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<document id="BAA8B3C081115E5A206988266188B71F" ID-CLB-Dataset="100954" ID-DOI="10.5281/zenodo.6887260" ID-GBIF-Dataset="a016af63-6437-427b-80b7-22bc9a002e20" ID-ISBN="978-84-16728-04-6" ID-Zenodo-Dep="6887260" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="guilherme" checkinTime="1656696812616" checkinUser="carolina" docAuthor="Don E. Wilson, Russell A. Mittermeier &amp; Thomas E. Lacher, Jr" docDate="2017" docId="1E30E27534DFFF6DE4952BDE75638B1D" docLanguage="en" docName="hbmw_7_Muridae_0536.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 7 Rodents II, Barcelona: Lynx Edicions" docTitle="Leopoldamys sabanus" docType="treatment" docVersion="11" lastPageNumber="855" masterDocId="E2099A0D3426FF97E1372C0977498313" masterDocTitle="Muridae" masterLastPageNumber="884" masterPageNumber="536" pageNumber="854" updateTime="1718983125921" updateUser="carolina">
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<mods:title id="9C4C4C3D07EA2AE082AB475514DCA567">Muridae</mods:title>
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<mods:namePart id="A238F571193E0C7889159801294234D7">Don E. Wilson</mods:namePart>
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<mods:namePart id="8E8F9553E12C9978DF76DBE162B3BB7A">Russell A. Mittermeier</mods:namePart>
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<mods:namePart id="A68742A1025E0EADE67A1D03FB8E52AC">Thomas E. Lacher, Jr</mods:namePart>
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<treatment id="1E30E27534DFFF6DE4952BDE75638B1D" ID-DOI="http://doi.org/10.5281/zenodo.6832963" ID-GBIF-Taxon="197826534" ID-Zenodo-Dep="6832963" LSID="urn:lsid:plazi:treatment:1E30E27534DFFF6DE4952BDE75638B1D" httpUri="http://treatment.plazi.org/id/1E30E27534DFFF6DE4952BDE75638B1D" lastPageId="250" lastPageNumber="855" pageId="249" pageNumber="854">
<subSubSection id="DE8300E834DFFF6EE4952BDE72BD8B1A" box="[1442,1524,2007,2057]" pageId="249" pageNumber="854" type="multiple">
<paragraph id="9626536334DFFF6EE4952BDE72BD8B1A" blockId="249.[1440,2589,2007,2219]" box="[1442,1524,2007,2057]" pageId="249" pageNumber="854">
<heading id="CD6EE40F34DFFF6EE4952BDE72BD8B1A" box="[1442,1524,2007,2057]" pageId="249" pageNumber="854">
<figureCitation id="0EA24FE634DFFF6EE4952BDE72BD8B1A" box="[1442,1524,2007,2057]" captionStart="Plate 56: Muridae" captionStartId="248.[116,147,3133,3158]" captionTargetBox="[14,2745,12,3649]" captionTargetPageId="247" captionText="724. Burmese Tree Rat (Chiromyscus chiropus), 725. Langbian Tree Rat (Chiromyscus langbianis), 726. Thomas's Tree Rat (Chiromyscus thomasi), 727. Millets Long-tailed Giant Rat (Leopoldamys milleti), 728. Edwardss Long-tailed Giant Rat (Leopoldamys edwardsi), 729. Indomalayan Long-tailed Giant Rat (Leopoldamys sabanus), 730. Herbert's Long-tailed Giant Rat (Leopoldamys herberti), 731. Neills Long-tailed Giant Rat (Leopoldamys neilli), 733. Diwangkaras Long-tailed Giant Rat (Leopoldamys diwangkarai), 734. Mentawai Archipelago Long-tailed Giant Rat (Leopoldamys siporanus), 735. Paulinas Limestone Rat (Saxatilomys paulinae), 736. Daovantiens Limestone Rat (Tonkinomys daovantieni), 737. Javan Bamboo Rat (Kadarsanomys sodyi), 739. Indomalayan Spiny Rat (Maxomys surifer), 740. Indochinese Mountain Spiny Rat (Maxomys moi), 741. Bornean Mountain Spiny Rat (Maxomys alticola), 742. Small Bornean Spiny Rat (Maxomys baeodon), 743. Bartelss Javan Spiny Rat (Maxomys bartelsii), 744. Sumatran Mountain Spiny Rat (Maxomys hylomyoides), 745. Malayan Mountain Spiny Rat (Maxomys inas), 746. Broad-nosed Sumatran Spiny Rat (Maxomys inflatus), 747. Ochraceous-bellied Bornean Spiny Rat (Maxomys ochracerventer), 748. Mentawai Archipelago Spiny Rat (Maxomys pagensis), 749. Palawan Spiny Rat (Maxomys panglima), 750. Rajah Sundaic Spiny Rat (Maxomys rajah), 751. Whiteheads Sundaic Spiny Rat (Maxomys whiteheadi), 752. Dollmans Sulawesi Spiny Rat (Maxomys dollmani), 753. Hellwalds Sulawesi Spiny Rat (Maxomys hellwaldii), 755. Tajuddins Spiny Rat (Maxomys tajuddinii)" figureDoi="http://doi.org/10.5281/zenodo.6888489" httpUri="https://zenodo.org/record/6888489/files/figure.png" pageId="249" pageNumber="854">729.</figureCitation>
</heading>
</paragraph>
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<subSubSection id="DE8300E834DFFF6EE7322BDE7FAE8B1A" box="[1541,2279,2007,2057]" pageId="249" pageNumber="854" type="vernacular_names">
<paragraph id="9626536334DFFF6EE7322BDE7FAE8B1A" blockId="249.[1440,2589,2007,2219]" box="[1541,2279,2007,2057]" pageId="249" pageNumber="854">
<heading id="CD6EE40F34DFFF6EE7322BDE7FAE8B1A" box="[1541,2279,2007,2057]" pageId="249" pageNumber="854">
<vernacularName id="189A234D34DFFF6EE7322BDE7FAE8B1A" ID-CoL="3T797" baseAuthorityName="Thomas" baseAuthorityYear="1887" box="[1541,2279,2007,2057]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" language="eng" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="sabanus">Indomalayan Long-tailed Giant Rat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DFFF6EE496240270718B2A" box="[1441,1848,2059,2105]" pageId="249" pageNumber="854" type="nomenclature">
<paragraph id="9626536334DFFF6EE496240270718B2A" blockId="249.[1440,2589,2007,2219]" box="[1441,1848,2059,2105]" pageId="249" pageNumber="854">
<heading id="CD6EE40F34DFFF6EE496240270718B2A" box="[1441,1848,2059,2105]" pageId="249" pageNumber="854">
<taxonomicName id="519928E034DFFF6EE496240270718B2A" ID-CoL="3T797" baseAuthorityName="Thomas" baseAuthorityYear="1887" box="[1441,1848,2059,2105]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="sabanus">
<emphasis id="A4ED8F7134DFFF6EE496240270718B2A" box="[1441,1848,2059,2105]" italics="true" pageId="249" pageNumber="854">Leopoldamys sabanus</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DFFF6EE496244E7FB18BB8" pageId="249" pageNumber="854" type="vernacular_names">
<paragraph id="9626536334DFFF6EE496244E715F8B90" blockId="249.[1440,2589,2007,2219]" pageId="249" pageNumber="854">
<heading id="CD6EE40F34DFFF6EE496244E715F8B90" pageId="249" pageNumber="854">
<emphasis id="A4ED8F7134DFFF6EE496244E72A58B4F" bold="true" box="[1441,1516,2119,2140]" pageId="249" pageNumber="854">French:</emphasis>
<vernacularName id="189A234D34DFFF6EE4C1244E71318B4F" ID-CoL="3T797" baseAuthorityName="Thomas" baseAuthorityYear="1887" box="[1526,1656,2119,2140]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" language="fra" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="sabanus">Rat de Sabah</vernacularName>
/
<emphasis id="A4ED8F7134DFFF6EE7B9244E71A18B4F" bold="true" box="[1678,1768,2119,2140]" pageId="249" pageNumber="854">German:</emphasis>
<vernacularName id="189A234D34DFFF6EE7C5244E7FCB8B4F" ID-CoL="3T797" baseAuthorityName="Thomas" baseAuthorityYear="1887" box="[1778,2178,2119,2140]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" language="deu" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="sabanus">Siidasiatische Langschwanz-Riesenratte</vernacularName>
/
<emphasis id="A4ED8F7134DFFF6EE9A0244E7FBA8B4F" bold="true" box="[2199,2291,2119,2140]" pageId="249" pageNumber="854">Spanish:</emphasis>
<vernacularName id="189A234D34DFFF6EE9CA244E715F8B90" ID-CoL="3T797" baseAuthorityName="Thomas" baseAuthorityYear="1887" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" language="esp" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="sabanus">Rata gigante de cola larga de Indomalasia</vernacularName>
</heading>
</paragraph>
<paragraph id="9626536334DFFF6EE497249F7FB18BB8" blockId="249.[1440,2589,2007,2219]" box="[1440,2296,2198,2219]" pageId="249" pageNumber="854">
<heading id="CD6EE40F34DFFF6EE497249F7FB18BB8" box="[1440,2296,2198,2219]" pageId="249" pageNumber="854">
<emphasis id="A4ED8F7134DFFF6EE497249F71D18BB8" bold="true" box="[1440,1688,2198,2219]" pageId="249" pageNumber="854">Other common names:</emphasis>
<vernacularName id="189A234D34DFFF6EE795249F70E58BB8" ID-CoL="3T797" baseAuthorityName="Thomas" baseAuthorityYear="1887" box="[1698,1964,2198,2219]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" language="eng" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="sabanus">Indomalayan Leopoldamys</vernacularName>
,
<vernacularName id="189A234D34DFFF6EE68D249F7FC58BB8" ID-CoL="3T797" baseAuthorityName="Thomas" baseAuthorityYear="1887" box="[1978,2188,2198,2219]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" language="eng" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="sabanus">Long-tailed Giant Rat</vernacularName>
,
<vernacularName id="189A234D34DFFF6EE9AD249F7FB18BB8" ID-CoL="3T797" baseAuthorityName="Thomas" baseAuthorityYear="1887" box="[2202,2296,2198,2219]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" language="eng" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="sabanus">Noisy Rat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DFFF6EE93324D47D1B8BE9" box="[2052,2642,2269,2298]" pageId="249" pageNumber="854" type="reference_group">
<paragraph id="9626536334DFFF6EE93324D47D1B8BE9" blockId="249.[2052,2645,2269,2691]" box="[2052,2642,2269,2298]" pageId="249" pageNumber="854">
<emphasis id="A4ED8F7134DFFF6EE93324D47FD68BE9" bold="true" box="[2052,2207,2269,2298]" pageId="249" pageNumber="854">Taxonomy.</emphasis>
<taxonomicName id="519928E034DFFF6EE98224D47D048BE9" authority="Thomas, 1887" authorityName="Thomas" authorityYear="1887" box="[2229,2637,2269,2298]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="sabanus">Mus sabanus Thomas, 1887</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DFFF6EE93225017EC58A63" pageId="249" pageNumber="854" type="materials_examined">
<paragraph id="9626536334DFFF6EE93225017E4D8A63" blockId="249.[2052,2645,2269,2691]" pageId="249" pageNumber="854">
<materialsCitation id="26F1593E34DFFF6EE93225017E4D8A63" ID-GBIF-Occurrence="3864424816" pageId="249" pageNumber="854">“Saba [= Sabah], the district of North Bor-neo in which Mount Kina-Balu [= Gunung Kinabalu] stands,”</materialsCitation>
</paragraph>
<paragraph id="9626536334DFFF6EE827255E7EC58A63" blockId="249.[2052,2645,2269,2691]" box="[2320,2444,2391,2416]" pageId="249" pageNumber="854">Malaysia.</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DFFF6EE93225737EF78863" pageId="249" pageNumber="854" type="discussion">
<paragraph id="9626536334DFFF6EE93225737EF78863" blockId="249.[2052,2645,2269,2691]" lastBlockId="249.[1438,2657,2697,3479]" pageId="249" pageNumber="854">
<taxonomicName id="519928E034DFFF6EE93225737FEF8A84" box="[2053,2214,2426,2455]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="genus">Leopoldamys</taxonomicName>
herberti was recently recognized as a distinct species from
<taxonomicName id="519928E034DFFF6EE8F025A87D1C8AAD" authorityName="Thomas" authorityYear="1887" box="[2503,2645,2465,2494]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="sabanus">L. sabanus</taxonomicName>
based on genetic and morphological data. Distribution of
<taxonomicName id="519928E034DFFF6EE80125E57E8E891D" authorityName="Thomas" authorityYear="1887" box="[2358,2503,2540,2574]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="sabanus">L. sabanus</taxonomicName>
has been restricted to the Sundaic region with a northern limit of Kanchanaburi Province, southern Thailand. Leopolamys
<taxonomicName id="519928E034DFFF6EE8FD266F7D7B8990" authorityName="Thomas" authorityYear="1887" box="[2506,2610,2662,2691]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="sabanus">sabanus</taxonomicName>
is in need of a thorough taxonomic revision because there have been many subspecies recognized and genetic data show that it probably represents a species complex. Remaining mainland populations in southern Thailand and Peninsular Malaysia were recently recognized as a distinct species,
<taxonomicName id="519928E034DFFF6EE6C426F67FD28832" box="[2035,2203,2815,2849]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="vociferans">L. vociferans</taxonomicName>
, which is not recognized here because a more thorough revision is needed. There are clearly more taxa to delineated within
<taxonomicName id="519928E034DFFF6EE736274771C78863" authorityName="Thomas" authorityYear="1887" box="[1537,1678,2894,2928]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="species" species="sabanus">L. sabanus</taxonomicName>
, pending additional research. Two subspecies recognized.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DFFF6EE497277371C28F27" pageId="249" pageNumber="854" type="distribution">
<caption id="C2E603EB34DFFF6EE497277371C28F27" ID-DOI="http://doi.org/10.5281/zenodo.6888473" ID-Zenodo-Dep="6888473" httpUri="https://zenodo.org/record/6888473/files/figure.png" inLine="true" pageId="249" pageNumber="854" startId="249.[1440,1594,2938,2967]" targetBox="[1438,2028,2275,2687]" targetPageId="249">
<paragraph id="9626536334DFFF6EE497277370798884" blockId="249.[1438,2657,2697,3479]" box="[1440,1840,2938,2967]" pageId="249" pageNumber="854">
<emphasis id="A4ED8F7134DFFF6EE497277370798884" bold="true" box="[1440,1840,2938,2967]" pageId="249" pageNumber="854">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph id="9626536334DFFF6EE49627AF715A88F5" blockId="249.[1438,2657,2697,3479]" pageId="249" pageNumber="854">
<taxonomicName id="519928E034DFFF6EE49627AF715A88F5" authority="Thomas, 1887" authorityName="Thomas" authorityYear="1887" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="subSpecies" species="sabanus" subSpecies="sabanus">L.s.sabanusThomas,1887—Sumatra,Java,andBorneo,aswellasvarioussmalloffshoreIs.</taxonomicName>
</paragraph>
<paragraph id="9626536334DFFF6EE49627FD71C28F27" blockId="249.[1438,2657,2697,3479]" pageId="249" pageNumber="854">
<taxonomicName id="519928E034DFFF6EE49627FD70118F1E" authority="G. S. Miller, 1900" authorityName="G. S. Miller" authorityYear="1900" box="[1441,1880,3060,3085]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="subSpecies" species="sabanus" subSpecies="vociferans">L. s. vociferans G. S. Miller, 1900</taxonomicName>
— S Myanmar, S Thailand, Malay Peninsula, and some small offshore Is.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DE8300E834DFFF6DE4A82037739482F5" lastPageId="250" lastPageNumber="855" pageId="249" pageNumber="854" type="description">
<paragraph id="9626536334DFFF6DE4A82037739482F5" blockId="249.[1438,2657,2697,3479]" lastBlockId="250.[113,1329,299,2062]" lastPageId="250" lastPageNumber="855" pageId="249" pageNumber="854">
<emphasis id="A4ED8F7134DFFF6EE4A8203771D58F48" bold="true" box="[1439,1692,3134,3163]" pageId="249" pageNumber="854">Descriptive notes.</emphasis>
Head-body 200-275 mm, tail 270-415 mm, hindfoot 42-52 mm; weight 250-500 g. The Indomalayan Long-tailed Giant Rat is very large, with soft, spiny, and sleek pelage. Dorsum is buffy brown to orangish brown, being darker along spine and brighter orangish brown onsides; guard hairs are shorter than most other species of
<taxonomicName id="519928E034DFFF6EE70120D171918FE9" box="[1590,1752,3288,3322]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" order="Rodentia" pageId="249" pageNumber="854" phylum="Chordata" rank="genus">Leopoldamys</taxonomicName>
. Venter is white to yellowish white, sometimes with more pronounced yellowish hue, and sharply demarcated from dorsum. Feet are brown on top with white toes; hindfeet are long. Ears are long, leathery, and brown; vibrissae are very long. Tail is 135% of head-body length, blotchy brown above and creamy white below, being clearly demarcated; tip of tail is white on some specimens. Skull is large and elongated, typical of the genus. There are four pairs of mammae: one pectoral, one post-axillary, and two inguinal pairs. Nematodes Filarioidea, Globocephalus, Heligmonoides, Mastopharus,
<taxonomicName id="519928E034DCFF6DE3072D7F7458828B" authorityName="Railliet" authorityYear="1893" box="[560,785,374,408]" class="Secernentea" family="Physalopteridae" kingdom="Animalia" order="Spirurida" pageId="250" pageNumber="855" phylum="Nematoda" rank="family">Physalopteridae</taxonomicName>
, etc., cestode species
<taxonomicName id="519928E034DCFF6DE57E2D7F73A1828B" box="[1097,1256,374,408]" class="Magnoliopsida" family="Asteraceae" genus="Hymenolepis" kingdom="Plantae" order="Asterales" pageId="250" pageNumber="855" phylum="Tracheophyta" rank="genus">Hymenolepis</taxonomicName>
and Rodentolepis, trematode Zonorchis, pentastomide Armillifer, and some ticks Ixodes have been recorded from this species. Chromosomal complement is 2n = 42, FN = 54.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DCFF6DE1452DE5765F81C1" pageId="250" pageNumber="855" type="biology_ecology">
<paragraph id="9626536334DCFF6DE1452DE5765F81C1" blockId="250.[113,1329,299,2062]" pageId="250" pageNumber="855">
<emphasis id="A4ED8F7134DCFF6DE1452DE577A8811D" bold="true" box="[114,225,492,526]" pageId="250" pageNumber="855">Habitat.</emphasis>
Primarily lowland tropical forests, sometimes montane wet tropical forests, and disturbed habitat (e.g. logged forests), preferring drier parts of forests, at elevations of 300-3100 m. The Indomalayan Long-tailed Giant Rat seems to be found at higher elevations on Borneo (3100 m) than in Malaysia (1400 m), and it isapparently found at a different elevation than the Sundaic Mountain Long-tailed Giant Rat (L. cthatus).
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DCFF6DE1452ED1755280AC" pageId="250" pageNumber="855" type="food_feeding">
<paragraph id="9626536334DCFF6DE1452ED1755280AC" blockId="250.[113,1329,299,2062]" pageId="250" pageNumber="855">
<emphasis id="A4ED8F7134DCFF6DE1452ED176CD81E9" bold="true" box="[114,388,728,762]" pageId="250" pageNumber="855">Food and Feeding.</emphasis>
The Indomalayan Long-tailed Giant Rat generally eats insects, snails, fruit, and other vegetation (including roots and leaves). Foraging occurs in the canopy and on the ground. Food hoarding occurs; in Malaysia, they would hide fruit given to them (oil palm fruit,
<taxonomicName id="519928E034DCFF6DE3542F4775E78063" box="[611,686,846,880]" class="Liliopsida" family="Arecaceae" genus="Elaeis" kingdom="Plantae" order="Arecales" pageId="250" pageNumber="855" phylum="Tracheophyta" rank="genus">Elaeis</taxonomicName>
guineensis,
<taxonomicName id="519928E034DCFF6DE27E2F4774A98063" box="[841,992,846,880]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="250" pageNumber="855" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
), caching it on ground covered with leaves. Camera traps showed that the Indomalayan Long-tailed Giant Rat was attracted to fish oil as bait.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DCFF6DE1442FC17483874E" pageId="250" pageNumber="855" type="breeding">
<paragraph id="9626536334DCFF6DE1442FC17483874E" blockId="250.[113,1329,299,2062]" pageId="250" pageNumber="855">
<emphasis id="A4ED8F7134DCFF6DE1442FC177B080F6" bold="true" box="[115,249,968,997]" pageId="250" pageNumber="855">Breeding.</emphasis>
Breeding occurs year-round, with peak of pregnant females in July—September, being less common in January-March. Litters have 1-7 young (mean3-1). Young are born in burrows with 1-2 entrances dug into hillsides. Nests are usually lined with broad leaves collected from nearby shrubs and undergrowth.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DCFF6DE146286B76E8868B" pageId="250" pageNumber="855" type="activity">
<paragraph id="9626536334DCFF6DE146286B76E8868B" blockId="250.[113,1329,299,2062]" pageId="250" pageNumber="855">
<emphasis id="A4ED8F7134DCFF6DE146286B762B8797" bold="true" box="[113,354,1122,1156]" pageId="250" pageNumber="855">Activity patterns.</emphasis>
The Indomalayan Long-tailed Giant Rat is mainly terrestrial and nocturnal. A recent study found 20 nests underground and four nests aboveground, and another study found 20 nests underground and 14 nests aboveground, indicating preference for nesting on the ground. Ground nests were 40-80 m apart, more commonly 50-80 m. Aboveground nests were located in tree cavities and logs with entrances 1-1-6 m aboveground. The species sleeps in nests during the day and comes out at night. Mean time spent moving at night in one study was 485 minutesin logged and unlogged forests.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DCFF6DE144299473398596" pageId="250" pageNumber="855" type="biology_ecology">
<paragraph id="9626536334DCFF6DE144299473398596" blockId="250.[113,1329,299,2062]" pageId="250" pageNumber="855">
<emphasis id="A4ED8F7134DCFF6DE1442994740686AC" bold="true" box="[115,847,1437,1471]" pageId="250" pageNumber="855">Movements, Home range and Social organization.</emphasis>
Indomalayan Long-tailed Giant Rats are known to traverse trees to forage. Females and males or females and juveniles are known to stay in the same nests. When moving around at night, paths (determined by radio-tracking data) are relatively straight, and movement averages 32 m/ten minutes, covering areas of 2083-9829 m®. Paths were mostlikely determined by location of resources rather than by sex or habitat type (logged vs. unlogged forest).
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DCFF6DE1432A8374938459" pageId="250" pageNumber="855" type="conservation">
<paragraph id="9626536334DCFF6DE1432A8374938459" blockId="250.[113,1329,299,2062]" pageId="250" pageNumber="855">
<emphasis id="A4ED8F7134DCFF6DE1432A83769E85BF" bold="true" box="[116,471,1674,1708]" pageId="250" pageNumber="855">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. The Indomalayan Long-tailed Giant Rat has a wide distribution and is fairly common, being the most common species found in a surveyat Mount Nuangin Selangor, Malaysia. Deforestation might be a threat in regions with prevalent logging, but it commonly occurs in logged forests and is found in many protected areas.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DCFF6DE1432B5575638B1D" pageId="250" pageNumber="855" type="bibRefCitation_list">
<paragraph id="9626536334DCFF6DE1432B5575638B1D" blockId="250.[113,1329,299,2062]" pageId="250" pageNumber="855">
<emphasis id="A4ED8F7134DCFF6DE1432B5576448462" bold="true" box="[116,269,1884,1905]" pageId="250" pageNumber="855">Bibliography.</emphasis>
Aplin, Ruedas &amp; Molur (2016), Balakirev, Abramov &amp; Rozhnov (2013), Francis (2008), Gorog et al. (2004), Lah et al. (2015), Latinne, Chaval et al. (2013), Latinne, Waengsothorn et al. (2013), Lim (1970), Madinah etal. (2011), Medway (1969), Musser (1981a), Musser &amp; Carleton (2005), Nor (2001), Ow-Yang (1971), Phillipps &amp; Phillipps (2016), Singh &amp; Cheong (1971), Thanee et al. (2009), Wells, Corlett et al. (2009), Wells, Kalko et al. (2008), Yasuda et al. (2000), Yong (1970).
</paragraph>
</subSubSection>
</treatment>
</document>