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<mods:title id="A82ADFEF10C58040A2D7E995EB6258F4">Calliaxina chalmasii (Brocchi, 1883) comb. nov. (Decapoda: Axiidea: Callianassidae: Eucalliacinae), a ghost shrimp from the Middle Miocene of Europe, with reappraisal of the fossil record of Eucalliacinae</mods:title>
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<heading id="6C5E6A4DFF887D49FF61A25D31B7FBAD" bold="true" box="[151,741,1123,1150]" fontSize="11" level="1" pageId="7" pageNumber="56" reason="1">
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<emphasis id="05DD0133FF887D49FF61A25D32C0FBAE" bold="true" box="[151,402,1124,1150]" italics="true" pageId="7" pageNumber="56">Calliaxina chalmasii</emphasis>
(
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)
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(
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AB, 4AR, 5AE, 6AE)
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<taxonomicName id="F0A9A6A2FF887D49FF61A2F6317DFB34" ID-CoL="f45854b2-a302-4361-bd40-0423151d5e6a" authority="Chalmasii Brocchi, 1883: 6" authorityName="Chalmasii Brocchi" authorityPageNumber="6" authorityYear="1883" box="[151,559,1230,1253]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="05DD0133FF887D49FF61A2F6322CFB35" box="[151,382,1231,1253]" italics="true" pageId="7" pageNumber="56">Callianassa Chalmasii</emphasis>
<treatmentCitation id="B608FB30FF887D49FE72A2F7317DFB34" author="Brocchi" box="[388,559,1230,1252]" page="6" pageId="7" pageNumber="56" year="1883">
<bibRefCitation id="5338A0D0FF887D49FE72A2F73145FB34" author="Brocchi" box="[388,535,1230,1252]" pageId="7" pageNumber="56" refString="Brocchi, P. (1883) Notes sur les Crustaces fossiles des terres tertiaires de la Hongrie. Annales des Sciences Geologiques, (2) 14, 1 - 8." type="journal article" year="1883">Brocchi, 1883</bibRefCitation>
: 6
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, 7, pl. 5, figs. 7, 8.
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<emphasis id="05DD0133FF887D49FF61A2D4322CFAD3" box="[151,382,1261,1283]" italics="true" pageId="7" pageNumber="56">Callianassa Chalmasi.</emphasis>
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<treatmentCitation id="B608FB30FF887D49FE6EA2D43130FAD3" author="Glaessner" box="[408,610,1261,1283]" page="77" pageId="7" pageNumber="56" year="1929">
<bibRefCitation id="5338A0D0FF887D49FE6EA2D4316FFAD3" author="Glaessner" box="[408,573,1261,1283]" pageId="7" pageNumber="56" refString="Glaessner, M. F. (1929) Crustacea Decapoda. In. Pompeckj, F. J. (ed.) Fossilium catalogus, 1: Animalium, 41, 1 - 464. W. Junk, Berlin." type="journal article" year="1929">Glaessner, 1929</bibRefCitation>
: 77
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.
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</paragraph>
<paragraph id="3716DD21FF887D49FF61A33530BCFAF1" blockId="7.[151,1436,1230,1743]" box="[151,1006,1291,1314]" pageId="7" pageNumber="56">
<treatmentCitationGroup id="17B9FA0FFF887D49FF61A33530BCFAF1" box="[151,1006,1291,1314]" pageId="7" pageNumber="56">
<emphasis id="05DD0133FF887D49FF61A33532D2FAF2" box="[151,384,1292,1314]" italics="true" pageId="7" pageNumber="56">
<taxonomicName id="F0A9A6A2FF887D49FF61A335322EFAF2" box="[151,380,1292,1314]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="chalmasii">Callianassa chalmasii</taxonomicName>
.
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<treatmentCitation id="B608FB30FF887D49FE6FA3323103FAF1" author="Muller" box="[409,593,1291,1313]" page="274" pageId="7" pageNumber="56" year="1979">
<bibRefCitation id="5338A0D0FF887D49FE6FA332314DFAF1" author="Muller" box="[409,543,1291,1313]" pageId="7" pageNumber="56" refString="Muller, P. (1979) Decapoda (Crustacea) fauna a budapesti miocenbol (5). Foldtani Kozlony, Bulletin of the Hungarian Geological Society, 108, 272 - 318." type="journal article" year="1979">Müller, 1979</bibRefCitation>
: 274
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, pl. 1, fig. 5;
<treatmentCitation id="B608FB30FF887D49FD2EA33230B8FAF1" author="Schweitzer" box="[728,1002,1291,1314]" page="34" pageId="7" pageNumber="56" year="2010">
<bibRefCitation id="5338A0D0FF887D49FD2EA3323096FAF1" author="Schweitzer" box="[728,964,1291,1314]" pageId="7" pageNumber="56" refString="Schweitzer, C. E., Feldmann, R. M., Garassino, A., Karasawa, H. &amp; Schweigert, G. (2010) Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10, 1 - 222." type="journal article" year="2010">
Schweitzer
<emphasis id="05DD0133FF887D49FCB8A3343029FAF2" box="[846,891,1292,1314]" italics="true" pageId="7" pageNumber="56">et al</emphasis>
., 2010
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: 34
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.
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</paragraph>
<paragraph id="3716DD21FF887D49FF61A3133223FAAE" blockId="7.[151,1436,1230,1743]" pageId="7" pageNumber="56">
<treatmentCitationGroup id="17B9FA0FFF887D49FF61A3133223FAAE" pageId="7" pageNumber="56">
<emphasis id="05DD0133FF887D49FF61A31332DDFA90" box="[151,399,1322,1344]" italics="true" pageId="7" pageNumber="56">
<taxonomicName id="F0A9A6A2FF887D49FF61A3133220FA90" box="[151,370,1322,1344]" class="Malacostraca" family="Callianassidae" genus="Calianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="chalmasii">Calianassa chalmasii</taxonomicName>
.—
</emphasis>
<treatmentCitation id="B608FB30FF887D49FE79A313313EFA90" author="Lorenthey" box="[399,620,1322,1344]" page="150" pageId="7" pageNumber="56" year="1897">
<bibRefCitation id="5338A0D0FF887D49FE79A313316BFA90" author="Lorenthey" box="[399,569,1322,1344]" pageId="7" pageNumber="56" refString="Lorenthey, E. (1897) Adatok Magyarorszag harmadkoru rak faunajahoz. Mathematikai es Termeszettudomanyi Ertesito, 15, 149 - 169." type="journal article" year="1897">Lőrenthey, 1897</bibRefCitation>
: 150
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, 160, 168;
<treatmentCitation id="B608FB30FF887D49FD28A31330E9FA90" author="Lorenthey" box="[734,955,1322,1344]" page="93" pageId="7" pageNumber="56" year="1898">
<bibRefCitation id="5338A0D0FF887D49FD28A31330C0FA90" author="Lorenthey" box="[734,914,1322,1344]" pageId="7" pageNumber="56" refString="Lorenthey, E. (1898 a) Beitrage zur Decapodenfauna des ungarischen Tertiars. Mathematischen und Naturwissenschaftlichen Berichte aus Ungarn, 14, 92 - 115." type="journal article" year="1898" yearSuffix="a">
Lőrenthey, 1898
<emphasis id="05DD0133FF887D49FC73A31230C0FA90" box="[901,914,1323,1344]" italics="true" pageId="7" pageNumber="56">a</emphasis>
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: 93
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, 105, 114;
<treatmentCitation id="B608FB30FF887D49FBDBA31337AEFA90" author="Lorenthey" box="[1069,1276,1322,1344]" page="9" pageId="7" pageNumber="56" year="1898">
<bibRefCitation id="5338A0D0FF887D49FBDBA31337B2FA90" author="Lorenthey" box="[1069,1248,1322,1344]" pageId="7" pageNumber="56" refString="Lorenthey, E. (1898 b) Adatok Magyarorszag harmadkoru rak-faunajahoz. Mathematikai es Termeszettudomanyi Kozllemenyek, 27 (2), 103 - 271." type="journal article" year="1898" yearSuffix="b">
Lőrenthey, 1898
<emphasis id="05DD0133FF887D49FB25A31337B2FA90" box="[1235,1248,1322,1344]" italics="true" pageId="7" pageNumber="56">b</emphasis>
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: 9
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, 129, 130, 155;
<treatmentCitation id="B608FB30FF887D49FF31A37032FCFA8F" author="Lorenthey" box="[199,430,1353,1375]" page="102" pageId="7" pageNumber="56" year="1898">
<bibRefCitation id="5338A0D0FF887D49FF31A3703225FA8F" author="Lorenthey" box="[199,375,1353,1375]" pageId="7" pageNumber="56" refString="Lorenthey, E. (1898 c) Beitrage zur Decapodenfauna des Ungarischen Tertiars. Termeszetrajzi Fuzetek, 21, 1 - 133." type="journal article" year="1898" yearSuffix="c">
Lőrenthey, 1898
<emphasis id="05DD0133FF887D49FE9AA3733225FA8F" box="[364,375,1354,1375]" italics="true" pageId="7" pageNumber="56">c</emphasis>
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: 102
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; Lőrenthey, 1904: 161; Lőrenthey, 1913: 332; Lőrenthey in
<treatmentCitation id="B608FB30FF887D49FBF9A3703602FA8F" author="Lorenthey" box="[1039,1360,1353,1375]" page="33" pageId="7" pageNumber="56" year="1929">
<bibRefCitation id="5338A0D0FF887D49FBF9A3703679FA8F" author="Lorenthey" box="[1039,1323,1353,1375]" pageId="7" pageNumber="56" refString="Lorenthey, E. &amp; Beurlen, K. (1929) Die fossilen Dekapoden der Lander der Ungarischen Krone. Geologica Hungarica, Series Palaeontologica, 3, 1 - 421." type="journal article" year="1929">Lőrenthey &amp; Beurlen, 1929</bibRefCitation>
: 33
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, 65, pl. 2, figs. 15a, 15b.
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<taxonomicName id="F0A9A6A2FF887D49FF69A3BE324AFA4D" ID-CoL="f45854b2-a302-4361-bd40-0423151d5e6a" box="[159,280,1415,1437]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="05DD0133FF887D49FF69A3BE324AFA4D" box="[159,280,1415,1437]" italics="true" pageId="7" pageNumber="56">Callianassa</emphasis>
</taxonomicName>
<taxonomicName id="F0A9A6A2FF887D49FED1A3BE32DBFA4D" box="[295,393,1415,1437]" class="Malacostraca" family="Callianassidae" genus="Calianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="chalmasii">
<emphasis id="05DD0133FF887D49FED1A3BE32DBFA4D" box="[295,393,1415,1437]" italics="true" pageId="7" pageNumber="56">chalmasii</emphasis>
</taxonomicName>
.—
<treatmentCitation id="B608FB30FF887D49FE51A3BF3104FA4C" author="Muller" box="[423,598,1414,1436]" page="50" pageId="7" pageNumber="56" year="1984">
<bibRefCitation id="5338A0D0FF887D49FE51A3BF3162FA4C" author="Muller" box="[423,560,1414,1436]" pageId="7" pageNumber="56" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller, 1984</bibRefCitation>
: 50
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, pl. 2, figs. 37.
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<taxonomicName id="F0A9A6A2FF887D49FF61A39C31F4FA6B" authority="Veiga Ferreira, 1961: 479" authorityName="Veiga Ferreira" authorityPageNumber="479" authorityYear="1961" box="[151,678,1445,1467]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="espichelensis">
<emphasis id="05DD0133FF887D49FF61A39C32CEFA6B" box="[151,412,1445,1467]" italics="true" pageId="7" pageNumber="56">Callianassa espichelensis</emphasis>
<treatmentCitation id="B608FB30FF887D49FE54A39C31F4FA6B" author="Veiga" box="[418,678,1445,1467]" page="479" pageId="7" pageNumber="56" year="1961">
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: 479
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481, pl. 1, figs. 111;
<treatmentCitation id="B608FB30FF887D49FC79A39C37F3FA6B" author="Schweitzer" box="[911,1185,1445,1467]" page="34" pageId="7" pageNumber="56" year="2010">
<bibRefCitation id="5338A0D0FF887D49FC79A39C3729FA6B" author="Schweitzer" box="[911,1147,1445,1467]" pageId="7" pageNumber="56" refString="Schweitzer, C. E., Feldmann, R. M., Garassino, A., Karasawa, H. &amp; Schweigert, G. (2010) Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10, 1 - 222." type="journal article" year="2010">
Schweitzer
<emphasis id="05DD0133FF887D49FBF0A39F3760FA6B" box="[1030,1074,1445,1467]" italics="true" pageId="7" pageNumber="56">et al</emphasis>
., 2010
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: 34
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<paragraph id="3716DD21FF887D49FF61A3FD30DDFA28" blockId="7.[151,1436,1230,1743]" pageId="7" pageNumber="56">
<treatmentCitationGroup id="17B9FA0FFF887D49FF61A3FD30DDFA28" pageId="7" pageNumber="56">
<taxonomicName id="F0A9A6A2FF887D49FF61A3FD313DFA09" authority="Lorenthey, 1897: 160" authorityName="Lorenthey" authorityPageNumber="160" authorityYear="1897" box="[151,623,1475,1498]" class="Malacostraca" family="Callianassidae" genus="Calianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="rakosiensis">
<emphasis id="05DD0133FF887D49FF61A3FD32D6FA0A" box="[151,388,1476,1498]" italics="true" pageId="7" pageNumber="56">Calianassa rákosiensis</emphasis>
<treatmentCitation id="B608FB30FF887D49FE7BA3FA313DFA09" author="Lorenthey" box="[397,623,1475,1497]" page="160" pageId="7" pageNumber="56" year="1897">
<bibRefCitation id="5338A0D0FF887D49FE7BA3FA3169FA09" author="Lorenthey" box="[397,571,1475,1497]" pageId="7" pageNumber="56" refString="Lorenthey, E. (1897) Adatok Magyarorszag harmadkoru rak faunajahoz. Mathematikai es Termeszettudomanyi Ertesito, 15, 149 - 169." type="journal article" year="1897">Lőrenthey, 1897</bibRefCitation>
: 160
</treatmentCitation>
</taxonomicName>
, 161, 168, 169 (new synonym);
<treatmentCitation id="B608FB30FF887D49FC3FA3FA37E5FA09" author="Lorenthey" box="[969,1207,1475,1498]" page="105" pageId="7" pageNumber="56" year="1898">
<bibRefCitation id="5338A0D0FF887D49FC3FA3FA372DFA0A" author="Lorenthey" box="[969,1151,1475,1498]" pageId="7" pageNumber="56" refString="Lorenthey, E. (1898 a) Beitrage zur Decapodenfauna des ungarischen Tertiars. Mathematischen und Naturwissenschaftlichen Berichte aus Ungarn, 14, 92 - 115." type="journal article" year="1898" yearSuffix="a">
Lőrenthey, 1898
<emphasis id="05DD0133FF887D49FB84A3FC372DFA0A" box="[1138,1151,1477,1498]" italics="true" pageId="7" pageNumber="56">a</emphasis>
</bibRefCitation>
: 105
</treatmentCitation>
, 114, 115;
<treatmentCitation id="B608FB30FF887D49FAD9A3FA326EFA28" author="Lorenthey" page="131" pageId="7" pageNumber="56" year="1898">
<bibRefCitation id="5338A0D0FF887D49FAD9A3FA3254FA28" author="Lorenthey" pageId="7" pageNumber="56" refString="Lorenthey, E. (1898 b) Adatok Magyarorszag harmadkoru rak-faunajahoz. Mathematikai es Termeszettudomanyi Kozllemenyek, 27 (2), 103 - 271." type="journal article" year="1898" yearSuffix="b">
Lőrenthey, 1898
<emphasis id="05DD0133FF887D49FF0FA3DB3254FA28" box="[249,262,1506,1528]" italics="true" pageId="7" pageNumber="56">b</emphasis>
</bibRefCitation>
: 131
</treatmentCitation>
, 132, pl. 9, fig. 4;
<treatmentCitation id="B608FB30FF887D49FE03A3DB318EFA28" author="Lorenthey" box="[501,732,1506,1528]" page="103" pageId="7" pageNumber="56" year="1898">
<bibRefCitation id="5338A0D0FF887D49FE03A3DB31F4FA28" author="Lorenthey" box="[501,678,1506,1528]" pageId="7" pageNumber="56" refString="Lorenthey, E. (1898 c) Beitrage zur Decapodenfauna des Ungarischen Tertiars. Termeszetrajzi Fuzetek, 21, 1 - 133." type="journal article" year="1898" yearSuffix="c">
Lőrenthey, 1898
<emphasis id="05DD0133FF887D49FD6DA3DA31F4FA28" box="[667,678,1507,1528]" italics="true" pageId="7" pageNumber="56">c</emphasis>
</bibRefCitation>
: 103
</treatmentCitation>
, 104, pl. 9, fig. 4;
</treatmentCitationGroup>
</paragraph>
<paragraph id="3716DD21FF887D49FF61A038376EF9C7" blockId="7.[151,1436,1230,1743]" box="[151,1084,1537,1559]" pageId="7" pageNumber="56">
<treatmentCitationGroup id="17B9FA0FFF887D49FF61A038376EF9C7" box="[151,1084,1537,1559]" pageId="7" pageNumber="56">
<taxonomicName id="F0A9A6A2FF887D49FF61A0383228F9C7" box="[151,378,1537,1559]" class="Malacostraca" family="Callianassidae" genus="Calianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="rakosensis">
<emphasis id="05DD0133FF887D49FF61A0383228F9C7" box="[151,378,1537,1559]" italics="true" pageId="7" pageNumber="56">Calianassa rákosensis</emphasis>
</taxonomicName>
.—Lőrenthey in
<treatmentCitation id="B608FB30FF887D49FDD4A0383030F9C7" author="Lorenthey" box="[546,866,1537,1559]" page="33" pageId="7" pageNumber="56" year="1929">
<bibRefCitation id="5338A0D0FF887D49FDD4A038306FF9C7" author="Lorenthey" box="[546,829,1537,1559]" pageId="7" pageNumber="56" refString="Lorenthey, E. &amp; Beurlen, K. (1929) Die fossilen Dekapoden der Lander der Ungarischen Krone. Geologica Hungarica, Series Palaeontologica, 3, 1 - 421." type="journal article" year="1929">Lőrenthey &amp; Beurlen, 1929</bibRefCitation>
: 33
</treatmentCitation>
, 66, 67, pl. 2, fig. 14.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3716DD21FF887D49FF61A019305EF9E5" blockId="7.[151,1436,1230,1743]" box="[151,780,1568,1590]" pageId="7" pageNumber="56">
<treatmentCitationGroup id="17B9FA0FFF887D49FF61A019305EF9E5" box="[151,780,1568,1590]" pageId="7" pageNumber="56">
<taxonomicName id="F0A9A6A2FF887D49FF61A01932D3F9E6" box="[151,385,1568,1590]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="rakosensis">
<emphasis id="05DD0133FF887D49FF61A01932D3F9E6" box="[151,385,1568,1590]" italics="true" pageId="7" pageNumber="56">Callianassa rákosensis</emphasis>
</taxonomicName>
.—
<treatmentCitation id="B608FB30FF887D49FE69A0193108F9E5" author="Muller" box="[415,602,1568,1590]" page="274" pageId="7" pageNumber="56" year="1979">
<bibRefCitation id="5338A0D0FF887D49FE69A019317AF9E5" author="Muller" box="[415,552,1568,1590]" pageId="7" pageNumber="56" refString="Muller, P. (1979) Decapoda (Crustacea) fauna a budapesti miocenbol (5). Foldtani Kozlony, Bulletin of the Hungarian Geological Society, 108, 272 - 318." type="journal article" year="1979">Müller, 1979</bibRefCitation>
: 274
</treatmentCitation>
, 276, pl. 1, fig. 4.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3716DD21FF887D49FF61A00630C6F984" blockId="7.[151,1436,1230,1743]" box="[151,916,1598,1621]" pageId="7" pageNumber="56">
<treatmentCitationGroup id="17B9FA0FFF887D49FF61A00630C6F984" box="[151,916,1598,1621]" pageId="7" pageNumber="56">
<taxonomicName id="F0A9A6A2FF887D49FF61A00632DAF985" box="[151,392,1599,1621]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="rakosiensis">
<emphasis id="05DD0133FF887D49FF61A00632DAF985" box="[151,392,1599,1621]" italics="true" pageId="7" pageNumber="56">Callianassa rakosiensis</emphasis>
</taxonomicName>
.—
<treatmentCitation id="B608FB30FF887D49FE50A0073121F984" author="Glaessner" box="[422,627,1598,1620]" page="89" pageId="7" pageNumber="56" year="1929">
<bibRefCitation id="5338A0D0FF887D49FE50A007311FF984" author="Glaessner" box="[422,589,1598,1620]" pageId="7" pageNumber="56" refString="Glaessner, M. F. (1929) Crustacea Decapoda. In. Pompeckj, F. J. (ed.) Fossilium catalogus, 1: Animalium, 41, 1 - 464. W. Junk, Berlin." type="journal article" year="1929">Glaessner, 1929</bibRefCitation>
: 89
</treatmentCitation>
;
<treatmentCitation id="B608FB30FF887D49FD88A00730C2F984" author="Schweitzer" box="[638,912,1598,1621]" page="36" pageId="7" pageNumber="56" year="2010">
<bibRefCitation id="5338A0D0FF887D49FD88A0073038F984" author="Schweitzer" box="[638,874,1598,1621]" pageId="7" pageNumber="56" refString="Schweitzer, C. E., Feldmann, R. M., Garassino, A., Karasawa, H. &amp; Schweigert, G. (2010) Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10, 1 - 222." type="journal article" year="2010">
Schweitzer
<emphasis id="05DD0133FF887D49FD02A0793072F985" box="[756,800,1599,1621]" italics="true" pageId="7" pageNumber="56">et al</emphasis>
., 2010
</bibRefCitation>
: 36
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3716DD21FF887D49FF61A0673053F9A3" blockId="7.[151,1436,1230,1743]" box="[151,769,1629,1651]" pageId="7" pageNumber="56">
<treatmentCitationGroup id="17B9FA0FFF887D49FF61A0673053F9A3" box="[151,769,1629,1651]" pageId="7" pageNumber="56">
<taxonomicName id="F0A9A6A2FF887D49FF69A064324AF9A3" ID-CoL="f45854b2-a302-4361-bd40-0423151d5e6a" box="[159,280,1629,1651]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="05DD0133FF887D49FF69A064324AF9A3" box="[159,280,1629,1651]" italics="true" pageId="7" pageNumber="56">Callianassa</emphasis>
</taxonomicName>
<taxonomicName id="F0A9A6A2FF887D49FED7A06432C1F9A3" box="[289,403,1629,1651]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="rakosiensis">
<emphasis id="05DD0133FF887D49FED7A06432C1F9A3" box="[289,403,1629,1651]" italics="true" pageId="7" pageNumber="56">rakosiensis</emphasis>
</taxonomicName>
.—
<treatmentCitation id="B608FB30FF887D49FE47A064310FF9A3" author="Muller" box="[433,605,1629,1651]" page="51" pageId="7" pageNumber="56" year="1984">
<bibRefCitation id="5338A0D0FF887D49FE47A0643168F9A3" author="Muller" box="[433,570,1629,1651]" pageId="7" pageNumber="56" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller, 1984</bibRefCitation>
: 51
</treatmentCitation>
, pl. 3, figs. 1, 2.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3716DD21FF887D49FF61A0453773F941" blockId="7.[151,1436,1230,1743]" box="[151,1057,1659,1682]" pageId="7" pageNumber="56">
<taxonomicName id="F0A9A6A2FF887D49FF61A04532F3F942" box="[151,417,1659,1682]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="rakosensis">
<emphasis id="05DD0133FF887D49FF61A0453242F942" box="[151,272,1660,1682]" italics="true" pageId="7" pageNumber="56">Callianassa</emphasis>
cf.
<emphasis id="05DD0133FF887D49FEC0A04532F3F942" box="[310,417,1660,1682]" italics="true" pageId="7" pageNumber="56">rákosensis</emphasis>
</taxonomicName>
.—Bachmayer, 1953: 242; Bachmayer &amp; Tollmann, 1953: 312.
</paragraph>
<paragraph id="3716DD21FF887D49FF61A0A231C9F960" blockId="7.[151,1436,1230,1743]" box="[151,667,1690,1712]" pageId="7" pageNumber="56">
?„
<taxonomicName id="F0A9A6A2FF887D49FF5BA0A33274F960" ID-CoL="f45854b2-a302-4361-bd40-0423151d5e6a" box="[173,294,1690,1712]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="05DD0133FF887D49FF5BA0A33274F960" box="[173,294,1690,1712]" italics="true" pageId="7" pageNumber="56">Callianassa</emphasis>
</taxonomicName>
“ sp. 2—
<bibRefCitation id="5338A0D0FF887D49FE74A0A33154F960" author="Hyzny" box="[386,518,1690,1712]" pageId="7" pageNumber="56" refString="Hyzny, M. (2011) In situ mud shrimps (Decapoda: Axiidea: Callianassidae) preserved within their burrows from the middle Miocene of the Central Paratethys. Bulletin of the Mizunami Fossil Museum, 37, 37 - 46." type="journal article" year="2011">Hyžný, 2011</bibRefCitation>
: 42, fig. 4E, F.
</paragraph>
<paragraph id="3716DD21FF887D49FF61A0833615F91F" blockId="7.[151,1436,1230,1743]" box="[151,1351,1721,1743]" pageId="7" pageNumber="56">
<treatmentCitationGroup id="17B9FA0FFF887D49FF61A0833615F91F" box="[151,1351,1721,1743]" pageId="7" pageNumber="56">
non
<emphasis id="05DD0133FF887D49FF35A08032FEF91F" box="[195,428,1721,1743]" italics="true" pageId="7" pageNumber="56">
<taxonomicName id="F0A9A6A2FF887D49FF35A08032FAF91F" box="[195,424,1721,1743]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="chalmasii">Callianassa chalmasii</taxonomicName>
.
</emphasis>
<treatmentCitation id="B608FB30FF887D49FE33A0833189F91F" author="De" box="[453,731,1721,1743]" page="168" pageId="7" pageNumber="56" year="2009">
<bibRefCitation id="5338A0D0FF887D49FE33A08331FBF91F" author="De" box="[453,681,1721,1743]" pageId="7" pageNumber="56" refString="De Angeli, A., Garassino, A. &amp; Pasini, G. (2009) New reports of anomurans and brachyurans from the Cenozoic of Tuscany (Italy). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 150 (2), 163 - 196." type="journal article" year="2009">
<collectingCountry id="4FBE9DB1FF887D49FE33A08332B0F91F" box="[453,482,1722,1743]" name="South Georgia and the South Sandwich Islands" pageId="7" pageNumber="56">De</collectingCountry>
Angeli
<emphasis id="05DD0133FF887D49FDC2A083310DF91F" box="[564,607,1721,1743]" italics="true" pageId="7" pageNumber="56">et al</emphasis>
., 2009
</bibRefCitation>
: 168
</treatmentCitation>
, figs. 2C, D;
<treatmentCitation id="B608FB30FF887D49FC94A080373BF91F" author="Garassino" box="[866,1129,1721,1743]" page="18" pageId="7" pageNumber="56" year="2012">
<bibRefCitation id="5338A0D0FF887D49FC94A0803716F91F" author="Garassino" box="[866,1092,1721,1743]" pageId="7" pageNumber="56" refString="Garassino, A., Pasini, G., De Angeli, A., Charbonnier, A., Famiani, F., Baldanza, A. &amp; Bizzarri, R. (2012) The decapod community from the Early Pliocene (Zanclean) of &quot; La Serra &quot; quarry (San Miniato, Pisa, Toscana, central Italy): sedimentology, systematics, and palaeoenvironmental implications. Annales de Paleontologie, 98, 1 - 61." type="journal article" year="2012">
Garassino
<emphasis id="05DD0133FF887D49FC3BA08330ABF91F" box="[973,1017,1721,1743]" italics="true" pageId="7" pageNumber="56">et al</emphasis>
., 2012
</bibRefCitation>
: 18
</treatmentCitation>
. (=
<taxonomicName id="F0A9A6A2FF887D49FB7FA0803612F91F" box="[1161,1344,1721,1743]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="punica">
<emphasis id="05DD0133FF887D49FB7FA0803612F91F" box="[1161,1344,1721,1743]" italics="true" pageId="7" pageNumber="56">Calliaxina punica</emphasis>
</taxonomicName>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="7FB38EAAFF887D49FF61A0C23186F88C" pageId="7" pageNumber="56" type="diagnosis">
<paragraph id="3716DD21FF887D49FF61A0C23186F88C" blockId="7.[151,1438,1787,2028]" pageId="7" pageNumber="56">
<emphasis id="05DD0133FF887D49FF61A0C23243F8C4" bold="true" box="[151,273,1787,1812]" pageId="7" pageNumber="56">Diagnosis.</emphasis>
<taxonomicName id="F0A9A6A2FF887D49FEEAA0C532C0F8C4" box="[284,402,1788,1812]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="05DD0133FF887D49FEEAA0C532C0F8C4" box="[284,402,1788,1812]" italics="true" pageId="7" pageNumber="56">Calliaxina</emphasis>
</taxonomicName>
with both P1 propodi quadrate, upper and lower margins keeled; fixed finger of major propodus with diagonally oriented ridge, occlusal margin toothed; minor propodus with well developed tuberculated depression on the base of fixed finger.
</paragraph>
</subSubSection>
<subSubSection id="7FB38EAAFF887D46FF31A15E31D6FA77" lastPageId="8" lastPageNumber="57" pageId="7" pageNumber="56" type="description">
<paragraph id="3716DD21FF887D49FF31A15E32B2F874" blockId="7.[151,1438,1787,2028]" pageId="7" pageNumber="56">
<emphasis id="05DD0133FF887D49FF31A15E320BF850" bold="true" box="[199,345,1895,1920]" pageId="7" pageNumber="56">Description.</emphasis>
First pereiopods (chelipeds) with major and minor cheliped strongly developed, subequal in size, similar, laterally compressed.
</paragraph>
<paragraph id="3716DD21FF887D49FF33A19636CCF83C" blockId="7.[151,1438,1787,2028]" pageId="7" pageNumber="56">Carpus unarmed, slightly higher than long, triangular in shape, upper and lower margin keeled, terminating in blunt corner at articulation with propodus, lower margin together with proximal margin forming a single rounded edge.</paragraph>
<paragraph id="3716DD21FF877D46FF33A6AE377FFEEF" blockId="8.[153,1436,151,644]" pageId="8" pageNumber="57">Both major and minor propodus quadrate, about as high as long, sometimes longer or shorter than high, converging distally; upper and lower margins keeled, terminating in blunt corners at articulation with carpus; keel on the upper margin forming sharp extension distally; keel on the lower margin distinctly broader proximally, forming flat edge; inner surface of propodi smooth with distinct setal pores; setal pores on the upper margin large, elongated in vertical aspect, i.e. perpendicular to the longitudinal axis of the limb.</paragraph>
<paragraph id="3716DD21FF877D46FF33A774329FFE1F" blockId="8.[153,1436,151,644]" pageId="8" pageNumber="57">
In major propodus (previously described as
<taxonomicName id="F0A9A6A2FF877D46FD26A775309DFEB4" box="[720,975,332,356]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="chalmasii">
<emphasis id="05DD0133FF877D46FD26A775309DFEB4" box="[720,975,332,356]" italics="true" pageId="8" pageNumber="57">Callianassa chalmasii</emphasis>
</taxonomicName>
), area at the articulation with dactylus covered with fine tubercles; fixed finger short, triangular, with distinct obliquely oriented ridge on the lateral surface; occlusal margin of fixed finger with distinct tooth followed by serrated edge; lower margin of propodus slightly convex proximally.
</paragraph>
<paragraph id="3716DD21FF877D46FF33A7E433A7FD8F" blockId="8.[153,1436,151,644]" pageId="8" pageNumber="57">
In minor propodus (previously described as
<taxonomicName id="F0A9A6A2FF877D46FD30A7E4303AFE24" box="[710,872,476,500]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="rakosiensis">
<emphasis id="05DD0133FF877D46FD30A7E4303AFE24" box="[710,872,476,500]" italics="true" pageId="8" pageNumber="57">C. rakosiensis</emphasis>
</taxonomicName>
), area at the articulation with dactylus distinctly depressed, covered with tubercles; depression forming sharp edge at its lower margin, which is parallel to the lower margin of propodus distally and distinctly rounded proximally; lower margin of propodus usually distinctly straight.
</paragraph>
<paragraph id="3716DD21FF877D46FF33A455311CFD54" blockId="8.[153,1436,151,644]" box="[197,590,620,644]" pageId="8" pageNumber="57">Dactylus slightly curved, unarmed.</paragraph>
<caption id="63D68DA9FF877D46FF61A2A43675FB64" box="[151,1319,1181,1204]" httpUri="https://zenodo.org/record/210056/files/figure.png" pageId="8" pageNumber="57" targetBox="[153,1435,686,1147]" targetPageId="8">
<paragraph id="3716DD21FF877D46FF61A2A43675FB64" blockId="8.[151,1319,1181,1204]" box="[151,1319,1181,1204]" pageId="8" pageNumber="57">
<emphasis id="05DD0133FF877D46FF61A2A43240FB63" bold="true" box="[151,274,1181,1204]" pageId="8" pageNumber="57">FIGURE 3.</emphasis>
Reconstruction of major (A) and minor (B) chelae of
<taxonomicName id="F0A9A6A2FF877D46FCC1A2A737E2FB64" authority="Brocchi, 1883" authorityName="Brocchi" authorityYear="1883" box="[823,1200,1182,1204]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="chalmasii" status="comb. nov.">
<emphasis id="05DD0133FF877D46FCC1A2A73759FB64" box="[823,1035,1182,1204]" italics="true" pageId="8" pageNumber="57">Calliaxina chalmasii</emphasis>
(Brocchi, 1883)
</taxonomicName>
<taxonomicNameLabel id="1EEEBC48FF877D46FB41A2A73675FB64" box="[1207,1319,1182,1204]" pageId="8" pageNumber="57">comb. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3716DD21FF877D46FF33A2E23764FA8F" blockId="8.[151,1437,1243,1988]" pageId="8" pageNumber="57">
<emphasis id="05DD0133FF877D46FF33A2E23216FB24" bold="true" box="[197,324,1243,1268]" pageId="8" pageNumber="57">Variations.</emphasis>
The material shows a degree of variation in certain characters present on propodus. The length/ height ratio is variable; both forms with distinctly longer (
<figureCitation id="AF92C1A4FF877D46FCC5A2C630D1FAC7" box="[819,899,1279,1303]" captionStart="FIGURE 4" captionStartId="10.[151,250,1580,1603]" captionTargetBox="[151,1435,481,1559]" captionTargetId="figure@10.[151,1436,481,1559]" captionTargetPageId="10" captionText="FIGURE 4. Calliaxina chalmasii (Brocchi, 1883) new combination, Budapest area (Great Hungarian Basin, Hungary): A, right major propodus (inner and outer view), PAL 2011.36; B, right major propodus (inner and outer view), KGP-MH RA- 033; C, right major propodus, PAL 2011.34; D, right major propodus, PAL 2011.35; E, left major propodus (outer and inner view), PAL 2011.37; F, left major propodus (outer and inner view), PAL 2011.33; G, left major propodus (outer and inner view), KGP- MH RA- 034; H, left major propodus, KGP-MH RA- 021; I, right major propodus, KGP-MH RA- 023; J, left major propodus, KGP-MH RA- 020; K, left minor propodus (“ short ” morphotype), KGP-MH RA- 036; L, left minor propodus, PAL 2011.30; M, left minor propodus, PAL 2011.32; N, left minor propodus articulated with dactylus (outer and inner view), PAL 2011.29; O, left minor propodus, KGP-MH RA- 035; P, left minor propodus (outer and inner view), PAL 2011.31; Q, right carpus, KGP-MH RA- 031; R, right carpus (outer and inner view), KGP-MH RA- 028. All specimens are to scale." httpUri="https://zenodo.org/record/210057/files/figure.png" pageId="8" pageNumber="57">Figs. 4</figureCitation>
E, I) or shorter propodus (
<figureCitation id="AF92C1A4FF877D46FB58A2C637A1FAC7" box="[1198,1267,1279,1303]" captionStart="FIGURE 4" captionStartId="10.[151,250,1580,1603]" captionTargetBox="[151,1435,481,1559]" captionTargetId="figure@10.[151,1436,481,1559]" captionTargetPageId="10" captionText="FIGURE 4. Calliaxina chalmasii (Brocchi, 1883) new combination, Budapest area (Great Hungarian Basin, Hungary): A, right major propodus (inner and outer view), PAL 2011.36; B, right major propodus (inner and outer view), KGP-MH RA- 033; C, right major propodus, PAL 2011.34; D, right major propodus, PAL 2011.35; E, left major propodus (outer and inner view), PAL 2011.37; F, left major propodus (outer and inner view), PAL 2011.33; G, left major propodus (outer and inner view), KGP- MH RA- 034; H, left major propodus, KGP-MH RA- 021; I, right major propodus, KGP-MH RA- 023; J, left major propodus, KGP-MH RA- 020; K, left minor propodus (“ short ” morphotype), KGP-MH RA- 036; L, left minor propodus, PAL 2011.30; M, left minor propodus, PAL 2011.32; N, left minor propodus articulated with dactylus (outer and inner view), PAL 2011.29; O, left minor propodus, KGP-MH RA- 035; P, left minor propodus (outer and inner view), PAL 2011.31; Q, right carpus, KGP-MH RA- 031; R, right carpus (outer and inner view), KGP-MH RA- 028. All specimens are to scale." httpUri="https://zenodo.org/record/210057/files/figure.png" pageId="8" pageNumber="57">Fig. 4</figureCitation>
K) are present, although the majority of material exhibits more-less equal ratio (e.i. quadrate manus) (
<tableCitation id="7A2BE89AFF877D46FB92A31A37EEFAEB" box="[1124,1212,1315,1339]" captionStart="TABLE 2" captionStartId="9.[151,235,152,173]" captionTargetBox="[151,1436,262,1814]" captionTargetPageId="9" captionText="TABLE 2. Measurements (mm) of Calliaxina chalmasii (Brocchi, 1883) comb. nov. Abbreviations: M = major, m = minor,? = unknown; handedness: L = left; R = right; max. L = maximum length of the element, max. H. = maximum height of the element; measurements: ~ = approximate value." httpUri="http://table.plazi.org/id/63D68DA9FF867D47FF61A6A13173FF3B" pageId="8" pageNumber="57" tableUuid="63D68DA9FF867D47FF61A6A13173FF3B">Table 2</tableCitation>
). Short morphotype has already been depicted, but not commented upon by
<bibRefCitation id="5338A0D0FF877D46FCFEA37E3760FA8F" author="Muller" box="[776,1074,1351,1375]" pageId="8" pageNumber="57" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984: pl. 2, fig. 3)</bibRefCitation>
.
</paragraph>
<paragraph id="3716DD21FF877D46FF61A35531D6FA77" blockId="8.[151,1437,1243,1988]" pageId="8" pageNumber="57">
Lower margin of minor propodus is sometimes concave at its distal end (
<figureCitation id="AF92C1A4FF877D46FC49A352375EFA53" box="[959,1036,1387,1411]" captionStart="FIGURE 4" captionStartId="10.[151,250,1580,1603]" captionTargetBox="[151,1435,481,1559]" captionTargetId="figure@10.[151,1436,481,1559]" captionTargetPageId="10" captionText="FIGURE 4. Calliaxina chalmasii (Brocchi, 1883) new combination, Budapest area (Great Hungarian Basin, Hungary): A, right major propodus (inner and outer view), PAL 2011.36; B, right major propodus (inner and outer view), KGP-MH RA- 033; C, right major propodus, PAL 2011.34; D, right major propodus, PAL 2011.35; E, left major propodus (outer and inner view), PAL 2011.37; F, left major propodus (outer and inner view), PAL 2011.33; G, left major propodus (outer and inner view), KGP- MH RA- 034; H, left major propodus, KGP-MH RA- 021; I, right major propodus, KGP-MH RA- 023; J, left major propodus, KGP-MH RA- 020; K, left minor propodus (“ short ” morphotype), KGP-MH RA- 036; L, left minor propodus, PAL 2011.30; M, left minor propodus, PAL 2011.32; N, left minor propodus articulated with dactylus (outer and inner view), PAL 2011.29; O, left minor propodus, KGP-MH RA- 035; P, left minor propodus (outer and inner view), PAL 2011.31; Q, right carpus, KGP-MH RA- 031; R, right carpus (outer and inner view), KGP-MH RA- 028. All specimens are to scale." httpUri="https://zenodo.org/record/210057/files/figure.png" pageId="8" pageNumber="57">Figs. 4</figureCitation>
M, P). In such cases the fixed finger bears a distinct tooth on its occlusal margin.
</paragraph>
</subSubSection>
<subSubSection id="7FB38EAAFF877D44FF33A38A3753FE7C" lastPageId="10" lastPageNumber="59" pageId="8" pageNumber="57" type="materials_examined">
<paragraph id="3716DD21FF877D46FF33A38A319CF8E3" blockId="8.[151,1437,1243,1988]" pageId="8" pageNumber="57">
<emphasis id="05DD0133FF877D46FF33A38A32E2FA1C" bold="true" box="[197,432,1459,1484]" pageId="8" pageNumber="57">Material examined.</emphasis>
<typeStatus id="E8126383FF877D46FE4EA38A3172FA1B" box="[440,544,1459,1483]" pageId="8" pageNumber="57" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="F0A9A6A2FF877D46FDBEA38D3007FA1C" box="[584,853,1460,1484]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="rakosiensis">
<emphasis id="05DD0133FF877D46FDBEA38D3007FA1C" box="[584,853,1460,1484]" italics="true" pageId="8" pageNumber="57">Callianassa rakosiensis</emphasis>
</taxonomicName>
representing right propodus (FI 29);
<bibRefCitation id="5338A0D0FF877D46FB08A38A33ECFA3F" author="Muller" pageId="8" pageNumber="57" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984: 51)</bibRefCitation>
stated that it had been probably lost, however, it was found during a visit of the Hungarian Geological Institute. The
<typeStatus id="E8126383FF877D46FF3DA3C2327FF9C3" box="[203,301,1531,1555]" pageId="8" pageNumber="57" type="holotype">holotype</typeStatus>
of
<taxonomicName id="F0A9A6A2FF877D46FEA2A3C532B0F9C4" box="[340,482,1532,1556]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="chalmasii">
<emphasis id="05DD0133FF877D46FEA2A3C532B0F9C4" box="[340,482,1532,1556]" italics="true" pageId="8" pageNumber="57">C. chalmasii</emphasis>
</taxonomicName>
has not been found and at present it should be considered to be lost as already stated by
<bibRefCitation id="5338A0D0FF877D46FF4AA02632DBF9E7" author="Muller" box="[188,393,1567,1591]" pageId="8" pageNumber="57" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984: 51)</bibRefCitation>
. Material from Rákos collected by P. Müller and deposited at the Hungarian Natural History Museum comprises the right propodus of
<taxonomicName id="F0A9A6A2FF877D46FD76A07D3070F98C" box="[640,802,1604,1628]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="rakosiensis">
<emphasis id="05DD0133FF877D46FD76A07D3070F98C" box="[640,802,1604,1628]" italics="true" pageId="8" pageNumber="57">C. rakosiensis</emphasis>
</taxonomicName>
(M.86.244, illustrated also by
<bibRefCitation id="5338A0D0FF877D46FB7DA07A364CF98B" author="Muller" box="[1163,1310,1603,1627]" pageId="8" pageNumber="57" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller 1984</bibRefCitation>
: pl. 3, figs 12.) and several isolated propodi of
<taxonomicName id="F0A9A6A2FF877D46FDC8A051319EF9AF" box="[574,716,1639,1663]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="chalmasii">
<emphasis id="05DD0133FF877D46FDC8A051319EF9AF" box="[574,716,1639,1663]" italics="true" pageId="8" pageNumber="57">C. chalmasii</emphasis>
</taxonomicName>
(M.86.275). Newer material collected by P. Müller is deposited under numbers PAL 2011.29, PAL 2011.3237 (locality Rákos), PAL 2011.30 (Örs vezér tere), and PAL 2011.31 (Gyakorló út) (
<figureCitation id="AF92C1A4FF877D46FEB8A09632C8F917" box="[334,410,1711,1735]" captionStart="FIGURE 4" captionStartId="10.[151,250,1580,1603]" captionTargetBox="[151,1435,481,1559]" captionTargetId="figure@10.[151,1436,481,1559]" captionTargetPageId="10" captionText="FIGURE 4. Calliaxina chalmasii (Brocchi, 1883) new combination, Budapest area (Great Hungarian Basin, Hungary): A, right major propodus (inner and outer view), PAL 2011.36; B, right major propodus (inner and outer view), KGP-MH RA- 033; C, right major propodus, PAL 2011.34; D, right major propodus, PAL 2011.35; E, left major propodus (outer and inner view), PAL 2011.37; F, left major propodus (outer and inner view), PAL 2011.33; G, left major propodus (outer and inner view), KGP- MH RA- 034; H, left major propodus, KGP-MH RA- 021; I, right major propodus, KGP-MH RA- 023; J, left major propodus, KGP-MH RA- 020; K, left minor propodus (“ short ” morphotype), KGP-MH RA- 036; L, left minor propodus, PAL 2011.30; M, left minor propodus, PAL 2011.32; N, left minor propodus articulated with dactylus (outer and inner view), PAL 2011.29; O, left minor propodus, KGP-MH RA- 035; P, left minor propodus (outer and inner view), PAL 2011.31; Q, right carpus, KGP-MH RA- 031; R, right carpus (outer and inner view), KGP-MH RA- 028. All specimens are to scale." httpUri="https://zenodo.org/record/210057/files/figure.png" pageId="8" pageNumber="57">Fig. 4</figureCitation>
). Material from Rákos collected by the author is deposited under numbers KGP-MH RA020037 (
<figureCitation id="AF92C1A4FF877D46FEC2A0EA322EF93B" box="[308,380,1747,1771]" captionStart="FIGURE 4" captionStartId="10.[151,250,1580,1603]" captionTargetBox="[151,1435,481,1559]" captionTargetId="figure@10.[151,1436,481,1559]" captionTargetPageId="10" captionText="FIGURE 4. Calliaxina chalmasii (Brocchi, 1883) new combination, Budapest area (Great Hungarian Basin, Hungary): A, right major propodus (inner and outer view), PAL 2011.36; B, right major propodus (inner and outer view), KGP-MH RA- 033; C, right major propodus, PAL 2011.34; D, right major propodus, PAL 2011.35; E, left major propodus (outer and inner view), PAL 2011.37; F, left major propodus (outer and inner view), PAL 2011.33; G, left major propodus (outer and inner view), KGP- MH RA- 034; H, left major propodus, KGP-MH RA- 021; I, right major propodus, KGP-MH RA- 023; J, left major propodus, KGP-MH RA- 020; K, left minor propodus (“ short ” morphotype), KGP-MH RA- 036; L, left minor propodus, PAL 2011.30; M, left minor propodus, PAL 2011.32; N, left minor propodus articulated with dactylus (outer and inner view), PAL 2011.29; O, left minor propodus, KGP-MH RA- 035; P, left minor propodus (outer and inner view), PAL 2011.31; Q, right carpus, KGP-MH RA- 031; R, right carpus (outer and inner view), KGP-MH RA- 028. All specimens are to scale." httpUri="https://zenodo.org/record/210057/files/figure.png" pageId="8" pageNumber="57">Fig. 4</figureCitation>
). Additional material comprises five specimens from the Fenk quarry (NHMW-2011-0167- 01160120) (
<figureCitation id="AF92C1A4FF877D46FED9A0CE3223F8DF" box="[303,369,1783,1807]" captionStart="FIGURE 5" captionStartId="11.[151,250,1190,1213]" captionTargetBox="[151,1435,337,1168]" captionTargetId="figure@11.[151,1436,337,1169]" captionTargetPageId="11" captionText="FIGURE 5. Calliaxina chalmasii (Brocchi, 1883) new combination, Fenk quarry (Vienna Basin, Austria): A, left minor propodus articulated with dactylus, NHMW- 2011 - 0167 - 0118; B, right major propodus, NHMW- 2011 - 0167 - 0120 a; C, left major propodus, NHMW- 2011 - 0167 - 0120 b; D, right minor propodus, NHMW- 2011 - 0167 - 0117; E, right carpus, NHMW- 2011 - 0167 - 0116. All specimens are to scale and were coated with ammonium chloride prior to photography." httpUri="https://zenodo.org/record/210058/files/figure.png" pageId="8" pageNumber="57">Fig. 5</figureCitation>
) and several propodal fragments (one articulated with carpus) from Dúbravská hlavica (SNM- Z 37570, 37571); KGP-MH DH074076) (
<figureCitation id="AF92C1A4FF877D46FD8DA1223193F8E3" box="[635,705,1819,1843]" captionStart="FIGURE 6" captionStartId="12.[151,250,1276,1299]" captionTargetBox="[151,1435,339,1254]" captionTargetId="figure@12.[151,1436,339,1255]" captionTargetPageId="12" captionText="FIGURE 6. Calliaxina chalmasii (Brocchi, 1883) new combination, Dúbravská hlavica (Vienna Basin, Slovakia): A, left major propodus, KGP-MH DH- 074; B, left major propodus, SNM-Z 37570; C, left minor propodus, KGP-MH DH- 075; D, right minor propodus articulated with carpus, SNM-Z 37571; E, fragmented right pollex (from right chela). All specimens are to scale and were coated with ammonium chloride prior to photography." httpUri="https://zenodo.org/record/210059/files/figure.png" pageId="8" pageNumber="57">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="3716DD21FF877D46FF31A1793758F887" blockId="8.[151,1437,1243,1988]" box="[199,1034,1855,1879]" pageId="8" pageNumber="57">
Measurements and details about the specimens are summarized in
<tableCitation id="7A2BE89AFF877D46FC46A1063754F887" box="[944,1030,1855,1879]" captionStart="TABLE 2" captionStartId="9.[151,235,152,173]" captionTargetBox="[151,1436,262,1814]" captionTargetPageId="9" captionText="TABLE 2. Measurements (mm) of Calliaxina chalmasii (Brocchi, 1883) comb. nov. Abbreviations: M = major, m = minor,? = unknown; handedness: L = left; R = right; max. L = maximum length of the element, max. H. = maximum height of the element; measurements: ~ = approximate value." httpUri="http://table.plazi.org/id/63D68DA9FF867D47FF61A6A13173FF3B" pageId="8" pageNumber="57" tableUuid="63D68DA9FF867D47FF61A6A13173FF3B">Table 2</tableCitation>
.
</paragraph>
<paragraph id="3716DD21FF877D46FF31A15A3005F813" blockId="8.[151,1437,1243,1988]" pageId="8" pageNumber="57">
<emphasis id="05DD0133FF877D46FF31A15A32DAF8AC" bold="true" box="[199,392,1891,1916]" pageId="8" pageNumber="57">Other material.</emphasis>
For comparative purposes, the extant material of several
<taxonomicName id="F0A9A6A2FF877D46FBB3A15A378EF8AB" box="[1093,1244,1891,1915]" class="Malacostraca" family="Callianassidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="subFamily" subFamily="Eucalliacinae">Eucalliacinae</taxonomicName>
taxa have been examined (all deposited in
<collectionCode id="51B845E4FF877D46FE29A1BE316DF84F" box="[479,575,1927,1951]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="57">NHMW</collectionCode>
), namely
<taxonomicName id="F0A9A6A2FF877D46FD4EA1BE30A0F84F" box="[696,1010,1927,1951]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="novaebritanniae">
<emphasis id="05DD0133FF877D46FD4EA1BE30A0F84F" box="[696,1010,1927,1951]" italics="true" pageId="8" pageNumber="57">Calliaxina novaebritanniae</emphasis>
</taxonomicName>
(
<collectionCode id="51B845E4FF877D46FBF1A1BE3737F84F" box="[1031,1125,1927,1951]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="57">NHMW</collectionCode>
25399, 25400);
<taxonomicName id="F0A9A6A2FF877D46FAC7A1BE3241F814" class="Malacostraca" family="Callianassidae" genus="Eucalliax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="aequimana">
<emphasis id="05DD0133FF877D46FAC7A1BE3241F814" italics="true" pageId="8" pageNumber="57">Eucalliax aequimana</emphasis>
</taxonomicName>
(
<collectionCode id="51B845E4FF877D46FED4A19232D2F813" box="[290,384,1963,1987]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="57">NHMW</collectionCode>
19365); and
<taxonomicName id="F0A9A6A2FF877D46FDE2A19431DFF814" box="[532,653,1964,1988]" class="Malacostraca" family="Callianassidae" genus="Eucalliax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="05DD0133FF877D46FDE2A19431DFF814" box="[532,653,1964,1988]" italics="true" pageId="8" pageNumber="57">E. kensleyi</emphasis>
</taxonomicName>
(
<collectionCode id="51B845E4FF877D46FD6AA19231A8F813" box="[668,762,1963,1987]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="57">NHMW</collectionCode>
16779).
</paragraph>
<caption id="63D68DA9FF867D47FF61A6A13173FF3B" ID-Table-UUID="63D68DA9FF867D47FF61A6A13173FF3B" httpUri="http://table.plazi.org/id/63D68DA9FF867D47FF61A6A13173FF3B" pageId="9" pageNumber="58" targetBox="[151,1436,262,1814]" targetIsTable="true" targetPageId="9">
<paragraph id="3716DD21FF867D47FF61A6A13173FF3B" blockId="9.[151,1436,152,235]" pageId="9" pageNumber="58">
<emphasis id="05DD0133FF867D47FF61A6A13256FF7D" bold="true" box="[151,260,152,173]" pageId="9" pageNumber="58">TABLE 2.</emphasis>
Measurements (mm) of
<taxonomicName id="F0A9A6A2FF867D47FE0BA6A13024FF7E" authority="Brocchi, 1883" authorityName="Brocchi" authorityYear="1883" box="[509,886,152,174]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="58" phylum="Arthropoda" rank="species" species="chalmasii" status="comb. nov.">
<emphasis id="05DD0133FF867D47FE0BA6A1313AFF7E" box="[509,616,152,174]" italics="true" pageId="9" pageNumber="58">Calliaxina</emphasis>
<emphasis id="05DD0133FF867D47FD98A6A13183FF7E" box="[622,721,152,174]" italics="true" pageId="9" pageNumber="58">chalmasii</emphasis>
(Brocchi, 1883)
</taxonomicName>
<taxonomicNameLabel id="1EEEBC48FF867D47FC8BA6A130BFFF7E" box="[893,1005,152,174]" pageId="9" pageNumber="58">comb. nov.</taxonomicNameLabel>
Abbreviations: M =
<taxonomicName id="F0A9A6A2FF867D47FB36A6A137AFFF7E" box="[1216,1277,152,174]" pageId="9" pageNumber="58" rank="species" species="major">major</taxonomicName>
, m = minor,? = unknown; handedness: L = left; R = right; max.L = maximum length of the element, max.H. = maximum height of the element; measurements: ~ = approximate value.
</paragraph>
</caption>
<paragraph id="3716DD21FF867D47FF6AA73F367CF8C5" pageId="9" pageNumber="58">
<table id="45A92F81FF8682B1FF61A73F36CEF8C6" box="[151,1436,262,1814]" gridcols="6" gridrows="9" pageId="9" pageNumber="58">
<tr id="8999DF63FF8682B1FF61A73F36CEFECC" box="[151,1436,262,284]" gridrow="0" pageId="9" pageNumber="58">
<th id="CA48B61FFF8682B1FF61A73F32F8FECC" box="[151,426,262,284]" gridcol="0" gridrow="0" pageId="9" pageNumber="58">Specimen</th>
<th id="CA48B61FFF8682B1FD95A73F31C7FECC" box="[611,661,262,284]" gridcol="1" gridrow="0" pageId="9" pageNumber="58">M/m</th>
<th id="CA48B61FFF8682B1FD2AA73F3003FECC" box="[732,849,262,284]" gridcol="2" gridrow="0" pageId="9" pageNumber="58">handedness</th>
<th id="CA48B61FFF8682B1FC35A73F3770FECC" box="[963,1058,262,284]" gridcol="3" gridrow="0" pageId="9" pageNumber="58">element</th>
<th id="CA48B61FFF8682B1FB8AA73F37EFFECC" box="[1148,1213,262,284]" gridcol="4" gridrow="0" pageId="9" pageNumber="58">max.L</th>
<th id="CA48B61FFF8682B1FAF9A73F36CEFECC" box="[1295,1436,262,284]" gridcol="5" gridrow="0" pageId="9" pageNumber="58">max.H</th>
</tr>
<tr id="8999DF63FF8682B1FF61A70936CEFDC4" box="[151,1436,304,532]" gridrow="1" pageId="9" pageNumber="58">
<th id="CA48B61FFF8682B1FF61A70932F8FDC4" box="[151,426,304,532]" gridcol="0" gridrow="1" pageId="9" pageNumber="58">KGP-MH DH-074 KGP-MH DH-075 KGP-MH RA-020 KGP-MH RA-021 KGP-MH RA-022 KGP-MH RA-023</th>
<td id="CA48B61FFF8682B1FD95A70931C7FDC4" box="[611,661,304,532]" gridcol="1" gridrow="1" pageId="9" pageNumber="58">M M M M M M</td>
<td id="CA48B61FFF8682B1FD2AA7093003FDC4" box="[732,849,304,532]" gridcol="2" gridrow="1" pageId="9" pageNumber="58">L L L L L R</td>
<td id="CA48B61FFF8682B1FC35A7093770FDC4" box="[963,1058,304,532]" gridcol="3" gridrow="1" pageId="9" pageNumber="58">propodus propodus propodus propodus propodus propodus</td>
<td id="CA48B61FFF8682B1FB8AA70937EFFDC4" box="[1148,1213,304,532]" gridcol="4" gridrow="1" pageId="9" pageNumber="58">8.0 6.2 10.0 ~7.5 6.0 4.4</td>
<td id="CA48B61FFF8682B1FAF9A70936CEFDC4" box="[1295,1436,304,532]" gridcol="5" gridrow="1" pageId="9" pageNumber="58">7.1 6.7 8.5 7.3 5.5 3.6</td>
</tr>
<tr id="8999DF63FF8682B1FF61A41136CEFCDC" box="[151,1436,552,780]" gridrow="2" pageId="9" pageNumber="58">
<th id="CA48B61FFF8682B1FF61A41132F8FCDC" box="[151,426,552,780]" gridcol="0" gridrow="2" pageId="9" pageNumber="58">KGP-MH RA-024 KGP-MH RA-025 KGP-MH RA-026 KGP-MH RA-027 KGP-MH RA-028 KGP-MH RA-029</th>
<td id="CA48B61FFF8682B1FD95A41131C7FCDC" box="[611,661,552,780]" gridcol="1" gridrow="2" pageId="9" pageNumber="58">M M M m??</td>
<td id="CA48B61FFF8682B1FD2AA4113003FCDC" box="[732,849,552,780]" gridcol="2" gridrow="2" pageId="9" pageNumber="58">L R R R R L</td>
<td id="CA48B61FFF8682B1FC35A4113770FCDC" box="[963,1058,552,780]" gridcol="3" gridrow="2" pageId="9" pageNumber="58">propodus propodus propodus propodus carpus propodus</td>
<td id="CA48B61FFF8682B1FB8AA41137EFFCDC" box="[1148,1213,552,780]" gridcol="4" gridrow="2" pageId="9" pageNumber="58">6.0 4.0 4.6 7.2 5.3 ~6.0</td>
<td id="CA48B61FFF8682B1FAF9A41136CEFCDC" box="[1295,1436,552,780]" gridcol="5" gridrow="2" pageId="9" pageNumber="58">4.3 3.4 4.5 7.0 5.8 ~6.3</td>
</tr>
<tr id="8999DF63FF8682B1FF61A51936CEFC8F" box="[151,1436,800,863]" gridrow="3" pageId="9" pageNumber="58">
<th id="CA48B61FFF8682B1FF61A51932F8FC8F" box="[151,426,800,863]" gridcol="0" gridrow="3" pageId="9" pageNumber="58">KGP-MH RA-030 KGP-MH RA-031</th>
<td id="CA48B61FFF8682B1FD95A51931C7FC8F" box="[611,661,800,863]" gridcol="1" gridrow="3" pageId="9" pageNumber="58">??</td>
<td id="CA48B61FFF8682B1FD2AA5193003FC8F" box="[732,849,800,863]" gridcol="2" gridrow="3" pageId="9" pageNumber="58">R R</td>
<td id="CA48B61FFF8682B1FC35A5193770FC8F" box="[963,1058,800,863]" gridcol="3" gridrow="3" pageId="9" pageNumber="58">carpus carpus</td>
<td id="CA48B61FFF8682B1FB8AA51937EFFC8F" box="[1148,1213,800,863]" gridcol="4" gridrow="3" pageId="9" pageNumber="58">5.0 5.5</td>
<td id="CA48B61FFF8682B1FAF9A51936CEFC8F" box="[1295,1436,800,863]" gridcol="5" gridrow="3" pageId="9" pageNumber="58">5.7 6.0</td>
</tr>
<tr id="8999DF63FF8682B1FF61A54A36CEFBD4" box="[151,1436,883,1028]" gridrow="4" pageId="9" pageNumber="58">
<th id="CA48B61FFF8682B1FF61A54A32F8FBD4" box="[151,426,883,1028]" gridcol="0" gridrow="4" pageId="9" pageNumber="58">KGP-MH RA-032 KGP-MH RA-033 KGP-MH RA-034 KGP-MH RA-035</th>
<td id="CA48B61FFF8682B1FD95A54A31C7FBD4" box="[611,661,883,1028]" gridcol="1" gridrow="4" pageId="9" pageNumber="58">? M M m</td>
<td id="CA48B61FFF8682B1FD2AA54A3003FBD4" box="[732,849,883,1028]" gridcol="2" gridrow="4" pageId="9" pageNumber="58">L R L L</td>
<td id="CA48B61FFF8682B1FC35A54A3770FBD4" box="[963,1058,883,1028]" gridcol="3" gridrow="4" pageId="9" pageNumber="58">carpus propodus propodus propodus</td>
<td id="CA48B61FFF8682B1FB8AA54A37EFFBD4" box="[1148,1213,883,1028]" gridcol="4" gridrow="4" pageId="9" pageNumber="58">3.2 6.0 5.6 7.6</td>
<td id="CA48B61FFF8682B1FAF9A54A36CEFBD4" box="[1295,1436,883,1028]" gridcol="5" gridrow="4" pageId="9" pageNumber="58">3.7 6.3 5.7 7.7</td>
</tr>
<tr id="8999DF63FF8682B1FF61A22136CEFB2C" box="[151,1436,1048,1276]" gridrow="5" pageId="9" pageNumber="58">
<th id="CA48B61FFF8682B1FF61A22132F8FB2C" box="[151,426,1048,1276]" gridcol="0" gridrow="5" pageId="9" pageNumber="58">KGP-MH RA-036 KGP-MH RA-037 NHMW-2011-0167-0116 NHMW-2011-0167-0117 NHMW-2011-0167-0118 NHMW-2011-0167-0120a</th>
<td id="CA48B61FFF8682B1FD95A22131C7FB2C" box="[611,661,1048,1276]" gridcol="1" gridrow="5" pageId="9" pageNumber="58">m m? m m M</td>
<td id="CA48B61FFF8682B1FD2AA2213003FB2C" box="[732,849,1048,1276]" gridcol="2" gridrow="5" pageId="9" pageNumber="58">L R R R L R</td>
<td id="CA48B61FFF8682B1FC35A2213770FB2C" box="[963,1058,1048,1276]" gridcol="3" gridrow="5" pageId="9" pageNumber="58">propodus propodus carpus propodus propodus propodus</td>
<td id="CA48B61FFF8682B1FB8AA22137EFFB2C" box="[1148,1213,1048,1276]" gridcol="4" gridrow="5" pageId="9" pageNumber="58">5.0 6.8 5.7 5.1 7.3 7.0</td>
<td id="CA48B61FFF8682B1FAF9A22136CEFB2C" box="[1295,1436,1048,1276]" gridcol="5" gridrow="5" pageId="9" pageNumber="58">6.0 6.2 5.6 5.0 7.3 6.7</td>
</tr>
<tr id="8999DF63FF8682B1FF61A32936CEFA24" box="[151,1436,1296,1524]" gridrow="6" pageId="9" pageNumber="58">
<th id="CA48B61FFF8682B1FF61A32932F8FA24" box="[151,426,1296,1524]" gridcol="0" gridrow="6" pageId="9" pageNumber="58">NHMW-2011-0167-0120b PAL 2011.29 PAL 2011.30 PAL 2011.31 PAL 2011.32 PAL 2011.33</th>
<td id="CA48B61FFF8682B1FD95A32931C7FA24" box="[611,661,1296,1524]" gridcol="1" gridrow="6" pageId="9" pageNumber="58">M m m m m M</td>
<td id="CA48B61FFF8682B1FD2AA3293003FA24" box="[732,849,1296,1524]" gridcol="2" gridrow="6" pageId="9" pageNumber="58">L L L L L L</td>
<td id="CA48B61FFF8682B1FC35A3293770FA24" box="[963,1058,1296,1524]" gridcol="3" gridrow="6" pageId="9" pageNumber="58">propodus propodus propodus propodus propodus propodus</td>
<td id="CA48B61FFF8682B1FB8AA32937EFFA24" box="[1148,1213,1296,1524]" gridcol="4" gridrow="6" pageId="9" pageNumber="58">3.2 6.3 ~4.5 8.6 5.0 5.6</td>
<td id="CA48B61FFF8682B1FAF9A32936CEFA24" box="[1295,1436,1296,1524]" gridcol="5" gridrow="6" pageId="9" pageNumber="58">3.0 6.4 4.2 8.3 4.5 5.4</td>
</tr>
<tr id="8999DF63FF8682B1FF61A03136CEF93C" box="[151,1436,1544,1772]" gridrow="7" pageId="9" pageNumber="58">
<th id="CA48B61FFF8682B1FF61A03132F8F93C" box="[151,426,1544,1772]" gridcol="0" gridrow="7" pageId="9" pageNumber="58">PAL 2011.34 PAL 2011.35 PAL 2011.36 PAL 2011.37 SNM Z-37570 SNM Z-37570</th>
<td id="CA48B61FFF8682B1FD95A03131C7F93C" box="[611,661,1544,1772]" gridcol="1" gridrow="7" pageId="9" pageNumber="58">M M M M m m</td>
<td id="CA48B61FFF8682B1FD2AA0313003F93C" box="[732,849,1544,1772]" gridcol="2" gridrow="7" pageId="9" pageNumber="58">R R R L R R</td>
<td id="CA48B61FFF8682B1FC35A0313770F93C" box="[963,1058,1544,1772]" gridcol="3" gridrow="7" pageId="9" pageNumber="58">propodus propodus propodus propodus propodus carpus</td>
<td id="CA48B61FFF8682B1FB8AA03137EFF93C" box="[1148,1213,1544,1772]" gridcol="4" gridrow="7" pageId="9" pageNumber="58">6.5 6.4 7.0 6.4 6.6 4.5</td>
<td id="CA48B61FFF8682B1FAF9A03136CEF93C" box="[1295,1436,1544,1772]" gridcol="5" gridrow="7" pageId="9" pageNumber="58">7.0 7.0 7.4 5.7 6.4 6.3</td>
</tr>
<tr id="8999DF63FF8682B1FF61A13936CEF8C6" box="[151,1436,1792,1814]" gridrow="8" pageId="9" pageNumber="58">
<th id="CA48B61FFF8682B1FF61A13932F8F8C6" box="[151,426,1792,1814]" gridcol="0" gridrow="8" pageId="9" pageNumber="58">SNM Z-375701</th>
<td id="CA48B61FFF8682B1FD95A13931C7F8C6" box="[611,661,1792,1814]" gridcol="1" gridrow="8" pageId="9" pageNumber="58">M</td>
<td id="CA48B61FFF8682B1FD2AA1393003F8C6" box="[732,849,1792,1814]" gridcol="2" gridrow="8" pageId="9" pageNumber="58">L</td>
<td id="CA48B61FFF8682B1FC35A1393770F8C6" box="[963,1058,1792,1814]" gridcol="3" gridrow="8" pageId="9" pageNumber="58">propodus</td>
<td id="CA48B61FFF8682B1FB8AA13937EFF8C6" box="[1148,1213,1792,1814]" gridcol="4" gridrow="8" pageId="9" pageNumber="58">8.5</td>
<td id="CA48B61FFF8682B1FAF9A13936CEF8C6" box="[1295,1436,1792,1814]" gridcol="5" gridrow="8" pageId="9" pageNumber="58">8.8</td>
</tr>
</table>
</paragraph>
<paragraph id="3716DD21FF867D44FF31A1633753FE7C" blockId="9.[151,1436,1882,2014]" lastBlockId="10.[151,1436,151,428]" lastPageId="10" lastPageNumber="59" pageId="9" pageNumber="58">
<emphasis id="05DD0133FF867D47FF31A163320BF8A3" bold="true" box="[199,345,1882,1907]" pageId="9" pageNumber="58">Occurrence.</emphasis>
The oldest occurrence of
<taxonomicName id="F0A9A6A2FF867D47FD7FA1623027F8A3" box="[649,885,1883,1907]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="58" phylum="Arthropoda" rank="species" species="chalmasii" status="comb. nov.">
<emphasis id="05DD0133FF867D47FD7FA1623027F8A3" box="[649,885,1883,1907]" italics="true" pageId="9" pageNumber="58">Calliaxina chalmasii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EEEBC48FF867D47FC88A16330ADF8A2" box="[894,1023,1882,1906]" pageId="9" pageNumber="58">comb. nov.</taxonomicNameLabel>
, as reported herein, is that from the Fenk quarry, Gross Höflein,
<collectingCountry id="4FBE9DB1FF867D47FE05A147311AF846" box="[499,584,1918,1942]" name="Austria" pageId="9" pageNumber="58">Austria</collectingCountry>
, which is middle 'Badenian' (upper Langhian) in age. From the upper 'Badenian' (lower Serravallian), the species is known from the surroundings of the Budapest area,
<collectingCountry id="4FBE9DB1FF867D47FB28A19A3612F86B" box="[1246,1344,1955,1979]" name="Hungary" pageId="9" pageNumber="58">Hungary</collectingCountry>
(
<bibRefCitation id="5338A0D0FF867D47FAB8A19B3381F80E" author="Muller" pageId="9" pageNumber="58" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller 1984</bibRefCitation>
). It was reported also from the Middle Miocene of
<collectingCountry id="4FBE9DB1FF867D47FCFFA1FF303AF80E" box="[777,872,1990,2014]" name="Portugal" pageId="9" pageNumber="58">Portugal</collectingCountry>
(
<bibRefCitation id="5338A0D0FF867D47FC83A1FF370AF80E" author="Veiga" box="[885,1112,1990,2014]" pageId="9" pageNumber="58" refString="Veiga Ferreira, O. da (1961) Nove especie de Callianassa no Miocenico da Bacia do Tejo. Comunicacoes dos Servicos Geologicos de Portugal, 45, 479 - 481." type="journal article" year="1961">Veiga Ferreira 1961</bibRefCitation>
). Thus, it appears that during Miocene the species was widespread in the Central Paratethys. Since, the Late Miocene, when the seaways between these two palaeogeographic areas were closed and the Central Paratethys became Lake Pannon (
<bibRefCitation id="5338A0D0FF857D44FA92A6853380FF27" author="Rogl" pageId="10" pageNumber="59" refString="Rogl, F. (1998) Palaeogeographic considerations for Mediterranean and Paratethys seaways (Oligocene to Miocene). Annalen des Naturhistorischen Museums in Wien, A 99, 279 - 310." type="journal article" year="1998">Rögl 1998</bibRefCitation>
,
<bibRefCitation id="5338A0D0FF857D44FF12A6E6324DFF27" author="Rogl" box="[228,287,223,247]" pageId="10" pageNumber="59" refString="Rogl, F. (1999) Mediterranean and Paratethys. Facts and hypotheses of an Oligocene to Miocene paleogeography (short overview). Geologica Carpathica, 50, 339 - 349." type="journal article" year="1999">1999</bibRefCitation>
;
<bibRefCitation id="5338A0D0FF857D44FEC4A6E6311AFF27" author="Harzhauser" box="[306,584,223,248]" pageId="10" pageNumber="59" refString="Harzhauser, M., Piller, W. E. &amp; Steininger, F. F. (2002) Circum-Mediterranean Oligo-Miocene biogeographic evolution - the gastropods' point of view. Palaeogeography, Palaeoclimatology, Palaeoecology, 183, 103 - 133." type="journal article" year="2002">
Harzhauser
<emphasis id="05DD0133FF857D44FE49A6D832AAFF28" box="[447,504,224,248]" italics="true" pageId="10" pageNumber="59">et al</emphasis>
. 2002
</bibRefCitation>
;
<bibRefCitation id="5338A0D0FF857D44FDACA6E630C0FF27" author="Harzhauser" box="[602,914,223,247]" pageId="10" pageNumber="59" refString="Harzhauser, M. &amp; Piller, W. E. (2007) Benchmark data of a changing sea - Palaeogeography, Palaeobiogeography and events in the Central Paratethys during the Miocene. Palaeogeography, Palaeoclimatology, Palaeoecology, 253, 8 - 31." type="journal article" year="2007">Harzhauser &amp; Piller 2007</bibRefCitation>
), the species might have migrated into the Mediterranean where it gave rise to extant
<taxonomicName id="F0A9A6A2FF857D44FD72A73D301CFECC" box="[644,846,260,284]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="59" phylum="Arthropoda" rank="species" species="punica">
<emphasis id="05DD0133FF857D44FD72A73D301CFECC" box="[644,846,260,284]" italics="true" pageId="10" pageNumber="59">Calliaxina punica</emphasis>
</taxonomicName>
. Following the chronological approach (see below) the material of
<taxonomicName id="F0A9A6A2FF857D44FEC9A711316BFE90" box="[319,569,296,320]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="59" phylum="Arthropoda" rank="species" species="chalmasii">
<emphasis id="05DD0133FF857D44FEC9A711316BFE90" box="[319,569,296,320]" italics="true" pageId="10" pageNumber="59">Callianassa chalmasii</emphasis>
</taxonomicName>
reported from the Early Pliocene of
<collectingCountry id="4FBE9DB1FF857D44FC26A71E3751FEEF" box="[976,1027,295,319]" name="Italy" pageId="10" pageNumber="59">Italy</collectingCountry>
(
<bibRefCitation id="5338A0D0FF857D44FBE4A71E3655FEEF" author="De" box="[1042,1287,295,320]" pageId="10" pageNumber="59" refString="De Angeli, A., Garassino, A. &amp; Pasini, G. (2009) New reports of anomurans and brachyurans from the Cenozoic of Tuscany (Italy). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 150 (2), 163 - 196." type="journal article" year="2009">
<collectingCountry id="4FBE9DB1FF857D44FBE4A71E3766FEEF" box="[1042,1076,295,319]" name="South Georgia and the South Sandwich Islands" pageId="10" pageNumber="59">De</collectingCountry>
Angeli
<emphasis id="05DD0133FF857D44FB7BA71037ECFE90" box="[1165,1214,296,320]" italics="true" pageId="10" pageNumber="59">et al</emphasis>
. 2009
</bibRefCitation>
;
<bibRefCitation id="5338A0D0FF857D44FAE7A71E33AAFEB4" author="Garassino" pageId="10" pageNumber="59" refString="Garassino, A., Pasini, G., De Angeli, A., Charbonnier, A., Famiani, F., Baldanza, A. &amp; Bizzarri, R. (2012) The decapod community from the Early Pliocene (Zanclean) of &quot; La Serra &quot; quarry (San Miniato, Pisa, Toscana, central Italy): sedimentology, systematics, and palaeoenvironmental implications. Annales de Paleontologie, 98, 1 - 61." type="journal article" year="2012">
Garassino
<emphasis id="05DD0133FF857D44FA7EA71033FFFEB4" italics="true" pageId="10" pageNumber="59">et al</emphasis>
. 2012
</bibRefCitation>
) may in fact represent
<taxonomicName id="F0A9A6A2FF857D44FDFDA7743129FEB4" box="[523,635,333,356]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="59" phylum="Arthropoda" rank="species" species="punica">
<emphasis id="05DD0133FF857D44FDFDA7743129FEB4" box="[523,635,333,356]" italics="true" pageId="10" pageNumber="59">C. punica</emphasis>
</taxonomicName>
, as numerous extant decapod species are known from their Pliocene occurrences. This conclusion may be supported by the reported co-occurrence of presumed
<taxonomicName id="F0A9A6A2FF857D44FB23A749363AFE58" box="[1237,1384,368,392]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="59" phylum="Arthropoda" rank="species" species="chalmasii">
<emphasis id="05DD0133FF857D44FB23A749363AFE58" box="[1237,1384,368,392]" italics="true" pageId="10" pageNumber="59">C. chalmasii</emphasis>
</taxonomicName>
and
<taxonomicName id="F0A9A6A2FF857D44FF61A7AD325FFE7C" box="[151,269,404,428]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="05DD0133FF857D44FF61A7AD325FFE7C" box="[151,269,404,428]" italics="true" pageId="10" pageNumber="59">Calliaxina</emphasis>
</taxonomicName>
cf.
<taxonomicName id="F0A9A6A2FF857D44FECEA7AC32F6FE7C" box="[312,420,405,428]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="59" phylum="Arthropoda" rank="species" species="punica">
<emphasis id="05DD0133FF857D44FECEA7AC32F6FE7C" box="[312,420,405,428]" italics="true" pageId="10" pageNumber="59">C. punica</emphasis>
</taxonomicName>
at one of the Italian localities (
<bibRefCitation id="5338A0D0FF857D44FCF6A7AD30A7FE7C" author="Garassino" box="[768,1013,404,428]" pageId="10" pageNumber="59" refString="Garassino, A., Pasini, G., De Angeli, A., Charbonnier, A., Famiani, F., Baldanza, A. &amp; Bizzarri, R. (2012) The decapod community from the Early Pliocene (Zanclean) of &quot; La Serra &quot; quarry (San Miniato, Pisa, Toscana, central Italy): sedimentology, systematics, and palaeoenvironmental implications. Annales de Paleontologie, 98, 1 - 61." type="journal article" year="2012">
Garassino
<emphasis id="05DD0133FF857D44FC8FA7AC30F8FE7C" box="[889,938,404,428]" italics="true" pageId="10" pageNumber="59">et al</emphasis>
. 2012
</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="63D68DA9FF857D44FF61A015370AF8E8" httpUri="https://zenodo.org/record/210057/files/figure.png" pageId="10" pageNumber="59" targetBox="[151,1435,481,1559]" targetPageId="10">
<paragraph id="3716DD21FF857D44FF61A015370AF8E8" blockId="10.[151,1436,1580,1848]" pageId="10" pageNumber="59">
<emphasis id="05DD0133FF857D44FF61A0153244F992" bold="true" box="[151,278,1580,1603]" pageId="10" pageNumber="59">FIGURE 4.</emphasis>
<taxonomicName id="F0A9A6A2FF857D44FEE9A01431F1F993" authority="Brocchi, 1883" authorityName="Brocchi" authorityYear="1883" box="[287,675,1581,1603]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="59" phylum="Arthropoda" rank="species" species="chalmasii" status="comb. nov.">
<emphasis id="05DD0133FF857D44FEE9A01432D8F993" box="[287,394,1581,1603]" italics="true" pageId="10" pageNumber="59">Calliaxina</emphasis>
<emphasis id="05DD0133FF857D44FE62A01432A5F993" box="[404,503,1581,1603]" italics="true" pageId="10" pageNumber="59">chalmasii</emphasis>
(Brocchi, 1883)
</taxonomicName>
<taxonomicNameLabel id="1EEEBC48FF857D44FD5BA0173030F993" box="[685,866,1581,1603]" pageId="10" pageNumber="59">new combination</taxonomicNameLabel>
, Budapest area (Great Hungarian Basin, Hungary): A, right
<taxonomicName id="F0A9A6A2FF857D44FF3BA072325AF9B1" box="[205,264,1611,1633]" pageId="10" pageNumber="59" rank="species" species="major">major</taxonomicName>
propodus (inner and outer view), PAL 2011.36; B, right
<taxonomicName id="F0A9A6A2FF857D44FCB0A07230D3F9B1" box="[838,897,1611,1633]" pageId="10" pageNumber="59" rank="species" species="major">major</taxonomicName>
propodus (inner and outer view), KGP-MH RA-033; C, right major propodus, PAL 2011.34; D, right
<taxonomicName id="F0A9A6A2FF857D44FD72A05331EDF950" box="[644,703,1642,1664]" pageId="10" pageNumber="59" rank="species" species="major">major</taxonomicName>
propodus, PAL 2011.35; E, left major propodus (outer and inner view), PAL 2011.37; F, left major propodus (outer and inner view), PAL 2011.33; G, left major propodus (outer and inner view), KGP- MH RA-034; H, left major propodus, KGP-MH RA-021; I, right major propodus, KGP-MH RA-023; J, left major propodus, KGP-MH RA-020; K, left minor propodus (“short” morphotype), KGP-MH RA-036; L, left minor propodus, PAL 2011.30; M, left minor propodus, PAL 2011.32; N, left minor propodus articulated with dactylus (outer and inner view), PAL 2011.29; O, left minor propodus, KGP-MH RA-035; P, left minor propodus (outer and inner view), PAL 2011.31; Q, right carpus, KGP-MH RA-031; R, right carpus (outer and inner view), KGP-MH RA-028. All specimens are to scale.
</paragraph>
</caption>
<subSubSection id="7FB38EAAFF857D45FF31A1593129FECC" lastPageId="11" lastPageNumber="60" pageId="10" pageNumber="59" type="discussion">
<paragraph id="3716DD21FF857D45FF31A1593129FECC" blockId="10.[153,1436,1887,2019]" lastBlockId="11.[153,1436,151,284]" lastPageId="11" lastPageNumber="60" pageId="10" pageNumber="59">
<emphasis id="05DD0133FF857D44FF31A1593269F8A8" bold="true" box="[199,315,1888,1912]" pageId="10" pageNumber="59">Remarks.</emphasis>
In description of
<taxonomicName id="F0A9A6A2FF857D44FDF0A1593040F8A8" box="[518,786,1888,1912]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="59" phylum="Arthropoda" rank="species" species="rakosiensis">
<emphasis id="05DD0133FF857D44FDF0A1593040F8A8" box="[518,786,1888,1912]" italics="true" pageId="10" pageNumber="59">Callianassa rakosiensis</emphasis>
</taxonomicName>
from the Middle Miocene of
<collectingCountry id="4FBE9DB1FF857D44FB91A159379BF8A8" box="[1127,1225,1888,1912]" name="Hungary" pageId="10" pageNumber="59">Hungary</collectingCountry>
<bibRefCitation id="5338A0D0FF857D44FB27A16636CEF8A7" author="Muller" box="[1233,1436,1887,1911]" pageId="10" pageNumber="59" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984: 51)</bibRefCitation>
stated: “The species is remarkably similar to
<taxonomicName id="F0A9A6A2FF857D44FD56A1BD307DF84C" box="[672,815,1924,1948]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="59" phylum="Arthropoda" rank="species" species="chalmasii">
<emphasis id="05DD0133FF857D44FD56A1BD307DF84C" box="[672,815,1924,1948]" italics="true" pageId="10" pageNumber="59">C. chalmasii</emphasis>
</taxonomicName>
in its general form, size and the distribution of pores. The form of the ridge on the finger and the position of the tooth is completely different in the two species and no trend was observed toward a transitional form. Thus their independence seems highly probable though a close relation is likely.” What
<bibRefCitation id="5338A0D0FF847D45FE5DA6AE311AFF7F" author="Muller" box="[427,584,151,175]" pageId="11" pageNumber="60" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984)</bibRefCitation>
recognized as two separate species in fact represents minor (
<taxonomicName id="F0A9A6A2FF847D45FB02A6A136C0FF60" box="[1268,1426,152,176]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="rakosiensis">
<emphasis id="05DD0133FF847D45FB02A6A136C0FF60" box="[1268,1426,152,176]" italics="true" pageId="11" pageNumber="60">C. rakosiensis</emphasis>
</taxonomicName>
) and major chelae (
<taxonomicName id="F0A9A6A2FF847D45FE7DA6843176FF04" box="[395,548,188,212]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="chalmasii">
<emphasis id="05DD0133FF847D45FE7DA6843176FF04" box="[395,548,188,212]" italics="true" pageId="11" pageNumber="60">C. chalmasii</emphasis>
</taxonomicName>
) of a single species. Müller's observation of no transitional form and simultaneously the presence of both morphotypes at the same localities (Rákos, Gyakorlo út and Örs vezér tere) support the synonymisation proposed here.
</paragraph>
</subSubSection>
<caption id="63D68DA9FF847D45FF61A29F37B0FAC8" httpUri="https://zenodo.org/record/210058/files/figure.png" pageId="11" pageNumber="60" targetBox="[151,1435,337,1168]" targetPageId="11">
<paragraph id="3716DD21FF847D45FF61A29F37B0FAC8" blockId="11.[151,1436,1190,1304]" pageId="11" pageNumber="60">
<emphasis id="05DD0133FF847D45FF61A29F3245FB6C" bold="true" box="[151,279,1190,1213]" pageId="11" pageNumber="60">FIGURE 5.</emphasis>
<taxonomicName id="F0A9A6A2FF847D45FED4A29E31F8FB6D" authority="Brocchi, 1883" authorityName="Brocchi" authorityYear="1883" box="[290,682,1190,1213]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="chalmasii" status="comb. nov.">
<emphasis id="05DD0133FF847D45FED4A29E32A8FB6D" box="[290,506,1191,1213]" italics="true" pageId="11" pageNumber="60">Calliaxina chalmasii</emphasis>
(Brocchi, 1883)
</taxonomicName>
<taxonomicNameLabel id="1EEEBC48FF847D45FD43A29E3039FB6C" box="[693,875,1190,1212]" pageId="11" pageNumber="60">new combination</taxonomicNameLabel>
, Fenk quarry (Vienna Basin, Austria): A, left minor propodus articulated with dactylus, NHMW-2011-0167-0118; B, right
<taxonomicName id="F0A9A6A2FF847D45FC75A2FC30ECFB0B" box="[899,958,1221,1243]" pageId="11" pageNumber="60" rank="species" species="major">major</taxonomicName>
propodus, NHMW-2011-0167-0120a; C, left
<taxonomicName id="F0A9A6A2FF847D45FF61A2DA3380FB29" box="[151,210,1251,1273]" pageId="11" pageNumber="60" rank="species" species="major">major</taxonomicName>
propodus, NHMW-2011-0167-0120b; D, right minor propodus, NHMW-2011-0167-0117; E, right carpus, NHMW- 2011-0167-0116. All specimens are to scale and were coated with ammonium chloride prior to photography.
</paragraph>
</caption>
<subSubSection id="7FB38EAAFF847D42FF33A3793240FECD" lastPageId="12" lastPageNumber="61" pageId="11" pageNumber="60" type="reference_group">
<paragraph id="3716DD21FF847D45FF33A3793664F9E0" blockId="11.[153,1436,1343,2016]" pageId="11" pageNumber="60">
<taxonomicName id="F0A9A6A2FF847D45FF33A37932B7FA88" box="[197,485,1344,1368]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="espichelensis">
<emphasis id="05DD0133FF847D45FF33A37932B7FA88" box="[197,485,1344,1368]" italics="true" pageId="11" pageNumber="60">Callianassa espichelensis</emphasis>
</taxonomicName>
described by
<bibRefCitation id="5338A0D0FF847D45FD77A3063023FA87" box="[641,881,1343,1367]" pageId="11" pageNumber="60" refString="Veiga Ferreira, O. da (1961) Nove especie de Callianassa no Miocenico da Bacia do Tejo. Comunicacoes dos Servicos Geologicos de Portugal, 45, 479 - 481." type="journal article">Veiga Ferreira (1961)</bibRefCitation>
has already been considered as a junior synonym of
<taxonomicName id="F0A9A6A2FF847D45FF4DA35C32A9FAAC" authority="Muller 1984" authorityName="Muller" authorityYear="1984" box="[187,507,1380,1404]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="chalmasii">
<emphasis id="05DD0133FF847D45FF4DA35C321EFAAC" box="[187,332,1380,1404]" italics="true" pageId="11" pageNumber="60">C. chalmasii</emphasis>
(
<bibRefCitation id="5338A0D0FF847D45FEA8A35D32A1FAAC" author="Muller" box="[350,499,1380,1404]" pageId="11" pageNumber="60" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller 1984</bibRefCitation>
)
</taxonomicName>
. Figures (
<bibRefCitation id="5338A0D0FF847D45FD99A35D3009FAAC" author="Veiga" box="[623,859,1380,1404]" pageId="11" pageNumber="60" refString="Veiga Ferreira, O. da (1961) Nove especie de Callianassa no Miocenico da Bacia do Tejo. Comunicacoes dos Servicos Geologicos de Portugal, 45, 479 - 481." type="journal article" year="1961">Veiga Ferreira 1961</bibRefCitation>
: fig. 1) clearly show the same morphotype. Most illustrated specimens represent major chelae (
<taxonomicName id="F0A9A6A2FF847D45FD60A3B13071FA70" box="[662,803,1416,1440]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="chalmasii">
<emphasis id="05DD0133FF847D45FD60A3B13071FA70" box="[662,803,1416,1440]" italics="true" pageId="11" pageNumber="60">C. chalmasii</emphasis>
</taxonomicName>
morphotype); however, one specimen (fig. 1.4) seems to represent the minor chela (
<taxonomicName id="F0A9A6A2FF847D45FE23A394312AFA14" box="[469,632,1452,1476]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="rakosiensis">
<emphasis id="05DD0133FF847D45FE23A394312AFA14" box="[469,632,1452,1476]" italics="true" pageId="11" pageNumber="60">C. rakosiensis</emphasis>
</taxonomicName>
morphotype). It is illustrated from the inner side, so, the depression present on the outer lateral surface is not visible.
<bibRefCitation id="5338A0D0FF847D45FD48A3F630FBFA37" author="Veiga" box="[702,937,1487,1511]" pageId="11" pageNumber="60" refString="Veiga Ferreira, O. da (1961) Nove especie de Callianassa no Miocenico da Bacia do Tejo. Comunicacoes dos Servicos Geologicos de Portugal, 45, 479 - 481." type="journal article" year="1961">
Veiga
<bibRefCitation id="5338A0D0FF847D45FCF2A3F630FBFA37" author="Ferreira" box="[772,937,1487,1511]" pageId="11" pageNumber="60" refString="Veiga Ferreira, O. da (1961) Nove especie de Callianassa no Miocenico da Bacia do Tejo. Comunicacoes dos Servicos Geologicos de Portugal, 45, 479 - 481." type="journal article" year="1961">Ferreira (1961</bibRefCitation>
</bibRefCitation>
: 480), however, mentioned the presence of a ridge, depression, and also faint tuberculation at the base of the fixed finger. I concur with
<bibRefCitation id="5338A0D0FF847D45FB64A3CD3660F9DC" author="Muller" box="[1170,1330,1524,1548]" pageId="11" pageNumber="60" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984)</bibRefCitation>
, contrary to
<bibRefCitation id="5338A0D0FF847D45FF43A02E3297F9FF" box="[181,453,1559,1584]" pageId="11" pageNumber="60" refString="Schweitzer, C. E., Feldmann, R. M., Garassino, A., Karasawa, H. &amp; Schweigert, G. (2010) Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10, 1 - 222." type="journal article">
Schweitzer
<emphasis id="05DD0133FF847D45FECFA0203238F9E0" box="[313,362,1560,1584]" italics="true" pageId="11" pageNumber="60">et al</emphasis>
. (2010)
</bibRefCitation>
, and consider
<taxonomicName id="F0A9A6A2FF847D45FD90A0213048F9E0" box="[614,794,1560,1584]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="espichelensis">
<emphasis id="05DD0133FF847D45FD90A0213048F9E0" box="[614,794,1560,1584]" italics="true" pageId="11" pageNumber="60">C. espichelensis</emphasis>
</taxonomicName>
as a junior subjective synonym of
<taxonomicName id="F0A9A6A2FF847D45FB57A021367DF9E0" box="[1185,1327,1560,1584]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="chalmasii">
<emphasis id="05DD0133FF847D45FB57A021367DF9E0" box="[1185,1327,1560,1584]" italics="true" pageId="11" pageNumber="60">C. chalmasii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3716DD21FF847D45FF31A0053071F8D8" blockId="11.[153,1436,1343,2016]" pageId="11" pageNumber="60">
<bibRefCitation id="5338A0D0FF847D45FF31A00532CAF984" author="Hyzny" box="[199,408,1596,1620]" pageId="11" pageNumber="60" refString="Hyzny, M. (2011) In situ mud shrimps (Decapoda: Axiidea: Callianassidae) preserved within their burrows from the middle Miocene of the Central Paratethys. Bulletin of the Mizunami Fossil Museum, 37, 37 - 46." type="journal article" year="2011">Hyžný (2011: 42)</bibRefCitation>
reported a burrow structure with the cheliped remains of two individuals identified as “
<taxonomicName id="F0A9A6A2FF847D45FF53A0593279F9A8" ID-CoL="f45854b2-a302-4361-bd40-0423151d5e6a" box="[165,299,1632,1656]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="05DD0133FF847D45FF53A0593279F9A8" box="[165,299,1632,1656]" italics="true" pageId="11" pageNumber="60">Callianassa</emphasis>
</taxonomicName>
” sp. 2 from the Gyakorló út locality. Although the material is insufficiently preserved, one individual clearly possesses equal or subequal chelipeds. Thus, it might represent
<taxonomicName id="F0A9A6A2FF847D45FC42A0BC3710F94C" box="[948,1090,1668,1692]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="chalmasii" status="comb. nov.">
<emphasis id="05DD0133FF847D45FC42A0BC3710F94C" box="[948,1090,1668,1692]" italics="true" pageId="11" pageNumber="60">C. chalmasii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EEEBC48FF847D45FBBFA0BD3795F94C" box="[1097,1223,1668,1692]" pageId="11" pageNumber="60">comb. nov.</taxonomicNameLabel>
, which has already been reported from that locality (
<bibRefCitation id="5338A0D0FF847D45FDFAA091304EF910" author="Muller" box="[524,796,1704,1728]" pageId="11" pageNumber="60" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller 1984; this paper</bibRefCitation>
). The material itself is, however, fragmentary and poorly preserved.
<collectingCountry id="4FBE9DB1FF847D45FEEEA0F53265F934" box="[280,311,1740,1764]" name="American Samoa" pageId="11" pageNumber="60">As</collectingCountry>
taxonomically important characters are discussed herein are not readily visible, I am hesitant to consider it to be conspecific with
<taxonomicName id="F0A9A6A2FF847D45FDE5A0C931F2F8D8" box="[531,672,1776,1800]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="chalmasii" status="comb. nov.">
<emphasis id="05DD0133FF847D45FDE5A0C931F2F8D8" box="[531,672,1776,1800]" italics="true" pageId="11" pageNumber="60">C. chalmasii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EEEBC48FF847D45FD51A0C93071F8D8" box="[679,803,1776,1800]" pageId="11" pageNumber="60">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="3716DD21FF847D42FF33A12D3240FECD" blockId="11.[153,1436,1343,2016]" lastBlockId="12.[153,1437,151,285]" lastPageId="12" lastPageNumber="61" pageId="11" pageNumber="60">
<taxonomicName id="F0A9A6A2FF847D45FF33A12D32FDF8FC" box="[197,431,1812,1836]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="chalmasii" status="comb. nov.">
<emphasis id="05DD0133FF847D45FF33A12D32FDF8FC" box="[197,431,1812,1836]" italics="true" pageId="11" pageNumber="60">Calliaxina chalmasii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EEEBC48FF847D45FE40A12D3161F8FC" box="[438,563,1812,1836]" pageId="11" pageNumber="60">comb. nov.</taxonomicNameLabel>
can be easily distinguished from all other Middle Miocene callianassid forms (for a review see
<bibRefCitation id="5338A0D0FF847D45FE9CA10132A9F880" author="Hyzny" box="[362,507,1848,1872]" pageId="11" pageNumber="60" refString="Hyzny, M. (2011) In situ mud shrimps (Decapoda: Axiidea: Callianassidae) preserved within their burrows from the middle Miocene of the Central Paratethys. Bulletin of the Mizunami Fossil Museum, 37, 37 - 46." type="journal article" year="2011">Hyžný 2011</bibRefCitation>
: table 2); none of them has a well developed tuberculated depression on minor propodus at the base of the fixed finger. Moreover,
<taxonomicName id="F0A9A6A2FF847D45FD16A164303CF8A4" box="[736,878,1884,1908]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="chalmasii" status="comb. nov.">
<emphasis id="05DD0133FF847D45FD16A164303CF8A4" box="[736,878,1884,1908]" italics="true" pageId="11" pageNumber="60">C. chalmasii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EEEBC48FF847D45FC80A16530A1F8A4" box="[886,1011,1884,1908]" pageId="11" pageNumber="60">comb. nov.</taxonomicNameLabel>
has laterally compressed chelipeds, a character that is uncommon in callianassid taxa co-occurring at the same localities. On the other hand,
<taxonomicName id="F0A9A6A2FF847D45FAF9A1B936CEF848" box="[1295,1436,1920,1944]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="chalmasii" status="comb. nov.">
<emphasis id="05DD0133FF847D45FAF9A1B936CEF848" box="[1295,1436,1920,1944]" italics="true" pageId="11" pageNumber="60">C. chalmasii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EEEBC48FF847D45FF6FA19D3244F86C" box="[153,278,1956,1980]" pageId="11" pageNumber="60">comb. nov.</taxonomicNameLabel>
is virtually indistinguishable from the extant forms, notably
<taxonomicName id="F0A9A6A2FF847D45FC31A19C3767F86C" box="[967,1077,1957,1980]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="punica">
<emphasis id="05DD0133FF847D45FC31A19C3767F86C" box="[967,1077,1957,1980]" italics="true" pageId="11" pageNumber="60">C. punica</emphasis>
</taxonomicName>
and
<taxonomicName id="F0A9A6A2FF847D45FB9BA19C3615F86C" box="[1133,1351,1956,1980]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="novaebritanniae">
<emphasis id="05DD0133FF847D45FB9BA19C3615F86C" box="[1133,1351,1956,1980]" italics="true" pageId="11" pageNumber="60">C. novaebritanniae</emphasis>
</taxonomicName>
, on the basis of propodus and carpus alone. Characters on merus might resolve this issue. I concur with
<bibRefCitation id="5338A0D0FF847D45FB25A1F136CEF830" author="Muller" box="[1235,1436,1992,2016]" pageId="11" pageNumber="60" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984: 48)</bibRefCitation>
that: “identification of 1215 milion year old species with extant ones should be done with caution”. Thus, a chronological approach (which operates with chronospecies rather than morphospecies) is followed here (see also
<bibRefCitation id="5338A0D0FF837D42FF6FA6E632D3FF27" author="Klaus" box="[153,385,223,247]" pageId="12" pageNumber="61" refString="Klaus, S. &amp; Gross, M. (2010) Synopsis of the fossil freshwater crabs of Europe (Brachyura: Potamoidea: Potamidae). Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen, 256, 39 - 59." type="journal article" year="2010">Klaus &amp; Gross 2010</bibRefCitation>
) and
<taxonomicName id="F0A9A6A2FF837D42FE49A6D931F8FF28" box="[447,682,224,248]" class="Malacostraca" family="Callianassidae" genus="Calliaxina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="chalmasii" status="comb. nov.">
<emphasis id="05DD0133FF837D42FE49A6D931F8FF28" box="[447,682,224,248]" italics="true" pageId="12" pageNumber="61">Calliaxina chalmasii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EEEBC48FF837D42FD47A6E6307FFF27" box="[689,813,223,247]" pageId="12" pageNumber="61">comb. nov.</taxonomicNameLabel>
is considered a separate species distinct from all extant congeners.
</paragraph>
</subSubSection>
</treatment>
</document>