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<document id="65427AF5F87BDE6806186D49D26AE6CE" ID-CLB-Dataset="299666" ID-DOI="10.11646/zootaxa.5474.5.1" ID-GBIF-Dataset="f6a45a12-d3b6-44a7-997e-75eb5ebe9901" ID-ISSN="1175-5326" ID-Zenodo-Dep="12712066" ID-ZooBank="3DDAA7F0-27D0-43E8-AAFB-E56F0E24FB3F" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1720373654954" checkinUser="plazi" docAuthor="Pisanty, Gideon &amp; Wood, Thomas James" docDate="2024" docId="077587FDB8546430FF6CDC34E83EF872" docLanguage="en" docName="zootaxa.5474.5.1.pdf" docOrigin="Zootaxa 5474 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5474.5.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Warnckandrena (Vellandrena) Pisanty &amp; Wood 2024, subgen. nov." docType="treatment" docVersion="5" lastPageNumber="478" masterDocId="FB4CFF85B84E642BFFFBDB78E95AFFCB" masterDocTitle="The early-diverging subgenera of the bee genus Andrena (Hymenoptera: Andrenidae) in the Old World" masterLastPageNumber="488" masterPageNumber="451" pageNumber="477" updateTime="1720652268470" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="883F99D01A40326E7D49B0723A44C0B4">The early-diverging subgenera of the bee genus Andrena (Hymenoptera: Andrenidae) in the Old World</mods:title>
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<mods:namePart id="7A4465C96FC5D8698E20EA662356F09F">Pisanty, Gideon</mods:namePart>
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<treatment id="077587FDB8546430FF6CDC34E83EF872" ID-DOI="http://doi.org/10.5281/zenodo.12710366" ID-GBIF-Taxon="235032108" ID-Zenodo-Dep="12710366" LSID="urn:lsid:plazi:treatment:077587FDB8546430FF6CDC34E83EF872" httpUri="http://treatment.plazi.org/id/077587FDB8546430FF6CDC34E83EF872" lastPageId="27" lastPageNumber="478" pageId="26" pageNumber="477">
<subSubSection id="C7C66560B8546431FF6CDC34EBF5F8AD" box="[151,687,1867,1894]" pageId="26" pageNumber="477" type="nomenclature">
<paragraph id="8F6336EBB8546431FF6CDC34EBF5F8AD" blockId="26.[151,687,1867,1894]" box="[151,687,1867,1894]" pageId="26" pageNumber="477">
<heading id="D42B8187B8546431FF6CDC34EBF5F8AD" bold="true" box="[151,687,1867,1894]" fontSize="11" level="1" pageId="26" pageNumber="477" reason="1">
<emphasis id="BDA8EAF9B8546431FF6CDC34EBF5F8AD" bold="true" box="[151,687,1867,1894]" pageId="26" pageNumber="477">
Subgenus
<taxonomicName id="48DC4D68B8546431FEECDC33EB54F8AD" authority="Pisanty" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[279,526,1867,1894]" class="Insecta" family="Andrenidae" genus="Warnckandrena" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Vellandrena">
<emphasis id="BDA8EAF9B8546431FEECDC33E8F0F8AD" bold="true" box="[279,426,1867,1894]" italics="true" pageId="26" pageNumber="477">Vellandrena</emphasis>
Pisanty
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<taxonomicNameLabel id="A69B5782B8546431FDEEDC34EBF5F8AD" box="[533,687,1868,1894]" pageId="26" pageNumber="477" rank="subGenus">subgen. nov.</taxonomicNameLabel>
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<subSubSection id="C7C66560B8546431FF6CDCECEA28F865" box="[151,882,1940,1966]" pageId="26" pageNumber="477" type="type_taxon">
<paragraph id="8F6336EBB8546431FF6CDCECEA28F865" blockId="26.[151,1436,1940,2038]" box="[151,882,1940,1966]" pageId="26" pageNumber="477">
<emphasis id="BDA8EAF9B8546431FF6CDCECE86FF865" bold="true" box="[151,309,1940,1966]" pageId="26" pageNumber="477">
<typeStatus id="50678849B8546431FF6CDCECE988F865" box="[151,210,1940,1966]" pageId="26" pageNumber="477">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="48DC4D68B8546431FEC7DCECEBEAF865" ID-CoL="DR4V" authority="Warncke, 1969" authorityName=", Warncke" authorityYear="1969" box="[316,688,1940,1966]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="bassana">
<emphasis id="BDA8EAF9B8546431FEC7DCECE8A4F865" box="[316,510,1940,1966]" italics="true" pageId="26" pageNumber="477">Andrena bassana</emphasis>
<bibRefCitation id="EB4D4B1AB8546431FDFFDCECEBEAF865" author="Warncke, K." box="[516,688,1940,1966]" pageId="26" pageNumber="477" pagination="377 - 408" refId="ref30210" refString="Warncke, K. (1969) A contribution to the knowledge of the genus Andrena (Apoidea) in Israel. Israel Journal of Entomology, 4, 377 - 408." type="journal article" year="1969">Warncke, 1969</bibRefCitation>
</taxonomicName>
, designated here.
</paragraph>
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<subSubSection id="C7C66560B8546430FF3CDCC0ED14FAC6" lastPageId="27" lastPageNumber="478" pageId="26" pageNumber="477" type="description">
<paragraph id="8F6336EBB8546430FF3CDCC0ED14FAC6" blockId="26.[151,1436,1940,2038]" lastBlockId="27.[151,1437,151,1977]" lastPageId="27" lastPageNumber="478" pageId="26" pageNumber="477">
<emphasis id="BDA8EAF9B8546431FF3CDCC0E803F819" bold="true" box="[199,345,1976,2002]" pageId="26" pageNumber="477">Description.</emphasis>
Small to medium-sized bees (
<quantity id="48249B0EB8546431FD4BDCC0EA6DF819" box="[688,823,1976,2003]" metricMagnitude="-2" metricUnit="m" metricValue="1.025" metricValueMax="1.25" metricValueMin="0.8" pageId="26" pageNumber="477" unit="mm" value="10.25" valueMax="12.5" valueMin="8.0">812.5 mm</quantity>
). Integument dark, male clypeus dark to partly yellow (
<figureCitation id="17E72A6EB8546431FF64DCA4E81FF83D" box="[159,325,2012,2038]" captionStart="FIGURES 100107" captionStartId="29.[151,264,1882,1906]" captionTargetBox="[165,1422,181,1858]" captionTargetId="figure-17@29.[165,1422,181,1858]" captionTargetPageId="29" captionText="FIGURES 100107. Andrena (Vellandrena) bassana Warncke, male (100, 102, 104, 106) and A. (Vellandrena) etesiaca Warncke stat. nov., male paratype (101, 103, 105, 107): 100101, head; 102103, vertex and mesosoma; 104105, metasoma; 106107, habitus." figureDoi="http://doi.org/10.5281/zenodo.12712104" httpUri="https://zenodo.org/record/12712104/files/figure.png" pageId="26" pageNumber="477">Figs. 100101</figureCitation>
). Head 1.2 times broader than long (
<figureCitation id="17E72A6EB8546431FD21DCA4EA6DF83D" box="[730,823,2012,2038]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="26" pageNumber="477">Figs. 94</figureCitation>
,
<figureCitation id="17E72A6EB8546431FCB9DCA4EAFDF83D" box="[834,935,2012,2038]" captionStart="FIGURES 100107" captionStartId="29.[151,264,1882,1906]" captionTargetBox="[165,1422,181,1858]" captionTargetId="figure-17@29.[165,1422,181,1858]" captionTargetPageId="29" captionText="FIGURES 100107. Andrena (Vellandrena) bassana Warncke, male (100, 102, 104, 106) and A. (Vellandrena) etesiaca Warncke stat. nov., male paratype (101, 103, 105, 107): 100101, head; 102103, vertex and mesosoma; 104105, metasoma; 106107, habitus." figureDoi="http://doi.org/10.5281/zenodo.12712104" httpUri="https://zenodo.org/record/12712104/files/figure.png" pageId="26" pageNumber="477">100, 101</figureCitation>
). Mandible bidentate, normally developed in both sexes (
<figureCitation id="17E72A6EB8556430FEE6DBEFE821FF7A" box="[285,379,151,177]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Figs. 94</figureCitation>
,
<figureCitation id="17E72A6EB8556430FE73DBEFE8ECFF7A" box="[392,438,151,177]" captionStart="FIGURES 100107" captionStartId="29.[151,264,1882,1906]" captionTargetBox="[165,1422,181,1858]" captionTargetId="figure-17@29.[165,1422,181,1858]" captionTargetPageId="29" captionText="FIGURES 100107. Andrena (Vellandrena) bassana Warncke, male (100, 102, 104, 106) and A. (Vellandrena) etesiaca Warncke stat. nov., male paratype (101, 103, 105, 107): 100101, head; 102103, vertex and mesosoma; 104105, metasoma; 106107, habitus." figureDoi="http://doi.org/10.5281/zenodo.12712104" httpUri="https://zenodo.org/record/12712104/files/figure.png" pageId="27" pageNumber="478">100</figureCitation>
). Condylar lamella of female mandible weakly developed. Galea shagreened, punctation sparse and very weak, apex more or less rounded. Maxillary palpus extending beyond galea, six-segmented, segment 2 longest, 1.3 times longer than 1. Glossa short, 2.7 times longer than broad, very slightly extending beyond galea. Labial palpus slightly longer than glossa, four-segmented (
<figureCitation id="17E72A6EB8556430FC8FDA7BEA8BFED6" box="[884,977,259,285]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Figs. 94</figureCitation>
,
<figureCitation id="17E72A6EB8556430FC26DA7BED51FED6" box="[989,1035,259,285]" captionStart="FIGURES 100107" captionStartId="29.[151,264,1882,1906]" captionTargetBox="[165,1422,181,1858]" captionTargetId="figure-17@29.[165,1422,181,1858]" captionTargetPageId="29" captionText="FIGURES 100107. Andrena (Vellandrena) bassana Warncke, male (100, 102, 104, 106) and A. (Vellandrena) etesiaca Warncke stat. nov., male paratype (101, 103, 105, 107): 100101, head; 102103, vertex and mesosoma; 104105, metasoma; 106107, habitus." figureDoi="http://doi.org/10.5281/zenodo.12712104" httpUri="https://zenodo.org/record/12712104/files/figure.png" pageId="27" pageNumber="478">100</figureCitation>
). Subgenal coronet present, coronet teeth limited to inner margin of paramandibular process. Labral process triangular in female (
<figureCitation id="17E72A6EB8556430FB25DA5FEC6FFE8A" box="[1246,1333,295,321]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Fig. 94</figureCitation>
), narrow and bidentate in male (
<figureCitation id="17E72A6EB8556430FE5BDA33EB59FEAE" box="[416,515,331,357]" captionStart="FIGURES 108111" captionStartId="30.[151,264,726,750]" captionTargetBox="[165,1421,181,703]" captionTargetId="figure-33@30.[165,1421,181,703]" captionTargetPageId="30" captionText="FIGURES 108111. Andrena (Vellandrena) bassana Warncke, male: 108, genitalia; 109, eighth sternum; 110, labral process; 111, seventh sternum." figureDoi="http://doi.org/10.5281/zenodo.12712108" httpUri="https://zenodo.org/record/12712108/files/figure.png" pageId="27" pageNumber="478">Fig. 110</figureCitation>
). Clypeus, mesonotum, scutellum, metanotum and terga mostly smooth, densely and strongly punctured in female (
<figureCitation id="17E72A6EB8556430FDD8DA17EBF5FE42" box="[547,687,367,393]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Figs. 8994</figureCitation>
), partly shagreened and more weakly, sparsely punctured in male (
<figureCitation id="17E72A6EB8556430FF64DAEBE81CFE66" box="[159,326,402,429]" captionStart="FIGURES 100107" captionStartId="29.[151,264,1882,1906]" captionTargetBox="[165,1422,181,1858]" captionTargetId="figure-17@29.[165,1422,181,1858]" captionTargetPageId="29" captionText="FIGURES 100107. Andrena (Vellandrena) bassana Warncke, male (100, 102, 104, 106) and A. (Vellandrena) etesiaca Warncke stat. nov., male paratype (101, 103, 105, 107): 100101, head; 102103, vertex and mesosoma; 104105, metasoma; 106107, habitus." figureDoi="http://doi.org/10.5281/zenodo.12712104" httpUri="https://zenodo.org/record/12712104/files/figure.png" pageId="27" pageNumber="478">Figs. 100105</figureCitation>
). Clypeus 1.7 times broader than long, in females almost completely flat (
<figureCitation id="17E72A6EB8556430FB84DAEBED88FE66" box="[1151,1234,403,429]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Fig. 94</figureCitation>
), in males weakly protuberant, flattened medially (
<figureCitation id="17E72A6EB8556430FDF1DACFEBE9FE1A" box="[522,691,439,465]" captionStart="FIGURES 100107" captionStartId="29.[151,264,1882,1906]" captionTargetBox="[165,1422,181,1858]" captionTargetId="figure-17@29.[165,1422,181,1858]" captionTargetPageId="29" captionText="FIGURES 100107. Andrena (Vellandrena) bassana Warncke, male (100, 102, 104, 106) and A. (Vellandrena) etesiaca Warncke stat. nov., male paratype (101, 103, 105, 107): 100101, head; 102103, vertex and mesosoma; 104105, metasoma; 106107, habitus." figureDoi="http://doi.org/10.5281/zenodo.12712104" httpUri="https://zenodo.org/record/12712104/files/figure.png" pageId="27" pageNumber="478">Figs. 100101</figureCitation>
). Malar space absent. Inner margins of compound eyes parallel-sided in females (
<figureCitation id="17E72A6EB8556430FE9EDAA3E8E1FE3E" box="[357,443,475,501]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Fig. 94</figureCitation>
), converging below in males (
<figureCitation id="17E72A6EB8556430FCE0DAA3EA9EFE3E" box="[795,964,475,501]" captionStart="FIGURES 100107" captionStartId="29.[151,264,1882,1906]" captionTargetBox="[165,1422,181,1858]" captionTargetId="figure-17@29.[165,1422,181,1858]" captionTargetPageId="29" captionText="FIGURES 100107. Andrena (Vellandrena) bassana Warncke, male (100, 102, 104, 106) and A. (Vellandrena) etesiaca Warncke stat. nov., male paratype (101, 103, 105, 107): 100101, head; 102103, vertex and mesosoma; 104105, metasoma; 106107, habitus." figureDoi="http://doi.org/10.5281/zenodo.12712104" httpUri="https://zenodo.org/record/12712104/files/figure.png" pageId="27" pageNumber="478">Figs. 100101</figureCitation>
). Facial foveae broad throughout, almost rectangular, separated from compound eye by smooth linear area, fovea width equals 0.65 times antennocular distance (
<figureCitation id="17E72A6EB8556430FEFAD95BE8EBFDF6" box="[257,433,547,573]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Figs. 89, 91, 94</figureCitation>
). Distance of fovea from lateral ocellus about 1.1 ocellar diameters. Female flagellomere 1 longer than 2+3, shorter than 2+3+4 (
<figureCitation id="17E72A6EB8556430FDC9D93FEBDEFDAA" box="[562,644,583,609]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Fig. 94</figureCitation>
), male flagellomere 1 as long as 2+3 or slightly shorter (
<figureCitation id="17E72A6EB8556430FB17D93FECCAFDAA" box="[1260,1424,583,609]" captionStart="FIGURES 100107" captionStartId="29.[151,264,1882,1906]" captionTargetBox="[165,1422,181,1858]" captionTargetId="figure-17@29.[165,1422,181,1858]" captionTargetPageId="29" captionText="FIGURES 100107. Andrena (Vellandrena) bassana Warncke, male (100, 102, 104, 106) and A. (Vellandrena) etesiaca Warncke stat. nov., male paratype (101, 103, 105, 107): 100101, head; 102103, vertex and mesosoma; 104105, metasoma; 106107, habitus." figureDoi="http://doi.org/10.5281/zenodo.12712104" httpUri="https://zenodo.org/record/12712104/files/figure.png" pageId="27" pageNumber="478">Figs. 100101</figureCitation>
). Genal area weakly broadened, especially in males, 11.3 times as broad as compound eye. Vertex well-developed, ocelloccipital distance 1.72.4 ocellar diameters (
<figureCitation id="17E72A6EB8556430FD2AD9F7EA04FD62" box="[721,862,655,681]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Figs. 89, 91</figureCitation>
,
<figureCitation id="17E72A6EB8556430FC97D9F7EA89FD62" box="[876,979,655,681]" captionStart="FIGURES 100107" captionStartId="29.[151,264,1882,1906]" captionTargetBox="[165,1422,181,1858]" captionTargetId="figure-17@29.[165,1422,181,1858]" captionTargetPageId="29" captionText="FIGURES 100107. Andrena (Vellandrena) bassana Warncke, male (100, 102, 104, 106) and A. (Vellandrena) etesiaca Warncke stat. nov., male paratype (101, 103, 105, 107): 100101, head; 102103, vertex and mesosoma; 104105, metasoma; 106107, habitus." figureDoi="http://doi.org/10.5281/zenodo.12712104" httpUri="https://zenodo.org/record/12712104/files/figure.png" pageId="27" pageNumber="478">102103</figureCitation>
). Preoccipital ridge moderately carinate dorsally, rounded laterally. Dorsolateral angle of pronotum distinctly elevated, forming a strong dorsolateral carina. Surface of mesepisternum and propodeum (excluding propodeal triangle) with strong, fine alveolation, overlayed by coarse, shallow punctures. Profile of propodeum separated into sloping basal region and vertical apical region, basal region about 1.1 times as long as metanotum. Propodeal triangle not to weakly rugose (
<figureCitation id="17E72A6EB8556430FBACD867EDF1FCF2" box="[1111,1195,799,825]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Fig. 97</figureCitation>
). Propodeal corbicula incomplete, upper half of corbicular surface with extremely dense, short plumose hair, with a texture reminiscent of a carpet (
<figureCitation id="17E72A6EB8556430FEE4D81FE82EFC4A" box="[287,372,871,897]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Fig. 98</figureCitation>
). Inner side of hind femur rounded. Inner hind tibial spur more or less straight and of uniform width. Femoral and tibial scopae composed of simple hairs. Flocculus complete (
<figureCitation id="17E72A6EB8556430FBDAD8F3ED2CFC6E" box="[1057,1142,906,933]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Fig. 95</figureCitation>
). Hind pretarsal claw with strong inner tooth. Forewing with three submarginal cells. Stigma 3.54 times longer than broad, 1.3 times broader than prestigma. Submarginal crossvein 1 enters marginal cell 46 vein widths from stigma. Recurrent vein 1 enters submarginal cell 2 distal to its middle. Recurrent vein 2 enters submarginal cell 3 at 0.6 of its length. Nervulus slightly postfurcal (
<figureCitation id="17E72A6EB8556430FE89DF63E89CFBFE" box="[370,454,1051,1077]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Fig. 99</figureCitation>
). Jugal lobe of hind wing 0.8 times shorter than vannal lobe. Profile of tergum 1 clearly separated into declivous basal region versus horizontal posterior region, basal region with medial longitudinal rim (
<figureCitation id="17E72A6EB8556430FF64DF1BE9A6FBB6" box="[159,252,1123,1149]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Figs. 93</figureCitation>
,
<figureCitation id="17E72A6EB8556430FEF3DF1AE86CFBB6" box="[264,310,1122,1149]" captionStart="FIGURES 100107" captionStartId="29.[151,264,1882,1906]" captionTargetBox="[165,1422,181,1858]" captionTargetId="figure-17@29.[165,1422,181,1858]" captionTargetPageId="29" captionText="FIGURES 100107. Andrena (Vellandrena) bassana Warncke, male (100, 102, 104, 106) and A. (Vellandrena) etesiaca Warncke stat. nov., male paratype (101, 103, 105, 107): 100101, head; 102103, vertex and mesosoma; 104105, metasoma; 106107, habitus." figureDoi="http://doi.org/10.5281/zenodo.12712104" httpUri="https://zenodo.org/record/12712104/files/figure.png" pageId="27" pageNumber="478">105</figureCitation>
). Tergal apical hair bands weakly developed (
<figureCitation id="17E72A6EB8556430FCC8DF1BEAE0FBB6" box="[819,954,1123,1149]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Figs. 90, 93</figureCitation>
,
<figureCitation id="17E72A6EB8556430FC3EDF1AED76FBB6" box="[965,1068,1122,1149]" captionStart="FIGURES 100107" captionStartId="29.[151,264,1882,1906]" captionTargetBox="[165,1422,181,1858]" captionTargetId="figure-17@29.[165,1422,181,1858]" captionTargetPageId="29" captionText="FIGURES 100107. Andrena (Vellandrena) bassana Warncke, male (100, 102, 104, 106) and A. (Vellandrena) etesiaca Warncke stat. nov., male paratype (101, 103, 105, 107): 100101, head; 102103, vertex and mesosoma; 104105, metasoma; 106107, habitus." figureDoi="http://doi.org/10.5281/zenodo.12712104" httpUri="https://zenodo.org/record/12712104/files/figure.png" pageId="27" pageNumber="478">104105</figureCitation>
). Female pygidial plate normally developed, without elevated medial area; male pygidial plate absent. Male sternum 7 normally developed, with two small medioapical lobes that are partly fused medially (
<figureCitation id="17E72A6EB8556430FCF6DFD3EA35FB0E" box="[781,879,1195,1221]" captionStart="FIGURES 108111" captionStartId="30.[151,264,726,750]" captionTargetBox="[165,1421,181,703]" captionTargetId="figure-33@30.[165,1421,181,703]" captionTargetPageId="30" captionText="FIGURES 108111. Andrena (Vellandrena) bassana Warncke, male: 108, genitalia; 109, eighth sternum; 110, labral process; 111, seventh sternum." figureDoi="http://doi.org/10.5281/zenodo.12712108" httpUri="https://zenodo.org/record/12712108/files/figure.png" pageId="27" pageNumber="478">Fig. 111</figureCitation>
). Male sternum 8 normally developed, columnar, apical process broadened, without strong emargination (
<figureCitation id="17E72A6EB8556430FCF2DFB7EA31FB22" box="[777,875,1231,1257]" captionStart="FIGURES 108111" captionStartId="30.[151,264,726,750]" captionTargetBox="[165,1421,181,703]" captionTargetId="figure-33@30.[165,1421,181,703]" captionTargetPageId="30" captionText="FIGURES 108111. Andrena (Vellandrena) bassana Warncke, male: 108, genitalia; 109, eighth sternum; 110, labral process; 111, seventh sternum." figureDoi="http://doi.org/10.5281/zenodo.12712108" httpUri="https://zenodo.org/record/12712108/files/figure.png" pageId="27" pageNumber="478">Fig. 109</figureCitation>
). Male genitalia with dorsal gonocoxite lobe well-developed, gonostylus blade strongly broadened, spatulate. Volsella small (
<figureCitation id="17E72A6EB8556430FC24DF8BED18FAC6" box="[991,1090,1267,1293]" captionStart="FIGURES 108111" captionStartId="30.[151,264,726,750]" captionTargetBox="[165,1421,181,703]" captionTargetId="figure-33@30.[165,1421,181,703]" captionTargetPageId="30" captionText="FIGURES 108111. Andrena (Vellandrena) bassana Warncke, male: 108, genitalia; 109, eighth sternum; 110, labral process; 111, seventh sternum." figureDoi="http://doi.org/10.5281/zenodo.12712108" httpUri="https://zenodo.org/record/12712108/files/figure.png" pageId="27" pageNumber="478">Fig. 108</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B8556430FF3CDE6EE8AAF9BE" pageId="27" pageNumber="478" type="diagnosis">
<paragraph id="8F6336EBB8556430FF3CDE6EE887FA2E" blockId="27.[151,1437,151,1977]" pageId="27" pageNumber="478">
<emphasis id="BDA8EAF9B8556430FF3CDE6EE818FAFB" bold="true" box="[199,322,1302,1328]" pageId="27" pageNumber="478">Diagnosis.</emphasis>
The most clearly defining character of
<taxonomicName id="48DC4D68B8556430FD0CDE6FEA27FAFA" box="[759,893,1303,1329]" pageId="27" pageNumber="477" rank="subGenus" subGenus="Vellandrena">
<emphasis id="BDA8EAF9B8556430FD0CDE6FEA27FAFA" box="[759,893,1303,1329]" italics="true" pageId="27" pageNumber="478">Vellandrena</emphasis>
</taxonomicName>
is the unique structure of the females propodeal corbicula, present nowhere else in
<taxonomicName id="48DC4D68B8556430FDD4DE43EBD4FA9E" authorityName="Confalonierii Guiglia" authorityYear="1929" box="[559,654,1339,1365]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="genus">
<emphasis id="BDA8EAF9B8556430FDD4DE43EBD4FA9E" box="[559,654,1339,1365]" italics="true" pageId="27" pageNumber="478">Andrena</emphasis>
</taxonomicName>
, in which the dorsal half of the corbicular surface is covered with an extremely dense and thick, carpet-like layer of short plumose hair (
<figureCitation id="17E72A6EB8556430FC3CDE27ED44FAB2" box="[967,1054,1375,1401]" captionStart="FIGURES 8999" captionStartId="28.[151,264,1834,1858]" captionTargetBox="[165,1421,181,1810]" captionTargetId="figure-33@28.[165,1421,181,1810]" captionTargetPageId="28" captionText="FIGURES 8999. Andrena (Vellandrena) bassana Warncke, female (8990, 9499) and A. (Vellandrena) etesiaca Warncke stat. nov., female holotype (9193): 89, vertex and mesosoma; 90, 93, metasoma; 91, vertex; 92, mesosoma; 94, head; 95, flocculus and scopa; 96, habitus; 97, propodeal triangle; 98, propodeal corbicula; 99, forewing venation. Figures 9193 courtesy of OLML, downloaded from https://www.europeana.eu/item/11603/ZOBODAT_LANDOOE_ AUSTRIA_6952193." figureDoi="http://doi.org/10.5281/zenodo.12712100" httpUri="https://zenodo.org/record/12712100/files/figure.png" pageId="27" pageNumber="478">Fig. 98</figureCitation>
). Additional important characters include: both sexes—carinate pronotum; female—smooth and coarsely punctured clypeus, mesonotum and terga, flat clypeus, triangular labral process; male—strongly bidentate labral process, long flagellomeres, broad vertex,
<taxonomicName id="48DC4D68B8556430FF6CDEB3E86FFA2E" box="[151,309,1483,1509]" pageId="27" pageNumber="478" rank="subGenus" subGenus="Truncandrena">
<emphasis id="BDA8EAF9B8556430FF6CDEB3E86FFA2E" box="[151,309,1483,1509]" italics="true" pageId="27" pageNumber="478">Truncandrena</emphasis>
</taxonomicName>
-
<typeStatus id="50678849B8556430FEC7DEB3E834FA2E" box="[316,366,1483,1509]" pageId="27" pageNumber="478">type</typeStatus>
genitalia.
</paragraph>
<paragraph id="8F6336EBB8556430FF3CDE97E8AAF9BE" blockId="27.[151,1437,151,1977]" pageId="27" pageNumber="478">
Species of
<taxonomicName id="48DC4D68B8556430FEB8DE97E893F9C2" box="[323,457,1519,1545]" pageId="27" pageNumber="477" rank="subGenus" subGenus="Vellandrena">
<emphasis id="BDA8EAF9B8556430FEB8DE97E893F9C2" box="[323,457,1519,1545]" italics="true" pageId="27" pageNumber="478">Vellandrena</emphasis>
</taxonomicName>
were previously associated with the subgenus
<taxonomicName id="48DC4D68B8556430FC20DE97ED23F9C2" box="[987,1145,1519,1545]" pageId="27" pageNumber="478" rank="subGenus" subGenus="Truncandrena">
<emphasis id="BDA8EAF9B8556430FC20DE97ED23F9C2" box="[987,1145,1519,1545]" italics="true" pageId="27" pageNumber="478">Truncandrena</emphasis>
</taxonomicName>
. However, aside from the similar male genital capsule, there is little similarity to typical members of the subgenus, especially in the females. Males are also readily differentiated from most
<taxonomicName id="48DC4D68B8556430FD54DD4FEA17F99A" box="[687,845,1591,1617]" pageId="27" pageNumber="478" rank="subGenus" subGenus="Truncandrena">
<emphasis id="BDA8EAF9B8556430FD54DD4FEA17F99A" box="[687,845,1591,1617]" italics="true" pageId="27" pageNumber="478">Truncandrena</emphasis>
</taxonomicName>
by the unusual labral process, broad vertex, carinate pronotum, and smoother terga.
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B8556430FF3CDD06E83EF976" pageId="27" pageNumber="478" type="biology_ecology">
<paragraph id="8F6336EBB8556430FF3CDD06E83EF976" blockId="27.[151,1437,151,1977]" pageId="27" pageNumber="478">
<emphasis id="BDA8EAF9B8556430FF3CDD06E87DF953" bold="true" box="[199,295,1662,1688]" pageId="27" pageNumber="478">Biology.</emphasis>
<taxonomicName id="48DC4D68B8556430FED4DD07E8EFF952" box="[303,437,1663,1689]" pageId="27" pageNumber="477" rank="subGenus" subGenus="Vellandrena">
<emphasis id="BDA8EAF9B8556430FED4DD07E8EFF952" box="[303,437,1663,1689]" italics="true" pageId="27" pageNumber="478">Vellandrena</emphasis>
</taxonomicName>
are active in springtime on a wide range of flowers. The two included species are likely pollen generalists.
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B8556430FF3CDDBEEA07F92A" box="[199,861,1734,1761]" pageId="27" pageNumber="478" type="distribution">
<paragraph id="8F6336EBB8556430FF3CDDBEEA07F92A" blockId="27.[151,1437,151,1977]" box="[199,861,1734,1761]" pageId="27" pageNumber="478">
<emphasis id="BDA8EAF9B8556430FF3CDDBEE839F92B" bold="true" box="[199,355,1734,1760]" pageId="27" pageNumber="478">Distribution:</emphasis>
Eastern Mediterranean (Levant and
<collectingCountry id="F7CB767BB8556430FD07DDBFEA0BF92A" box="[764,849,1735,1761]" name="Turkey" pageId="27" pageNumber="478">Turkey</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B8556430FF3CDD93E83EF872" pageId="27" pageNumber="478" type="etymology">
<paragraph id="8F6336EBB8556430FF3CDD93EA21F886" blockId="27.[151,1437,151,1977]" pageId="27" pageNumber="478">
<emphasis id="BDA8EAF9B8556430FF3CDD93E814F8CE" bold="true" box="[199,334,1771,1797]" pageId="27" pageNumber="478">Etymology.</emphasis>
The prefix
<emphasis id="BDA8EAF9B8556430FE21DD93EB5EF8CE" box="[474,516,1771,1797]" italics="true" pageId="27" pageNumber="478">Vell</emphasis>
- is from the Latin
<emphasis id="BDA8EAF9B8556430FD1CDD93EA72F8CE" box="[743,808,1771,1797]" italics="true" pageId="27" pageNumber="478">vellus</emphasis>
, which means fleece or wool, in combination with
<taxonomicName id="48DC4D68B8556430FF6CDC77E9ACF8E2" authorityName="Confalonierii Guiglia" authorityYear="1929" box="[151,246,1807,1833]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="genus">
<emphasis id="BDA8EAF9B8556430FF6CDC77E9ACF8E2" box="[151,246,1807,1833]" italics="true" pageId="27" pageNumber="478">Andrena</emphasis>
</taxonomicName>
, the name of the higher taxon. The name refers to the unique dense pubescence on the dorsal half of the propodeal corbicular surface. The gender of the name is feminine.
</paragraph>
<paragraph id="8F6336EBB8556430FF3CDC2FE83EF872" blockId="27.[151,1437,151,1977]" pageId="27" pageNumber="478">
<emphasis id="BDA8EAF9B8556430FF3CDC2FE8C8F8BB" bold="true" box="[199,402,1878,1905]" pageId="27" pageNumber="478">Included species:</emphasis>
<taxonomicName id="48DC4D68B8556430FE63DC2FEA5FF8BA" authority="Warncke, 1969" authorityName=", Warncke" authorityYear="1969" box="[408,773,1879,1905]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="bassana">
<emphasis id="BDA8EAF9B8556430FE63DC2FEB03F8BA" box="[408,601,1879,1905]" italics="true" pageId="27" pageNumber="478">Andrena bassana</emphasis>
<bibRefCitation id="EB4D4B1AB8556430FDA5DC2FEA5FF8BA" author="Warncke, K." box="[606,773,1879,1905]" pageId="27" pageNumber="478" pagination="377 - 408" refId="ref30210" refString="Warncke, K. (1969) A contribution to the knowledge of the genus Andrena (Apoidea) in Israel. Israel Journal of Entomology, 4, 377 - 408." type="journal article" year="1969">Warncke, 1969</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F7CB767BB8556430FCE8DC2FEA0FF8BA" box="[787,853,1879,1905]" name="Israel" pageId="27" pageNumber="478">Israel</collectingCountry>
,
<collectingCountry id="F7CB767BB8556430FCA4DC2FEAF7F8BA" box="[863,941,1879,1905]" name="Jordan" pageId="27" pageNumber="478">Jordan</collectingCountry>
,
<collectingCountry id="F7CB767BB8556430FC4DDC2FEAAFF8BA" box="[950,1013,1879,1905]" name="Syria" pageId="27" pageNumber="478">Syria</collectingCountry>
),
<taxonomicName id="48DC4D68B8556430FBFCDC2FEC77F8BA" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" baseAuthorityName="Warncke" baseAuthorityYear="1975" box="[1031,1325,1878,1905]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="etesiaca" status="stat. nov.">
<emphasis id="BDA8EAF9B8556430FBFCDC2FEDDBF8BB" box="[1031,1153,1879,1905]" italics="true" pageId="27" pageNumber="478">A. etesiaca</emphasis>
<bibRefCitation id="EB4D4B1AB8556430FB7DDC2FEC77F8BA" author="Warncke, K." box="[1158,1325,1878,1905]" pageId="27" pageNumber="478" pagination="29 - 102" refId="ref30277" refString="Warncke, K. (1975) Die Sandbienen der Turkei (Hymenoptera, Apoidea, Andrena), Teil B. Beschreibungen der neuen Andrena Arten bzw. Unterarten. Mitteilungen der Munchner Entomologischen Gesellschaft, 65, 29 - 102." type="journal article" year="1975">Warncke, 1975</bibRefCitation>
</taxonomicName>
<emphasis id="BDA8EAF9B8556430FAC8DC2FECC6F8BA" bold="true" box="[1331,1436,1879,1905]" pageId="27" pageNumber="478">
<taxonomicNameLabel id="A69B5782B8556430FAC8DC2FECC6F8BA" box="[1331,1436,1879,1905]" pageId="27" pageNumber="478">stat. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F7CB767BB8556430FF64DC03E9ACF85E" box="[159,246,1915,1941]" name="Turkey" pageId="27" pageNumber="478">Turkey</collectingCountry>
). Species belonging to
<taxonomicName id="48DC4D68B8556430FE0FDC03EB20F85E" box="[500,634,1915,1941]" pageId="27" pageNumber="477" rank="subGenus" subGenus="Vellandrena">
<emphasis id="BDA8EAF9B8556430FE0FDC03EB20F85E" box="[500,634,1915,1941]" italics="true" pageId="27" pageNumber="478">Vellandrena</emphasis>
</taxonomicName>
were originally assigned to
<taxonomicName id="48DC4D68B8556430FC49DC03ED0AF85E" box="[946,1104,1915,1941]" pageId="27" pageNumber="478" rank="subGenus" subGenus="Truncandrena">
<emphasis id="BDA8EAF9B8556430FC49DC03ED0AF85E" box="[946,1104,1915,1941]" italics="true" pageId="27" pageNumber="478">Truncandrena</emphasis>
</taxonomicName>
(
<bibRefCitation id="EB4D4B1AB8556430FBA7DC03EC58F85E" author="Warncke, K." box="[1116,1282,1915,1941]" pageId="27" pageNumber="478" pagination="377 - 408" refId="ref30210" refString="Warncke, K. (1969) A contribution to the knowledge of the genus Andrena (Apoidea) in Israel. Israel Journal of Entomology, 4, 377 - 408." type="journal article" year="1969">Warncke 1969</bibRefCitation>
;
<bibRefCitation id="EB4D4B1AB8556430FAF6DC03E802F872" author="Gusenleitner, F. &amp; Schwarz, M." pageId="27" pageNumber="478" pagination="1 - 1280" refId="ref28812" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena: mit Bemerkungen und Erganzungen ¨ zu palaarktischen ¨ Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>