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<document id="0B349E26DFB20A6026142A0F1A1954E3" ID-CLB-Dataset="100954" ID-DOI="10.5281/zenodo.6887260" ID-GBIF-Dataset="a016af63-6437-427b-80b7-22bc9a002e20" ID-ISBN="978-84-16728-04-6" ID-Zenodo-Dep="6887260" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="guilherme" checkinTime="1656696812616" checkinUser="carolina" docAuthor="Don E. Wilson, Russell A. Mittermeier &amp; Thomas E. Lacher, Jr" docDate="2017" docId="1E30E27534DDFF6BE49326CD7359867F" docLanguage="en" docName="hbmw_7_Muridae_0536.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 7 Rodents II, Barcelona: Lynx Edicions" docTitle="Saxatilomys paulinae Musser et al. 2005" docType="treatment" docVersion="11" lastPageNumber="857" masterDocId="E2099A0D3426FF97E1372C0977498313" masterDocTitle="Muridae" masterLastPageNumber="884" masterPageNumber="536" pageNumber="856" updateTime="1718983125921" updateUser="carolina">
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<mods:title id="1A54D706AD309F6DDCA22DF42A07F9D6">Muridae</mods:title>
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<mods:namePart id="EAA346A696F4FC28201C08F7D235CB02">Don E. Wilson</mods:namePart>
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<mods:namePart id="7110F64BC163AD34FA557D47E1E0FFB9">Russell A. Mittermeier</mods:namePart>
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<mods:namePart id="6A641455E20BD8ECC922C6373DDC6B33">Thomas E. Lacher, Jr</mods:namePart>
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<mods:title id="FF7DF8A3B061EF87F9E35A028835EEAE">Handbook of the Mammals of the World Volume 7 Rodents II</mods:title>
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<treatment id="1E30E27534DDFF6BE49326CD7359867F" ID-DOI="http://doi.org/10.5281/zenodo.6869037" ID-GBIF-Taxon="197826217" ID-Zenodo-Dep="6869037" LSID="urn:lsid:plazi:treatment:1E30E27534DDFF6BE49326CD7359867F" httpUri="http://treatment.plazi.org/id/1E30E27534DDFF6BE49326CD7359867F" lastPageId="252" lastPageNumber="857" pageId="251" pageNumber="856">
<subSubSection id="DE8300E834DDFF6CE49326CD72BC89E5" box="[1444,1525,2756,2806]" pageId="251" pageNumber="856" type="multiple">
<paragraph id="9626536334DDFF6CE49326CD72BC89E5" blockId="251.[1442,2498,2756,2845]" box="[1444,1525,2756,2806]" pageId="251" pageNumber="856">
<figureCitation id="0EA24FE634DDFF6CE49326CD72BC89E5" box="[1444,1525,2756,2806]" captionStart="Plate 56: Muridae" captionStartId="248.[116,147,3133,3158]" captionTargetBox="[14,2745,12,3649]" captionTargetPageId="247" captionText="724. Burmese Tree Rat (Chiromyscus chiropus), 725. Langbian Tree Rat (Chiromyscus langbianis), 726. Thomas's Tree Rat (Chiromyscus thomasi), 727. Millets Long-tailed Giant Rat (Leopoldamys milleti), 728. Edwardss Long-tailed Giant Rat (Leopoldamys edwardsi), 729. Indomalayan Long-tailed Giant Rat (Leopoldamys sabanus), 730. Herbert's Long-tailed Giant Rat (Leopoldamys herberti), 731. Neills Long-tailed Giant Rat (Leopoldamys neilli), 733. Diwangkaras Long-tailed Giant Rat (Leopoldamys diwangkarai), 734. Mentawai Archipelago Long-tailed Giant Rat (Leopoldamys siporanus), 735. Paulinas Limestone Rat (Saxatilomys paulinae), 736. Daovantiens Limestone Rat (Tonkinomys daovantieni), 737. Javan Bamboo Rat (Kadarsanomys sodyi), 739. Indomalayan Spiny Rat (Maxomys surifer), 740. Indochinese Mountain Spiny Rat (Maxomys moi), 741. Bornean Mountain Spiny Rat (Maxomys alticola), 742. Small Bornean Spiny Rat (Maxomys baeodon), 743. Bartelss Javan Spiny Rat (Maxomys bartelsii), 744. Sumatran Mountain Spiny Rat (Maxomys hylomyoides), 745. Malayan Mountain Spiny Rat (Maxomys inas), 746. Broad-nosed Sumatran Spiny Rat (Maxomys inflatus), 747. Ochraceous-bellied Bornean Spiny Rat (Maxomys ochracerventer), 748. Mentawai Archipelago Spiny Rat (Maxomys pagensis), 749. Palawan Spiny Rat (Maxomys panglima), 750. Rajah Sundaic Spiny Rat (Maxomys rajah), 751. Whiteheads Sundaic Spiny Rat (Maxomys whiteheadi), 752. Dollmans Sulawesi Spiny Rat (Maxomys dollmani), 753. Hellwalds Sulawesi Spiny Rat (Maxomys hellwaldii), 755. Tajuddins Spiny Rat (Maxomys tajuddinii)" figureDoi="http://doi.org/10.5281/zenodo.6888489" httpUri="https://zenodo.org/record/6888489/files/figure.png" pageId="251" pageNumber="856">735.</figureCitation>
</paragraph>
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<paragraph id="9626536334DDFF6CE73126CD70B489E5" blockId="251.[1442,2498,2756,2845]" box="[1542,2045,2756,2806]" pageId="251" pageNumber="856">
<vernacularName id="189A234D34DDFF6CE73126CD70B489E5" ID-CoL="79QYQ" authorityName="Musser et al." authorityYear="2005" box="[1542,2045,2756,2806]" class="Mammalia" family="Muridae" genus="Saxatilomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="251" pageNumber="856" phylum="Chordata" rank="species" species="paulinae">Paulinas Limestone Rat</vernacularName>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DDFF6CE90626CD7E8B89E5" box="[2097,2498,2756,2806]" pageId="251" pageNumber="856" type="nomenclature">
<paragraph id="9626536334DDFF6CE90626CD7E8B89E5" blockId="251.[1442,2498,2756,2845]" box="[2097,2498,2756,2806]" pageId="251" pageNumber="856">
<taxonomicName id="519928E034DDFF6CE90626CD7E8B89E5" ID-CoL="79QYQ" authorityName="Musser et al." authorityYear="2005" box="[2097,2498,2756,2806]" class="Mammalia" family="Muridae" genus="Saxatilomys" kingdom="Animalia" order="Rodentia" pageId="251" pageNumber="856" phylum="Chordata" rank="species" species="paulinae">
<emphasis id="A4ED8F7134DDFF6CE90626CD7E8B89E5" box="[2097,2498,2756,2806]" italics="true" pageId="251" pageNumber="856">Saxatilomys paulinae</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DDFF6CE49527017E54880E" box="[1442,2333,2824,2845]" pageId="251" pageNumber="856" type="vernacular_names">
<paragraph id="9626536334DDFF6CE49527017E54880E" blockId="251.[1442,2498,2756,2845]" box="[1442,2333,2824,2845]" pageId="251" pageNumber="856">
<emphasis id="A4ED8F7134DDFF6CE495270172A7880E" bold="true" box="[1442,1518,2824,2845]" pageId="251" pageNumber="856">French:</emphasis>
<vernacularName id="189A234D34DDFF6CE4C0270171C0880E" ID-CoL="79QYQ" authorityName="Musser et al." authorityYear="2005" box="[1527,1673,2824,2845]" class="Mammalia" family="Muridae" genus="Saxatilomys" kingdom="Animalia" language="fra" order="Rodentia" pageId="251" pageNumber="856" phylum="Chordata" rank="species" species="paulinae">Rat de Jenkins</vernacularName>
/
<emphasis id="A4ED8F7134DDFF6CE7A9270171B0880E" bold="true" box="[1694,1785,2824,2845]" pageId="251" pageNumber="856">German:</emphasis>
<vernacularName id="189A234D34DDFF6CE635270170E7880E" ID-CoL="79QYQ" authorityName="Musser et al." authorityYear="2005" box="[1794,1966,2824,2845]" class="Mammalia" family="Muridae" genus="Saxatilomys" kingdom="Animalia" language="deu" order="Rodentia" pageId="251" pageNumber="856" phylum="Chordata" rank="species" species="paulinae">Paulina-Karstratte</vernacularName>
/
<emphasis id="A4ED8F7134DDFF6CE6F427017F56880E" bold="true" box="[1987,2079,2824,2845]" pageId="251" pageNumber="856">Spanish:</emphasis>
<vernacularName id="189A234D34DDFF6CE91E27017E54880E" ID-CoL="79QYQ" authorityName="Musser et al." authorityYear="2005" box="[2089,2333,2824,2845]" class="Mammalia" family="Muridae" genus="Saxatilomys" kingdom="Animalia" language="esp" order="Rodentia" pageId="251" pageNumber="856" phylum="Chordata" rank="species" species="paulinae">Rata de caliza de Paulina</vernacularName>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DDFF6CE93227467FEA8887" pageId="251" pageNumber="856" type="reference_group">
<paragraph id="9626536334DDFF6CE93227467FEA8887" blockId="251.[2053,2647,2895,3318]" pageId="251" pageNumber="856">
<emphasis id="A4ED8F7134DDFF6CE93227467FE9887F" bold="true" box="[2053,2208,2895,2924]" pageId="251" pageNumber="856">Taxonomy.</emphasis>
<taxonomicName id="519928E034DDFF6CE98A27467FD78887" ID-CoL="79QYQ" authorityName="Musser et al." authorityYear="2005" class="Mammalia" family="Muridae" genus="Saxatilomys" kingdom="Animalia" order="Rodentia" pageId="251" pageNumber="856" phylum="Chordata" rank="species" species="paulinae">Saxatilomys paulinae Musser et al., 2005</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DDFF6CE985277B7E768FBB" pageId="251" pageNumber="856" type="materials_examined">
<paragraph id="9626536334DDFF6CE985277B7E768FBB" blockId="251.[2053,2647,2895,3318]" pageId="251" pageNumber="856">
<materialsCitation id="26F1593E34DDFF6CE985277B7E768FBB" ID-GBIF-Occurrence="3864424839" pageId="251" pageNumber="856">“Ban (5 village) Mauang and Ban Doy (178339150N/1048499300E), Thakhet District, on the southwest margin of the Khammouan Limestone National Biodiversity Conservation Area (178269-188059N/1048259-1058109E), Lao PDR, approximately 18 km north of the town of Thakhek.”</materialsCitation>
</paragraph>
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<subSubSection id="DE8300E834DDFF6CE93120A47D188E0D" pageId="251" pageNumber="856" type="discussion">
<paragraph id="9626536334DDFF6CE93120A47D188E0D" blockId="251.[2053,2647,2895,3318]" lastBlockId="251.[1440,2644,3324,3476]" pageId="251" pageNumber="856">
<taxonomicName id="519928E034DDFF6CE93120A47FD28FDC" authorityName="Musser, Smith, Robinson &amp; Lunde" authorityYear="2005" box="[2054,2203,3245,3279]" class="Mammalia" family="Muridae" genus="Saxatilomys" kingdom="Animalia" order="Rodentia" pageId="251" pageNumber="856" phylum="Chordata" rank="genus">Saxatilomys</taxonomicName>
is phylogenetically, morphologically and ecologically closest to
<taxonomicName id="519928E034DDFF6CE8C120DD72968E0D" authorityName="Musser, Lunde &amp; Nguyen Truong Son" authorityYear="2006" class="Mammalia" family="Muridae" genus="Tonkinomys" kingdom="Animalia" order="Rodentia" pageId="251" pageNumber="856" phylum="Chordata" rank="genus">Tonkinomys</taxonomicName>
, and they are sister to the rest of the
<taxonomicName id="519928E034DDFF6CE6D120F57F218E0D" authorityName="Thomas" authorityYear="1916" box="[2022,2152,3324,3358]" class="Mammalia" family="Muridae" genus="Dacnomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Rodentia" pageId="251" pageNumber="856" phylum="Chordata" rank="genus">Dacnomys</taxonomicName>
division within
<taxonomicName id="519928E034DDFF6CE87220F57EEC8E0D" box="[2373,2469,3324,3358]" pageId="251" pageNumber="883" rank="tribe" tribe="Rattini">Rattini</taxonomicName>
. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DDFF6CE496212A7F638E78" pageId="251" pageNumber="856" type="distribution">
<caption id="C2E603EB34DDFF6CE496212A7F638E78" ID-DOI="http://doi.org/10.5281/zenodo.6888485" ID-Zenodo-Dep="6888485" httpUri="https://zenodo.org/record/6888485/files/figure.png" inLine="true" pageId="251" pageNumber="856" startId="251.[1441,1613,3363,3397]" targetBox="[1439,2029,2905,3318]" targetPageId="251">
<paragraph id="9626536334DDFF6CE496212A7F638E78" blockId="251.[1440,2644,3324,3476]" pageId="251" pageNumber="856">
<emphasis id="A4ED8F7134DDFF6CE496212A71188E56" bold="true" box="[1441,1617,3363,3397]" pageId="251" pageNumber="856">Distribution.</emphasis>
C Laos (Khammouan Limestone National Biodiversity Conservation Area) and C Vietnam (Quang Binh Province).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DE8300E834DDFF6BE496217B759F8056" lastPageId="252" lastPageNumber="857" pageId="251" pageNumber="856" type="description">
<paragraph id="9626536334DDFF6BE496217B759F8056" blockId="251.[1440,2644,3324,3476]" lastBlockId="252.[103,1311,257,1393]" lastPageId="252" lastPageNumber="857" pageId="251" pageNumber="856">
<emphasis id="A4ED8F7134DDFF6CE496217B71D18E87" bold="true" box="[1441,1688,3442,3476]" pageId="251" pageNumber="856">Descriptive notes.</emphasis>
Head-body 157-164 mm,tail 180-203 mm, ear 24-25 mm, hindfoot 29-30 mm; weight 110-130 g. Paulinas Limestone Rat is medium-sized, with short, dense, and semi-spinous pelage (less spiny on side). Dorsum is dark gray-black, with slightly burnished highlights, being darker gray on head, shoulders,sides, and limbs; dorsal pelage is indistinctly demarcated from ventral pelage. Guard hairs are pale gray at bases and tipped with black or brown. Venteris dark frosted gray and soft. Eyes are relatively small; ears are relatively large, oval, and grayish brown, covered sparsely with small brown hair. Vibrissae are fairly long and dark brown or unpigmented. Tail is slim, tapered, 113-129% of head-body length, dark brown dorsally, and unpigmented ventrally, with hair length increasing near tip. Claws are pale brown on forefeet, unpigmented on hindfeet, moderately curved, and long. Dorsal part of feet is brownish and has six large bulbous pads on bottom. Skull is elongated, with narrow rostrum and wide braincase. Mites Afrolistrophoroides saxatilomys and Listrophoroides oconnori have been recorded on Paulinas Limestone Rat. There are four pairs of mammae: one pectoral, one post-axillary, and two inguinal.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DAFF6BE15E2F43763380AF" pageId="252" pageNumber="857" type="biology_ecology">
<paragraph id="9626536334DAFF6BE15E2F43763380AF" blockId="252.[103,1311,257,1393]" pageId="252" pageNumber="857">
<emphasis id="A4ED8F7134DAFF6BE15E2F437791807F" bold="true" box="[105,216,842,876]" pageId="252" pageNumber="857">Habitat.</emphasis>
Rocky habitats covered in degraded deciduous forest mixed with scrub and bamboo and complex karst towers under limestone humid evergreen forest at elevations of 295-315 m.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DAFF6BE15D2FC9737F80F1" box="[106,1078,960,994]" pageId="252" pageNumber="857" type="food_feeding">
<paragraph id="9626536334DAFF6BE15D2FC9737F80F1" blockId="252.[103,1311,257,1393]" box="[106,1078,960,994]" pageId="252" pageNumber="857">
<emphasis id="A4ED8F7134DAFF6BE15D2FC9762680F1" bold="true" box="[106,367,960,994]" pageId="252" pageNumber="857">Food and Feeding.</emphasis>
Paulinas Limestone Rat is probably insectivorous.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DAFF6BE15D2FE57696871A" box="[106,479,1004,1033]" pageId="252" pageNumber="857" type="breeding">
<paragraph id="9626536334DAFF6BE15D2FE57696871A" blockId="252.[103,1311,257,1393]" box="[106,479,1004,1033]" pageId="252" pageNumber="857">
<emphasis id="A4ED8F7134DAFF6BE15D2FE577B9871A" bold="true" box="[106,240,1004,1033]" pageId="252" pageNumber="857">Breeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DAFF6BE15F280674218722" box="[104,872,1039,1073]" pageId="252" pageNumber="857" type="activity">
<paragraph id="9626536334DAFF6BE15F280674218722" blockId="252.[103,1311,257,1393]" box="[104,872,1039,1073]" pageId="252" pageNumber="857">
<emphasis id="A4ED8F7134DAFF6BE15F2806761A8722" bold="true" box="[104,339,1039,1073]" pageId="252" pageNumber="857">Activity patterns.</emphasis>
Paulinas Limestone Rat is nocturnal.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DAFF6BE15D283F735C874B" box="[106,1045,1078,1112]" pageId="252" pageNumber="857" type="biology_ecology">
<paragraph id="9626536334DAFF6BE15D283F735C874B" blockId="252.[103,1311,257,1393]" box="[106,1045,1078,1112]" pageId="252" pageNumber="857">
<emphasis id="A4ED8F7134DAFF6BE15D283F746F874B" bold="true" box="[106,806,1078,1112]" pageId="252" pageNumber="857">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DAFF6BE15D2854749D860E" pageId="252" pageNumber="857" type="conservation">
<paragraph id="9626536334DAFF6BE15D2854749D860E" blockId="252.[103,1311,257,1393]" pageId="252" pageNumber="857">
<emphasis id="A4ED8F7134DAFF6BE15D28547682876C" bold="true" box="[106,459,1117,1151]" pageId="252" pageNumber="857">Status and Conservation.</emphasis>
Classified as Data Deficient on The IUCN Red List. Hunting in Vietnam is the only
<taxonomicName id="519928E034DAFF6BE0982880754B87B5" authorityName="Hatt" authorityYear="1934" box="[431,514,1161,1190]" class="Mammalia" family="Muridae" genus="Thamnomys" kingdom="Animalia" order="Rodentia" pageId="252" pageNumber="857" phylum="Chordata" rank="species" species="major">major</taxonomicName>
threat to Paulinas Limestone Rat. Limestone karsts are relatively safe from large scale destruction, and it occurs in Khammouan Limestone National Biodiversity Conservation Area in Laos. Additional research is needed to understand its natural history and potential conservation threats.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834DAFF6BE15C29227359867F" pageId="252" pageNumber="857" type="bibRefCitation_list">
<paragraph id="9626536334DAFF6BE15C29227359867F" blockId="252.[103,1311,257,1393]" pageId="252" pageNumber="857">
<emphasis id="A4ED8F7134DAFF6BE15C2922764D8657" bold="true" box="[107,260,1323,1348]" pageId="252" pageNumber="857">Bibliography.</emphasis>
Balakirev, Abramov et al. (2012), Balakirev, Aniskin et al. (2013), Francis (2008), Kennerley &amp; Correia (2016), Musseret al. (2005), Nguyen Nghia Xuan et al. (2015), Pagés et al. (2016).
</paragraph>
</subSubSection>
</treatment>
</document>