dataset="15acdccd-830c-40dd-84e3-b793f92827a4" ID-PMC="PMC6367307" ID-Pensoft-Pub="1314-2003-116-1" ID-Pensoft-UUID="182FFF91FFCDFF9CFF811552FFCCFFCF" ID-PubMed="30740023" ID-Zenodo-Dep="2559703" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2003-116-1" ModsDocOrigin="PhytoKeys 116" ModsDocTitle="Taxonomic revision of Chenopodiaceae in Himalaya and Tibet" checkinTime="1555330571688" checkinUser="pensoft" docAuthor="Sukhorukov, Alexander P., Liu, Pei-Liang & Kushunina, Maria" docDate="2019" docId="8FCE091D5A0E48E7419F6501556BAAB5" docLanguage="en" docName="PhytoKeys 116: 1-141" docOrigin="PhytoKeys 116" docSource="http://dx.doi.org/10.3897/phytokeys.116.27301" docTitle="Salsola hartmannii Sukhorukov & Liu & Kushunina 2019, sp. nov." docType="treatment" docVersion="6" id="182FFF91FFCDFF9CFF811552FFCCFFCF" lastPageNumber="109" masterDocId="182FFF91FFCDFF9CFF811552FFCCFFCF" masterDocTitle="Taxonomic revision of Chenopodiaceae in Himalaya and Tibet" masterLastPageNumber="141" masterPageNumber="1" pageNumber="108" updateTime="1668139168965" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of Chenopodiaceae in Himalaya and Tibet</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sukhorukov, Alexander P.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liu, Pei-Liang</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kushunina, Maria</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>116</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>141</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.116.27301</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.116.27301</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-116-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">182FFF91FFCDFF9CFF811552FFCCFFCF</mods:identifier>
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<mods:identifier type="Zenodo-Dep">2559703</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="156201177" LSID="urn:lsid:plazi:treatment:8FCE091D5A0E48E7419F6501556BAAB5" httpUri="http://treatment.plazi.org/id/8FCE091D5A0E48E7419F6501556BAAB5" lastPageId="108" lastPageNumber="109" pageId="107" pageNumber="108">
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<subSubSection pageId="107" pageNumber="108" type="nomenclature">
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<paragraph pageId="107" pageNumber="108">
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1.
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<taxonomicName LSID="8FCE091D-5A0E-48E7-419F-6501556BAAB5" authority="Sukhor." authorityName="Sukhorukov & Liu & Kushunina" authorityYear="2019" class="Magnoliopsida" family="Chenopodiacea" genus="Salsola" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salsola hartmannii" order="Caryophyllales" pageId="107" pageNumber="108" phylum="Tracheophyta" rank="species" species="hartmannii" status="sp. nov.">Salsola hartmannii Sukhor.</taxonomicName>
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<taxonomicNameLabel pageId="107" pageNumber="108">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Holotype of Salsola hartmannii Sukhor. sp. nov. (G)." figureDoi="10.3897/phytokeys.114.27301.figure45" httpUri="https://binary.pensoft.net/fig/260786" pageId="107" pageNumber="108">Fig. 45</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="107" pageNumber="108" type="holotype">
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<paragraph pageId="107" pageNumber="108">Holotype.</paragraph>
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<paragraph pageId="107" pageNumber="108">
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[INDIA, Jammu & Kashmir] E-Ladakh, Alpine Steppe mit
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<taxonomicName class="Dicotyledoneae" family="Asteraceae" genus="Artemisia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Artemisia minor" order="Campanulales" pageId="107" pageNumber="108" phylum="Angiospermae" rank="species" species="minor">
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<emphasis italics="true" pageId="107" pageNumber="108">Artemisia minor</emphasis>
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</taxonomicName>
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, ca. 1 km W oberhalb Kiagar Tso [alpine steppe with
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<taxonomicName class="Dicotyledoneae" family="Asteraceae" genus="Artemisia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Artemisia minor" order="Campanulales" pageId="107" pageNumber="108" phylum="Angiospermae" rank="species" species="minor">
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<emphasis italics="true" pageId="107" pageNumber="108">Artemisia minor</emphasis>
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</taxonomicName>
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, ca. 1 km W above Kiagar Tso], 4720 m a.s.l., 27 Aug 1997,
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<emphasis italics="true" pageId="107" pageNumber="108">H. Hartmann 6080</emphasis>
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(G00410003! isotype - MSB160560!).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="107" pageNumber="108" type="taxonomic notes">
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<paragraph pageId="107" pageNumber="108">Taxonomic notes.</paragraph>
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<paragraph pageId="107" pageNumber="108">
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The new species was formerly identified as
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. jacquemontii" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="jacquemontii">
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<emphasis italics="true" pageId="107" pageNumber="108">S. jacquemontii</emphasis>
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</taxonomicName>
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or
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. collina" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="collina">
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<emphasis italics="true" pageId="107" pageNumber="108">S. collina</emphasis>
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</taxonomicName>
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. In many characters (e.g. tiny anthers, small outgrowths of the perianth segments),
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<taxonomicName class="Magnoliopsida" family="Chenopodiaceae" genus="Salsola" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salsola hartmannii" order="Caryophyllales" pageId="107" pageNumber="108" phylum="Tracheophyta" rank="species" species="hartmannii">
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<emphasis italics="true" pageId="107" pageNumber="108">Salsola hartmannii</emphasis>
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</taxonomicName>
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resembles
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. monoptera" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="monoptera">
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<emphasis italics="true" pageId="107" pageNumber="108">S. monoptera</emphasis>
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</taxonomicName>
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, but the latter species has only one well-expressed, wing-like projection on a segment closest to the bract, whereas projections on other segments are much smaller (in the form of tubercles). Additionally, the gibbous bracts and bracteoles are exceptionally rare in
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. monoptera" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="monoptera">
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<emphasis italics="true" pageId="107" pageNumber="108">S. monoptera</emphasis>
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</taxonomicName>
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. In contrast,
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. hartmannii" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="hartmannii">
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<emphasis italics="true" pageId="107" pageNumber="108">S. hartmannii</emphasis>
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</taxonomicName>
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possesses gibbous bracts and bracteoles and two or three perianth segments develop tiny, but clearly visible wings. The new species is also similar to
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. jacquemontii" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="jacquemontii">
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<emphasis italics="true" pageId="107" pageNumber="108">S. jacquemontii</emphasis>
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</taxonomicName>
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but has smaller anthers and a smaller diameter of the entire diaspore (winged perianth enclosing the fruit), which is hidden within the subtending bract and bracteoles.
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</paragraph>
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<paragraph pageId="107" pageNumber="108">
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The gibbous bract and bracteoles cannot be considered as strong characteristic features of
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. hartmannii" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="hartmannii">
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<emphasis italics="true" pageId="107" pageNumber="108">S. hartmannii</emphasis>
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</taxonomicName>
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; some specimens of
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. jacquemontii" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="jacquemontii">
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<emphasis italics="true" pageId="107" pageNumber="108">S. jacquemontii</emphasis>
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</taxonomicName>
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may possess such a shape of the bracts and bracteoles. All three species,
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. monoptera" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="monoptera">
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<emphasis italics="true" pageId="107" pageNumber="108">S. monoptera</emphasis>
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</taxonomicName>
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,
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. hartmannii" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="hartmannii">
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<emphasis italics="true" pageId="107" pageNumber="108">S. hartmannii</emphasis>
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</taxonomicName>
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and
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. jacquemontii" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="jacquemontii">
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<emphasis italics="true" pageId="107" pageNumber="108">S. jacquemontii</emphasis>
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</taxonomicName>
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, are morphologically related.
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<taxonomicName authorityName="Pall., Ill. Pl.: 34" authorityYear="1803" class="Magnoliopsida" family="Chenopodiaceae" genus="Salsola" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salsola collina" order="Caryophyllales" pageId="107" pageNumber="108" phylum="Tracheophyta" rank="species" species="collina">
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<emphasis italics="true" pageId="107" pageNumber="108">Salsola collina</emphasis>
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</taxonomicName>
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is morphologically distant from these species, having appressed bracts without clearly expressed wings on the perianth at fruiting. The distribution patterns are also different:
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. monoptera" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="monoptera">
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<emphasis italics="true" pageId="107" pageNumber="108">S. monoptera</emphasis>
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</taxonomicName>
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is widespread in Central Asia and Tibet,
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. hartmannii" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="hartmannii">
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<emphasis italics="true" pageId="107" pageNumber="108">S. hartmannii</emphasis>
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</taxonomicName>
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seems to be localised in North Himalaya and Karakoram and
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. jacquemontii" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="jacquemontii">
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<emphasis italics="true" pageId="107" pageNumber="108">S. jacquemontii</emphasis>
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</taxonomicName>
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is found in Central Himalaya and Tibet. The presence of
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. jacquemontii" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="jacquemontii">
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<emphasis italics="true" pageId="107" pageNumber="108">S. jacquemontii</emphasis>
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</taxonomicName>
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and
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. collina" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="collina">
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<emphasis italics="true" pageId="107" pageNumber="108">S. collina</emphasis>
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</taxonomicName>
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in North Himalaya (
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<bibRefCitation author="Rilke, S" journalOrPublisher="Bibliotheca Botanica" pageId="129" pageNumber="130" pagination="1 - 190" refId="B86" refString="Rilke, S, 1999. Revision der Sektion Salsola s.l. der Gattung Salsola (Chenopodiaceae). Bibliotheca Botanica 149: 1 - 190" title="Revision der Sektion Salsola s. l. der Gattung Salsola (Chenopodiaceae)." volume="149" year="1999">Rilke 1999</bibRefCitation>
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) is not confirmed.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="107" pageNumber="108" type="description">
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<paragraph pageId="107" pageNumber="108">Description.</paragraph>
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<paragraph pageId="107" pageNumber="108">
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Annual with prostrate or ascending stems up to 15 cm long, glabrous or scarcely papillate. Leaves terete, 10-20
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<normalizedToken originalValue="×">x</normalizedToken>
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1.0-1.5 mm. Flowers in axillary clusters and arranged in the main inflorescence. Flower clusters (located below the main inflorescence) abundant and consist of two flowers covered with bracts and bracteoles that are connate almost to the top and basally gibbous; perianth segments membranous, fimbriate, r-shaped, their tips more or less appressed to each other, not hardened and not forming a tight conus, with white, small tubercles 0.3-0.5 mm long in the flexure; styles with stigmas ca. 0.5 mm; fruits 1.2-1.6 mm, depressed roundish. Flowers in the main inflorescence with (almost) free, basally gibbous bract and bracteoles, hermaphrodite; perianth segments 5, membranous, fimbriate, r-shaped at the fruiting stage, <2.5 mm long, above the flexure soft, hyaline, horizontally orientated and not forming a distinct conus; 2 or 3 (of 5) segments develop wing-like appendages (up to 1.5 mm long) that are located almost horizontally or obliquely, two other segments develop tubercles only; the whole diaspore with the perianth wings is 2.0-3.0 mm across and is hidden within the bract and bracteoles; anthers 0.3-0.5 mm long; styles with stigmas ca. 1 mm; fruits lenticular, 1.1-1.5 mm. Seeds with horizontal or obliquely orientated embryo.
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</paragraph>
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<caption ID-Zenodo-Dep="2559821" doi="10.3897/phytokeys.114.27301.figure45" httpUri="https://binary.pensoft.net/fig/260786" pageId="107" pageNumber="108" start="Figure 45" startId="F45">
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<paragraph pageId="107" pageNumber="108">
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<emphasis bold="true" pageId="107" pageNumber="108">Figure 45.</emphasis>
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Holotype of
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<taxonomicName class="Magnoliopsida" family="Chenopodiaceae" genus="Salsola" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salsola hartmannii" order="Caryophyllales" pageId="107" pageNumber="108" phylum="Tracheophyta" rank="species" species="hartmannii">
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<emphasis italics="true" pageId="107" pageNumber="108">Salsola hartmannii</emphasis>
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</taxonomicName>
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Sukhor. sp. nov. (G).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="107" pageNumber="108" type="habitat">
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<paragraph pageId="107" pageNumber="108">Habitat.</paragraph>
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<paragraph pageId="107" pageNumber="108">Gravelly or sandy substrates at altitudes of 3500-5200 m. The species seems to be common in Ladakh and Spiti Valley.</paragraph>
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</subSubSection>
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<subSubSection pageId="107" pageNumber="108" type="phenology">
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<paragraph pageId="107" pageNumber="108">Phenology.</paragraph>
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<paragraph pageId="107" pageNumber="108">Flowering: August-September; fruiting: September-October.</paragraph>
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<paragraph pageId="107" pageNumber="108">
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<emphasis bold="true" pageId="107" pageNumber="108">Etymology</emphasis>
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: The specific epithet is after Hans Hartmann, a Swiss geobotanist who provided significant contributions to the flora and vegetation of North India.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="107" pageNumber="108" type="distribution">
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<paragraph pageId="107" pageNumber="108">Distribution.</paragraph>
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<paragraph pageId="107" pageNumber="108">
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See Fig.
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<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Distribution map of Salsola hartmannii (circles), S. collina (stars) and S. jacquemontii (triangles)." figureDoi="10.3897/phytokeys.114.27301.figure46" httpUri="https://binary.pensoft.net/fig/260787" pageId="107" pageNumber="108">46</figureCitation>
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.
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</paragraph>
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<caption ID-Zenodo-Dep="2559823" doi="10.3897/phytokeys.114.27301.figure46" httpUri="https://binary.pensoft.net/fig/260787" pageId="107" pageNumber="108" start="Figure 46" startId="F46">
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<paragraph pageId="107" pageNumber="108">
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<emphasis bold="true" pageId="107" pageNumber="108">Figure 46.</emphasis>
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Distribution map of
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<taxonomicName class="Magnoliopsida" family="Chenopodiaceae" genus="Salsola" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salsola hartmannii" order="Caryophyllales" pageId="107" pageNumber="108" phylum="Tracheophyta" rank="species" species="hartmannii">
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<emphasis italics="true" pageId="107" pageNumber="108">Salsola hartmannii</emphasis>
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</taxonomicName>
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(circles),
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. collina" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="collina">
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<emphasis italics="true" pageId="107" pageNumber="108">S. collina</emphasis>
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</taxonomicName>
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(stars) and
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<taxonomicName family="Chenopodiacea" kingdom="Plantae" lsidName="S. jacquemontii" order="Caryophyllales" pageId="107" pageNumber="108" rank="species" species="jacquemontii">
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<emphasis italics="true" pageId="107" pageNumber="108">S. jacquemontii</emphasis>
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</taxonomicName>
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(triangles).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="108" lastPageNumber="109" pageId="107" pageNumber="108" type="materials_examined">
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<paragraph pageId="107" pageNumber="108">Specimens examined.</paragraph>
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<paragraph pageId="107" pageNumber="108">
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<emphasis bold="true" pageId="107" pageNumber="108">CHINA</emphasis>
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:
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<emphasis bold="true" pageId="107" pageNumber="108">Xizang</emphasis>
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:
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<emphasis bold="true" pageId="107" pageNumber="108">Ngari Prefecture</emphasis>
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: [Zanda County] Tisum, 15000 ft a.s.l., [without date]
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<emphasis italics="true" pageId="107" pageNumber="108">R. Strachey & J.E. Winterbottom</emphasis>
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s.n. (BR);
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</paragraph>
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<paragraph lastPageId="108" lastPageNumber="109" pageId="107" pageNumber="108">
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<emphasis bold="true" pageId="107" pageNumber="108">INDIA: Jammu & Kashmir</emphasis>
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: Naini valley to Srinagar, [without date and collector] (BM); [Ladakh] Parang valley, 11 Sep 1847,
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<emphasis italics="true" pageId="107" pageNumber="108">T.T. Thomson</emphasis>
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s.n. (K); Ladakh, Muklip to Lukung, ca. 4000 m a.s.l., 29 Aug 1976,
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<emphasis italics="true" pageId="107" pageNumber="108">B.M. Wadhwa 60152</emphasis>
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(BSD); [Ladakh] Debring to Tanglang La, 4800-5200 m a.s.l., 3 Sep 1989,
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<emphasis italics="true" pageId="107" pageNumber="108">
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L.
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<normalizedToken originalValue="Klimeš">Klimes</normalizedToken>
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& M.
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<normalizedToken originalValue="Šrůtek">Srutek</normalizedToken>
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28/37
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</emphasis>
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(PRA); Rupshu, N of Tso Kar, ca. 4600 m a.s.l., 22 Aug 1995,
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<emphasis italics="true" pageId="107" pageNumber="108">H. Hartmann 5036</emphasis>
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(G455293, MSB160561); Ladakh, Zanskar Region, Zara, N of Kurio, 33°35'30"N, 77°28'E, 4250 m a.s.l., 23 Aug 1998, code 98-11-2,
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<emphasis italics="true" pageId="107" pageNumber="108">
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L.
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<normalizedToken originalValue="Klimeš">Klimes</normalizedToken>
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161
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</emphasis>
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(PRA); Ladakh, Indus valley, Stot (E) [Stod River valley], between Angkung & Puga,
|
||
<geoCoordinate degrees="33" direction="north" minutes="14" orientation="latitude" precision="925" value="33.233334">33°14'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="78" direction="east" minutes="16" orientation="longitude" precision="925" value="78.26667">78°16'E</geoCoordinate>
|
||
, 4550 m a.s.l., 8 Sep 1999,
|
||
<emphasis italics="true" pageId="107" pageNumber="108">
|
||
L.
|
||
<normalizedToken originalValue="Klimeš">Klimes</normalizedToken>
|
||
776
|
||
</emphasis>
|
||
(PRA); Ladakh, Rupshu Region, Tso Moriri, Lema to Peldo,
|
||
<geoCoordinate degrees="33" direction="north" minutes="1" orientation="latitude" precision="15" seconds="53" value="33.031387">33°1'53"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="78" direction="east" minutes="16" orientation="longitude" precision="15" seconds="34" value="78.276115">78°16'34"E</geoCoordinate>
|
||
, 4550-4560 m a.s.l., 13 Sep 2005,
|
||
<emphasis italics="true" pageId="107" pageNumber="108">
|
||
L.
|
||
<normalizedToken originalValue="Klimeš">Klimes</normalizedToken>
|
||
6301
|
||
</emphasis>
|
||
(PRA);
|
||
<emphasis bold="true" pageId="107" pageNumber="108">Himachal Pradesh</emphasis>
|
||
: Parang La, 11 Sep 1847,
|
||
<emphasis italics="true" pageId="107" pageNumber="108">anonym</emphasis>
|
||
s.n. (K); Spiti valley, Samdo to Kaurik, 21 Aug 1994,
|
||
<emphasis italics="true" pageId="108" pageNumber="109">
|
||
<pageBreakToken pageId="108" pageNumber="109" start="start">S</pageBreakToken>
|
||
.K. Murti & S. Singh 81588
|
||
</emphasis>
|
||
(BSD); Pin Valley National Park, Mane vill., 3700 m a.s.l., Sep 2002,
|
||
<emphasis italics="true" pageId="108" pageNumber="109">K.C. Sekar 100736</emphasis>
|
||
(BSD).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="108" pageNumber="109" type="distribution">
|
||
<paragraph pageId="108" pageNumber="109">General distribution.</paragraph>
|
||
<paragraph pageId="108" pageNumber="109">
|
||
North India, NW China (Xizang) and North Pakistan ([Gilgit-Baltistan] Nagar, dry stony slope, 9000 ft a.s.l., 22 Aug 1987,
|
||
<emphasis italics="true" pageId="108" pageNumber="109">Jamshed 239</emphasis>
|
||
(E!)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |