treatments-xml/data/3B/5E/07/3B5E0726356AFFFBFF0730BF3734FE72.xml

<document id="BE1664164B11A2F533B2F3D47CE217F2" ID-DOI="10.11646/phytotaxa.329.3.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="13723030" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725666487150" checkinUser="felipe" docAuthor="Glushchenko, Anton, Kulikovskiy, Maxim &amp; Kociolek, John Patrick" docDate="2017" docId="3B5E0726356AFFFBFF0730BF3734FE72" docLanguage="en" docName="phytotaxa.329.3.3.pdf" docOrigin="Phytotaxa 329 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.329.3.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Gomphonema dalatica Glushchenko, Kulikovskiy &amp; Kociolek 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="229" masterDocId="C7677F5E356EFFFDFF8F3022364AFFDD" masterDocTitle="New diatom species from the Gomphonema subtile group in Southeast Asia" masterLastPageNumber="232" masterPageNumber="223" pageNumber="227" updateTime="1725677996708" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0CA39B196A4299012F1DE5E3C05BEBAD">New diatom species from the Gomphonema subtile group in Southeast Asia</mods:title>
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<mods:affiliation id="BB928D8E70EF6B7C6DA85FD15B9609AA">Kaluga State University, 248023, Kaluga District, Stepan Rasin street 26, Russia &amp; Institute of Plant Physiology, Russian Academy of Sciences, Botanical Street 35, 127276, Moscow, Russia</mods:affiliation>
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<mods:affiliation id="FB0B774E4344FE0BBC80A0EB784C1133">Department of Water Plants Taxonomy and Geography, I. D. Papanin Institute for Biology of Inland Waters, Russian Academy of Sciences, 152742 Yaroslavl, Nekouz, Borok, Russia &amp; Institute of Plant Physiology, Russian Academy of Sciences, Botanical Street 35, 127276, Moscow, Russia</mods:affiliation>
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<mods:namePart id="761DFF42B5CA74A5C95D12056E4AC534">Kociolek, John Patrick</mods:namePart>
<mods:affiliation id="13CF3C4DA685BAF16D73475F8022978D">Museum of Natural History and Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, 80309, USA.</mods:affiliation>
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<treatment id="3B5E0726356AFFFBFF0730BF3734FE72" ID-DOI="http://doi.org/10.5281/zenodo.13722974" ID-Zenodo-Dep="13722974" LSID="urn:lsid:plazi:treatment:3B5E0726356AFFFBFF0730BF3734FE72" httpUri="http://treatment.plazi.org/id/3B5E0726356AFFFBFF0730BF3734FE72" lastPageId="6" lastPageNumber="229" pageId="4" pageNumber="227">
<subSubSection id="FBEDE5BB356AFFF9FF0730BF321DFF65" box="[136,1111,157,184]" pageId="4" pageNumber="227" type="nomenclature">
<paragraph id="B348B630356AFFF9FF0730BF321DFF65" blockId="4.[136,1111,157,184]" box="[136,1111,157,184]" pageId="4" pageNumber="227">
<heading id="E800015C356AFFF9FF0730BF321DFF65" bold="true" box="[136,1111,157,184]" fontSize="11" level="1" pageId="4" pageNumber="227" reason="1">
<emphasis id="81836A22356AFFF9FF0730BF321DFF65" bold="true" box="[136,1111,157,184]" pageId="4" pageNumber="227">
<taxonomicName id="74F7CDB3356AFFF9FF0730BF352AFF65" authority="Glushchenko, Kulikovskiy &amp; Kociolek" authorityName="Glushchenko, Kulikovskiy &amp; Kociolek" authorityYear="2017" box="[136,864,157,184]" class="Bacillariophyceae" family="Gomphonemaceae" genus="Gomphonema" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="4" pageNumber="227" phylum="Ochrophyta" rank="species" species="dalatica" status="sp. nov.">
<emphasis id="81836A22356AFFF9FF0730BF37C4FF65" bold="true" box="[136,398,157,184]" italics="true" pageId="4" pageNumber="227">Gomphonema dalatica</emphasis>
Glushchenko, Kulikovskiy &amp; Kociolek
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<taxonomicNameLabel id="9AB0D759356AFFF9FCE830BC35F4FF65" box="[871,958,158,184]" pageId="4" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="2BCCAAB5356AFFF9FC4230BD3205FF65" box="[973,1103,159,184]" captionStart-0="FIGURES 34–55" captionStart-1="FIGURES 56–63" captionStartId-0="4.[136,243,2030,2051]" captionStartId-1="5.[136,243,1786,1807]" captionTargetBox-0="[221,1366,733,2007]" captionTargetBox-1="[195,1391,175,1762]" captionTargetId-0="figure-247@4.[221,1366,733,2007]" captionTargetId-1="figure-14@5.[195,1391,175,1762]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 34–55: Gomphonema dalatica Glushchenko, Kulikovskiy &amp; Kociolek sp. nov. LM. Size diminution series, valve views. Figure 39 is of the holotype. Scale bar = 10 μm." captionText-1="FIGURES 56–63: Gomphonema dalatica Glushchenko, Kulikovskiy &amp; Kociolek sp. nov. SEM. Figs 56, 58, 60, 62 are external views; 57, 59, 61, 63 are internal views. Fig. 56. Whole valve illustrating valve shape. Fig. 57. Whole valve, showing small pseudoseptum at each pole. Fig. 58. Valve center with slightly dilated proximal raphe ends, small, round stigma opening and areolae occluded by flaps that create C-shaped openings. Fig. 59. Valve center with small central nodule bearing recurved proximal raphe ends and slit-like stigmal opening. Striae are in troughs, with thickened rims around each areola. Fig. 60. Headpole, with the hooked-shaped distal raphe end curving onto the valve mantle. C-shaped areolar openings are evident. Fig. 61. Headpole, showing prominent helictoglossa and pseudoseptum. Fig. 62. Footpole, with round porelli positioned mostly on the valve mantle, physically separated, and morphologically distinct from the areolae. Fig. 63. Footpole, with large helictoglossa and pseudoseptum evident. Porelli are occluded internally. Small siliceous projections can be seen around the areolae. Scale bars = 5 μm (Figs 56, 57); 1 μm (Figs 58–60, 62); 0.5 μm (Figs 61, 63)." figureDoi-0="http://doi.org/10.5281/zenodo.13723036" figureDoi-1="http://doi.org/10.5281/zenodo.13723038" httpUri-0="https://zenodo.org/record/13723036/files/figure.png" httpUri-1="https://zenodo.org/record/13723038/files/figure.png" pageId="4" pageNumber="227">Figs 34–63</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FBEDE5BB356AFFF9FF0730C535A9FDA2" pageId="4" pageNumber="227" type="description">
<paragraph id="B348B630356AFFF9FF0730C53750FEDD" blockId="4.[136,1452,231,708]" box="[136,282,231,256]" pageId="4" pageNumber="227">
<emphasis id="81836A22356AFFF9FF0730C53750FEDD" bold="true" box="[136,282,231,256]" pageId="4" pageNumber="227">Description.</emphasis>
</paragraph>
<paragraph id="B348B630356AFFF9FF32312937C2FE72" blockId="4.[136,1452,231,708]" pageId="4" pageNumber="227">
<emphasis id="81836A22356AFFF9FF32312936BBFEF9" bold="true" box="[189,241,267,292]" pageId="4" pageNumber="227">LM.</emphasis>
Valves lanceolate-clavate with headpole protracted, capitate and footpole rounded. Length 40–64 μm, breadth 6.5–8.0 μm. Axial area narrowly lanceolate, forming a small, rectangular central area possessing a single, enlarged stigma positioned at the end of a central stria. Raphe more or less lateral, straight, with external proximal ends dilated slightly and distal ends deflected onto the valve mantle. Striae parallel to radiate, 
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10 μm, slightly coarser at the center of the valve.
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<paragraph id="B348B630356AFFF9FF32319B35A9FDA2" blockId="4.[136,1452,231,708]" pageId="4" pageNumber="227">
<emphasis id="81836A22356AFFF9FF32319B374BFE0F" bold="true" box="[189,257,441,466]" pageId="4" pageNumber="227">SEM.</emphasis>
The valve exterior has areolae occluded by flaps, giving the openings the appearance of being c-shaped (
<figureCitation id="2BCCAAB5356AFFF9FF1F31F93728FE29" box="[144,354,475,500]" captionStart="FIGURES 56–63" captionStartId="5.[136,243,1786,1807]" captionTargetBox="[195,1391,175,1762]" captionTargetId="figure-14@5.[195,1391,175,1762]" captionTargetPageId="5" captionText="FIGURES 56–63: Gomphonema dalatica Glushchenko, Kulikovskiy &amp; Kociolek sp. nov. SEM. Figs 56, 58, 60, 62 are external views; 57, 59, 61, 63 are internal views. Fig. 56. Whole valve illustrating valve shape. Fig. 57. Whole valve, showing small pseudoseptum at each pole. Fig. 58. Valve center with slightly dilated proximal raphe ends, small, round stigma opening and areolae occluded by flaps that create C-shaped openings. Fig. 59. Valve center with small central nodule bearing recurved proximal raphe ends and slit-like stigmal opening. Striae are in troughs, with thickened rims around each areola. Fig. 60. Headpole, with the hooked-shaped distal raphe end curving onto the valve mantle. C-shaped areolar openings are evident. Fig. 61. Headpole, showing prominent helictoglossa and pseudoseptum. Fig. 62. Footpole, with round porelli positioned mostly on the valve mantle, physically separated, and morphologically distinct from the areolae. Fig. 63. Footpole, with large helictoglossa and pseudoseptum evident. Porelli are occluded internally. Small siliceous projections can be seen around the areolae. Scale bars = 5 μm (Figs 56, 57); 1 μm (Figs 58–60, 62); 0.5 μm (Figs 61, 63)." figureDoi="http://doi.org/10.5281/zenodo.13723038" httpUri="https://zenodo.org/record/13723038/files/figure.png" pageId="4" pageNumber="227">Figs 56, 58, 60, 62</figureCitation>
). The center of the valve has dilated proximal raphe ends and enlarged, round stigmal opening (
<figureCitation id="2BCCAAB5356AFFF9FAF131F936E2FDCA" captionStart="FIGURES 56–63" captionStartId="5.[136,243,1786,1807]" captionTargetBox="[195,1391,175,1762]" captionTargetId="figure-14@5.[195,1391,175,1762]" captionTargetPageId="5" captionText="FIGURES 56–63: Gomphonema dalatica Glushchenko, Kulikovskiy &amp; Kociolek sp. nov. SEM. Figs 56, 58, 60, 62 are external views; 57, 59, 61, 63 are internal views. Fig. 56. Whole valve illustrating valve shape. Fig. 57. Whole valve, showing small pseudoseptum at each pole. Fig. 58. Valve center with slightly dilated proximal raphe ends, small, round stigma opening and areolae occluded by flaps that create C-shaped openings. Fig. 59. Valve center with small central nodule bearing recurved proximal raphe ends and slit-like stigmal opening. Striae are in troughs, with thickened rims around each areola. Fig. 60. Headpole, with the hooked-shaped distal raphe end curving onto the valve mantle. C-shaped areolar openings are evident. Fig. 61. Headpole, showing prominent helictoglossa and pseudoseptum. Fig. 62. Footpole, with round porelli positioned mostly on the valve mantle, physically separated, and morphologically distinct from the areolae. Fig. 63. Footpole, with large helictoglossa and pseudoseptum evident. Porelli are occluded internally. Small siliceous projections can be seen around the areolae. Scale bars = 5 μm (Figs 56, 57); 1 μm (Figs 58–60, 62); 0.5 μm (Figs 61, 63)." figureDoi="http://doi.org/10.5281/zenodo.13723038" httpUri="https://zenodo.org/record/13723038/files/figure.png" pageId="4" pageNumber="227">Fig. 58</figureCitation>
). At the headpole, the distal raphe end is deflected onto the mantle (
<figureCitation id="2BCCAAB5356AFFF9FC2631DC35B4FDCA" box="[937,1022,510,535]" captionStart="FIGURES 56–63" captionStartId="5.[136,243,1786,1807]" captionTargetBox="[195,1391,175,1762]" captionTargetId="figure-14@5.[195,1391,175,1762]" captionTargetPageId="5" captionText="FIGURES 56–63: Gomphonema dalatica Glushchenko, Kulikovskiy &amp; Kociolek sp. nov. SEM. Figs 56, 58, 60, 62 are external views; 57, 59, 61, 63 are internal views. Fig. 56. Whole valve illustrating valve shape. Fig. 57. Whole valve, showing small pseudoseptum at each pole. Fig. 58. Valve center with slightly dilated proximal raphe ends, small, round stigma opening and areolae occluded by flaps that create C-shaped openings. Fig. 59. Valve center with small central nodule bearing recurved proximal raphe ends and slit-like stigmal opening. Striae are in troughs, with thickened rims around each areola. Fig. 60. Headpole, with the hooked-shaped distal raphe end curving onto the valve mantle. C-shaped areolar openings are evident. Fig. 61. Headpole, showing prominent helictoglossa and pseudoseptum. Fig. 62. Footpole, with round porelli positioned mostly on the valve mantle, physically separated, and morphologically distinct from the areolae. Fig. 63. Footpole, with large helictoglossa and pseudoseptum evident. Porelli are occluded internally. Small siliceous projections can be seen around the areolae. Scale bars = 5 μm (Figs 56, 57); 1 μm (Figs 58–60, 62); 0.5 μm (Figs 61, 63)." figureDoi="http://doi.org/10.5281/zenodo.13723038" httpUri="https://zenodo.org/record/13723038/files/figure.png" pageId="4" pageNumber="227">Fig. 60</figureCitation>
). At the footpole, the apical pore field (APF) is positioned mostly on the valve mantle (
<figureCitation id="2BCCAAB5356AFFF9FD2A320334B3FDE7" box="[677,761,545,570]" captionStart="FIGURES 56–63" captionStartId="5.[136,243,1786,1807]" captionTargetBox="[195,1391,175,1762]" captionTargetId="figure-14@5.[195,1391,175,1762]" captionTargetPageId="5" captionText="FIGURES 56–63: Gomphonema dalatica Glushchenko, Kulikovskiy &amp; Kociolek sp. nov. SEM. Figs 56, 58, 60, 62 are external views; 57, 59, 61, 63 are internal views. Fig. 56. Whole valve illustrating valve shape. Fig. 57. Whole valve, showing small pseudoseptum at each pole. Fig. 58. Valve center with slightly dilated proximal raphe ends, small, round stigma opening and areolae occluded by flaps that create C-shaped openings. Fig. 59. Valve center with small central nodule bearing recurved proximal raphe ends and slit-like stigmal opening. Striae are in troughs, with thickened rims around each areola. Fig. 60. Headpole, with the hooked-shaped distal raphe end curving onto the valve mantle. C-shaped areolar openings are evident. Fig. 61. Headpole, showing prominent helictoglossa and pseudoseptum. Fig. 62. Footpole, with round porelli positioned mostly on the valve mantle, physically separated, and morphologically distinct from the areolae. Fig. 63. Footpole, with large helictoglossa and pseudoseptum evident. Porelli are occluded internally. Small siliceous projections can be seen around the areolae. Scale bars = 5 μm (Figs 56, 57); 1 μm (Figs 58–60, 62); 0.5 μm (Figs 61, 63)." figureDoi="http://doi.org/10.5281/zenodo.13723038" httpUri="https://zenodo.org/record/13723038/files/figure.png" pageId="4" pageNumber="227">Fig. 62</figureCitation>
). Internally, there is a small central nodule, recurved proximal raphe ends, and a slit-like stigma opening (
<figureCitation id="2BCCAAB5356AFFF9FDE9326134A2FD81" box="[614,744,579,604]" captionStart="FIGURES 56–63" captionStartId="5.[136,243,1786,1807]" captionTargetBox="[195,1391,175,1762]" captionTargetId="figure-14@5.[195,1391,175,1762]" captionTargetPageId="5" captionText="FIGURES 56–63: Gomphonema dalatica Glushchenko, Kulikovskiy &amp; Kociolek sp. nov. SEM. Figs 56, 58, 60, 62 are external views; 57, 59, 61, 63 are internal views. Fig. 56. Whole valve illustrating valve shape. Fig. 57. Whole valve, showing small pseudoseptum at each pole. Fig. 58. Valve center with slightly dilated proximal raphe ends, small, round stigma opening and areolae occluded by flaps that create C-shaped openings. Fig. 59. Valve center with small central nodule bearing recurved proximal raphe ends and slit-like stigmal opening. Striae are in troughs, with thickened rims around each areola. Fig. 60. Headpole, with the hooked-shaped distal raphe end curving onto the valve mantle. C-shaped areolar openings are evident. Fig. 61. Headpole, showing prominent helictoglossa and pseudoseptum. Fig. 62. Footpole, with round porelli positioned mostly on the valve mantle, physically separated, and morphologically distinct from the areolae. Fig. 63. Footpole, with large helictoglossa and pseudoseptum evident. Porelli are occluded internally. Small siliceous projections can be seen around the areolae. Scale bars = 5 μm (Figs 56, 57); 1 μm (Figs 58–60, 62); 0.5 μm (Figs 61, 63)." figureDoi="http://doi.org/10.5281/zenodo.13723038" httpUri="https://zenodo.org/record/13723038/files/figure.png" pageId="4" pageNumber="227">Figs 57, 59</figureCitation>
). Helictoglossae are present, and small pseudosepta are evident at both apices (
<figureCitation id="2BCCAAB5356AFFF9FEBE324437F9FDA2" box="[305,435,614,639]" captionStart="FIGURES 56–63" captionStartId="5.[136,243,1786,1807]" captionTargetBox="[195,1391,175,1762]" captionTargetId="figure-14@5.[195,1391,175,1762]" captionTargetPageId="5" captionText="FIGURES 56–63: Gomphonema dalatica Glushchenko, Kulikovskiy &amp; Kociolek sp. nov. SEM. Figs 56, 58, 60, 62 are external views; 57, 59, 61, 63 are internal views. Fig. 56. Whole valve illustrating valve shape. Fig. 57. Whole valve, showing small pseudoseptum at each pole. Fig. 58. Valve center with slightly dilated proximal raphe ends, small, round stigma opening and areolae occluded by flaps that create C-shaped openings. Fig. 59. Valve center with small central nodule bearing recurved proximal raphe ends and slit-like stigmal opening. Striae are in troughs, with thickened rims around each areola. Fig. 60. Headpole, with the hooked-shaped distal raphe end curving onto the valve mantle. C-shaped areolar openings are evident. Fig. 61. Headpole, showing prominent helictoglossa and pseudoseptum. Fig. 62. Footpole, with round porelli positioned mostly on the valve mantle, physically separated, and morphologically distinct from the areolae. Fig. 63. Footpole, with large helictoglossa and pseudoseptum evident. Porelli are occluded internally. Small siliceous projections can be seen around the areolae. Scale bars = 5 μm (Figs 56, 57); 1 μm (Figs 58–60, 62); 0.5 μm (Figs 61, 63)." figureDoi="http://doi.org/10.5281/zenodo.13723038" httpUri="https://zenodo.org/record/13723038/files/figure.png" pageId="4" pageNumber="227">Figs 61, 63</figureCitation>
). At the footpole, porelli of the APF are occluded.
</paragraph>
</subSubSection>
<subSubSection id="FBEDE5BB356AFFF9FF3232AB32CBFD19" pageId="4" pageNumber="227" type="diagnosis">
<paragraph id="B348B630356AFFF9FF3232AB32CBFD19" blockId="4.[136,1452,231,708]" pageId="4" pageNumber="227">
<emphasis id="81836A22356AFFF9FF3232AB378DFD7F" bold="true" box="[189,455,649,674]" pageId="4" pageNumber="227">Differential diagnosis.</emphasis>
Differ from morphologically similar species 
<taxonomicName id="74F7CDB3356AFFF9FC5B32AB32B5FD7F" authority="Reichardt" authorityName="Reichardt. In" authorityYear="2015" box="[980,1279,649,674]" class="Bacillariophyceae" family="Gomphonemaceae" genus="Gomphonema" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="4" pageNumber="227" phylum="Ochrophyta" rank="species" species="pantropicum">
<emphasis id="81836A22356AFFF9FC5B32AB32C2FD7F" box="[980,1160,649,674]" italics="true" pageId="4" pageNumber="227">G. pantropicum</emphasis>
Reichardt
</taxonomicName>
by much finer stria density (
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10 μm, not 
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10 μm) and less radiate orientation of the striae.
</paragraph>
</subSubSection>
<caption id="E788E6B8356AFFF9FF0737CC341CF7FB" ID-DOI="http://doi.org/10.5281/zenodo.13723036" ID-Zenodo-Dep="13723036" httpUri="https://zenodo.org/record/13723036/files/figure.png" pageId="4" pageNumber="227" startId="4.[136,243,2030,2051]" targetBox="[221,1366,733,2007]" targetPageId="4" targetType="figure">
<paragraph id="B348B630356AFFF9FF0737CC341CF7FB" blockId="4.[136,1452,2030,2086]" pageId="4" pageNumber="227">
<emphasis id="81836A22356AFFF9FF0737CC3775F7DE" bold="true" box="[136,319,2030,2051]" pageId="4" pageNumber="227">FIGURES 34–55:</emphasis>
<taxonomicName id="74F7CDB3356AFFF9FEC637CC35F9F7DE" authority="Glushchenko, Kulikovskiy &amp; Kociolek" authorityName="Glushchenko, Kulikovskiy &amp; Kociolek" authorityYear="2017" box="[329,947,2030,2051]" class="Bacillariophyceae" family="Gomphonemaceae" genus="Gomphonema" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="4" pageNumber="227" phylum="Ochrophyta" rank="species" species="dalatica" status="sp. nov.">
<emphasis id="81836A22356AFFF9FEC637CC3462F7DE" box="[329,552,2030,2051]" italics="true" pageId="4" pageNumber="227">Gomphonema dalatica</emphasis>
Glushchenko, Kulikovskiy &amp; Kociolek
</taxonomicName>
<taxonomicNameLabel id="9AB0D759356AFFF9FC3337CC3240F7DE" box="[956,1034,2030,2051]" pageId="4" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
LM. Size diminution series, valve views. Figure 39 is of the holotype. Scale bar = 10 μm.
</paragraph>
</caption>
<caption id="E788E6B8356BFFF8FF0736D83229F7F4" ID-DOI="http://doi.org/10.5281/zenodo.13723038" ID-Zenodo-Dep="13723038" httpUri="https://zenodo.org/record/13723038/files/figure.png" pageId="5" pageNumber="228" startId="5.[136,243,1786,1807]" targetBox="[195,1391,175,1762]" targetPageId="5" targetType="figure">
<paragraph id="B348B630356BFFF8FF0736D83229F7F4" blockId="5.[136,1452,1786,2089]" pageId="5" pageNumber="228">
<emphasis id="81836A22356BFFF8FF0736D83774F8D2" bold="true" box="[136,318,1786,1807]" pageId="5" pageNumber="228">FIGURES 56–63:</emphasis>
<taxonomicName id="74F7CDB3356BFFF8FECB36D835EBF8D2" authority="Glushchenko, Kulikovskiy &amp; Kociolek" authorityName="Glushchenko, Kulikovskiy &amp; Kociolek" authorityYear="2017" box="[324,929,1786,1807]" class="Bacillariophyceae" family="Gomphonemaceae" genus="Gomphonema" higherTaxonomySource="GBIF" kingdom="Chromista" order="Naviculales" pageId="5" pageNumber="228" phylum="Ochrophyta" rank="species" species="dalatica" status="sp. nov.">
<emphasis id="81836A22356BFFF8FECB36D8346BF8D2" box="[324,545,1786,1807]" italics="true" pageId="5" pageNumber="228">Gomphonema dalatica</emphasis>
Glushchenko, Kulikovskiy &amp; Kociolek
</taxonomicName>
<taxonomicNameLabel id="9AB0D759356BFFF8FC2736D835BEF8D2" box="[936,1012,1786,1807]" pageId="5" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
SEM. Figs 56, 58, 60, 62 are external views; 57, 59, 61, 63 are internal views. Fig. 56. Whole valve illustrating valve shape. Fig. 57. Whole valve, showing small pseudoseptum at each pole. Fig. 58. Valve center with slightly dilated proximal raphe ends, small, round stigma opening and areolae occluded by flaps that create C-shaped openings. Fig. 59. Valve center with small central nodule bearing recurved proximal raphe ends and slit-like stigmal opening. Striae are in troughs, with thickened rims around each areola. Fig. 60. Headpole, with the hooked-shaped distal raphe end curving onto the valve mantle. C-shaped areolar openings are evident. Fig. 61. Headpole, showing prominent helictoglossa and pseudoseptum. Fig. 62. Footpole, with round porelli positioned mostly on the valve mantle, physically separated, and morphologically distinct from the areolae. Fig. 63. Footpole, with large helictoglossa and pseudoseptum evident. Porelli are occluded internally. Small siliceous projections can be seen around the areolae. Scale bars = 5 μm (Figs 56, 57); 1 μm (Figs 58 
<emphasis id="81836A22356BFFF8FCB53836350CF7F4" bold="true" box="[826,838,2068,2089]" pageId="5" pageNumber="228">–</emphasis>
60, 62); 0.5 μm (Figs 61, 63).
</paragraph>
</caption>
<subSubSection id="FBEDE5BB3568FFFBFF07308232AEFF22" pageId="6" pageNumber="229" type="materials_examined">
<paragraph id="B348B6303568FFFBFF07308232AEFF22" blockId="6.[136,1452,160,431]" pageId="6" pageNumber="229">
<emphasis id="81836A223568FFFBFF0730823686FF64" bold="true" box="[136,204,160,185]" pageId="6" pageNumber="229">Type:</emphasis>
— 
<collectingCountry id="CBE0F6A03568FFFBFF6630823727FF64" box="[233,365,160,185]" name="Vietnam" pageId="6" pageNumber="229">VIETNAM</collectingCountry>
. Lâm Đồng Province, Ðà Lạt Town, Đả Tiên Reservoir, 
<geoCoordinate id="D6C3D0F73568FFFBFC6130823236FF64" box="[1006,1148,160,185]" degrees="11" direction="north" minutes="58.816" orientation="latitude" pageId="6" pageNumber="229" precision="1" value="11.980267">N11°58.816’</geoCoordinate>
<geoCoordinate id="D6C3D0F73568FFFBFB0A30823369FF64" box="[1157,1315,160,185]" degrees="108" direction="east" minutes="26.987" orientation="longitude" pageId="6" pageNumber="229" precision="1" value="108.44978">E108°26.987’</geoCoordinate>
, 
<emphasis id="81836A223568FFFBFABF3082346BFF01" italics="true" pageId="6" pageNumber="229">
E.S. Gusev &amp; M.S. Kulikovskiy, 21 
<superScript id="44821B783568FFFBFE0030E337D6FF12" attach="left" box="[399,412,193,207]" fontSize="6" pageId="6" pageNumber="229">th</superScript>
<date id="C74990F03568FFFBFE2A30E13457FF01" box="[421,541,195,220]" pageId="6" pageNumber="229" value="2012-06">June 2012</date>
,
</emphasis>
(collection of Maxim Kulikovskiy, I.D. Papanin Institute for Biology of Inland Waters, Russian Academy of Sciences (
<collectionCode id="D5E62EF53568FFFBFDCC30C434C2FF22" box="[579,648,230,255]" country="Russia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15311" lsid="urn:lsid:biocol.org:col:15311" name="I. D. Papanin Institute for Biology of Inland Waters, Russian Academy of Sciences" pageId="6" pageNumber="229" type="Herbarium">IBIW</collectionCode>
), 
<typeStatus id="6C4C08923568FFFBFD1430C434B7FF22" box="[667,765,230,255]" pageId="6" pageNumber="229" type="holotype">holotype</typeStatus>
here designated, slide no 00269 = 
<figureCitation id="2BCCAAB53568FFFBFB0B30C4329DFF22" box="[1156,1239,230,255]" captionStart="FIGURES 34–55" captionStartId="4.[136,243,2030,2051]" captionTargetBox="[221,1366,733,2007]" captionTargetId="figure-247@4.[221,1366,733,2007]" captionTargetPageId="4" captionText="FIGURES 34–55: Gomphonema dalatica Glushchenko, Kulikovskiy &amp; Kociolek sp. nov. LM. Size diminution series, valve views. Figure 39 is of the holotype. Scale bar = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.13723036" httpUri="https://zenodo.org/record/13723036/files/figure.png" pageId="6" pageNumber="229">Fig. 39</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FBEDE5BB3568FFFBFF32312B3490FEFF" box="[189,730,265,290]" pageId="6" pageNumber="229" type="etymology">
<paragraph id="B348B6303568FFFBFF32312B3490FEFF" blockId="6.[136,1452,160,431]" box="[189,730,265,290]" pageId="6" pageNumber="229">
<emphasis id="81836A223568FFFBFF32312B372EFEFF" bold="true" box="[189,356,265,290]" pageId="6" pageNumber="229">Etymology:—</emphasis>
Epithet refers to the 
<typeStatus id="6C4C08923568FFFBFDC7312B3432FEFF" box="[584,632,265,290]" pageId="6" pageNumber="229">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="FBEDE5BB3568FFFBFF32310E3734FE72" pageId="6" pageNumber="229" type="distribution">
<paragraph id="B348B6303568FFFBFF32310E3734FE72" blockId="6.[136,1452,160,431]" pageId="6" pageNumber="229">
<emphasis id="81836A223568FFFBFF32310E371DFE98" bold="true" box="[189,343,300,325]" pageId="6" pageNumber="229">Distribution.</emphasis>
This species is known from the 
<typeStatus id="6C4C08923568FFFBFD4C310E34B9FE98" box="[707,755,300,325]" pageId="6" pageNumber="229">type</typeStatus>
locality and from Da Tanla Waterfall, waterbodies of Mỹ Sõn (Province Ninh Thuận), Ðà Lạt reservoir in 
<collectingCountry id="CBE0F6A03568FFFBFD0E317234AAFEB4" box="[641,736,336,361]" name="Vietnam" pageId="6" pageNumber="229">Vietnam</collectingCountry>
(as “ 
<taxonomicName id="74F7CDB33568FFFBFC91317235B5FEB4" box="[798,1023,336,361]" class="Bacillariophyceae" family="Gomphonemaceae" genus="Gomphonema" kingdom="Chromista" order="Naviculales" pageId="6" pageNumber="229" phylum="Ochrophyta" rank="species" species="undetermined">
<emphasis id="81836A223568FFFBFC91317235F1FEB4" box="[798,955,336,361]" italics="true" pageId="6" pageNumber="229">Gomphonema</emphasis>
spec.
</taxonomicName>
” Figs 117–120, 
<bibRefCitation id="D766CBC13568FFFBFB3331723339FEB4" author="Reichardt, E." box="[1212,1395,336,361]" pageId="6" pageNumber="229" pagination="27 - 38" refId="ref5093" refString="Reichardt, E. (2015) Taxonomy and distribution of Gomphonema subtile Ehrenberg (Bacillariophyceae) and six related taxa. Fottea 15 (1): 27 - 38. https: // doi. org / 10.5507 / fot. 2015.003" type="journal article" year="2015">Reichardt, 2015</bibRefCitation>
) and from Nam Leuk River in 
<collectingCountry id="CBE0F6A03568FFFBFE21315137AEFE51" box="[430,484,371,396]" name="Laos" pageId="6" pageNumber="229">Laos</collectingCountry>
(this investigation). Specimens of this species were found in samples 00269, 00984, 02269, 02406, 03817.
</paragraph>
</subSubSection>
</treatment>
</document>