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<mods:title id="6C301BEFC0FDAB71FA8E4B06D9EA1ABF">A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species</mods:title>
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<subSubSection id="26966522FFB6FFD5FF7E42E190BB62E6" box="[151,735,1754,1781]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="6E3336A9FFB6FFD5FF7E42E190BB62E6" blockId="40.[151,735,1754,1781]" box="[151,735,1754,1781]" pageId="40" pageNumber="41">
<heading id="357B81C5FFB6FFD5FF7E42E190BB62E6" bold="true" box="[151,735,1754,1781]" fontSize="11" level="2" pageId="40" pageNumber="41" reason="4">
<emphasis id="5CF8EABBFFB6FFD5FF7E42E190BB62E6" bold="true" box="[151,735,1754,1781]" pageId="40" pageNumber="41">
<taxonomicName id="A98C4D2AFFB6FFD5FF7E42E1900962E7" ID-CoL="3NYWL" authority="(Holmgren, 1860)" authorityName="Galsworthy &amp; Shaw &amp; Haraldseide" authorityYear="2023" baseAuthorityName="Holmgren" baseAuthorityYear="1860" box="[151,621,1754,1781]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="dubitatus" status="stat. nov.">
<emphasis id="5CF8EABBFFB6FFD5FF7E42E193EE62E6" bold="true" box="[151,394,1754,1781]" italics="true" pageId="40" pageNumber="41">Hyposoter dubitatus</emphasis>
(
<bibRefCitation id="0A1D4B58FFB6FFD5FE7342E0900162E7" author="Holmgren, A. E." box="[410,613,1754,1780]" pageId="40" pageNumber="41" pagination="1 - 158" refId="ref53302" refString="Holmgren, A. E. (1860) F ˆ rs ˆ k till uppstallning och beskrifning af de i Sverige funna Ophionider (Monographia Ophionidum Sueciae). Kongliga Svenska Vetenskapsakademiens Handlingar, 2 (8), 1 - 158." type="journal article" year="1860">Holmgren, 1860</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="47CB57C0FFB6FFD5FD9C42E190BB62E6" box="[629,735,1755,1781]" pageId="40" pageNumber="41">stat. rev.</taxonomicNameLabel>
</emphasis>
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<paragraph id="6E3336A9FFB6FFD5FF7E431A96176324" blockId="40.[151,1139,1823,1847]" box="[151,1139,1823,1847]" pageId="40" pageNumber="41">
<taxonomicName id="A98C4D2AFFB6FFD5FF7E431A939B6324" authority="Holmgren, 1860" authorityName="Holmgren" authorityYear="1860" box="[151,511,1823,1847]" class="Gastropoda" family="Velutinidae" genus="Limneria" kingdom="Animalia" order="Littorinimorpha" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="dubitata">
<emphasis id="5CF8EABBFFB6FFD5FF7E431A932A6324" box="[151,334,1823,1847]" italics="true" pageId="40" pageNumber="41">Limneria dubitata</emphasis>
<materialsCitation id="DEE43CF4FFB6FFD5FEBC4325939B6324" ID-GBIF-Occurrence="4116948339" box="[341,511,1823,1847]" collectionCode="NHRS" pageId="40" pageNumber="41" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<bibRefCitation id="0A1D4B58FFB6FFD5FEBC4325939B6324" author="Holmgren, A. E." box="[341,511,1823,1847]" pageId="40" pageNumber="41" pagination="1 - 158" refId="ref53302" refString="Holmgren, A. E. (1860) F ˆ rs ˆ k till uppstallning och beskrifning af de i Sverige funna Ophionider (Monographia Ophionidum Sueciae). Kongliga Svenska Vetenskapsakademiens Handlingar, 2 (8), 1 - 158." type="journal article" year="1860">Holmgren, 1860</bibRefCitation>
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</taxonomicName>
:
<typeStatus id="B137880BFFB6FFD5FDE0432590026324" box="[521,614,1823,1847]" pageId="40" pageNumber="41" type="lectotype">lectotype</typeStatus>
<specimenCount id="788AFD20FFB6FFD5FD85431A901B6324" box="[620,639,1824,1847]" pageId="40" pageNumber="41" type="female"></specimenCount>
in NHRS, designated by Hinz (1962), examined.
</paragraph>
<paragraph id="6E3336A9FFB6FFD4FF7E435C90DE6576" blockId="40.[151,1436,1893,2028]" lastBlockId="41.[151,1437,151,573]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<taxonomicName id="A98C4D2AFFB6FFD5FF7E435C9384636C" authority="Holmgren" authorityName="Holmgren" authorityYear="1860" box="[151,480,1893,1919]" class="Gastropoda" family="Velutinidae" genus="Limneria" kingdom="Animalia" order="Littorinimorpha" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="dubitata">
<emphasis id="5CF8EABBFFB6FFD5FF7E435C9300636C" box="[151,356,1893,1919]" italics="true" pageId="40" pageNumber="41">Limneria dubitata</emphasis>
Holmgren
</taxonomicName>
was synonymized with
<taxonomicName id="A98C4D2AFFB6FFD5FD1E435F96D1636C" authority="Gravenhorst, 1829" authorityName="Gravenhorst" authorityYear="1829" box="[759,1205,1893,1919]" class="Insecta" family="Ichneumonidae" genus="Campoplex" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="sicarius">
<emphasis id="5CF8EABBFFB6FFD5FD1E435F91BD636C" box="[759,985,1893,1919]" italics="true" pageId="40" pageNumber="41">Campoplex sicarius</emphasis>
<bibRefCitation id="0A1D4B58FFB6FFD5FC0B435F96D1636C" author="Gravenhorst, J. L. C." box="[994,1205,1893,1919]" pageId="40" pageNumber="41" refId="ref53031" refString="Gravenhorst, J. L. C. (1829) Ichneumonologia Europaea. Pars III. Sumtibus auctoris, Vratislaviae, 1097 pp. https: // doi. org / 10.5962 / bhl. title. 65750" type="book" year="1829">Gravenhorst, 1829</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="0A1D4B58FFB6FFD5FB0A435F928063B0" author="Schmiedeknecht, O." pageId="40" pageNumber="41" refId="ref55070" refString="Schmiedeknecht, O. (1909) Opuscula Ichneumonologica. IV. Band. Fasc. XXI - XXIII. Ophioninae. s. n., Blankenburg in Th ¸ ringen. [unknown pagination]" type="book" year="1909">Schmiedeknecht (1909)</bibRefCitation>
, though without explanation.
<bibRefCitation id="0A1D4B58FFB6FFD5FDDE43B390BF63B0" author="Aubert, J. - F." box="[567,731,1929,1955]" pageId="40" pageNumber="41" pagination="145 - 152" refId="ref52179" refString="Aubert, J. - F. (1972) Etude commentee de nouveaux lectotypes choisis dans les collections HOLMGREN et THOMSON (Hym. Ichneumonidae). Entomologica Scandinavica, 3, 145 - 152. https: // doi. org / 10.1163 / 187631272 X 00229" type="journal article" year="1972">Aubert (1972)</bibRefCitation>
, after examining the
<typeStatus id="B137880BFFB6FFD5FC2643B3965D63B0" box="[975,1081,1929,1955]" pageId="40" pageNumber="41" type="lectotype">lectotype</typeStatus>
, stated that the synonymy with
<taxonomicName id="A98C4D2AFFB6FFD5FF7E4394929463D4" authorityName="Gravenhorst" authorityYear="1829" box="[151,240,1966,1991]" class="Insecta" family="Ichneumonidae" genus="Campoplex" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="sicarius">
<emphasis id="5CF8EABBFFB6FFD5FF7E4394929463D4" box="[151,240,1966,1991]" italics="true" pageId="40" pageNumber="41">sicarius</emphasis>
</taxonomicName>
was doubtful. Horstmanns draft key makes it clear that he had intended to separate
<taxonomicName id="A98C4D2AFFB6FFD5FB594397977C63D4" ID-CoL="3NYWL" authorityName="Galsworthy &amp; Shaw &amp; Haraldseide" authorityYear="2023" baseAuthorityName="Holmgren" baseAuthorityYear="1860" box="[1200,1304,1965,1991]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="dubitatus">
<emphasis id="5CF8EABBFFB6FFD5FB594397977C63D4" box="[1200,1304,1965,1991]" italics="true" pageId="40" pageNumber="41">dubitatus</emphasis>
</taxonomicName>
from
<taxonomicName id="A98C4D2AFFB6FFD5FAB2439492DC63F8" authorityName="Gravenhorst" authorityYear="1829" class="Insecta" family="Ichneumonidae" genus="Campoplex" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="sicarius">
<emphasis id="5CF8EABBFFB6FFD5FAB2439492DC63F8" italics="true" pageId="40" pageNumber="41">sicarius</emphasis>
</taxonomicName>
, but we have not found evidence in his notes that he had seen the
<typeStatus id="B137880BFFB6FFD5FC7D43EB919F63F8" box="[916,1019,2001,2027]" pageId="40" pageNumber="41" type="lectotype">lectotype</typeStatus>
of
<emphasis id="5CF8EABBFFB6FFD5FBF643EB96EB63F8" box="[1055,1167,2001,2027]" italics="true" pageId="40" pageNumber="41">
<taxonomicName id="A98C4D2AFFB6FFD5FBF643EB96EF63F8" ID-CoL="3NYWL" authorityName="Galsworthy &amp; Shaw &amp; Haraldseide" authorityYear="2023" baseAuthorityName="Holmgren" baseAuthorityYear="1860" box="[1055,1163,2001,2027]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="dubitatus">dubitatus</taxonomicName>
.
</emphasis>
However, we have seen the
<typeStatus id="B137880BFFB7FFD4FF5644AD934D64A2" box="[191,297,151,177]" pageId="41" pageNumber="42" type="lectotype">lectotype</typeStatus>
, and confirm that it is quite different from the specimen of
<taxonomicName id="A98C4D2AFFB7FFD4FC5D44AD966964A3" authorityName="Gravenhorst" authorityYear="1829" box="[948,1037,151,176]" class="Insecta" family="Ichneumonidae" genus="Campoplex" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sicarius">
<emphasis id="5CF8EABBFFB7FFD4FC5D44AD966964A3" box="[948,1037,151,176]" italics="true" pageId="41" pageNumber="42">sicarius</emphasis>
</taxonomicName>
in Horstmanns collection which he had compared with the type of the latter. We therefore formalise here Horstmanns intention to restore
<taxonomicName id="A98C4D2AFFB7FFD4FAFE4481971B64C6" ID-CoL="3NYWL" authorityName="Galsworthy &amp; Shaw &amp; Haraldseide" authorityYear="2023" baseAuthorityName="Holmgren" baseAuthorityYear="1860" box="[1303,1407,187,213]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="dubitatus">
<emphasis id="5CF8EABBFFB7FFD4FAFE4481971B64C6" box="[1303,1407,187,213]" italics="true" pageId="41" pageNumber="42">dubitatus</emphasis>
</taxonomicName>
to valid species status
<emphasis id="5CF8EABBFFB7FFD4FE9A44E593B364EA" bold="true" box="[371,471,223,249]" pageId="41" pageNumber="42">stat. rev.</emphasis>
Three specimens (
<specimenCount id="788AFD20FFB7FFD4FD4144E5916A64EA" box="[680,782,223,249]" pageId="41" pageNumber="42" type="male">one male</specimenCount>
and
<specimenCount id="788AFD20FFB7FFD4FCAD44E591A964EA" box="[836,973,223,249]" pageId="41" pageNumber="42" type="female">two females</specimenCount>
) in Horstmanns collection from northern
<collectingCountry id="169B7639FFB7FFD4FF7E4539929B650E" box="[151,255,259,285]" name="Germany" pageId="41" pageNumber="42">Germany</collectingCountry>
had originally been identified by Horstmann as
<taxonomicName id="A98C4D2AFFB7FFD4FCCE453996F8650E" authority="Schmiedeknecht" authorityName="Schmiedeknecht" authorityYear="1909" box="[807,1180,259,285]" class="Insecta" family="Ichneumonidae" genus="Anilastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="thuringiacus">
<emphasis id="5CF8EABBFFB7FFD4FCCE453991BC650E" box="[807,984,259,285]" italics="true" pageId="41" pageNumber="42">H. thuringiacus</emphasis>
Schmiedeknecht
</taxonomicName>
, but bear Horstmanns labels correcting this to
<emphasis id="5CF8EABBFFB7FFD4FE4D451D90536552" box="[420,567,295,321]" italics="true" pageId="41" pageNumber="42">
<taxonomicName id="A98C4D2AFFB7FFD4FE4D451D90576552" ID-CoL="3NYWL" authorityName="Galsworthy &amp; Shaw &amp; Haraldseide" authorityYear="2023" baseAuthorityName="Holmgren" baseAuthorityYear="1860" box="[420,563,295,321]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="dubitatus">H. dubitatus</taxonomicName>
.
</emphasis>
They match the type of
<taxonomicName id="A98C4D2AFFB7FFD4FCA1451D91D46552" ID-CoL="3NYWL" authorityName="Galsworthy &amp; Shaw &amp; Haraldseide" authorityYear="2023" baseAuthorityName="Holmgren" baseAuthorityYear="1860" box="[840,944,295,321]" class="Insecta" family="Ichneumonidae" genus="Hyposoter" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="dubitatus">
<emphasis id="5CF8EABBFFB7FFD4FCA1451D91D46552" box="[840,944,295,321]" italics="true" pageId="41" pageNumber="42">dubitatus</emphasis>
</taxonomicName>
very well (see above under
<taxonomicName id="A98C4D2AFFB7FFD4FB05451D97F96552" authorityName="Schmiedeknecht" authorityYear="1909" box="[1260,1437,295,321]" class="Insecta" family="Ichneumonidae" genus="Anilastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="thuringiacus">
<emphasis id="5CF8EABBFFB7FFD4FB05451D97F96552" box="[1260,1437,295,321]" italics="true" pageId="41" pageNumber="42">H. thuringiacus</emphasis>
</taxonomicName>
in the section on species excluded from the key).
</paragraph>
<paragraph id="6E3336A9FFB7FFD4FF2E45559765659A" blockId="41.[151,1437,151,573]" box="[199,1281,367,393]" pageId="41" pageNumber="42">
The species is very similar to
<taxonomicName id="A98C4D2AFFB7FFD4FDFF455590F8659A" authorityName="Gravenhorst" authorityYear="1829" box="[534,668,367,393]" class="Insecta" family="Ichneumonidae" genus="Campoplex" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="caedator">
<emphasis id="5CF8EABBFFB7FFD4FDFF455590F8659A" box="[534,668,367,393]" italics="true" pageId="41" pageNumber="42">H. caedator</emphasis>
</taxonomicName>
: see notes under that species for separation of the two.
</paragraph>
</subSubSection>
<subSubSection id="26966522FFB7FFD4FF2E45A990F4662E" pageId="41" pageNumber="42" type="description">
<paragraph id="6E3336A9FFB7FFD4FF2E45A990F4662E" blockId="41.[151,1437,151,573]" pageId="41" pageNumber="42">
The
<typeStatus id="B137880BFFB7FFD4FF1045A9934D65BE" box="[249,297,403,429]" pageId="41" pageNumber="42">type</typeStatus>
material is from
<collectingCountry id="169B7639FFB7FFD4FE0045A9902165BE" box="[489,581,403,429]" name="Sweden" pageId="41" pageNumber="42">Sweden</collectingCountry>
. The
<specimenCount id="788AFD20FFB7FFD4FD6945A9909965BE" box="[640,765,403,429]" pageId="41" pageNumber="42" type="female">two female</specimenCount>
specimens in Horstmanns collection mentioned above bear labels ex
<taxonomicName id="A98C4D2AFFB7FFD4FEE3458D933D65C2" authorityName="Duponchel" authorityYear="1829" box="[266,345,439,465]" class="Insecta" family="Geometridae" genus="Epione" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">Epione</taxonomicName>
parallelaria, i.e.
<taxonomicName id="A98C4D2AFFB7FFD4FDF6458D911265C2" authority="(Linnaeus)" baseAuthorityName="Linnaeus" box="[543,886,439,465]" class="Insecta" family="Geometridae" genus="Epione" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="vespertaria">
<emphasis id="5CF8EABBFFB7FFD4FDF6458D909265C3" box="[543,758,439,464]" italics="true" pageId="41" pageNumber="42">Epione vespertaria</emphasis>
(Linnaeus)
</taxonomicName>
(
<taxonomicName id="A98C4D2AFFB7FFD4FC6D458D967465C2" authorityName="Linnaeus" authorityYear="1758" box="[900,1040,439,465]" class="Insecta" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
,
<taxonomicName id="A98C4D2AFFB7FFD4FBF2458D96D465C2" baseAuthorityName="Leach" baseAuthorityYear="1815" box="[1051,1200,439,465]" class="Insecta" family="Geometridae" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="family">Geometridae</taxonomicName>
), though there are no cocoons or larval remains. One specimen in NMS was reared in
<collectingCountry id="169B7639FFB7FFD4FC9745E191D565E6" box="[894,945,475,501]" name="Italy" pageId="41" pageNumber="42">Italy</collectingCountry>
from an unknown geometrid and a further specimen in the French Alps also from an indeterminate geometrid, possibly
<taxonomicName id="A98C4D2AFFB7FFD4FC1345C59635660A" authorityName="Hubner" authorityYear="1821" box="[1018,1105,511,537]" class="Insecta" family="Geometridae" genus="Eulithis" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="5CF8EABBFFB7FFD4FC1345C59635660A" box="[1018,1105,511,537]" italics="true" pageId="41" pageNumber="42">Eulithis</emphasis>
</taxonomicName>
. There are also further specimens in NMS from
<collectingCountry id="169B7639FFB7FFD4FE9E461993A7662E" box="[375,451,547,573]" name="France" pageId="41" pageNumber="42">France</collectingCountry>
and from the
<collectingCountry id="169B7639FFB7FFD4FD89461990E8662E" box="[608,652,547,573]" name="United Kingdom" pageId="41" pageNumber="42">UK</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>