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<document id="0DBC38E7FD38491EEEE46F037DA112E2" ID-CLB-Dataset="21047" ID-DOI="10.1080/00222933.2013.861938" ID-GBIF-Dataset="ac743694-3c75-4efe-82b1-1c49797b3323" ID-ISSN="1464-5262" ID-Zenodo-Dep="4006759" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1598645811345" checkinUser="carolina" docAuthor="Fenglong, J. I. A." docDate="2014" docId="59670F47CC5CFF8D4FE7FE0DFED9FA11" docLanguage="en" docName="JNATHIST.48.17-18.1085-1101.pdf" docOrigin="Journal of Natural History 48 (17)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Amphiops coelopunctatus Fenglong 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="1092" masterDocId="A55E773FCC58FF8A4E1FFA56FF99FF9D" masterDocTitle="A revisional study of the Chinese species of Amphiops Erichson (Coleoptera, Hydrophilidae, Chaetarthriini)" masterLastPageNumber="1101" masterPageNumber="1085" pageNumber="1089" updateTime="1698845745751" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="5850C197A666A3425FDAFFB0FDD2756D">A revisional study of the Chinese species of Amphiops Erichson (Coleoptera, Hydrophilidae, Chaetarthriini)</mods:title>
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<mods:namePart id="4432252AC216BC0D03DFB4CC805E4D4E">Fenglong, J. I. A.</mods:namePart>
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<mods:title id="5A5975D2F283ECEB2E681FB9B7063B03">Journal of Natural History</mods:title>
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<mods:date id="F8370B7956C8DE0130601474DAD48C2A">2014</mods:date>
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<mods:number id="5B170B9DCA9852963C516E98880D1BA3">2014-02-05</mods:number>
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<mods:number id="A63C2A4DD6DC14F6774D47FEBD577EA9">48</mods:number>
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<mods:classification id="944C492EDEE509A3E610CA384CB895AA">journal article</mods:classification>
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<mods:identifier id="23C70F92F7657A4CA0843F15F3C22DCE" type="DOI">10.1080/00222933.2013.861938</mods:identifier>
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<treatment id="59670F47CC5CFF8D4FE7FE0DFED9FA11" ID-DOI="http://doi.org/10.5281/zenodo.4328987" ID-GBIF-Taxon="167375511" ID-Zenodo-Dep="4328987" LSID="urn:lsid:plazi:treatment:59670F47CC5CFF8D4FE7FE0DFED9FA11" httpUri="http://treatment.plazi.org/id/59670F47CC5CFF8D4FE7FE0DFED9FA11" lastPageId="7" lastPageNumber="1092" pageId="4" pageNumber="1089">
<subSubSection id="99D4EDDACC5CFF8E4FE7FE0DFCF6FBEE" box="[504,879,1115,1139]" pageId="4" pageNumber="1089" type="nomenclature">
<paragraph id="D171BE51CC5CFF8E4FE7FE0DFCF6FBEE" blockId="4.[504,879,1115,1171]" box="[504,879,1115,1139]" pageId="4" pageNumber="1089">
<heading id="8A39093DCC5CFF8E4FE7FE0DFCF6FBEE" box="[504,879,1115,1139]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="1089" reason="2">
<emphasis id="E3BA6243CC5CFF8E4FE7FE0DFCF6FBEE" bold="true" box="[504,879,1115,1139]" pageId="4" pageNumber="1089">
<taxonomicName id="16CEC5D2CC5CFF8E4FE7FE0DFC88FBEE" ID-CoL="D4R7" authority="Fenglong, 2014" authorityName="Fenglong" authorityYear="2014" box="[504,785,1115,1139]" class="Insecta" family="Hydrophilidae" genus="Amphiops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1089" phylum="Arthropoda" rank="species" species="coelopunctatus" status="sp. nov.">
<emphasis id="E3BA6243CC5CFF8E4FE7FE0DFC88FBEE" bold="true" box="[504,785,1115,1139]" italics="true" pageId="4" pageNumber="1089">Amphiops coelopunctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F889DF38CC5CFF8E4D05FE0DFCF6FBEE" box="[794,879,1115,1139]" pageId="4" pageNumber="1089" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="99D4EDDACC5CFF8E4C2FFE2DFCA1FB0E" box="[560,824,1147,1171]" pageId="4" pageNumber="1089" type="description">
<paragraph id="D171BE51CC5CFF8E4C2FFE2DFCA1FB0E" blockId="4.[504,879,1115,1171]" box="[560,824,1147,1171]" pageId="4" pageNumber="1089">
(
<figureCitation id="49F5A2D4CC5CFF8E4C28FE2DFD49FB0E" box="[567,720,1147,1171]" captionStart="Figure 2" captionStartId="5.[144,211,1487,1509]" captionTargetBox="[158,1090,168,1380]" captionTargetId="figure@5.[157,1091,168,1380]" captionTargetPageId="5" captionText="Figure 2. Amphiops caristripus Jia. (A) Head and pronotum; (B) elytra, lateral margin; (C) elytral apex; (D) elytra. Amphiops coelopunctatus sp. nov. (E) Habitus; (F) head, pronotum and elytra base, lateral view; (G) elytra; (H) head; (I) pronotum and elytral base; (J) metasternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4006763" httpUri="https://zenodo.org/record/4006763/files/figure.png" pageId="4" pageNumber="1089">Figures 2EJ</figureCitation>
, and
<figureCitation id="49F5A2D4CC5CFF8E4D0FFE2DFCB6FB0E" box="[784,815,1147,1171]" captionStart="Figure 5" captionStartId="13.[144,211,1397,1419]" captionTargetBox="[145,1061,171,1302]" captionTargetId="figure@13.[145,1103,168,1307]" captionTargetPageId="13" captionText="Figure 5. Aedeagi. (A) Amphiops coomani; (B) Amphiops coelopunctatus; (CF Amphiops mirabilis, from (C) Guangzhou, (E) Hainan and (D, F) Yunnan; (G, H) Amphiops mater from (G) Guangdong and (H) Yunnan." figureDoi="http://doi.org/10.5281/zenodo.4006769" httpUri="https://zenodo.org/record/4006769/files/figure.png" pageId="4" pageNumber="1089">5B</figureCitation>
)
</paragraph>
</subSubSection>
<paragraph id="D171BE51CC5CFF8E4ECBFE8FFEFFFB6D" blockId="4.[212,1172,1240,1304]" box="[212,358,1240,1264]" pageId="4" pageNumber="1089">
<heading id="8A39093DCC5CFF8E4ECBFE8FFEFFFB6D" box="[212,358,1240,1264]" fontSize="10" level="3" pageId="4" pageNumber="1089" reason="5">
<emphasis id="E3BA6243CC5CFF8E4ECBFE8FFEFFFB6D" box="[212,358,1240,1264]" italics="true" pageId="4" pageNumber="1089">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="D171BE51CC5CFF8E4ECBFF56FB0DFA8A" blockId="4.[212,1172,1240,1304]" box="[212,1172,1280,1304]" pageId="4" pageNumber="1089">
<materialsCitation id="61A6B40CCC5CFF8E4ECBFF56FCC8FA8A" ID-GBIF-Occurrence="2850099367" box="[212,849,1280,1304]" country="China" elevation="650" location="Limushan Mountains" municipality="Island" pageId="4" pageNumber="1089" specimenCount="1" stateProvince="Hainan">
<collectingCountry id="A9D9FEC1CC5CFF8E4ECBFF56FE82FA85" box="[212,283,1280,1304]" name="China" pageId="4" pageNumber="1089">China</collectingCountry>
,
<collectingRegion id="130A70B3CC5CFF8E4F39FF56FEE3FA85" box="[294,378,1280,1304]" country="China" name="Hainan" pageId="4" pageNumber="1089">Hainan</collectingRegion>
<collectingMunicipality id="3115242BCC5CFF8E4F9EFF56FE50FA85" box="[385,457,1280,1304]" pageId="4" pageNumber="1089">Island</collectingMunicipality>
,
<location id="D411E88ACC5CFF8E4FCCFF56FD50FA85" LSID="urn:lsid:plazi:treatment:59670F47CC5CFF8D4FE7FE0DFED9FA11:D411E88ACC5CFF8E4FCCFF56FD50FA85" box="[467,713,1280,1304]" country="China" municipality="Island" name="Limushan Mountains" pageId="4" pageNumber="1089" stateProvince="Hainan">Limushan Mountains</location>
,
<quantity id="163613B4CC5CFF8E4CCBFF56FCC8FA8A" box="[724,849,1280,1304]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="4" pageNumber="1089" unit="m" value="650.0" valueMax="750.0" valueMin="550.0">
<elevation id="5AE35962CC5CFF8E4CCBFF56FCC8FA8A" box="[724,849,1280,1304]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="4" pageNumber="1089" unit="m" value="650.0" valueMax="750.0" valueMin="550.0">550750 m</elevation>
</quantity>
</materialsCitation>
, 19°9.19.2
<emphasis id="E3BA6243CC5CFF8E4DCEFF56FC4EFA85" bold="true" box="[977,983,1280,1304]" italics="true" pageId="4" pageNumber="1089"></emphasis>
N, 109°4546
<emphasis id="E3BA6243CC5CFF8E4A69FF56FBE5FA85" bold="true" box="[1142,1148,1280,1304]" italics="true" pageId="4" pageNumber="1089"></emphasis>
E
</paragraph>
<paragraph id="D171BE51CC5CFF8E4ECBFF0BFEE9FAE9" blockId="4.[212,1173,1372,1692]" box="[212,368,1372,1396]" pageId="4" pageNumber="1089">
<heading id="8A39093DCC5CFF8E4ECBFF0BFEE9FAE9" box="[212,368,1372,1396]" fontSize="10" level="3" pageId="4" pageNumber="1089" reason="5">
<emphasis id="E3BA6243CC5CFF8E4ECBFF0BFEE9FAE9" box="[212,368,1372,1396]" italics="true" pageId="4" pageNumber="1089">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="D171BE51CC5CFF8E4ECBFFD2FE5BF901" blockId="4.[212,1173,1372,1692]" pageId="4" pageNumber="1089">
<materialsCitation id="61A6B40CCC5CFF8E4ECBFFD2FC63FA21" ID-GBIF-Occurrence="2850099382" collectingDate="2011-05-05" collectionCode="SYSU" country="China" elevation="650" location="Limushan Mountains" pageId="4" pageNumber="1089" specimenCount="1" stateProvince="Hainan" typeStatus="holotype">
<typeStatus id="0E7500F3CC5CFF8E4ECBFFD2FED9FA01" box="[212,320,1412,1436]" pageId="4" pageNumber="1089" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="B7DF2694CC5CFF8E4F8EFFD2FE47FA01" box="[401,478,1412,1436]" pageId="4" pageNumber="1089">SYSU</collectionCode>
):
<collectingCountry id="A9D9FEC1CC5CFF8E4FECFFD2FDA5FA01" box="[499,572,1412,1436]" name="China" pageId="4" pageNumber="1089">China</collectingCountry>
,
<collectingRegion id="130A70B3CC5CFF8E4C55FFD2FD39FA01" box="[586,672,1412,1436]" country="China" name="Hainan" pageId="4" pageNumber="1089">Hainan</collectingRegion>
island
<location id="D411E88ACC5CFF8E4CE9FFD2FC6AFA01" LSID="urn:lsid:plazi:treatment:59670F47CC5CFF8D4FE7FE0DFED9FA11:D411E88ACC5CFF8E4CE9FFD2FC6AFA01" box="[758,1011,1412,1436]" country="China" name="Limushan Mountains" pageId="4" pageNumber="1089" stateProvince="Hainan">Limushan Mountains</location>
,
<date id="A5709891CC5CFF8E4A1EFFD2FB09FA01" box="[1025,1168,1412,1437]" pageId="4" pageNumber="1089" value="2011-05-05">
<collectingDate id="B5346179CC5CFF8E4A1EFFD2FB09FA01" box="[1025,1168,1412,1437]" pageId="4" pageNumber="1089" value="2011-05-05">5 May 2011</collectingDate>
</date>
, 19°9.19.2
<emphasis id="E3BA6243CC5CFF8E4F52FFF2FECAFA21" bold="true" box="[333,339,1444,1468]" italics="true" pageId="4" pageNumber="1089"></emphasis>
N, 109°4546
<emphasis id="E3BA6243CC5CFF8E4C17FFF2FD97FA21" bold="true" box="[520,526,1444,1468]" italics="true" pageId="4" pageNumber="1089"></emphasis>
E,
<quantity id="163613B4CC5CFF8E4C58FFF2FD49FA21" box="[583,720,1444,1468]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="4" pageNumber="1089" unit="m" value="650.0" valueMax="750.0" valueMin="550.0">
<elevation id="5AE35962CC5CFF8E4C58FFF2FD49FA21" box="[583,720,1444,1468]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="4" pageNumber="1089" unit="m" value="650.0" valueMax="750.0" valueMin="550.0">550750 m</elevation>
</quantity>
, Fikáček and Zhao lgt
</materialsCitation>
.
<materialsCitation id="61A6B40CCC5CFF8E4A11FFF2FC38FA41" ID-GBIF-Occurrence="2850099373" collectingDate="2011-05-05" collectionCode="SYSU" country="China" elevation="650" location="Limushan Mountains" pageId="4" pageNumber="1089" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<typeStatus id="0E7500F3CC5CFF8E4A11FFF2FB1EFA21" box="[1038,1159,1444,1468]" pageId="4" pageNumber="1089" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="C7C875D8CC5CFF8E4A94FFF2FE92FA41" pageId="4" pageNumber="1089" type="male">1 male</specimenCount>
and
<specimenCount id="C7C875D8CC5CFF8E4F7BFF92FE46FA41" box="[356,479,1476,1500]" pageId="4" pageNumber="1089" type="female">3 females</specimenCount>
(
<collectionCode id="B7DF2694CC5CFF8E4FE1FF92FDD2FA41" box="[510,587,1476,1500]" pageId="4" pageNumber="1089">SYSU</collectionCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="61A6B40CCC5CFF8E4DA3FF92FEA6F9C6" ID-GBIF-Occurrence="2850099363" collectingDate="2011-05-05" collectionCode="NMPC" collectorName="Fikacek &amp; Zhao" country="China" elevation="650" location="Limushan Mountaint" pageId="4" pageNumber="1089" specimenCount="3" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C7C875D8CC5CFF8E4DA3FF92FB82FA41" box="[956,1051,1476,1500]" pageId="4" pageNumber="1089" type="male">1 male</specimenCount>
,
<specimenCount id="C7C875D8CC5CFF8E4A28FF92FB0CFA46" box="[1079,1173,1476,1500]" pageId="4" pageNumber="1089" type="generic">2 spec.</specimenCount>
(
<collectionCode id="B7DF2694CC5CFF8E4EC3FFB2FEACFA61" box="[220,309,1508,1532]" country="Czech Republic" name="National Museum Prague" pageId="4" pageNumber="1089">NMPC</collectionCode>
):
<collectingCountry id="A9D9FEC1CC5CFF8E4F4FFFB2FE00FA61" box="[336,409,1508,1532]" name="China" pageId="4" pageNumber="1089">China</collectingCountry>
,
<collectingRegion id="130A70B3CC5CFF8E4FB4FFB2FD98FA61" box="[427,513,1508,1532]" country="China" name="Hainan" pageId="4" pageNumber="1089">Hainan</collectingRegion>
island,
<location id="D411E88ACC5CFF8E4C75FFB2FCF2FA61" LSID="urn:lsid:plazi:treatment:59670F47CC5CFF8D4FE7FE0DFED9FA11:D411E88ACC5CFF8E4C75FFB2FCF2FA61" box="[618,875,1508,1532]" country="China" name="Limushan Mountaint" pageId="4" pageNumber="1089" stateProvince="Hainan">Limushan Mountaint</location>
, 19°9.19.2
<emphasis id="E3BA6243CC5CFF8E4DE8FFB2FC64FA61" bold="true" box="[1015,1021,1508,1532]" italics="true" pageId="4" pageNumber="1089"></emphasis>
N, 109°45 46
<emphasis id="E3BA6243CC5CFF8E4EF0FC52FF6CF981" bold="true" box="[239,245,1540,1564]" italics="true" pageId="4" pageNumber="1089"></emphasis>
E,
<quantity id="163613B4CC5CFF8E4F30FC55FE21F986" box="[303,440,1539,1564]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="4" pageNumber="1089" unit="m" value="650.0" valueMax="750.0" valueMin="550.0">
<elevation id="5AE35962CC5CFF8E4F30FC55FE21F986" box="[303,440,1539,1564]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="4" pageNumber="1089" unit="m" value="650.0" valueMax="750.0" valueMin="550.0">550750 m</elevation>
</quantity>
, along the road, small streamlet in secondary forest with bamboo: among gravel and stones in a small pool,
<date id="A5709891CC5CFF8E4D49FC75FC74F9A6" box="[854,1005,1571,1596]" pageId="4" pageNumber="1089" value="2011-05-05">
<collectingDate id="B5346179CC5CFF8E4D49FC75FC74F9A6" box="[854,1005,1571,1596]" pageId="4" pageNumber="1089" value="2011-05-05">5 May 2011</collectingDate>
</date>
,
<collectorName id="7C3BDB87CC5CFF8E4DE0FC75FBC2F9A6" box="[1023,1115,1571,1595]" pageId="4" pageNumber="1089">Fikáček</collectorName>
and
<collectorName id="7C3BDB87CC5CFF8E4ECBFC15FE88F9C6" box="[212,273,1603,1627]" pageId="4" pageNumber="1089">Zhao</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="61A6B40CCC5CFF8E4F4CFC15FD59F9C6" ID-GBIF-Occurrence="2850099409" box="[339,704,1603,1627]" collectingDate="2011-05-05" collectionCode="NMPC" collectorName="Fikacek &amp; Zhao" country="China" elevation="650" location="Limushan Mountaint" pageId="4" pageNumber="1089" specimenCount="5" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C7C875D8CC5CFF8E4F4CFC15FE3DF9C6" box="[339,420,1603,1627]" pageId="4" pageNumber="1089" type="generic">5 spec.</specimenCount>
(
<collectionCode id="B7DF2694CC5CFF8E4FA9FC15FD96F9C6" box="[438,527,1603,1627]" country="Czech Republic" name="National Museum Prague" pageId="4" pageNumber="1089">NMPC</collectionCode>
): same locality
</materialsCitation>
data, but with habitat data as follows: exposed pools along the road partly connected to the stream; sparsely to densely overgrown by grass.
</paragraph>
<paragraph id="D171BE51CC5DFF8F4F5EFF2AFC59FA34" blockId="5.[321,960,1400,1453]" box="[321,960,1400,1453]" pageId="5" pageNumber="1090">H I J</paragraph>
<caption id="85B1EED9CC5DFF8F4E8FFF99FEBAF9A0" ID-DOI="http://doi.org/10.5281/zenodo.4006763" ID-Zenodo-Dep="4006763" httpUri="https://zenodo.org/record/4006763/files/figure.png" pageId="5" pageNumber="1090" startId="5.[144,211,1487,1509]" targetBox="[158,1090,168,1380]" targetPageId="5">
<paragraph id="D171BE51CC5DFF8F4E8FFF99FEBAF9A0" blockId="5.[144,1104,1487,1597]" pageId="5" pageNumber="1090">
Figure 2.
<taxonomicName id="16CEC5D2CC5DFF8F4EE4FF99FE64FA78" authority="Jia." authorityName="Jia." box="[251,509,1487,1509]" class="Insecta" family="Hydrophilidae" genus="Amphiops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1090" phylum="Arthropoda" rank="species" species="caristripus">
<emphasis id="E3BA6243CC5DFF8F4EE4FF99FE56FA79" box="[251,463,1487,1508]" italics="true" pageId="5" pageNumber="1090">Amphiops caristripus</emphasis>
Jia.
</taxonomicName>
(A) Head and pronotum; (B) elytra, lateral margin; (C) elytral apex; (D) elytra.
<taxonomicName id="16CEC5D2CC5DFF8F4F9DFFBBFDE4F99F" authority="Fenglong, 2014" authorityName="Fenglong" authorityYear="2014" box="[386,637,1517,1538]" class="Insecta" family="Hydrophilidae" genus="Amphiops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1090" phylum="Arthropoda" rank="species" species="coelopunctatus" status="sp. nov.">
<emphasis id="E3BA6243CC5DFF8F4F9DFFBBFDE4F99F" box="[386,637,1517,1538]" italics="true" pageId="5" pageNumber="1090">Amphiops coelopunctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F889DF38CC5DFF8F4C9CFFBBFD4AF99F" box="[643,723,1517,1538]" pageId="5" pageNumber="1090" rank="species">sp. nov.</taxonomicNameLabel>
(E) Habitus; (F) head, pronotum and elytra base, lateral view; (G) elytra; (H) head; (I) pronotum and elytral base; (J) metasternum and abdomen.
</paragraph>
</caption>
<subSubSection id="99D4EDDACC5EFF8C4ECBFAFFFE3AFDFA" pageId="6" pageNumber="1091" type="diagnosis">
<paragraph id="D171BE51CC5EFF8C4ECBFAFFFED9FF5D" blockId="6.[212,1172,169,615]" box="[212,320,169,192]" pageId="6" pageNumber="1091">
<heading id="8A39093DCC5EFF8C4ECBFAFFFED9FF5D" box="[212,320,169,192]" fontSize="10" level="3" pageId="6" pageNumber="1091" reason="5">
<emphasis id="E3BA6243CC5EFF8C4ECBFAFFFED9FF5D" box="[212,320,169,192]" italics="true" pageId="6" pageNumber="1091">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="D171BE51CC5EFF8C4ECBFA87FE3AFDFA" blockId="6.[212,1172,169,615]" pageId="6" pageNumber="1091">
Dorsum glabrous, black. Anterior margin of clypeus broadly truncate; head with sparse punctation, all punctures of the same size; lateral portions of pronotum with punctation identical to those on head, punctures on median portion finer; elytra without serial punctures, ground punctation similar to head, lateral punctures somewhat coarser than those on inner parts. Systematic punctures on head, pronotum and elytra of the same size as ground punctures. Lateral margin continuously curved to posterior margin, (
<figureCitation id="49F5A2D4CC5EFF8C4FB8FBD9FDABFE3A" box="[423,562,399,423]" captionStart="Figure 2" captionStartId="5.[144,211,1487,1509]" captionTargetBox="[158,1090,168,1380]" captionTargetId="figure@5.[157,1091,168,1380]" captionTargetPageId="5" captionText="Figure 2. Amphiops caristripus Jia. (A) Head and pronotum; (B) elytra, lateral margin; (C) elytral apex; (D) elytra. Amphiops coelopunctatus sp. nov. (E) Habitus; (F) head, pronotum and elytra base, lateral view; (G) elytra; (H) head; (I) pronotum and elytral base; (J) metasternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4006763" httpUri="https://zenodo.org/record/4006763/files/figure.png" pageId="6" pageNumber="1091">Figure 2F, I</figureCitation>
), hence posterolateral angle absent. Mesofemora very narrowly separated. Metaventrite without a transverse ridge anteriorly. Mesofemora densely pubescent on basal third, metafemora glabrous, only pubescent along anterior margin on basal half. Aedeagus with median lobe broad, triangular apically, broadly notched behind apex on each side, gonopore situated apically; parameres as long as median lobe, almost parallel-sided, rounded apically, distinctly curved medially on inner face.
</paragraph>
</subSubSection>
<subSubSection id="99D4EDDACC5EFF8D4ECBF8ECFC50FE78" lastPageId="7" lastPageNumber="1092" pageId="6" pageNumber="1091" type="description">
<paragraph id="D171BE51CC5EFF8C4ECBF8ECFECAFD4C" blockId="6.[212,1172,698,888]" box="[212,339,698,721]" pageId="6" pageNumber="1091">
<heading id="8A39093DCC5EFF8C4ECBF8ECFECAFD4C" box="[212,339,698,721]" fontSize="10" level="3" pageId="6" pageNumber="1091" reason="5">
<emphasis id="E3BA6243CC5EFF8C4ECBF8ECFECAFD4C" box="[212,339,698,721]" italics="true" pageId="6" pageNumber="1091">Description</emphasis>
</heading>
</paragraph>
<paragraph id="D171BE51CC5EFF8C4ECBF8B4FCA8FCE5" blockId="6.[212,1172,698,888]" pageId="6" pageNumber="1091">
<emphasis id="E3BA6243CC5EFF8C4ECBF8B4FE0EFD64" box="[212,407,737,761]" italics="true" pageId="6" pageNumber="1091">Form and colour.</emphasis>
Body strongly convex, broadly oval. Length
<quantity id="163613B4CC5EFF8C4DBAF8B7FBDEFD64" box="[933,1095,737,761]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.65" metricValueMin="3.55" pageId="6" pageNumber="1091" unit="mm" value="3.5999999999999996" valueMax="3.65" valueMin="3.55">3.553.65 mm</quantity>
(
<typeStatus id="0E7500F3CC5EFF8C4A47F8B7FE9DFC87" pageId="6" pageNumber="1091" type="holotype">holotype</typeStatus>
<quantity id="163613B4CC5EFF8C4F11F957FEE1FC84" box="[270,376,769,793]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="1091" unit="mm" value="3.6">3.60 mm</quantity>
), width
<quantity id="163613B4CC5EFF8C4FC5F957FDE5FC84" box="[474,636,769,793]" metricMagnitude="-3" metricUnit="m" metricValue="2.625" metricValueMax="2.65" metricValueMin="2.6" pageId="6" pageNumber="1091" unit="mm" value="2.625" valueMax="2.65" valueMin="2.6">2.602.65 mm</quantity>
(
<typeStatus id="0E7500F3CC5EFF8C4C91F957FD68FC84" box="[654,753,769,793]" pageId="6" pageNumber="1091" type="holotype">holotype</typeStatus>
<quantity id="163613B4CC5EFF8C4CE2F957FCFEFC84" box="[765,871,769,793]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" pageId="6" pageNumber="1091" unit="mm" value="2.65">2.65 mm</quantity>
). Black, lateral margin of pronotum yellowish brown. Maxillary palpomeres, labial palpomeres, antennae, legs and ventrites yellowish brown, ventral surface of mesoventrite and metaventrite yellowish brown to dark brown. Antennal club dark.
</paragraph>
<paragraph id="D171BE51CC5EFF8C4ECBF9E5FC8BFB37" blockId="6.[212,1172,947,1194]" pageId="6" pageNumber="1091">
<emphasis id="E3BA6243CC5EFF8C4ECBF9E5FE81FC56" box="[212,280,947,971]" italics="true" pageId="6" pageNumber="1091">Head.</emphasis>
About 1.75 × as wide as long. Glabrous, without microsculpture between punctures, with punctation consisting of single size of ground punctures mixed with some systematic punctures of the same size as ground punctures, intervals between punctures c.1.251.50 × the diameter of a puncture. Anterior margin of clypeus broadly truncate or slightly concave anteriorly. Frontoclypeal suture indistinct, sometimes undetectable.
<collectingRegion id="130A70B3CC5EFF8C4FA5FE05FD82FBF7" box="[442,539,1107,1130]" country="Myanmar" name="Chin" pageId="6" pageNumber="1091">Mentum</collectingRegion>
c.0.75 × as long as wide, with sparsely arranged systematic punctures, slightly emarginate anteriorly. Apical maxillary palpomere asymmetrical, longer than penultimate palpomere.
</paragraph>
<paragraph id="D171BE51CC5EFF8D4ECBFEB3FE9CFF7D" blockId="6.[212,1172,1253,1691]" lastBlockId="7.[144,1103,168,224]" lastPageId="7" lastPageNumber="1092" pageId="6" pageNumber="1091">
<emphasis id="E3BA6243CC5EFF8C4ECBFEB3FEB4FB60" box="[212,301,1253,1277]" italics="true" pageId="6" pageNumber="1091">Thorax.</emphasis>
Pronotum c.2.25 × as wide as long, median portion with finer and sparser punctures than those on head, lateral portion with coarser punctation consisting of single size of ground punctures intermixed with systematic punctures of the same size; intervals between punctures about c.1.51.7 × the diameter of a puncture mesally, c.1.21.4 × laterally. Posterolateral margin rounded, posterior angle unclear. Prosternum short, with a strong median carina. Scutellum longer than wide, with same punctures mesally as on pronotum. Elytra almost as high as long, ground punctures similar to head but somewhat coarser laterally, intervals between punctures c.1.31.5 × the diameter of a puncture mesally, c.1.11.2 × laterally; elytron with five series of setiferous punctures (systematic punctures) that are the same size as ground punctures, serial punctures absent. Mesoventrite gradually raised posteriorly to form a narrowly notched protuberance, mesocoxae very narrowly separated. Metaventrite slightly convex medially, without transverse carina anteriorly. Profemora and mesofemora densely pubescent on basal half, metafemora only pubescent anteriorly on basal half, with fine and sparse punctures beneath, without microsculpture between punctures.
</paragraph>
<paragraph id="D171BE51CC5FFF8D4E8FFB5DFCE6FEDF" blockId="7.[144,1103,266,323]" pageId="7" pageNumber="1092">
<emphasis id="E3BA6243CC5FFF8D4E8FFB5DFE99FEBE" box="[144,256,267,291]" italics="true" pageId="7" pageNumber="1092">Abdomen.</emphasis>
Covered with hyaline mass. Visible part of first ventrite very short, without median carina, with marginal short and very fine setae anteriorly.
</paragraph>
<paragraph id="D171BE51CC5FFF8D4E8FFB3BFC50FE78" blockId="7.[144,1104,365,485]" pageId="7" pageNumber="1092">
<emphasis id="E3BA6243CC5FFF8D4E8FFB3BFE98FE18" box="[144,257,365,389]" italics="true" pageId="7" pageNumber="1092">Aedeagus.</emphasis>
Median lobe broad, triangular apically, broadly notched behind apex on each side, gonopore situated apically. Parameres as long as median lobe, almost parallel-sided, rounded apically, distinctly curved mesad on inner face. Phallobase slightly longer than parameres, abruptly narrowed medially (
<figureCitation id="49F5A2D4CC5FFF8D4D59FB9BFC25FE78" box="[838,956,461,485]" captionStart="Figure 5" captionStartId="13.[144,211,1397,1419]" captionTargetBox="[145,1061,171,1302]" captionTargetId="figure@13.[145,1103,168,1307]" captionTargetPageId="13" captionText="Figure 5. Aedeagi. (A) Amphiops coomani; (B) Amphiops coelopunctatus; (CF Amphiops mirabilis, from (C) Guangzhou, (E) Hainan and (D, F) Yunnan; (G, H) Amphiops mater from (G) Guangdong and (H) Yunnan." figureDoi="http://doi.org/10.5281/zenodo.4006769" httpUri="https://zenodo.org/record/4006769/files/figure.png" pageId="7" pageNumber="1092">Figure 5B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="99D4EDDACC5FFF8D4E8FF871FD80FCDB" pageId="7" pageNumber="1092" type="diagnosis">
<paragraph id="D171BE51CC5FFF8D4E8FF871FEE4FDA2" blockId="7.[144,1104,551,838]" box="[144,381,551,575]" pageId="7" pageNumber="1092">
<heading id="8A39093DCC5FFF8D4E8FF871FEE4FDA2" box="[144,381,551,575]" fontSize="10" level="3" pageId="7" pageNumber="1092" reason="5">
<emphasis id="E3BA6243CC5FFF8D4E8FF871FEE4FDA2" box="[144,381,551,575]" italics="true" pageId="7" pageNumber="1092">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="D171BE51CC5FFF8D4E8FF819FD80FCDB" blockId="7.[144,1104,551,838]" pageId="7" pageNumber="1092">
The new species is close to
<taxonomicName id="16CEC5D2CC5FFF8D4FD6F819FDD9FDFA" authorityName="d' Orchymont" authorityYear="1926" box="[457,576,591,615]" class="Insecta" family="Hydrophilidae" genus="Amphiops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1092" phylum="Arthropoda" rank="species" species="coomani">
<emphasis id="E3BA6243CC5FFF8D4FD6F819FDD9FDFA" box="[457,576,591,615]" italics="true" pageId="7" pageNumber="1092">A.coomani</emphasis>
</taxonomicName>
and
<taxonomicName id="16CEC5D2CC5FFF8D4C63F819FD44FDFA" baseAuthorityName="d' Orchymont" baseAuthorityYear="1926" box="[636,733,591,615]" class="Insecta" family="Hydrophilidae" genus="Amphiops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1092" phylum="Arthropoda" rank="species" species="globus">
<emphasis id="E3BA6243CC5FFF8D4C63F819FD44FDFA" box="[636,733,591,615]" italics="true" pageId="7" pageNumber="1092">A.globus</emphasis>
</taxonomicName>
based on the pronotum without distinct posterolateral corners, elytra without series of coarse punctures, head, pronotum and elytra with ground punctures of single size; metaventrite without a transverse carina anteriorly, and metafemora glabrous with fine punctures. It can be easily distinguished from these species by the dorsum with denser and coarser punctures, pronotum and elytra with systematic punctures of the same size as ground punctures, and median lobe of the aedeagus narrower, triangular apically, broadly notched behind apex on each side.
</paragraph>
</subSubSection>
<subSubSection id="99D4EDDACC5FFF8D4E8FF9DFFD95FBB5" pageId="7" pageNumber="1092" type="etymology">
<paragraph id="D171BE51CC5FFF8D4E8FF9DFFE90FC3C" blockId="7.[144,1104,905,1065]" box="[144,265,905,929]" pageId="7" pageNumber="1092">
<heading id="8A39093DCC5FFF8D4E8FF9DFFE90FC3C" box="[144,265,905,929]" fontSize="10" level="3" pageId="7" pageNumber="1092" reason="5">
<emphasis id="E3BA6243CC5FFF8D4E8FF9DFFE90FC3C" box="[144,265,905,929]" italics="true" pageId="7" pageNumber="1092">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="D171BE51CC5FFF8D4E8FF9E6FD95FBB5" blockId="7.[144,1104,905,1065]" pageId="7" pageNumber="1092">
The species name is a combination of
<taxonomicName id="16CEC5D2CC5FFF8D4C52F9E7FD56FC54" box="[589,719,945,969]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1092" phylum="Arthropoda" rank="genus">
<emphasis id="E3BA6243CC5FFF8D4C52F9E7FD56FC54" box="[589,719,945,969]" italics="true" pageId="7" pageNumber="1092">Coelostoma</emphasis>
</taxonomicName>
, a genus of the hydrophilid tribe
<taxonomicName id="16CEC5D2CC5FFF8D4E8FF986FED9FC75" box="[144,320,976,1000]" class="Insecta" family="Hydrophilidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1092" phylum="Arthropoda" rank="tribe" tribe="Coelostomatini">Coelostomatini</taxonomicName>
, and the Latin
<emphasis id="E3BA6243CC5FFF8D4FECF987FDC5FC75" box="[499,604,977,1000]" italics="true" pageId="7" pageNumber="1092">punctatus</emphasis>
. The name refers to the fact that the head, pronotum and elytral punctation of this new species are similar to the dorsal punctation of many
<taxonomicName id="16CEC5D2CC5FFF8D4F32FE46FE36FBB5" box="[301,431,1040,1064]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1092" phylum="Arthropoda" rank="genus">
<emphasis id="E3BA6243CC5FFF8D4F32FE46FE36FBB5" box="[301,431,1040,1064]" italics="true" pageId="7" pageNumber="1092">Coelostoma</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="99D4EDDACC5FFF8D4E8FFE3DFC87FA97" pageId="7" pageNumber="1092" type="biology_ecology">
<paragraph id="D171BE51CC5FFF8D4E8FFE3DFF7CFB1E" blockId="7.[144,1104,1131,1290]" box="[144,229,1131,1155]" pageId="7" pageNumber="1092">
<heading id="8A39093DCC5FFF8D4E8FFE3DFF7CFB1E" box="[144,229,1131,1155]" fontSize="10" level="3" pageId="7" pageNumber="1092" reason="5">
<emphasis id="E3BA6243CC5FFF8D4E8FFE3DFF7CFB1E" box="[144,229,1131,1155]" italics="true" pageId="7" pageNumber="1092">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="D171BE51CC5FFF8D4E8FFEC4FC87FA97" blockId="7.[144,1104,1131,1290]" pageId="7" pageNumber="1092">
Aquatic. Part of
<typeStatus id="0E7500F3CC5FFF8D4F50FEC5FEE6FB36" box="[335,383,1171,1195]" pageId="7" pageNumber="1092">type</typeStatus>
series was collected in an exposed pool connected to the stream and overgrown by grass, while the second part of the series was collected from the gravel on the side of a small streamlet shaded streamlet in the secondary forest c.
<quantity id="163613B4CC5FFF8D4A0AFE84FBC9FB77" box="[1045,1104,1234,1258]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="1092" unit="m" value="50.0">50 m</quantity>
from the exposed pool (Fikáček and Zhao, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="99D4EDDACC5FFF8D4E8FFF1BFED9FA11" box="[144,320,1357,1420]" pageId="7" pageNumber="1092" type="distribution">
<paragraph id="D171BE51CC5FFF8D4E8FFF1BFE8DFAF8" blockId="7.[144,320,1357,1420]" box="[144,276,1357,1381]" pageId="7" pageNumber="1092">
<heading id="8A39093DCC5FFF8D4E8FFF1BFE8DFAF8" box="[144,276,1357,1381]" fontSize="10" level="3" pageId="7" pageNumber="1092" reason="5">
<emphasis id="E3BA6243CC5FFF8D4E8FFF1BFE8DFAF8" box="[144,276,1357,1381]" italics="true" pageId="7" pageNumber="1092">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="D171BE51CC5FFF8D4E8FFF22FED9FA11" blockId="7.[144,320,1357,1420]" box="[144,320,1396,1420]" pageId="7" pageNumber="1092">
<collectingCountry id="A9D9FEC1CC5FFF8D4E8FFF22FF41FA11" box="[144,216,1396,1420]" name="China" pageId="7" pageNumber="1092">China</collectingCountry>
:
<collectingRegion id="130A70B3CC5FFF8D4EFBFF22FEA5FA11" box="[228,316,1396,1420]" country="China" name="Hainan" pageId="7" pageNumber="1092">Hainan</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>