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White-footed Indochinese Rat (Rattus nitidus), 674. Osgood’s Vietnamese Rat (Rattus osgoodi), 675. Little Indochinese Field Rat (Rattus sakeratensis), 676. Burnished Enggano Rat (Rattus adustus), 677. Kinabalu Rat (Rattus baluensis), 678. Aceh Rat (Rattus blangorum), 679. Enggano Island Rat (Rattus enganus), 680. Hoogerwert’s Sumatran Rat (Rattus hoogerwerfi), 682. Mentawai Archipelago Rat (Rattus lugens), 683. Gag Island Rat (Rattus nikenu), 684. Simalur Archipelago Rat (Rattus simalurensis), 685. Malaysian Field Rat (Rattus tiomanicus), 686. South-western Xanthurus Rat (Rattus bontanus), 687. Hoffmann’s Sulawesi Rat (Rattus hoffmanni), 688. Koopman' ’ s Peleng Island Rat (Rattus koopmani), 689. Marmoset Xanthurus Rat (Rattus marmosurus), 690. Lampobatang Sulawesi Rat (Rattus mollicomulus)" figureDoi="http://doi.org/10.5281/zenodo.6888389" httpUri="https://zenodo.org/record/6888389/files/figure.png" pageId="231" pageNumber="836">676.</figureCitation>
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<emphasis id="A4ED8F7134C1FF70E20B2870731087B4" box="[828,1113,1145,1191]" italics="true" pageId="231" pageNumber="836">Rattus adustus</emphasis>
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<heading id="CD6EE40F34C1FF70E19928B0731B87DD" box="[174,1106,1209,1230]" pageId="231" pageNumber="836">
<emphasis id="A4ED8F7134C1FF70E19928B077B087DD" bold="true" box="[174,249,1209,1230]" pageId="231" pageNumber="836">French:</emphasis>
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<emphasis id="A4ED8F7134C1FF70E0B128B076A987DD" bold="true" box="[390,480,1209,1230]" pageId="231" pageNumber="836">German:</emphasis>
<vernacularName id="189A234D34C1FF70E0DC28B075A487DD" ID-CoL="4RM3Q" authorityName="Sody" authorityYear="1940" box="[491,749,1209,1230]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" language="deu" order="Rodentia" pageId="231" pageNumber="836" phylum="Chordata" rank="species" species="adustus">Indonesische Glattfellratte</vernacularName>
/ 
<emphasis id="A4ED8F7134C1FF70E23528B0741487DD" bold="true" box="[770,861,1209,1230]" pageId="231" pageNumber="836">Spanish:</emphasis>
<vernacularName id="189A234D34C1FF70E25028B0731B87DD" ID-CoL="4RM3Q" authorityName="Sody" authorityYear="1940" box="[871,1106,1209,1230]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" language="esp" order="Rodentia" pageId="231" pageNumber="836" phylum="Chordata" rank="species" species="adustus">Rata de Enggano pulida</vernacularName>
</heading>
</paragraph>
<paragraph id="9626536334C1FF70E19928E9758487E6" blockId="231.[174,1113,1145,1269]" box="[174,717,1248,1269]" pageId="231" pageNumber="836">
<heading id="CD6EE40F34C1FF70E19928E9758487E6" box="[174,717,1248,1269]" pageId="231" pageNumber="836">
<emphasis id="A4ED8F7134C1FF70E19928E976ED87E6" bold="true" box="[174,420,1248,1269]" pageId="231" pageNumber="836">Other common names:</emphasis>
<vernacularName id="189A234D34C1FF70E09828E9757487E6" ID-CoL="4RM3Q" authorityName="Sody" authorityYear="1940" box="[431,573,1248,1269]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" language="eng" order="Rodentia" pageId="231" pageNumber="836" phylum="Chordata" rank="species" species="adustus">Burnished Rat</vernacularName>
, 
<vernacularName id="189A234D34C1FF70E37D28E9758487E6" ID-CoL="4RM3Q" authorityName="Sody" authorityYear="1940" box="[586,717,1248,1269]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" language="eng" order="Rodentia" pageId="231" pageNumber="836" phylum="Chordata" rank="species" species="adustus">Sunburnt Rat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834C1FF70E224292E72608657" box="[787,1321,1319,1348]" pageId="231" pageNumber="836" type="reference_group">
<paragraph id="9626536334C1FF70E224292E72608657" blockId="231.[787,1379,1319,1742]" box="[787,1321,1319,1348]" pageId="231" pageNumber="836">
<emphasis id="A4ED8F7134C1FF70E224292E74E78657" bold="true" box="[787,942,1319,1348]" pageId="231" pageNumber="836">Taxonomy.</emphasis>
<taxonomicName id="519928E034C1FF70E289292E726D8657" ID-CoL="4RM3Q" authority="Sody, 1940" authorityName="Sody" authorityYear="1940" box="[958,1316,1319,1348]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="231" pageNumber="836" phylum="Chordata" rank="species" species="adustus">Rattus adustus Sody, 1940</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834C1FF70E400292E74B586A9" pageId="231" pageNumber="836" type="materials_examined">
<materialsCitation id="26F1593E34C1FF70E400292E74B586A9" ID-GBIF-Occurrence="3864424335" pageId="231" pageNumber="836">
<paragraph id="9626536334C1FF70E400292E72288678" blockId="231.[787,1379,1319,1742]" pageId="231" pageNumber="836">sea level, Kiojoh, off the coast of W Sumatra</paragraph>
<paragraph id="9626536334C1FF70E220297C74B586A9" blockId="231.[787,1379,1319,1742]" pageId="231" pageNumber="836">(and offthe continental shelf), Pulau Enggano, Indonesia.</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="DE8300E834C1FF70E22229CD724D8544" pageId="231" pageNumber="836" type="discussion">
<paragraph id="9626536334C1FF70E22229CD724D8544" blockId="231.[787,1379,1319,1742]" pageId="231" pageNumber="836">
Phylogenetic position of 
<taxonomicName id="519928E034C1FF70E54829CD720886F2" authorityName="Sody" authorityYear="1940" box="[1151,1345,1476,1505]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="231" pageNumber="836" phylum="Chordata" rank="species" species="adustus">Rattus adustus</taxonomicName>
is uncertain, but itis probably related to 
<taxonomicName id="519928E034C1FF70E42829E574008523" baseAuthorityName="Miller" baseAuthorityYear="1903" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="231" pageNumber="836" phylum="Chordata" rank="species" species="lugens">R. lugens</taxonomicName>
and 
<taxonomicName id="519928E034C1FF70E2AB2A1A73298523" baseAuthorityName="Miller" baseAuthorityYear="1903" box="[924,1120,1555,1584]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="231" pageNumber="836" phylum="Chordata" rank="species" species="simalurensis">R. simalurensis</taxonomicName>
. It is only known from the type specimen. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834C1FF70E2242A6B721A856C" box="[787,1363,1634,1663]" pageId="231" pageNumber="836" type="distribution">
<caption id="C2E603EB34C1FF70E2242A6B721A856C" ID-DOI="http://doi.org/10.5281/zenodo.6888361" ID-Zenodo-Dep="6888361" box="[787,1363,1634,1663]" httpUri="https://zenodo.org/record/6888361/files/figure.png" inLine="true" pageId="231" pageNumber="836" startId="231.[787,959,1634,1663]" targetBox="[173,763,1326,1738]" targetPageId="231">
<paragraph id="9626536334C1FF70E2242A6B721A856C" blockId="231.[787,1379,1319,1742]" box="[787,1363,1634,1663]" pageId="231" pageNumber="836">
<emphasis id="A4ED8F7134C1FF70E2242A6B748A856C" bold="true" box="[787,963,1634,1663]" pageId="231" pageNumber="836">Distribution.</emphasis>
EngganoI, off SW Sumatra.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DE8300E834C1FF70E2242A8075F484A9" pageId="231" pageNumber="836" type="description">
<paragraph id="9626536334C1FF70E2242A8075F484A9" blockId="231.[787,1379,1319,1742]" lastBlockId="231.[172,1378,1752,2411]" pageId="231" pageNumber="836">
<emphasis id="A4ED8F7134C1FF70E2242A80735C85B5" bold="true" box="[787,1045,1673,1702]" pageId="231" pageNumber="836">Descriptive notes.</emphasis>
Head-body 180 mm, tail 148 mm, ear 22 mm, hindfoot 42 mm (from type specimen). Nospecific data are available for body weight. The Burnished Enggano Rat is medium-sized, with short and harsh pelage having many hard spines mixed throughout. Dorsum is very dark brown (hence the common name); venter is dark brownish gray. Ears are dark. Tail is black and 82% of head-body length. Skull is larger than in the EngganoIsland Rat (
<taxonomicName id="519928E034C1FF70E3E82B7C74308481" baseAuthorityName="Miller" baseAuthorityYear="1906" box="[735,889,1909,1938]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="231" pageNumber="836" phylum="Chordata" rank="species" species="enganus">R. enganus</taxonomicName>
). There are five pairs of mammae: two pectoral and three inguinal pairs.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834C1FF70E19A2BC9753184F1" box="[173,632,1984,2018]" pageId="231" pageNumber="836" type="biology_ecology">
<paragraph id="9626536334C1FF70E19A2BC9753184F1" blockId="231.[172,1378,1752,2411]" box="[173,632,1984,2018]" pageId="231" pageNumber="836">
<emphasis id="A4ED8F7134C1FF70E19A2BC9765584F1" bold="true" box="[173,284,1984,2018]" pageId="231" pageNumber="836">Habitat.</emphasis>
Probably forest habitats.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834C1FF70E19A2BEE75EB8B1A" box="[173,674,2023,2057]" pageId="231" pageNumber="836" type="food_feeding">
<paragraph id="9626536334C1FF70E19A2BEE75EB8B1A" blockId="231.[172,1378,1752,2411]" box="[173,674,2023,2057]" pageId="231" pageNumber="836">
<emphasis id="A4ED8F7134C1FF70E19A2BEE76FA8B1A" bold="true" box="[173,435,2023,2057]" pageId="231" pageNumber="836">Food and Feeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834C1FF70E19A241A756A8B23" box="[173,547,2067,2096]" pageId="231" pageNumber="836" type="breeding">
<paragraph id="9626536334C1FF70E19A241A756A8B23" blockId="231.[172,1378,1752,2411]" box="[173,547,2067,2096]" pageId="231" pageNumber="836">
<emphasis id="A4ED8F7134C1FF70E19A241A767D8B23" bold="true" box="[173,308,2067,2096]" pageId="231" pageNumber="836">Breeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834C1FF70E19B243375CF8B44" box="[172,646,2106,2135]" pageId="231" pageNumber="836" type="activity">
<paragraph id="9626536334C1FF70E19B243375CF8B44" blockId="231.[172,1378,1752,2411]" box="[172,646,2106,2135]" pageId="231" pageNumber="836">
<emphasis id="A4ED8F7134C1FF70E19B243376DE8B44" bold="true" box="[172,407,2106,2135]" pageId="231" pageNumber="836">Activity patterns.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834C1FF70E19A246B73118B6C" box="[173,1112,2146,2175]" pageId="231" pageNumber="836" type="biology_ecology">
<paragraph id="9626536334C1FF70E19A246B73118B6C" blockId="231.[172,1378,1752,2411]" box="[173,1112,2146,2175]" pageId="231" pageNumber="836">
<emphasis id="A4ED8F7134C1FF70E19A246B74238B6C" bold="true" box="[173,874,2146,2175]" pageId="231" pageNumber="836">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834C1FF70E199248C72168A50" pageId="231" pageNumber="836" type="conservation">
<paragraph id="9626536334C1FF70E199248C72168A50" blockId="231.[172,1378,1752,2411]" pageId="231" pageNumber="836">
<emphasis id="A4ED8F7134C1FF70E199248C76B38BB4" bold="true" box="[174,506,2181,2215]" pageId="231" pageNumber="836">Status and Conservation.</emphasis>
Classified as Data Deficient on The IUCN Red List. The Burnished Enggano Rat is in critical need for research because it is only known from the type specimen collected in 1920 and virtually nothing is known ofits ecology. It (and the EngganoIsland Rat) could be extinct. It is probably threatened by habitat loss because Enggano’s forests have been almost completely destroyed and competition with the Roof Rat (
<taxonomicName id="519928E034C1FF70E5D6252F721B8A50" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1249,1362,2342,2371]" class="Mammalia" family="Muridae" genus="Epimys" kingdom="Animalia" order="Rodentia" pageId="231" pageNumber="836" phylum="Chordata" rank="species" species="rattus">R. rattus</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834C1FF70E199255B72298A78" box="[174,1376,2386,2411]" pageId="231" pageNumber="836" type="bibRefCitation_list">
<paragraph id="9626536334C1FF70E199255B72298A78" blockId="231.[172,1378,1752,2411]" box="[174,1376,2386,2411]" pageId="231" pageNumber="836">
<emphasis id="A4ED8F7134C1FF70E199255B760E8A78" bold="true" box="[174,327,2386,2411]" pageId="231" pageNumber="836">Bibliography.</emphasis>
Aplin &amp; Lunde (2008b), Musser &amp; Carleton (2005), Musser &amp; Heaney (1985), Sody (1940, 1941).
</paragraph>
</subSubSection>
</treatment>
</document>