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<document id="C07195245885D4D4020225B34AF1F035" ID-CLB-Dataset="100954" ID-DOI="10.5281/zenodo.6887260" ID-GBIF-Dataset="a016af63-6437-427b-80b7-22bc9a002e20" ID-ISBN="978-84-16728-04-6" ID-Zenodo-Dep="6887260" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="guilherme" checkinTime="1656696812616" checkinUser="carolina" docAuthor="Don E. Wilson, Russell A. Mittermeier &amp; Thomas E. Lacher, Jr" docDate="2017" docId="1E30E275341CFFADE1802DA4720A8BB6" docLanguage="en" docName="hbmw_7_Muridae_0536.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 7 Rodents II, Barcelona: Lynx Edicions" docTitle="Carpomys phaeurus Thomas 1895" docType="treatment" docVersion="14" lastPageNumber="658" masterDocId="E2099A0D3426FF97E1372C0977498313" masterDocTitle="Muridae" masterLastPageNumber="884" masterPageNumber="536" pageNumber="658" updateTime="1718983125921" updateUser="carolina">
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<mods:title id="00FFAB5A6EA76186AF8EFBFB930B9576">Muridae</mods:title>
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<mods:namePart id="BB94C79E11E60788D6F2D7AADFA2EA01">Don E. Wilson</mods:namePart>
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<mods:namePart id="16FF8F46D190F7D693008457D2E11D8A">Russell A. Mittermeier</mods:namePart>
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<mods:namePart id="CF028FA92341CFB306D6FACFA32788CD">Thomas E. Lacher, Jr</mods:namePart>
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<subSubSection id="DE8300E8341CFFADE1802DA4764F82CC" box="[183,262,429,479]" pageId="58" pageNumber="658" type="multiple">
<paragraph id="96265363341CFFADE1802DA4764F82CC" blockId="58.[178,1352,429,558]" box="[183,262,429,479]" pageId="58" pageNumber="658">
<heading id="CD6EE40F341CFFADE1802DA4764F82CC" box="[183,262,429,479]" pageId="58" pageNumber="658">
<figureCitation id="0EA24FE6341CFFADE1802DA4764F82CC" box="[183,262,429,479]" captionStart="Plate 38: Muridae" captionStartId="53.[89,119,3298,3319]" captionTargetBox="[11,2716,13,3646]" captionTargetPageId="52" captionText="161. Lesser Marmoset Rat (Hapalomys delacouri), 162. Greater Marmoset Rat (Hapalomys longicaudatus), 163. Suntsovs Marmoset Rat (Hapalomys suntsovt), 164. Mount Anacuao Tree Mouse (Musseromys anacuao), 165. Mount Pulag Tree Mouse (Musseromys beneficus), 166. Mount Banahaw Tree Mouse (Musseromys gulantang), 167. Mount Amuyao Tree Mouse (Musseromys inopinatus), 168. Southern Luzon Giant Cloud Rat (Phloecomys cumingi), 169. Northern Luzon Giant Cloud Rat (Phloeomys pallidus), 170. Large-toothed Hairy-tailed Rat (Batomys dentatus), 171. Luzon Cordillera Hairy-tailed Rat (Batomys grantii), 172. Hamiguitan Hairy-tailed Rat (Batomys hamiguitan), 173. Russet Hairy-tailed Rat (Batomys russatus), 174. Mindanao Hairy-tailed Rat (Batomys salomonseni), 175. Mount Isarog Hairy-tailed Rat (Batomys uragon), 176. Black-tailed Dwarf Cloud Rat (Carpomys melanurus), 177. Brown-tailed Dwarf Cloud Rat (Carpomys phaeurus), 178. Dinagat Bushy-tailed Cloud Rat (Crateromys australis), 179. Panay Bushy-tailed Cloud Rat (Crateromys heaneyi), 180. Ilin Bushy-tailed Cloud Rat (Crateromys paulus), 181. Luzon Bushy-tailed Cloud Rat (Crateromys schadenbergu)" figureDoi="http://doi.org/10.5281/zenodo.6887392" httpUri="https://zenodo.org/record/6887392/files/figure.png" pageId="58" pageNumber="658">177.</figureCitation>
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<subSubSection id="DE8300E8341CFFADE0202DA474C582CC" box="[279,908,429,479]" pageId="58" pageNumber="658" type="vernacular_names">
<paragraph id="96265363341CFFADE0202DA474C582CC" blockId="58.[178,1352,429,558]" box="[279,908,429,479]" pageId="58" pageNumber="658">
<heading id="CD6EE40F341CFFADE0202DA474C582CC" box="[279,908,429,479]" pageId="58" pageNumber="658">
<vernacularName id="189A234D341CFFADE0202DA474C582CC" ID-CoL="RGF4" authorityName="Thomas" authorityYear="1895" box="[279,908,429,479]" class="Mammalia" family="Muridae" genus="Carpomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" language="eng" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="species" species="phaeurus">Brown-tailed Dwarf Cloud Rat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E8341CFFADE2E22DA4727982CC" box="[981,1328,429,479]" pageId="58" pageNumber="658" type="nomenclature">
<paragraph id="96265363341CFFADE2E22DA4727982CC" blockId="58.[178,1352,429,558]" box="[981,1328,429,479]" pageId="58" pageNumber="658">
<heading id="CD6EE40F341CFFADE2E22DA4727982CC" box="[981,1328,429,479]" pageId="58" pageNumber="658">
<taxonomicName id="519928E0341CFFADE2E22DA4727982CC" ID-CoL="RGF4" authorityName="Thomas" authorityYear="1895" box="[981,1328,429,479]" class="Mammalia" family="Muridae" genus="Carpomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="species" species="phaeurus">
<emphasis id="A4ED8F71341CFFADE2E22DA4727982CC" box="[981,1328,429,479]" italics="true" pageId="58" pageNumber="658">Carpomys phaeurus</emphasis>
</taxonomicName>
</heading>
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<subSubSection id="DE8300E8341CFFADE1842DFB73E1813D" pageId="58" pageNumber="658" type="vernacular_names">
<paragraph id="96265363341CFFADE1842DFB72018114" blockId="58.[178,1352,429,558]" box="[179,1352,498,519]" pageId="58" pageNumber="658">
<heading id="CD6EE40F341CFFADE1842DFB72018114" box="[179,1352,498,519]" pageId="58" pageNumber="658">
<emphasis id="A4ED8F71341CFFADE1842DFB77B68114" bold="true" box="[179,255,498,519]" pageId="58" pageNumber="658">French:</emphasis>
<vernacularName id="189A234D341CFFADE0302DFB754B8114" ID-CoL="RGF4" authorityName="Thomas" authorityYear="1895" box="[263,514,498,519]" class="Mammalia" family="Muridae" genus="Carpomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" language="fra" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="species" species="phaeurus">Carpomys a queue brune</vernacularName>
/
<emphasis id="A4ED8F71341CFFADE3202DFB75388114" bold="true" box="[535,625,498,519]" pageId="58" pageNumber="658">German:</emphasis>
<vernacularName id="189A234D341CFFADE34B2DFB748C8114" ID-CoL="RGF4" authorityName="Thomas" authorityYear="1895" box="[636,965,498,519]" class="Mammalia" family="Muridae" genus="Carpomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" language="deu" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="species" species="phaeurus">Braunschwanz-Zwergborkenratte</vernacularName>
/
<emphasis id="A4ED8F71341CFFADE2ED2DFB737C8114" bold="true" box="[986,1077,498,519]" pageId="58" pageNumber="658">Spanish:</emphasis>
<vernacularName id="189A234D341CFFADE5082DFB72018114" ID-CoL="RGF4" authorityName="Thomas" authorityYear="1895" box="[1087,1352,498,519]" class="Mammalia" family="Muridae" genus="Carpomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" language="esp" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="species" species="phaeurus">Rata enana de cola marrén</vernacularName>
</heading>
</paragraph>
<paragraph id="96265363341CFFADE1852E1073E1813D" blockId="58.[178,1352,429,558]" box="[178,1192,537,558]" pageId="58" pageNumber="658">
<heading id="CD6EE40F341CFFADE1852E1073E1813D" box="[178,1192,537,558]" pageId="58" pageNumber="658">
<emphasis id="A4ED8F71341CFFADE1852E1076E3813D" bold="true" box="[178,426,537,558]" pageId="58" pageNumber="658">Other common names:</emphasis>
<vernacularName id="189A234D341CFFADE0842E1075D1813D" ID-CoL="RGF4" authorityName="Thomas" authorityYear="1895" box="[435,664,537,558]" class="Mammalia" family="Muridae" genus="Carpomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" language="eng" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="species" species="phaeurus">Small Luzon Carpomys</vernacularName>
,
<vernacularName id="189A234D341CFFADE3932E10748A813D" ID-CoL="RGF4" authorityName="Thomas" authorityYear="1895" box="[676,963,537,558]" class="Mammalia" family="Muridae" genus="Carpomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" language="eng" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="species" species="phaeurus">White-bellied Luzon Tree Rat</vernacularName>
,
<vernacularName id="189A234D341CFFADE2FA2E1073E1813D" ID-CoL="RGF4" authorityName="Thomas" authorityYear="1895" box="[973,1192,537,558]" class="Mammalia" family="Muridae" genus="Carpomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" language="eng" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="species" species="phaeurus">White-bellied Tree Rat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E8341CFFADE2212E6C742981B6" pageId="58" pageNumber="658" type="reference_group">
<paragraph id="96265363341CFFADE2212E6C742981B6" blockId="58.[790,1381,613,1032]" pageId="58" pageNumber="658">
<emphasis id="A4ED8F71341CFFADE2212E6C74F8816D" bold="true" box="[790,945,613,638]" pageId="58" pageNumber="658">Taxonomy.</emphasis>
<taxonomicName id="519928E0341CFFADE2F82E6C741281B6" ID-CoL="RGF4" authority="Thomas, 1895" authorityName="Thomas" authorityYear="1895" class="Mammalia" family="Muridae" genus="Carpomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="species" species="phaeurus">Carpomys phaeurus Thomas, 1895</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="DE8300E8341CFFADE25B2E81722A81B6" box="[876,1379,648,677]" pageId="58" pageNumber="658" type="materials_examined">
<paragraph id="96265363341CFFADE25B2E81722A81B6" blockId="58.[790,1381,613,1032]" box="[876,1379,648,677]" pageId="58" pageNumber="658">
<materialsCitation id="26F1593E341CFFADE25B2E81722A81B6" ID-GBIF-Occurrence="3864424766" box="[876,1379,648,677]" pageId="58" pageNumber="658">Mt. Data, Luzon Island, Philippines.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DE8300E8341CFFADE2202EA274FA8051" pageId="58" pageNumber="658" type="discussion">
<paragraph id="96265363341CFFADE2202EA274FA8051" blockId="58.[790,1381,613,1032]" pageId="58" pageNumber="658">
Different studies place it as sister to a clade including
<taxonomicName id="519928E0341CFFADE2992EDE735F81E7" box="[942,1046,727,756]" class="Mammalia" family="Muridae" genus="Batomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="genus">Batomys</taxonomicName>
and
<taxonomicName id="519928E0341CFFADE55D2EDE73B881E7" box="[1130,1265,727,756]" class="Mammalia" family="Muridae" genus="Crateromys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="genus">Crateromys</taxonomicName>
or to a clade including
<taxonomicName id="519928E0341CFFADE53C2EF7733B8008" box="[1035,1138,766,795]" class="Mammalia" family="Muridae" genus="Batomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="genus">Batomys</taxonomicName>
and
<taxonomicName id="519928E0341CFFADE5FF2EF772168008" authorityName="Heaney, Balete, Rickart, Veluz &amp; Jansa" authorityYear="2009" box="[1224,1375,766,795]" class="Mammalia" family="Muridae" genus="Musseromys" kingdom="Animalia" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="genus">Musseromys</taxonomicName>
. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E8341CFFADE2212F44749B8082" pageId="58" pageNumber="658" type="distribution">
<caption id="C2E603EB341CFFADE2212F44749B8082" ID-DOI="http://doi.org/10.5281/zenodo.6898795" ID-Zenodo-Dep="6898795" httpUri="https://zenodo.org/record/6898795/files/figure.png" inLine="true" pageId="58" pageNumber="658" startId="58.[790,958,845,874]" targetBox="[176,766,617,1030]" targetPageId="58">
<paragraph id="96265363341CFFADE2212F44749B8082" blockId="58.[790,1381,613,1032]" pageId="58" pageNumber="658">
<emphasis id="A4ED8F71341CFFADE2212F44748B8079" bold="true" box="[790,962,845,874]" pageId="58" pageNumber="658">Distribution.</emphasis>
Cordillera Central of N Luzon I, Philippines.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DE8300E8341CFFADE2212F9F727B853C" pageId="58" pageNumber="658" type="description">
<paragraph id="96265363341CFFADE2212F9F727B853C" blockId="58.[790,1381,613,1032]" lastBlockId="58.[174,1383,1045,2213]" pageId="58" pageNumber="658">
<emphasis id="A4ED8F71341CFFADE2212F9F736380AB" bold="true" box="[790,1066,918,952]" pageId="58" pageNumber="658">Descriptive notes.</emphasis>
Head-body 166-175 mm,tail 156-161 mm,ear 19 mm, hindfoot 31-32 mm; weight 123 g. The Browntailed Dwarf Cloud Rat is a small rat with abroad, blunt head and soft, fluffy, and thick fur and being significantly smaller than the Black-tailed Dwarf Cloud Rat (
<taxonomicName id="519928E0341CFFADE59C283D721F8745" authorityName="Thomas" authorityYear="1895" box="[1195,1366,1076,1110]" class="Mammalia" family="Muridae" genus="Carpomys" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Rodentia" pageId="58" pageNumber="658" phylum="Chordata" rank="species" species="melanurus">C. melanurus</taxonomicName>
). Dorsal pelage is dark brown with coarse black lining throughout and is well demarcated from ventral pelage with some shading between them. Head is a lighter brown. Ventral pelage is nearly white, being slightly yellowish with a slate gray base to the hairs. There is a dark ring offur surrounding the eye that gives a masked appearance. Feet are broad with white digits and a silvery brown stripe down dorsal surface of the hindfeet. Ears are relatively short, rounded, and a lighter brown, being less hairy than in the Black-tailed Dwarf Cloud Rat. Tail is long (c.93% of head-body length), hairy, and a dark brown coloration without the dorsal pelage extending to the base of the tail as it does in the Black-tailed Dwarf Cloud Rat—also being less hairy on the tail than its congener. Skull is robust with a rounded braincase and short face, being smaller with smaller molars and incisors when compared to the Brown-tailed Dwarf Cloud Rat. Females have two pairs of mammae, both of which are inguinal, and an elongated urinary papilla that looks like a penis sheath.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E8341CFFADE1872A3D73AB856E" pageId="58" pageNumber="658" type="biology_ecology">
<paragraph id="96265363341CFFADE1872A3D73AB856E" blockId="58.[174,1383,1045,2213]" pageId="58" pageNumber="658">
<emphasis id="A4ED8F71341CFFADE1872A3D76568545" bold="true" box="[176,287,1588,1622]" pageId="58" pageNumber="658">Habitat.</emphasis>
Mossy forest at elevations of 2150-2500 m. One specimen was captured on the trunk of a large moss covered oak tree (
<taxonomicName id="519928E0341CFFADE3E82A69743E856E" box="[735,887,1632,1661]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="58" pageNumber="658" phylum="Tracheophyta" rank="species" species="undetermined">Quercus sp.</taxonomicName>
) ¢.5 m above the ground.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E8341CFFADE1872A8A738C85B6" box="[176,1221,1667,1701]" pageId="58" pageNumber="658" type="food_feeding">
<paragraph id="96265363341CFFADE1872A8A738C85B6" blockId="58.[174,1383,1045,2213]" box="[176,1221,1667,1701]" pageId="58" pageNumber="658">
<emphasis id="A4ED8F71341CFFADE1872A8A76FC85B6" bold="true" box="[176,437,1667,1701]" pageId="58" pageNumber="658">Food and Feeding.</emphasis>
The Brown-tailed Dwarf Cloud Rat probably eats seeds.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E8341CFFADE1872AA7756C85D8" box="[176,549,1710,1739]" pageId="58" pageNumber="658" type="breeding">
<paragraph id="96265363341CFFADE1872AA7756C85D8" blockId="58.[174,1383,1045,2213]" box="[176,549,1710,1739]" pageId="58" pageNumber="658">
<emphasis id="A4ED8F71341CFFADE1872AA7767F85D8" bold="true" box="[176,310,1710,1739]" pageId="58" pageNumber="658">Breeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E8341CFFADE1992AD3741D8409" pageId="58" pageNumber="658" type="activity">
<paragraph id="96265363341CFFADE1992AD3741D8409" blockId="58.[174,1383,1045,2213]" pageId="58" pageNumber="658">
<emphasis id="A4ED8F71341CFFADE1992AD376D085E0" bold="true" box="[174,409,1754,1779]" pageId="58" pageNumber="658">Activity patterns.</emphasis>
Brown-tailed Dwarf Cloud Rats are probably nocturnal and arboreal, although they apparently nest among tree roots.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E8341CFFADE1872B2D73138452" box="[176,1114,1828,1857]" pageId="58" pageNumber="658" type="biology_ecology">
<paragraph id="96265363341CFFADE1872B2D73138452" blockId="58.[174,1383,1045,2213]" box="[176,1114,1828,1857]" pageId="58" pageNumber="658">
<emphasis id="A4ED8F71341CFFADE1872B2D74228452" bold="true" box="[176,875,1828,1857]" pageId="58" pageNumber="658">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E8341CFFADE1872B45738E8B45" pageId="58" pageNumber="658" type="conservation">
<paragraph id="96265363341CFFADE1872B45738E8B45" blockId="58.[174,1383,1045,2213]" pageId="58" pageNumber="658">
<emphasis id="A4ED8F71341CFFADE1872B457542847A" bold="true" box="[176,523,1868,1897]" pageId="58" pageNumber="658">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. Although the Brown-tailed Dwarf Cloud Rat is known from few specimens, probably due to the difficulty of trapping this canopy dependent species,it is seemingly widespread in mossy forest and further sampling using more focused capture techniques is needed. The species may be locally extinct on Mount Data. It is probably threatened by urbanization and agricultural expansion that is destroying its habitat throughout Luzon, although thisis occurring mostly in lowland habitats. It is found in Balbasung National Park.
</paragraph>
</subSubSection>
<subSubSection id="DE8300E8341CFFADE187246C720A8BB6" pageId="58" pageNumber="658" type="bibRefCitation_list">
<paragraph id="96265363341CFFADE187246C720A8BB6" blockId="58.[174,1383,1045,2213]" pageId="58" pageNumber="658">
<emphasis id="A4ED8F71341CFFADE187246C76008B6D" bold="true" box="[176,329,2149,2174]" pageId="58" pageNumber="658">Bibliography.</emphasis>
Balete et al. (2015), Heaney (2016g), Heaney, Balete, Gee et al. (2005), Heaney, Balete &amp; Rickart (2016), Heaney, Balete, Rickart, Utzurrum &amp; Gonzales (1999), Largen (1985), Rabor (1955), Sanborn (1952a).
</paragraph>
</subSubSection>
</treatment>
</document>