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<document id="51F7AC197207CBDED59D1B953C5638CB" ID-DOI="10.5281/zenodo.4619317" ID-ISSN="2345-7600" ID-Zenodo-Dep="4619317" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616095026154" checkinUser="carolina" docAuthor="H. - L., Chen &amp; Ng, Peter K. L." docDate="2003" docId="4C3B87DF7C2CFFC2FC1BF9EA538AFB4A" docLanguage="en" docName="RafflesBZool.51.1.61-69.pdf" docOrigin="Raffles Bulletin of Zoology 51 (1)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Leucosia major H.-L. &amp; Ng 2003, new species" docType="treatment" docVersion="3" lastPageNumber="67" masterDocId="B002FFA77C2FFFC4FFD2FF8B574DFFE7" masterDocTitle="On New Species Of Leucosiidae (Crustacea: Decapoda: Brachyura) From Singapore And The South China Sea" masterLastPageNumber="69" masterPageNumber="61" pageNumber="64" updateTime="1698890842442" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6E51B97A551D6CCAC08F7D98764529C3">On New Species Of Leucosiidae (Crustacea: Decapoda: Brachyura) From Singapore And The South China Sea</mods:title>
</mods:titleInfo>
<mods:name id="C7DC8D8FC133A2945CA5F2E5BA1BA114" type="personal">
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<mods:namePart id="E9CADAAB1507C3CCAB0060CE2C3E33DD">H. - L., Chen</mods:namePart>
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<mods:namePart id="33812AE054877D14E00F617AE260741F">Ng, Peter K. L.</mods:namePart>
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<mods:typeOfResource id="97E7C3A78E934C24EA36EF3603B33C38">text</mods:typeOfResource>
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<mods:title id="BBC0F639D0FC01FC390FC7ADD0295A84">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="105D8EEE716EBC2826FD07523CFE21A4">2003</mods:date>
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<mods:number id="67FE0043B2C37337E38732CA30EAB8B2">51</mods:number>
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<mods:identifier id="5DFCE8C13CD2EC34B972C39C6F99F7F3" type="ISSN">2345-7600</mods:identifier>
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<treatment id="4C3B87DF7C2CFFC2FC1BF9EA538AFB4A" LSID="urn:lsid:plazi:treatment:4C3B87DF7C2CFFC2FC1BF9EA538AFB4A" httpUri="http://treatment.plazi.org/id/4C3B87DF7C2CFFC2FC1BF9EA538AFB4A" lastPageId="6" lastPageNumber="67" pageId="3" pageNumber="64">
<subSubSection id="8C8865427C2CFFC7FC1BF9EA5240F99F" box="[969,1293,1632,1657]" pageId="3" pageNumber="64" type="nomenclature">
<paragraph id="C42D36C97C2CFFC7FC1BF9EA5240F99F" blockId="3.[969,1293,1632,1688]" box="[969,1293,1632,1657]" pageId="3" pageNumber="64">
<heading id="9F6581A57C2CFFC7FC1BF9EA5240F99F" bold="true" box="[969,1293,1632,1657]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="64" reason="2">
<emphasis id="F6E6EADB7C2CFFC7FC1BF9EA5240F99F" bold="true" box="[969,1293,1632,1657]" pageId="3" pageNumber="64">
<taxonomicName id="03924D4A7C2CFFC7FC1BF9EA5335F99E" ID-CoL="8YXR8" authority="H.-L. &amp; Ng, 2003" authorityName="H.-L. &amp; Ng" authorityYear="2003" box="[969,1144,1633,1657]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="major" status="sp. nov.">
<emphasis id="F6E6EADB7C2CFFC7FC1BF9EA5335F99E" bold="true" box="[969,1144,1633,1657]" italics="true" pageId="3" pageNumber="64">Leucosia major</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="EDD557A07C2CFFC7FB55F9EB5240F99F" box="[1159,1293,1632,1656]" pageId="3" pageNumber="64" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8865427C2CFFC7FBCBF90A53F3F97F" box="[1049,1214,1665,1688]" pageId="3" pageNumber="64" type="description">
<paragraph id="C42D36C97C2CFFC7FBCBF90A53F3F97F" blockId="3.[969,1293,1632,1688]" box="[1049,1214,1665,1688]" pageId="3" pageNumber="64">
(
<figureCitation id="5CA92A4C7C2CFFC7FBF3F90A5324F97F" box="[1057,1129,1665,1688]" captionStart="Fig" captionStartId="1.[114,150,1980,2001]" captionTargetBox="[152,1415,190,1942]" captionTargetId="figure-102@1.[782,1414,636,1070]" captionTargetPageId="1" captionText="Fig. 1. A, Heteronucia margaritata, new species: holotype male (3.0 by 3.5mm) (ZRC 1993.263); B, Pseudophilyra punctulata, new species: paratype female (6.0 by 5.4 mm) (ZRC 1995.550); C-F, Leucosia major, new species, C: holotype male (38.0 by 34.0mm) (ZRC 1984.50), D, paratype male (37.1 by 32.0 mm) (ZRC 1984.6325), E, F, paratype female (36.0 by 31.9 mm) (ZRC 1984.6326); G, H, Leucosia singaporensis, new species: holotype male (28.0 by 24.0 mm) (ZRC 1965.10.13.49). A-C, E, G = dorsal views; D, F, H = ventral views." figureDoi="http://doi.org/10.5281/zenodo.4619319" httpUri="https://zenodo.org/record/4619319/files/figure.png" pageId="3" pageNumber="64">Figs. 1</figureCitation>
C-F, 4)
</paragraph>
</subSubSection>
<subSubSection id="8C8865427C2CFFC7FCF9F94B5255F83E" pageId="3" pageNumber="64" type="materials_examined">
<paragraph id="C42D36C97C2CFFC7FCF9F94B54DEF8E8" blockId="3.[811,1452,1728,1807]" pageId="3" pageNumber="64">
<emphasis id="F6E6EADB7C2CFFC7FCF9F94B5348F932" bold="true" box="[811,1029,1728,1749]" italics="true" pageId="3" pageNumber="64">Material examined. </emphasis>
<materialsCitation id="74FA3C947C2CFFC7FBDCF94B54DEF8E8" collectingDate="1982-11-26" collectionCode="ZRC" collectorName="H. Huat" country="China" location="China" pageId="3" pageNumber="64" specimenCode="ZRC 1984.50" specimenCount="1" typeStatus="holotype">
<typeStatus id="1B29886B7C2CFFC7FBDCF94B5324F932" box="[1038,1129,1728,1749]" pageId="3" pageNumber="64" type="holotype">Holotype</typeStatus>
- male (38.0 by 34.0mm) (
<specimenCode id="94349EB27C2CFFC7FAA9F94B54CFF915" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="64">ZRC 1984.50</specimenCode>
), South
<collectingCountry id="BC8576597C2CFFC7FC0EF9565355F915" box="[988,1048,1757,1778]" name="China" pageId="3" pageNumber="64">China</collectingCountry>
Sea, off
<collectingCountry id="BC8576597C2CFFC7FBADF95653A4F915" box="[1151,1257,1757,1778]" name="Singapore" pageId="3" pageNumber="64">Singapore</collectingCountry>
, coll.
<collectorName id="6967531F7C2CFFC7FAFCF95652C8F915" box="[1326,1413,1757,1778]" pageId="3" pageNumber="64">H. Huat</collectorName>
,
<date id="B02C10097C2CFFC7FA41F95654C2F8E8" pageId="3" pageNumber="64" value="1982-11-26">
<collectingDate id="A068E9E17C2CFFC7FA41F95654C2F8E8" pageId="3" pageNumber="64" value="1982-11-26">26 Nov.1982</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="C42D36C97C2CFFC7FCF9F8BF5255F83E" blockId="3.[811,1452,1844,2009]" pageId="3" pageNumber="64">
<materialsCitation id="74FA3C947C2CFFC7FCF9F8BF5350F878" collectionCode="ZRC" collectorName="South &amp; Sea &amp; Horsburgh Lighthouse &amp; H. Huat" country="China" location="China" pageId="3" pageNumber="64" specimenCode="ZRC 1984.6325, ZRC 1984.6326" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="1B29886B7C2CFFC7FCF9F8BF54C2F8AE" box="[811,911,1844,1865]" pageId="3" pageNumber="64" type="paratype">Paratypes</typeStatus>
<specimenCount id="D294FD407C2CFFC7FC62F8BF54BAF8AE" box="[944,1015,1844,1865]" pageId="3" pageNumber="64" type="male">1 male</specimenCount>
(37.1 by 32.0 mm) (
<specimenCode id="94349EB27C2CFFC7FB0EF8BF52C7F8AE" box="[1244,1418,1844,1865]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="64">ZRC 1984.6325</specimenCode>
), 1 ovigerous female (38.5 by 39.0 mm) (
<specimenCode id="94349EB27C2CFFC7FB64F8DB5210F882" box="[1206,1373,1872,1893]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="64">ZRC 1984.6326</specimenCode>
),
<collectorName id="6967531F7C2CFFC7FAA0F8DB52E6F882" box="[1394,1451,1872,1893]" pageId="3" pageNumber="64">South</collectorName>
<collectingCountry id="BC8576597C2CFFC7FCF9F8E65424F865" box="[811,873,1901,1922]" name="China" pageId="3" pageNumber="64">China</collectingCountry>
<collectorName id="6967531F7C2CFFC7FCA6F8E654D1F865" box="[884,924,1901,1922]" pageId="3" pageNumber="64">Sea</collectorName>
, off
<collectingCountry id="BC8576597C2CFFC7FC07F8E6530EF865" box="[981,1091,1901,1922]" name="Singapore" pageId="3" pageNumber="64">Singapore</collectingCountry>
, about
<quantity id="036A9B2C7C2CFFC7FB44F8E653B9F865" box="[1174,1268,1901,1922]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="3" pageNumber="64" unit="mi" value="30.0">30 miles</quantity>
from
<collectorName id="6967531F7C2CFFC7FAEEF8E654D4F878" pageId="3" pageNumber="64">Horsburgh Lighthouse</collectorName>
, coll.
<collectorName id="6967531F7C2CFFC7FC02F8015350F878" box="[976,1053,1930,1951]" pageId="3" pageNumber="64">H. Huat</collectorName>
</materialsCitation>
, 1980s;
<materialsCitation id="74FA3C947C2CFFC7FBB8F801525EF83E" collectionCode="ZRC" country="China" location="China" pageId="3" pageNumber="64" specimenCode="ZRC 1985.79" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="D294FD407C2CFFC7FBB8F80153F1F878" box="[1130,1212,1930,1951]" pageId="3" pageNumber="64" type="female">1 female</specimenCount>
(36.0 by
<quantity id="036A9B2C7C2CFFC7FAC6F8015227F878" box="[1300,1386,1930,1951]" metricMagnitude="-2" metricUnit="m" metricValue="3.19" pageId="3" pageNumber="64" unit="mm" value="31.9">31.9 mm</quantity>
) (
<specimenCode id="94349EB27C2CFFC7FAA9F8015433F85B" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="64">ZRC 1985.79</specimenCode>
), South
<collectingCountry id="BC8576597C2CFFC7FC03F82C5346F85B" box="[977,1035,1959,1980]" name="China" pageId="3" pageNumber="64">China</collectingCountry>
Sea, station B77, 6-7 fathoms, mud, coll.
<collectingCountry id="BC8576597C2CFFC7FCF9F84F54C3F83E" box="[811,910,1988,2009]" name="Singapore" pageId="3" pageNumber="64">Singapore</collectingCountry>
Fisheries Research Center, 1955-1956
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8C8865427C2CFFC2FCF9F78A5388FC6A" lastPageId="6" lastPageNumber="67" pageId="3" pageNumber="64" type="diagnosis">
<paragraph id="C42D36C97C2CFFC7FCF9F78A52E6F7DF" blockId="3.[811,1451,2049,2104]" pageId="3" pageNumber="64">
<emphasis id="F6E6EADB7C2CFFC7FCF9F78A54F6F7FE" bold="true" box="[811,955,2049,2073]" italics="true" pageId="3" pageNumber="64">Diagnosis. </emphasis>
Carapace somewhat rhomboidal, 1.12-1.16 times as long as broad. Median part, from front to intestinal
</paragraph>
<caption id="90ED66417C2BFFC0FF5AF87F56EEF7A5" ID-DOI="http://doi.org/10.5281/zenodo.4619325" ID-Zenodo-Dep="4619325" httpUri="https://zenodo.org/record/4619325/files/figure.png" pageId="4" pageNumber="65" startId="4.[136,172,2036,2057]" targetBox="[156,1458,329,1896]" targetPageId="4">
<paragraph id="C42D36C97C2BFFC0FF5AF87F56EEF7A5" blockId="4.[136,1475,2036,2114]" pageId="4" pageNumber="65">
Fig. 3.
<taxonomicName id="03924D4A7C2BFFC0FF03F87F5682F7EE" authority="H.-L. &amp; Ng, 2003" authorityName="H.-L. &amp; Ng" authorityYear="2003" box="[209,463,2036,2057]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="punctulata" status="sp. nov.">
<emphasis id="F6E6EADB7C2BFFC0FF03F87F5682F7EE" box="[209,463,2036,2057]" italics="true" pageId="4" pageNumber="65">Pseudophilyra punctulata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="EDD557A07C2BFFC0FE0FF87F551AF7EE" box="[477,599,2036,2057]" pageId="4" pageNumber="65" rank="species">new species</taxonomicNameLabel>
. A, C, F, G, paratype female (6.0 by 5.4 mm) (ZRC 1995.550); B, D, E, holotype male (5.8 by 5.0 mm) (ZRC 1993.92). A, B, carapaces; C, left cheliped; D, right chela; E, fourth left ambulatory leg; F, fourth right ambulatory leg; G, left third maxilliped.
</paragraph>
</caption>
<caption id="90ED66417C2AFFC1FFA0F85C56DAF7A4" ID-DOI="http://doi.org/10.5281/zenodo.4619329" ID-Zenodo-Dep="4619329" httpUri="https://zenodo.org/record/4619329/files/figure.png" pageId="5" pageNumber="66" startId="5.[114,150,2007,2028]" targetBox="[141,1422,285,1918]" targetPageId="5">
<paragraph id="C42D36C97C2AFFC1FFA0F85C56DAF7A4" blockId="5.[114,1452,2007,2115]" pageId="5" pageNumber="66">
Fig. 4.
<taxonomicName id="03924D4A7C2AFFC1FF65F853561DF80A" authority="H.-L. &amp; Ng, 2003" authorityName="H.-L. &amp; Ng" authorityYear="2003" box="[183,336,2008,2029]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="major" status="sp. nov.">
<emphasis id="F6E6EADB7C2AFFC1FF65F853561DF80A" box="[183,336,2008,2029]" italics="true" pageId="5" pageNumber="66">Leucosia major</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="EDD557A07C2AFFC1FE8FF85C569BF80B" box="[349,470,2007,2028]" pageId="5" pageNumber="66" rank="species">new species</taxonomicNameLabel>
. A-C, E-H, paratype female (36.0 by 31.9 mm) (ZRC 1985.79); D, I-L, holotype male (38.0 by 34.0 mm) (ZRC 19984.50). A, carapace; B, front; C, thoracic sinus; D, right cheliped; E, fourth left ambulatory leg; F, female genital opening; G, right third maxilliped; H, female abdomen; I, male abdomen; J, lateral view of male abdominal segment 6 and telson; K, G1; L, distal part of G1. Scales = 5.0 mm.
</paragraph>
</caption>
<paragraph id="C42D36C97C29FFC2FF5AFCBD56B9F82A" blockId="6.[136,776,822,1997]" pageId="6" pageNumber="67">region more convex than lateral sides. Hepatic region slightly elevated, other regions indistinct. Front divided into 3 teeth: median tooth produced anteriorly downwards, broad, long, lateral teeth bluntly narrow, short, distal ends of all 3 teeth bluntly rounded. Anterolateral margin and distal half of posterolateral margin with fine granules. Posterior carapace margin with distinct granules. Thoracic sinus shallow, with setae, base of sinus with 7 granular tubercles, of which first and seventh tubercles smaller than others, with 2-3 fine granules behind seventh tubercle. Exopod of third maxilliped slender, long, 4.5 times as long as broad, distal end reaching distal part of merus of endopod; merus narrowly triangular, 2.5 times as long as broad, longer than ischium; distal part of female ischium elevated, inner side and base of merus with longitudinal row of setae; male ischium not elevated, glabrous. Chelipeds stout, merus twice as long as broad, distal end broader than basal half, median part of distal one-third elevated, dorsum of distal two-thirds smooth, without granules, dorsal surface of basal half near outer border with patch of 10 fine granules, anterior part with 2 longitudinal rows of 2 or 3 similarly shaped granules; carpus subglobate; palm flattened, 1.5 times as long as broad, borders thin, inner border with coarse granules; movable finger shorter than palm, cutting edges of both fingers with indistinct teeth. First pair of ambulatory legs longest, other legs gradually shorter, last pair shortest. Male abdomen with 5 free segments; segments 3-5 fused, stout, short, borders sinuous; segment 1 short, broad; segment 2 very small; segment 6 broad, flattened, distal part of borders narrowed; telson equilaterally triangular. Female abdomen longitudinally ovate, segments 3-6 fused, stout, large, suture between fused third and fourth segments visible; segment 1 very short, visible only laterally; median part pf segment 2 protruding posteriorly; telson triangular, 1.18 times as long as broad, borders with short setae. G1 stout, sinuous, distal part corneous, hard, yellow (in preservative), surrounded by long setae, finger-like process slightly curving inwards.</paragraph>
<paragraph id="C42D36C97C29FFC2FF5AF87D5388FC6A" blockId="6.[136,775,2038,2093]" lastBlockId="6.[835,1474,822,909]" pageId="6" pageNumber="67">
<emphasis id="F6E6EADB7C29FFC2FF5AF87D578BF7E9" bold="true" box="[136,198,2038,2062]" italics="true" pageId="6" pageNumber="67">Color</emphasis>
(in alcohol).
<emphasis id="F6E6EADB7C29FFC2FE83F87D5613F7E9" bold="true" box="[337,350,2038,2062]" pageId="6" pageNumber="67"></emphasis>
Anterior two-thirds of carapace grayish brown, posterior one-third pale brown, each side of meso- gastric region with 4 white spots in 2 oblique rows followed by longitudinal rows of white stripes. Each side of cardiac region with indistinct black blotch.
</paragraph>
</subSubSection>
<subSubSection id="8C8865427C29FFC2FC91FC3D5226FC0A" pageId="6" pageNumber="67" type="etymology">
<paragraph id="C42D36C97C29FFC2FC91FC3D5226FC0A" blockId="6.[835,1474,950,1005]" pageId="6" pageNumber="67">
<emphasis id="F6E6EADB7C29FFC2FC91FC3D5499FC29" bold="true" box="[835,980,950,974]" italics="true" pageId="6" pageNumber="67">Etymology. </emphasis>
The name is derived from the Latin for large, alluding to the relatively large size of the species.
</paragraph>
</subSubSection>
<subSubSection id="8C8865427C29FFC2FC91FB9D538AFB4A" pageId="6" pageNumber="67" type="discussion">
<paragraph id="C42D36C97C29FFC2FC91FB9D538AFB4A" blockId="6.[835,1474,1046,1197]" pageId="6" pageNumber="67">
<emphasis id="F6E6EADB7C29FFC2FC91FB9D548EFBC9" bold="true" box="[835,963,1046,1070]" italics="true" pageId="6" pageNumber="67">Remarks. </emphasis>
This new species closely resembles
<taxonomicName id="03924D4A7C29FFC2FAB3FB9D5218FBAA" authority="Tyndale-Biscoe &amp; George, 1962" authorityName="Tyndale-Biscoe &amp; George" authorityYear="1962" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="F6E6EADB7C29FFC2FAB3FB9D54DEFBAA" italics="true" pageId="6" pageNumber="67">Leucosia magna</emphasis>
Tyndale-Biscoe &amp; George, 1962
</taxonomicName>
, and
<taxonomicName id="03924D4A7C29FFC2FA79FBBD54ADFB8A" authority="H.-L. &amp; Ng, 2003" authorityName="H.-L. &amp; Ng" authorityYear="2003" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="singaporensis" status="sp. nov.">
<emphasis id="F6E6EADB7C29FFC2FA79FBBD54ADFB8A" italics="true" pageId="6" pageNumber="67">L. singaporensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="EDD557A07C29FFC2FC26FBDD53CFFB8A" box="[1012,1154,1110,1133]" pageId="6" pageNumber="67" rank="species">new species</taxonomicNameLabel>
, in the carapace shape and structure of the thoracic sinus. The differences between these species are summarised in Table 1.
</paragraph>
</subSubSection>
</treatment>
</document>