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<document id="109B18D8836F46E898D876AA8F8E7FDA" ID-CLB-Dataset="100954" ID-DOI="10.5281/zenodo.6887260" ID-GBIF-Dataset="a016af63-6437-427b-80b7-22bc9a002e20" ID-ISBN="978-84-16728-04-6" ID-Zenodo-Dep="6887260" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="guilherme" checkinTime="1656696812616" checkinUser="carolina" docAuthor="Don E. Wilson, Russell A. Mittermeier &amp; Thomas E. Lacher, Jr" docDate="2017" docId="1E30E27534B4FF05E16A26FF7F308459" docLanguage="en" docName="hbmw_7_Muridae_0536.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 7 Rodents II, Barcelona: Lynx Edicions" docTitle="Otomys unisulcatus F. Cuvier 1829" docType="treatment" docVersion="10" lastPageNumber="747" masterDocId="E2099A0D3426FF97E1372C0977498313" masterDocTitle="Muridae" masterLastPageNumber="884" masterPageNumber="536" pageNumber="747" updateTime="1718983125921" updateUser="carolina">
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<mods:title id="5D0FDBC793F6F7F4942D1DE3B60B62EC">Muridae</mods:title>
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<mods:namePart id="0ECF8328EBC431E3D14D1C2B84B80F09">Don E. Wilson</mods:namePart>
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<mods:namePart id="E881860AB9FA2A0A0ADE0BE54018D4CF">Russell A. Mittermeier</mods:namePart>
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<mods:namePart id="8A4A0344EEF376B71B03E09511525970">Thomas E. Lacher, Jr</mods:namePart>
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<mods:title id="E09875F2428B0E2261C351392CD17435">Handbook of the Mammals of the World Volume 7 Rodents II</mods:title>
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<treatment id="1E30E27534B4FF05E16A26FF7F308459" ID-DOI="http://doi.org/10.5281/zenodo.6788328" ID-GBIF-Taxon="197826076" ID-Zenodo-Dep="6788328" LSID="urn:lsid:plazi:treatment:1E30E27534B4FF05E16A26FF7F308459" httpUri="http://treatment.plazi.org/id/1E30E27534B4FF05E16A26FF7F308459" lastPageNumber="747" pageId="146" pageNumber="747">
<subSubSection id="DE8300E834B4FF05E16A26FF77F98837" box="[93,176,2806,2852]" pageId="146" pageNumber="747" type="multiple">
<paragraph id="9626536334B4FF05E16A26FF77F98837" blockId="146.[92,1045,2806,2930]" box="[93,176,2806,2852]" pageId="146" pageNumber="747">
<heading id="CD6EE40F34B4FF05E16A26FF77F98837" box="[93,176,2806,2852]" pageId="146" pageNumber="747">
<figureCitation id="0EA24FE634B4FF05E16A26FF77F98837" box="[93,176,2806,2852]" captionStart="On" captionStartId="140.[94,124,3255,3280]" captionTargetBox="[12,2724,14,3649]" captionTargetPageId="139" captionText="On following pages: 385. Yalden's Vlei Rat (Otomys yaldeni); 386. Burton's Vlei Rat (Otomys burton); 387. Western Vlei Rat (Otomys occidentalis); 388. Barbours Vlei Rat (Otomys barbouri); 389. Rwenzori Vlei Rat (Otomys dartmouthi): 390. Dent's Vlei Rat (Otomys denti); 391. Dollimans Vlei Rat (Otomys dollmani); 392. Mount Elgon Vlei Rat (Otomys jacksoni); 393. Afroalpine Vlei Rat (Otomys orestes); 394. Thomas's Vlei Rat (Otomys thomas); 395. EastAfrican Vlei Rat (Otomys tropicalis); 396. Tanzanian Vlei Rat (Otomys lacustris); 397. Eastern Arc Vlei Rat (Otomys sungae); 398. Udzungwa Vlei Rat (Otomys uzungwensis); 399. MountKilimanjaro Vlei Rat (Otomys zinki): 400. Angolan Vlei Rat (Otomys anchietae); 401. Cuanza Vlei Rat (Otomys cuanzensis); 402. Angoni Vlei Rat (Otomys angoniensis); 403. South East African Vlei Rat (Otomys auratus); 404. Southern African Vlei Rat (Otomys irroratus): 405. Fynbos Vlei Rat (Otomys karoensis); 406. KwaZulu Natal Vlei Rat (Otomys laminatus); 407. Sloggett's Vlei Rat (Otomys sloggett)); 408. Karoo Vlei Rat (Otomys unisulcatus); 409. Brants's Whistling Rat (Parotomys brantsii); 410. Littledales Whistling Rat (Parotomyslittledale)." figureDoi="http://doi.org/10.5281/zenodo.6887778" httpUri="https://zenodo.org/record/6887778/files/figure.png" pageId="146" pageNumber="747">408.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E1F726FF76B88837" box="[192,497,2806,2852]" pageId="146" pageNumber="747" type="vernacular_names">
<paragraph id="9626536334B4FF05E1F726FF76B88837" blockId="146.[92,1045,2806,2930]" box="[192,497,2806,2852]" pageId="146" pageNumber="747">
<heading id="CD6EE40F34B4FF05E1F726FF76B88837" box="[192,497,2806,2852]" pageId="146" pageNumber="747">
<vernacularName id="189A234D34B4FF05E1F726FF76B88837" ID-CoL="75DKP" authorityName="F. Cuvier" authorityYear="1829" box="[192,497,2806,2852]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="146" pageNumber="747" phylum="Chordata" rank="species" species="unisulcatus">Karoo Vlei Rat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E31026FF74DC8837" box="[551,917,2806,2852]" pageId="146" pageNumber="747" type="nomenclature">
<paragraph id="9626536334B4FF05E31026FF74DC8837" blockId="146.[92,1045,2806,2930]" box="[551,917,2806,2852]" pageId="146" pageNumber="747">
<heading id="CD6EE40F34B4FF05E31026FF74DC8837" box="[551,917,2806,2852]" pageId="146" pageNumber="747">
<taxonomicName id="519928E034B4FF05E31026FF74DC8837" ID-CoL="75DKP" authorityName="F. Cuvier" authorityYear="1829" box="[551,917,2806,2852]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="146" pageNumber="747" phylum="Chordata" rank="species" species="unisulcatus">
<emphasis id="A4ED8F7134B4FF05E31026FF74DC8837" box="[551,917,2806,2852]" italics="true" pageId="146" pageNumber="747">Otomys unisulcatus</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E16A273F75C08861" pageId="146" pageNumber="747" type="vernacular_names">
<paragraph id="9626536334B4FF05E16A273F735C8858" blockId="146.[92,1045,2806,2930]" box="[93,1045,2870,2891]" pageId="146" pageNumber="747">
<heading id="CD6EE40F34B4FF05E16A273F735C8858" box="[93,1045,2870,2891]" pageId="146" pageNumber="747">
<emphasis id="A4ED8F7134B4FF05E16A273F77E18858" bold="true" box="[93,168,2870,2891]" pageId="146" pageNumber="747">French:</emphasis>
<vernacularName id="189A234D34B4FF05E186273F76718858" ID-CoL="75DKP" authorityName="F. Cuvier" authorityYear="1829" box="[177,312,2870,2891]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" language="fra" order="Rodentia" pageId="146" pageNumber="747" phylum="Chordata" rank="species" species="unisulcatus">Otomys cafre</vernacularName>
/
<emphasis id="A4ED8F7134B4FF05E07A273F76EE8858" bold="true" box="[333,423,2870,2891]" pageId="146" pageNumber="747">German:</emphasis>
<vernacularName id="189A234D34B4FF05E086273F75EE8858" ID-CoL="75DKP" authorityName="F. Cuvier" authorityYear="1829" box="[433,679,2870,2891]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" language="deu" order="Rodentia" pageId="146" pageNumber="747" phylum="Chordata" rank="species" species="unisulcatus">Karoo-Lamellenzahnratte</vernacularName>
/
<emphasis id="A4ED8F7134B4FF05E38A273F745E8858" bold="true" box="[701,791,2870,2891]" pageId="146" pageNumber="747">Spanish:</emphasis>
<vernacularName id="189A234D34B4FF05E215273F735C8858" ID-CoL="75DKP" authorityName="F. Cuvier" authorityYear="1829" box="[802,1045,2870,2891]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" language="esp" order="Rodentia" pageId="146" pageNumber="747" phylum="Chordata" rank="species" species="unisulcatus">Rata de laguna de Karoo</vernacularName>
</heading>
</paragraph>
<paragraph id="9626536334B4FF05E16B275475C08861" blockId="146.[92,1045,2806,2930]" box="[92,649,2909,2930]" pageId="146" pageNumber="747">
<heading id="CD6EE40F34B4FF05E16B275475C08861" box="[92,649,2909,2930]" pageId="146" pageNumber="747">
<emphasis id="A4ED8F7134B4FF05E16B2754761A8861" bold="true" box="[92,339,2909,2930]" pageId="146" pageNumber="747">Other common names:</emphasis>
<vernacularName id="189A234D34B4FF05E06A275476AB8861" ID-CoL="75DKP" authorityName="F. Cuvier" authorityYear="1829" box="[349,482,2909,2930]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="146" pageNumber="747" phylum="Chordata" rank="species" species="unisulcatus">Bush Vlei Rat</vernacularName>
,
<vernacularName id="189A234D34B4FF05E0C7275475C08861" ID-CoL="75DKP" authorityName="F. Cuvier" authorityYear="1829" box="[496,649,2909,2930]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="146" pageNumber="747" phylum="Chordata" rank="species" species="unisulcatus">Karoo Bush Rat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E3F627AD744288FB" pageId="146" pageNumber="747" type="reference_group">
<paragraph id="9626536334B4FF05E3F627AD744288FB" blockId="146.[705,1298,2980,3403]" pageId="146" pageNumber="747">
Taxonomy.
<taxonomicName id="519928E034B4FF05E24327AD744F88FB" ID-CoL="75DKP" authority="F. Cuvier" authorityName="F. Cuvier" authorityYear="1829" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="146" pageNumber="747" phylum="Chordata" rank="species" species="unisulcatus">Otomys unisulcatus F. Cuvier, 1829</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E22D27CF73C48F4D" pageId="146" pageNumber="747" type="materials_examined">
<paragraph id="9626536334B4FF05E22D27CF73C48F4D" blockId="146.[705,1298,2980,3403]" pageId="146" pageNumber="747">
<materialsCitation id="26F1593E34B4FF05E22D27CF73C38F4D" ID-GBIF-Occurrence="3864424403" pageId="146" pageNumber="747">type locality not given. Fixed by A. Roberts in 1946 as “Matjiesfontein on the southern edge of the upper Karoo,” Western Cape Province, South Africa</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E3F5206174178E30" pageId="146" pageNumber="747" type="discussion">
<paragraph id="9626536334B4FF05E3F5206174178E30" blockId="146.[705,1298,2980,3403]" pageId="146" pageNumber="747">
Some authors have included O. wunisulcatus and
<taxonomicName id="519928E034B4FF05E26F209974AF8FBE" authorityName="Thomas" authorityYear="1902" box="[856,998,3216,3245]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="146" pageNumber="747" phylum="Chordata" rank="species" species="sloggetti">O. sloggetti</taxonomicName>
in a separate genus Mpyotomys, but phylogenetic position of these primitive species remains elusive. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E3F6212074778E89" pageId="146" pageNumber="747" type="distribution">
<caption id="C2E603EB34B4FF05E3F6212074778E89" ID-DOI="http://doi.org/10.5281/zenodo.6887764" ID-Zenodo-Dep="6887764" httpUri="https://zenodo.org/record/6887764/files/figure.png" inLine="true" pageId="146" pageNumber="747" startId="146.[705,877,3369,3403]" targetBox="[91,681,2990,3402]" targetPageId="146">
<paragraph id="9626536334B4FF05E3F6212074778E89" blockId="146.[705,1298,2980,3403]" lastBlockId="146.[93,1296,3416,3482]" pageId="146" pageNumber="747">Distribution. Occurs throughout the semi-arid Succulent Karoo and Nama Karoo biomes within parts of Northern Cape, Western Cape, and Eastern Cape provinces, South Africa.</paragraph>
</caption>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E4612D397E91811C" pageId="146" pageNumber="747" type="description">
<paragraph id="9626536334B4FF05E4612D397E91811C" blockId="146.[1364,2583,279,1868]" pageId="146" pageNumber="747">Descriptive notes. Head-body 110-205 mm, tail 63-106 mm, ear 20-33 mm, hindfoot 22-32 mm; weight 61-156 g. The Karoo Vlei Ratis large and robust, with large blunt head, short tail, and shaggy fur. Fur is speckled bright tawny brown above and gray below. Tail is short (c.51% of head-body length) and bicolored. Females have four pairs of nipples. Upper incisors have single groove, and lower incisors have two deep grooves. M, has four laminae, and M” has 8-9 laminae. Diploid numberis 2n = 28.</paragraph>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E4612E1D71DC80D3" pageId="146" pageNumber="747" type="biology_ecology">
<paragraph id="9626536334B4FF05E4612E1D71338196" blockId="146.[1364,2583,279,1868]" pageId="146" pageNumber="747">Habitat. Shrublands typically indry river beds and banks and rocky and non-rocky terrain. Near the west coast, Karoo Vlei Rats are associatedwith protected lower lying lees of coastal dunes.</paragraph>
<paragraph id="9626536334B4FF05E4612E8271DC80D3" blockId="146.[1364,2583,279,1868]" pageId="146" pageNumber="747">
Food and Feeding. Diet includes plant stems and leaves, mostly ofshrubs. Species composition of diets varies geographically, with
<taxonomicName id="519928E034B4FF05E93F2EBB7E2D81C7" box="[2056,2404,690,724]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="species" species="karroo">Acacia karroo (Fabaceae)</taxonomicName>
,
<taxonomicName id="519928E034B4FF05E8412EBB711881E9" class="Magnoliopsida" family="Amaranthaceae" genus="Atriplex" kingdom="Plantae" order="Caryophyllales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="species" species="undetermined">Atriplex sp. (Amaranthaceae)</taxonomicName>
,
<taxonomicName id="519928E034B4FF05E7552ED4718881E9" box="[1634,1729,733,762]" class="Magnoliopsida" family="Solanaceae" genus="Lycium" kingdom="Plantae" order="Solanales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="genus">Lycium</taxonomicName>
austrinum (
<taxonomicName id="519928E034B4FF05E6542ED47F4E81E9" box="[1891,2055,733,762]" class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" order="Solanales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="family">Solanaceae</taxonomicName>
),
<taxonomicName id="519928E034B4FF05E9162ED47D4D81E9" box="[2081,2564,733,762]" class="Magnoliopsida" family="Aizoaceae" genus="Mesembryanthemum" kingdom="Plantae" order="Caryophyllales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="species" species="undetermined">Mesembryanthemum sp. (Aizoaceae)</taxonomicName>
, and
<taxonomicName id="519928E034B4FF05E4AA2F08707F8030" box="[1437,1846,769,803]" class="Magnoliopsida" family="Zygophyllaceae" genus="Tribulus" kingdom="Plantae" order="Zygophyllales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="species" species="undetermined">Tribulus sp. (Zygophyllaceae)</taxonomicName>
being important in Eastern Cape;
<taxonomicName id="519928E034B4FF05E8762F087EE98030" box="[2369,2464,769,803]" class="Magnoliopsida" family="Solanaceae" genus="Lycium" kingdom="Plantae" order="Solanales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="genus">Lycium</taxonomicName>
bushes and Galeria
<taxonomicName id="519928E034B4FF05E7322F257139805A" box="[1541,1648,812,841]" class="Magnoliopsida" family="Burseraceae" genus="Commiphora" kingdom="Plantae" order="Sapindales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="species" species="africana">africana</taxonomicName>
,
<taxonomicName id="519928E034B4FF05E7B72F257039805A" box="[1664,1904,812,841]" class="Magnoliopsida" family="Aizoaceae" genus="Drosanthemum" kingdom="Plantae" order="Caryophyllales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="species" species="undetermined">Drosanthemum sp.</taxonomicName>
, and Psilocaulon sp. (all three
<taxonomicName id="519928E034B4FF05E8202F257EEC805A" box="[2327,2469,812,841]" class="Magnoliopsida" family="Aizoaceae" kingdom="Plantae" order="Caryophyllales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="family">Aizoaceae</taxonomicName>
) being important in southern Karoo; and
<taxonomicName id="519928E034B4FF05E6662F467FD68062" box="[1873,2207,847,881]" class="Magnoliopsida" family="Aizoaceae" genus="Ruschia" kingdom="Plantae" order="Caryophyllales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="species" species="undetermined">Ruschia sp. (Aizoaceae)</taxonomicName>
,
<taxonomicName id="519928E034B4FF05E9852F46714B808A" class="Magnoliopsida" family="Amaranthaceae" genus="Exomis" kingdom="Plantae" order="Caryophyllales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="species" species="microphylla">Exomis microphylla (Amaranthaceae)</taxonomicName>
, and
<taxonomicName id="519928E034B4FF05E76F2F7E71B1808A" box="[1624,1784,887,921]" class="Magnoliopsida" family="Zygophyllaceae" genus="Zygophyllum" kingdom="Plantae" order="Zygophyllales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="genus">Zygophyllum</taxonomicName>
flexuosum (
<taxonomicName id="519928E034B4FF05E6AB2F7E7F3F808A" box="[1948,2166,887,921]" class="Magnoliopsida" family="Zygophyllaceae" kingdom="Plantae" order="Zygophyllales" pageId="146" pageNumber="747" phylum="Tracheophyta" rank="family">Zygophyllaceae</taxonomicName>
) being important in coastal areas of Western Cape.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E4612FCF72B287BE" pageId="146" pageNumber="747" type="breeding">
<paragraph id="9626536334B4FF05E4612FCF72B287BE" blockId="146.[1364,2583,279,1868]" pageId="146" pageNumber="747">
<emphasis id="A4ED8F7134B4FF05E4612FCF729480FB" bold="true" box="[1366,1501,966,1000]" pageId="146" pageNumber="747">Breeding.</emphasis>
Karoo Vlei Rats seem to be opportunist breeders, responding to rainfall patterns. Breeding occurs year-round in areas that receive summer and winter rainfall, but it occurs in August-September following winter rains in Namaqualand. Gestation is 37 days, and inter-litter interval is 40 days, suggesting postpartum estrus. Litters in captivity have 1-3 young. Young are semi-precocious and nipple-cling until they are six days old.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E46228BB71E58659" pageId="146" pageNumber="747" type="activity">
<paragraph id="9626536334B4FF05E46228BB71E58659" blockId="146.[1364,2583,279,1868]" pageId="146" pageNumber="747">
<emphasis id="A4ED8F7134B4FF05E46228BB717687C7" bold="true" box="[1365,1599,1202,1236]" pageId="146" pageNumber="747">Activity patterns.</emphasis>
The Karoo Vlei Rat is terrestrial, diurnal, and crepuscular. In Eastern Cape, activity peaks during cooler hours in summer and warmer hours in winter. In Western Cape, this species is crepuscular. Activity periods outside last 5-60 minutes, totaling c.3 periods/day.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E4612946718885C7" pageId="146" pageNumber="747" type="biology_ecology">
<paragraph id="9626536334B4FF05E4612946718885C7" blockId="146.[1364,2583,279,1868]" pageId="146" pageNumber="747">
<emphasis id="A4ED8F7134B4FF05E46129467F458662" bold="true" box="[1366,2060,1359,1393]" pageId="146" pageNumber="747">Movements, Home range and Social organization.</emphasis>
Karoo Vlei Rats construct dome-like “lodges” of interwoven sticks around bases of shrubs, connected with nearby lodges and shrubs by radiating runways. Lodges have systems of tunnels leading to nests, latrines, and entrances. There can be underground tunnels underneath lodges. Groups of 2-8 individuals share lodges. Karoo Vlei Rats forage individually or in groups that move up to 50 m from their lodges. They forage in shrubs, climbing up to 1 m. Aggression between group members is low and ritualized, but between individuals of different lodges, higher levels of aggression can lead to fatalities among males. In the Eastern Cape, density of 64 ind/ha was recorded, but in Western Cape, up to 155 lodges/ha were recorded.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E4602AD07E3B85E8" box="[1367,2418,1753,1787]" pageId="146" pageNumber="747" type="conservation">
<paragraph id="9626536334B4FF05E4602AD07E3B85E8" blockId="146.[1364,2583,279,1868]" box="[1367,2418,1753,1787]" pageId="146" pageNumber="747">
<emphasis id="A4ED8F7134B4FF05E4602AD071FD85E8" bold="true" box="[1367,1716,1753,1787]" pageId="146" pageNumber="747">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red Lust.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E834B4FF05E4602B037F308459" pageId="146" pageNumber="747" type="bibRefCitation_list">
<paragraph id="9626536334B4FF05E4602B037F308459" blockId="146.[1364,2583,279,1868]" pageId="146" pageNumber="747">
<emphasis id="A4ED8F7134B4FF05E4602B0372B98430" bold="true" box="[1367,1520,1802,1827]" pageId="146" pageNumber="747">Bibliography.</emphasis>
Brown (1987), Brown &amp; Willan (1991), Monadjem et al. (2015), Pillay (2001), du Plessis etal. (1991), Roberts (1946), Robinson &amp; Elder (1987), Vermeulen &amp; Nel (1988).
</paragraph>
</subSubSection>
</treatment>
</document>