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<document id="B84FA3C397756FE54F2124EF9F8AACC4" ID-CLB-Dataset="100954" ID-DOI="10.5281/zenodo.6887260" ID-GBIF-Dataset="a016af63-6437-427b-80b7-22bc9a002e20" ID-ISBN="978-84-16728-04-6" ID-Zenodo-Dep="6887260" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="guilherme" checkinTime="1656696812616" checkinUser="carolina" docAuthor="Don E. Wilson, Russell A. Mittermeier &amp; Thomas E. Lacher, Jr" docDate="2017" docId="1E30E2753532FE82E44424AC76F68630" docLanguage="en" docName="hbmw_7_Muridae_0536.pdf.imf" docOrigin="Handbook of the Mammals of the World – Volume 7 Rodents II, Barcelona: Lynx Edicions" docTitle="Limnomys bryophilus Rickart, Heaney &amp; Tabaranza 2003" docType="treatment" docVersion="11" lastPageNumber="882" masterDocId="E2099A0D3426FF97E1372C0977498313" masterDocTitle="Muridae" masterLastPageNumber="884" masterPageNumber="536" pageNumber="881" updateTime="1718983125921" updateUser="carolina">
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<mods:titleInfo id="3F978174874BACDE75A725DBDE1A93F3">
<mods:title id="1078BE8719023DF4B9736E3E9345959A">Muridae</mods:title>
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<mods:namePart id="4DE3935EA296E35667156EFBD33E5FF1">Don E. Wilson</mods:namePart>
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<mods:namePart id="C9243176A5F1E5BA0A3E9A8FE585E5D7">Russell A. Mittermeier</mods:namePart>
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<mods:namePart id="8A56991DF965DE221979BD05FAE79735">Thomas E. Lacher, Jr</mods:namePart>
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<mods:dateIssued id="5462C8C6A5B86FD0DDC6DCACBD2CF92D">2017</mods:dateIssued>
<mods:dateOther id="8120607616F94A24AEAB55641A30C1F0" type="pubDate">2017-11-30</mods:dateOther>
<mods:publisher id="BE5B593F43C96605F0D052105CF5F0B9">Lynx Edicions</mods:publisher>
<mods:place id="7661BE86AA382BB816B9CD187A2027B6">
<mods:placeTerm id="A7B2CECBF5277822AE705E840C34B98E">Barcelona</mods:placeTerm>
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<mods:titleInfo id="465B933FD775ADCA3E7DE4B5BD37CE1C">
<mods:title id="8BC99F1FCF259C14FD851E0F155A4963">Handbook of the Mammals of the World – Volume 7 Rodents II</mods:title>
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<mods:classification id="45D76A07B212A405BD304780BE51FE64">book chapter</mods:classification>
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<treatment id="1E30E2753532FE82E44424AC76F68630" ID-DOI="http://doi.org/10.5281/zenodo.6834787" ID-GBIF-Taxon="197826527" ID-Zenodo-Dep="6834787" LSID="urn:lsid:plazi:treatment:1E30E2753532FE82E44424AC76F68630" httpUri="http://treatment.plazi.org/id/1E30E2753532FE82E44424AC76F68630" lastPageId="277" lastPageNumber="882" pageId="276" pageNumber="881">
<subSubSection id="DE8300E83532FE83E44424AC728F8BC0" box="[1395,1478,2213,2259]" pageId="276" pageNumber="881" type="multiple">
<paragraph id="962653633532FE83E44424AC728F8BC0" blockId="276.[1394,2465,2213,2337]" box="[1395,1478,2213,2259]" pageId="276" pageNumber="881">
<heading id="CD6EE40F3532FE83E44424AC728F8BC0" box="[1395,1478,2213,2259]" pageId="276" pageNumber="881">
<figureCitation id="0EA24FE63532FE83E44424AC728F8BC0" box="[1395,1478,2213,2259]" captionStart="Plate 58: Muridae" captionStartId="270.[111,141,3217,3238]" captionTargetBox="[1382,1974,2548,2959]" captionTargetPageId="270" captionText="784. Salokko Rat (Taeromys arcuatus), 786. Celebes Rat (Taeromys celebensis), 788. Small-eared Rat (Taeromys microbullatus), 789. Sulawesi Forest Rat (Taeromys punicans), 790. Tondano Rat (Taeromys taerae), 791. Long-tailed Sulawesian Shrew Rat (Tateomys macrocercus), 792. Tate’s Sulawesian Shrew Rat (Tateomys rhinogradoides), 793. Mamasa Water Rat (Waiomys mamasae), 795. Komodo Rat (Komodomys rintjanus), 796. Seram Island Mountain Rat (Nesoromys ceramicus), 797. Armandbville’s Giant Tree Rat (Papagomys armanduillei), 798. Paula’s Long-nosed Rat (Paulamys naso), 799. [MISSING], 801. Mindanao Forest Rat (Bullimus bagobus), 803. Luzon Forest Rat (Bullimus luzonicus), 804. Sulawesi Shrew Mouse (Crunomys celebensis), 805. Luzon Shrew Mouse (Crunomys fallax), 806. Mindanao Shrew Mouse (Crunomys melanius), 807. Katanglad Shrew Mouse (Crunomys suncoides), 808. Gray-bellied Moss Mouse (Limnomys bryophilus), 809. Long-tailed Moss Mouse (Limnomys sibuanus), 810. Dusky Long-footed Rat (Tarsomys apoensis), 812. Luzon Short-nosed Rat (Tryphomys adustus), 814. Hume’s Hadromys (Hadromys humei), 816. Vernay’s Climbing Mouse (Vernaya fulva)" figureDoi="http://doi.org/10.5281/zenodo.6888661" httpUri="https://zenodo.org/record/6888661/files/figure.png" pageId="276" pageNumber="881">808.</figureCitation>
</heading>
</paragraph>
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<subSubSection id="DE8300E83532FE83E4E024AC70948BC0" box="[1495,2013,2213,2259]" pageId="276" pageNumber="881" type="vernacular_names">
<paragraph id="962653633532FE83E4E024AC70948BC0" blockId="276.[1394,2465,2213,2337]" box="[1495,2013,2213,2259]" pageId="276" pageNumber="881">
<heading id="CD6EE40F3532FE83E4E024AC70948BC0" box="[1495,2013,2213,2259]" pageId="276" pageNumber="881">
<vernacularName id="189A234D3532FE83E4E024AC70948BC0" ID-CoL="6QDJK" authorityName="Rickart, Heaney &amp; Tabaranza" authorityYear="2003" box="[1495,2013,2213,2259]" class="Mammalia" family="Muridae" genus="Limnomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="species" species="bryophilus">Gray-bellied Moss Mouse</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E83532FE83E91124AC7EE88BC0" box="[2086,2465,2213,2259]" pageId="276" pageNumber="881" type="nomenclature">
<paragraph id="962653633532FE83E91124AC7EE88BC0" blockId="276.[1394,2465,2213,2337]" box="[2086,2465,2213,2259]" pageId="276" pageNumber="881">
<heading id="CD6EE40F3532FE83E91124AC7EE88BC0" box="[2086,2465,2213,2259]" pageId="276" pageNumber="881">
<taxonomicName id="519928E03532FE83E91124AC7EE88BC0" ID-CoL="6QDJK" authorityName="Rickart, Heaney &amp; Tabaranza" authorityYear="2003" box="[2086,2465,2213,2259]" class="Mammalia" family="Muridae" genus="Limnomys" kingdom="Animalia" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="species" species="bryophilus">
<emphasis id="A4ED8F713532FE83E91124AC7EE88BC0" box="[2086,2465,2213,2259]" italics="true" pageId="276" pageNumber="881">Limnomys bryophilus</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E83532FE83E44524EC701F8A32" pageId="276" pageNumber="881" type="vernacular_names">
<paragraph id="962653633532FE83E44524EC7E068BE9" blockId="276.[1394,2465,2213,2337]" box="[1394,2383,2277,2298]" pageId="276" pageNumber="881">
<heading id="CD6EE40F3532FE83E44524EC7E068BE9" box="[1394,2383,2277,2298]" pageId="276" pageNumber="881">
<emphasis id="A4ED8F713532FE83E44524EC72F78BE9" bold="true" box="[1394,1470,2277,2298]" pageId="276" pageNumber="881">French:</emphasis>
<vernacularName id="189A234D3532FE83E4FF24EC71008BE9" ID-CoL="6QDJK" authorityName="Rickart, Heaney &amp; Tabaranza" authorityYear="2003" box="[1480,1609,2277,2298]" class="Mammalia" family="Muridae" genus="Limnomys" kingdom="Animalia" language="fra" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="species" species="bryophilus">Rat bryophile</vernacularName>
/
<emphasis id="A4ED8F713532FE83E76824EC71F08BE9" bold="true" box="[1631,1721,2277,2298]" pageId="276" pageNumber="881">German:</emphasis>
<vernacularName id="189A234D3532FE83E7F524EC70D38BE9" ID-CoL="6QDJK" authorityName="Rickart, Heaney &amp; Tabaranza" authorityYear="2003" box="[1730,1946,2277,2298]" class="Mammalia" family="Muridae" genus="Limnomys" kingdom="Animalia" language="deu" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="species" species="bryophilus">Graubauch-Moosratte</vernacularName>
/
<emphasis id="A4ED8F713532FE83E69824EC7F438BE9" bold="true" box="[1967,2058,2277,2298]" pageId="276" pageNumber="881">Spanish:</emphasis>
<vernacularName id="189A234D3532FE83E92324EC7E068BE9" ID-CoL="6QDJK" authorityName="Rickart, Heaney &amp; Tabaranza" authorityYear="2003" box="[2068,2383,2277,2298]" class="Mammalia" family="Muridae" genus="Limnomys" kingdom="Animalia" language="esp" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="species" species="bryophilus">Ratéon de musgo de vientre gris</vernacularName>
</heading>
</paragraph>
<paragraph id="962653633532FE83E4452505701F8A32" blockId="276.[1394,2465,2213,2337]" box="[1394,1878,2316,2337]" pageId="276" pageNumber="881">
<heading id="CD6EE40F3532FE83E4452505701F8A32" box="[1394,1878,2316,2337]" pageId="276" pageNumber="881">
<emphasis id="A4ED8F713532FE83E445250571208A32" bold="true" box="[1394,1641,2316,2337]" pageId="276" pageNumber="881">Other common names:</emphasis>
<vernacularName id="189A234D3532FE83E7452505701F8A32" ID-CoL="6QDJK" authorityName="Rickart, Heaney &amp; Tabaranza" authorityYear="2003" box="[1650,1878,2316,2337]" class="Mammalia" family="Muridae" genus="Limnomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="species" species="bryophilus">Gray-bellied Limnomys</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E83532FE83E6E125467E268A8B" pageId="276" pageNumber="881" type="reference_group">
<paragraph id="962653633532FE83E6E125467E268A8B" blockId="276.[2005,2610,2383,2811]" pageId="276" pageNumber="881">
<emphasis id="A4ED8F713532FE83E6E125467F388A62" bold="true" box="[2006,2161,2383,2417]" pageId="276" pageNumber="881">Taxonomy.</emphasis>
<taxonomicName id="519928E03532FE83E9BD25467E228A8B" ID-CoL="6QDJK" authority="Rickart, Heaney &amp; Tabaranza, 2003" authorityName="Rickart, Heaney &amp; Tabaranza" authorityYear="2003" class="Mammalia" family="Muridae" genus="Limnomys" kingdom="Animalia" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="species" species="bryophilus">Limnomys bryophilus Rickart, Heaney &amp; Tabaranza, 2003</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="DE8300E83532FE83E8B2257F7EAF891D" pageId="276" pageNumber="881" type="materials_examined">
<paragraph id="962653633532FE83E8B2257F7EAF891D" blockId="276.[2005,2610,2383,2811]" pageId="276" pageNumber="881">
<materialsCitation id="26F1593E3532FE83E8B2257F7EAF891D" ID-GBIF-Occurrence="3864424683" pageId="276" pageNumber="881">18-5 km S, 4 km E Camp Phillips, 2250 m, Mount Kitanglad Range, Bukidnon Province, north-central Mindanao Island, Philippines.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E83532FE83E6E026117147885B" pageId="276" pageNumber="881" type="discussion">
<paragraph id="962653633532FE83E6E026117D7B8926" blockId="276.[2005,2610,2383,2811]" box="[2007,2610,2584,2613]" pageId="276" pageNumber="881">
<taxonomicName id="519928E03532FE83E6E026117FAE8926" authorityName="Rickart, Heaney &amp; Tabaranza" authorityYear="2003" box="[2007,2279,2584,2613]" class="Mammalia" family="Muridae" genus="Limnomys" kingdom="Animalia" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="species" species="bryophilus">Limnomys bryophilus</taxonomicName>
was believed to be an
</paragraph>
<paragraph id="962653633532FE83E6E126327147885B" blockId="276.[2005,2610,2383,2811]" lastBlockId="276.[1393,2598,2815,3479]" pageId="276" pageNumber="881">
undescribed species of
<taxonomicName id="519928E03532FE83E82C26327EC5894E" authorityName="Mearns" authorityYear="1905" box="[2331,2444,2619,2653]" class="Mammalia" family="Muridae" genus="Tarsomys" kingdom="Animalia" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="genus">Tarsomys</taxonomicName>
, according to L. R. Heaney and colleagues in 1998. According to the most recent phylogeny by S. J. Steppan and J. J. Schenk in 2017,
<taxonomicName id="519928E03532FE83E6E026D07FCB89E8" authorityName="Rickart, Heaney &amp; Tabaranza" authorityYear="2003" box="[2007,2178,2777,2811]" class="Mammalia" family="Muridae" genus="Limnomys" kingdom="Animalia" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="species" species="bryophilus">L. bryophilus</taxonomicName>
is a close ally of
<taxonomicName id="519928E03532FE83E8B426D07D5689E8" authorityName="Mearns" authorityYear="1905" box="[2435,2591,2777,2811]" class="Mammalia" family="Muridae" genus="Limnomys" kingdom="Animalia" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="species" species="sibuanus">L. sibuanus</taxonomicName>
, nested in a well-supported clade containing
<taxonomicName id="519928E03532FE83E6C726F67F3A8832" authorityName="Mearns" authorityYear="1905" box="[2032,2163,2815,2849]" class="Mammalia" family="Muridae" genus="Limnomys" kingdom="Animalia" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="genus">Limnomys</taxonomicName>
, 1.
<taxonomicName id="519928E03532FE83E99A26F67E5F8832" authorityName="Mearns" authorityYear="1905" box="[2221,2326,2815,2849]" class="Mammalia" family="Muridae" genus="Tarsomys" kingdom="Animalia" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="species" species="apoensis">apoensis</taxonomicName>
, and
<taxonomicName id="519928E03532FE83E85D26F67D698832" baseAuthorityName="Gunther" baseAuthorityYear="1879" box="[2410,2592,2815,2849]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="276" pageNumber="881" phylum="Chordata" rank="species" species="everetti">Rattus everetti</taxonomicName>
. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E83532FE83E446274770478884" pageId="276" pageNumber="881" type="distribution">
<caption id="C2E603EB3532FE83E446274770478884" ID-DOI="http://doi.org/10.5281/zenodo.6888643" ID-Zenodo-Dep="6888643" httpUri="https://zenodo.org/record/6888643/files/figure.png" inLine="true" pageId="276" pageNumber="881" startId="276.[1393,1565,2894,2928]" targetBox="[1392,1982,2397,2809]" targetPageId="276">
<paragraph id="962653633532FE83E446274770478884" blockId="276.[1393,2598,2815,3479]" pageId="276" pageNumber="881">
<emphasis id="A4ED8F713532FE83E446274771688863" bold="true" box="[1393,1569,2894,2928]" pageId="276" pageNumber="881">Distribution.</emphasis>
Kitanglad Mountain Range, Mindanao I, Philippines (perhaps other high peaks of C Mindanao I).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DE8300E83532FE82E446279474A88263" lastPageId="277" lastPageNumber="882" pageId="276" pageNumber="881" type="description">
<paragraph id="962653633532FE82E446279474A88263" blockId="276.[1393,2598,2815,3479]" lastBlockId="277.[164,1372,295,1315]" lastPageId="277" lastPageNumber="882" pageId="276" pageNumber="881">
<emphasis id="A4ED8F713532FE83E4462794712488AC" bold="true" box="[1393,1645,2973,3007]" pageId="276" pageNumber="881">Descriptive notes.</emphasis>
Head-body 121-140 mm, tail 152-181 mm, ear 20-24 mm, hindfoot 30-35 mm; weight 48-90 g. The Gray-bellied Moss Mouse is slightly larger than the Long-tailed Moss Mouse (L. sitbuanus) in most external and cranial measurements. Pelage is long, thick, and soft. Upperparts are yellowish brown, darkest mid-dorsally and on rump, and paler on sides. Underparts are grayish white, with moderately demarcation against dorsum. There is usually a band of bufty fur acrosslower neck. Ears are brown and sparsely covered with hair. Tail is substantially longer than head-body length and relatively well-haired. It is dark brown for most of its length, with white tail tip (5—=20 mm), which is not present in most Long-tailed Moss Mice or much smaller. Forefeet and hindfeet are unpigmented and sparsely covered with short, light brown hair on top and silvery white laterally and on digits. The Gray-bellied Moss Mouse differs from the Long-tailed Moss Mouse in having relatively longer tail, larger and wider hindfeet, softer pelage with grayish white venter, larger skull with longer rostrum, larger auditory bullae, larger molars, and other cranial and dental features. There are three pairs of mammae: one post-axillary and two inguinal.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E83533FE82E1922D7F732B81BE" pageId="277" pageNumber="882" type="biology_ecology">
<paragraph id="962653633533FE82E1922D7F732B81BE" blockId="277.[164,1372,295,1315]" pageId="277" pageNumber="882">
<emphasis id="A4ED8F713533FE82E1922D7F765D828B" bold="true" box="[165,276,374,408]" pageId="277" pageNumber="882">Habitat.</emphasis>
Primary mossy montane forest at elevations from 2250 m to 2800 m. At the type locality on a steep ridgeon northern flank of Mount Kitanglad (2250 m elevation), Gray-bellied Moss Mice were captured in a transition zone between lower montane and upper montane rainforest, dominated by several species of gymnosperms, laurels, and epiphytes (moss, ferns, lichen, and orchids). Soil was covered with deep humus layer and leaf litter. Understory plants included broadleaf saplings,
<taxonomicName id="519928E03533FE82E5E12E3276C38197" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="277" pageNumber="882" phylum="Tracheophyta" rank="subGenus" subGenus="Ericaceae">Rhododendron (Ericaceae)</taxonomicName>
and other shrubs, ferns, and sedges. The Gray-bellied Moss Mouse and the Long-tailed Moss Mouse are sympatric on Mount Kitanglad.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E83533FE82E1922EBB74C981EF" pageId="277" pageNumber="882" type="food_feeding">
<paragraph id="962653633533FE82E1922EBB74C981EF" blockId="277.[164,1372,295,1315]" pageId="277" pageNumber="882">
<emphasis id="A4ED8F713533FE82E1922EBB76FB81C7" bold="true" box="[165,434,690,724]" pageId="277" pageNumber="882">Food and Feeding.</emphasis>
The Gray-bellied Moss Mouse eats seeds, fruits, arthropods, and earthworms on the ground and often in steep areas.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E83533FE82E1922F0C724D8031" box="[165,1284,773,802]" pageId="277" pageNumber="882" type="breeding">
<paragraph id="962653633533FE82E1922F0C724D8031" blockId="277.[164,1372,295,1315]" box="[165,1284,773,802]" pageId="277" pageNumber="882">
<emphasis id="A4ED8F713533FE82E1922F0C76658031" bold="true" box="[165,300,773,802]" pageId="277" pageNumber="882">Breeding.</emphasis>
Gray-bellied Moss Mice have small litters (1-2 young) once each year.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E83533FE82E1932F39725580AC" pageId="277" pageNumber="882" type="activity">
<paragraph id="962653633533FE82E1932F39725580AC" blockId="277.[164,1372,295,1315]" pageId="277" pageNumber="882">
<emphasis id="A4ED8F713533FE82E1932F3976D9805A" bold="true" box="[164,400,816,841]" pageId="277" pageNumber="882">Activity patterns.</emphasis>
Gray-bellied Moss Mice are nocturnal, possibly crepuscular, and terrestrial-scansorial. Most specimens were trapped on steeply sloping ground with abundant moss cover. On Mount Kitanglad and Mount Dulang-dulang (elevations 2375— 2800 m), they were trapped in association with nine other small mammal species.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E83533FE82E1922FCF731880FB" box="[165,1105,966,1000]" pageId="277" pageNumber="882" type="biology_ecology">
<paragraph id="962653633533FE82E1922FCF731880FB" blockId="277.[164,1372,295,1315]" box="[165,1105,966,1000]" pageId="277" pageNumber="882">
<emphasis id="A4ED8F713533FE82E1922FCF742B80FB" bold="true" box="[165,866,966,1000]" pageId="277" pageNumber="882">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E83533FE82E1912FF876038796" pageId="277" pageNumber="882" type="conservation">
<paragraph id="962653633533FE82E1912FF876038796" blockId="277.[164,1372,295,1315]" pageId="277" pageNumber="882">
<emphasis id="A4ED8F713533FE82E1912FF8755C871D" bold="true" box="[166,533,1009,1038]" pageId="277" pageNumber="882">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. The Gray-bellied Moss Mouse is common in high-elevation forests but might be restricted geographically. It occurs in Mount Kitanglad Nature Park, and its habitat is not threatened.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E83533FE82E190289176F68630" pageId="277" pageNumber="882" type="bibRefCitation_list">
<paragraph id="962653633533FE82E190289176F68630" blockId="277.[164,1372,295,1315]" pageId="277" pageNumber="882">
<emphasis id="A4ED8F713533FE82E1902891760987BE" bold="true" box="[167,320,1176,1197]" pageId="277" pageNumber="882">Bibliography.</emphasis>
Boitani et al. (2006), Ellerman (1941), Fabre, Hautier et al. (2012), Fabre, Pages et al. (2013), Heaney &amp; Tabaranza (2008), Heaney, Balete et al. (1998), Heaney, Dolar et al. (2010), Heaney, Tabaranza, Rickart et al. (2006), Jansa et al. (2006), Musser &amp; Carleton (2005), Musser &amp; Heaney (1992), Rickart &amp; Heaney (2002), Steppan &amp; Schenk (2017).
</paragraph>
</subSubSection>
</treatment>
</document>
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