195 lines
20 KiB
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195 lines
20 KiB
XML
<document id="195D0D7D353C49D83D17AD965562B02C" ID-DOI="10.11646/zootaxa.5319.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8182108" ID-ZooBank="45CD8E29-3320-42E2-A8A8-3A36BC0F41CE" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1690276500814" checkinUser="plazi" docAuthor="Hashizume, Takuto, Yamamoto, Shûhei & Maruyama, Munetoshi" docDate="2023" docId="090C879DFFEF8E11899AFF4DADE7FD4A" docLanguage="en" docName="zootaxa.5319.1.2.pdf" docOrigin="Zootaxa 5319 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5319.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phymatura Sahlberg 1876" docType="treatment" docVersion="1" lastPageNumber="42" masterDocId="F535FFE5FFE08E1E890DFFDAA87BFFDA" masterDocTitle="Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="47" masterPageNumber="27" pageNumber="42" updateTime="1690313925549" updateUser="jonas" zenodo-license-document="CLOSED">
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<mods:title id="2DA786F16359E01868434445D67B198E">Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
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<mods:namePart id="37E165B9A3C942A3EBC9803CE83B5A55">Hashizume, Takuto</mods:namePart>
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<mods:affiliation id="711E56AC27F42A4D6422027845F3B331">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395 Japan.</mods:affiliation>
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<mods:namePart id="EAE1DA634BA5A0372AFC7787814A6321">Yamamoto, Shûhei</mods:namePart>
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<mods:affiliation id="3E82248C98A2C4BE9E883DDA3BA313CC">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo, 060 - 0810 Japan. s. yamamoto. 64 @ gmail. com; https: // orcid. org / 0000 - 0002 - 4162 - 8457</mods:affiliation>
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<mods:nameIdentifier id="E7ADFF6C8C4D8BEB03045AA29B5020A2" type="email">s.yamamoto.64@gmail.com</mods:nameIdentifier>
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<mods:namePart id="1B1F1FEC6BE8BF005B5033106A9F1C5C">Maruyama, Munetoshi</mods:namePart>
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<mods:affiliation id="A2DF0C59D8439BA3C487C37A5FF4F496">The Kyushu University Museum, Fukuoka, 812 - 8581 Japan.</mods:affiliation>
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<mods:title id="A63F3931A2C78A8B0F00159A4E66240C">Zootaxa</mods:title>
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<mods:date id="41037DEE841DCA6D5C1233C091837803">2023</mods:date>
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<mods:number id="0F141ECC2056DEDC0E164F68DB57CF49">2023-07-24</mods:number>
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<heading id="DA5281E7FFEF8E11899AFF4DAA4CFF6B" bold="true" box="[151,567,151,178]" fontSize="11" level="1" pageId="15" pageNumber="42" reason="1">
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<emphasis id="B3D1EA99FFEF8E11899AFF4DAA4CFF6B" bold="true" box="[151,567,151,178]" pageId="15" pageNumber="42">
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Genus
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<taxonomicName id="46A54D08FFEF8E1189E3FF4DAA4CFF6B" ID-CoL="8VRZ4" authority="Sahlberg, 1876" authorityName="Sahlberg" authorityYear="1876" box="[238,567,151,178]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="42" phylum="Arthropoda" rank="genus">
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<emphasis id="B3D1EA99FFEF8E1189E3FF4DA90EFF68" bold="true" box="[238,373,151,178]" italics="true" pageId="15" pageNumber="42">Phymatura</emphasis>
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<bibRefCitation id="E5344B7AFFEF8E118870FF4DAA4CFF6B" author="Sahlberg, J." box="[381,567,151,177]" pageId="15" pageNumber="42" pagination="1 - 248" refId="ref16559" refString="Sahlberg, J. (1876) Enumeratio Coleopterorum brachelytrorum Fenniae. Systematisk f ˆ rteckning ˆ fver de inom Finlands naturalhistoriska omrade hittills funna Coleoptera Brachelytra jemte uppgift om arternas utbredning och beskrifningar af nya och mindre kanda species. I. Staphylinidae. Acta Societatis pro Fauna et Flora Fennica, 1 (3), 1 - 248." type="journal article" year="1876">Sahlberg, 1876</bibRefCitation>
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</emphasis>
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<paragraph id="811A368BFFEF8E11899AFF66AA84FF0F" blockId="15.[151,767,151,214]" box="[151,767,187,214]" pageId="15" pageNumber="42">[Japanese name: Kinokotsuyakeshi-hanekakushi-zoku]</paragraph>
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<taxonomicName id="46A54D08FFEF8E11899AFF23A9A8FECB" ID-CoL="8VRZ4" authority="Sahlberg, 1876: 85" authorityName="Sahlberg" authorityPageNumber="85" authorityYear="1876" box="[151,467,249,273]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="42" phylum="Arthropoda" rank="genus">
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<emphasis id="B3D1EA99FFEF8E11899AFF23A973FECB" box="[151,264,249,273]" italics="true" pageId="15" pageNumber="42">Phymatura</emphasis>
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<bibRefCitation id="E5344B7AFFEF8E11881DFF23A9A8FECB" author="Sahlberg, J." box="[272,467,249,273]" pageId="15" pageNumber="42" pagination="1 - 248" refId="ref16559" refString="Sahlberg, J. (1876) Enumeratio Coleopterorum brachelytrorum Fenniae. Systematisk f ˆ rteckning ˆ fver de inom Finlands naturalhistoriska omrade hittills funna Coleoptera Brachelytra jemte uppgift om arternas utbredning och beskrifningar af nya och mindre kanda species. I. Staphylinidae. Acta Societatis pro Fauna et Flora Fennica, 1 (3), 1 - 248." type="journal article" year="1876">Sahlberg, 1876: 85</bibRefCitation>
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(original description;
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<typeStatus id="5E1E8829FFEF8E118BB6FF23AA9CFECB" box="[699,743,249,273]" pageId="15" pageNumber="42">type</typeStatus>
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species:
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<taxonomicName id="46A54D08FFEF8E118A4AFF23ACB9FECB" authority="Kraatz, 1856" authorityName="Kraatz" authorityYear="1856" box="[839,1218,249,273]" class="Insecta" family="Staphylinidae" genus="Bolitochara" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="42" phylum="Arthropoda" rank="species" species="brevicollis">
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<emphasis id="B3D1EA99FFEF8E118A4AFF23AC49FECB" box="[839,1074,249,273]" italics="true" pageId="15" pageNumber="42">Bolitochara brevicollis</emphasis>
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Kraatz, 1856
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</taxonomicName>
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, fixed by subsequent designation by
|
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<treatmentCitation id="0004109AFFEF8E11886FFEC2AA6EFEF5" author="Casey, T. L." box="[354,533,279,304]" page="264" pageId="15" pageNumber="42" year="1906">
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<bibRefCitation id="E5344B7AFFEF8E11886FFEC2AA6EFEF5" author="Casey, T. L." box="[354,533,279,304]" pageId="15" pageNumber="42" pagination="125 - 434" refId="ref15341" refString="Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Science of St. Louis, 16, 125 - 434. https: // doi. org / 10.5962 / bhl. title. 48654" type="journal article" year="1906">Casey, 1906: 264</bibRefCitation>
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</treatmentCitation>
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).
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</paragraph>
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<subSubSection id="C9BF6500FFEF8E11899AFEEDAA06FE94" box="[151,637,310,334]" pageId="15" pageNumber="42" type="reference_group">
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<paragraph id="811A368BFFEF8E11899AFEEDAA06FE94" blockId="15.[151,1437,249,365]" box="[151,637,310,334]" pageId="15" pageNumber="42">
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<treatmentCitationGroup id="A1B511A5FFEF8E11899AFEEDAA06FE94" box="[151,637,310,334]" pageId="15" pageNumber="42">
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<treatmentCitation id="0004109AFFEF8E11899AFEEDA9E7FE94" author="Casey, T. L." box="[151,412,310,334]" page="272" pageId="15" pageNumber="42" year="1906">
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<taxonomicName id="46A54D08FFEF8E11899AFEEDA9E7FE94" ID-CoL="8XQNH" authority="Casey, 1906: 272" authorityName="Casey" authorityPageNumber="272" authorityYear="1906" box="[151,412,310,334]" class="Insecta" family="Staphylinidae" genus="Venusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="42" phylum="Arthropoda" rank="genus">
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<emphasis id="B3D1EA99FFEF8E11899AFEEDA8A4FE94" box="[151,223,311,334]" italics="true" pageId="15" pageNumber="42">Venusa</emphasis>
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<bibRefCitation id="E5344B7AFFEF8E1189E5FEECA9E7FE94" author="Casey, T. L." box="[232,412,310,334]" pageId="15" pageNumber="42" pagination="125 - 434" refId="ref15341" refString="Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Science of St. Louis, 16, 125 - 434. https: // doi. org / 10.5962 / bhl. title. 48654" type="journal article" year="1906">Casey, 1906: 272</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(original description;
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C9BF6500FFEF8E118B88FEEDAB11FEB7" pageId="15" pageNumber="42" type="synonymic_list">
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<paragraph id="811A368BFFEF8E118B88FEEDAB11FEB7" blockId="15.[151,1437,249,365]" pageId="15" pageNumber="42">
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<typeStatus id="5E1E8829FFEF8E118B88FEEDAACAFE95" box="[645,689,311,335]" pageId="15" pageNumber="42">type</typeStatus>
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species:
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||
<taxonomicName id="46A54D08FFEF8E118A1EFEEDAC5BFE94" authority="Casey, 1906" authorityName="Casey" authorityYear="1906" box="[787,1056,310,334]" class="Insecta" family="Staphylinidae" genus="Venusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="42" phylum="Arthropoda" rank="species" species="picta">
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||
<emphasis id="B3D1EA99FFEF8E118A1EFEEDABECFE94" box="[787,919,311,334]" italics="true" pageId="15" pageNumber="42">Venusa picta</emphasis>
|
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<bibRefCitation id="E5344B7AFFEF8E118A92FEECAC5BFE94" author="Casey, T. L." box="[927,1056,310,334]" pageId="15" pageNumber="42" pagination="125 - 434" refId="ref15341" refString="Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Science of St. Louis, 16, 125 - 434. https: // doi. org / 10.5962 / bhl. title. 48654" type="journal article" year="1906">Casey, 1906</bibRefCitation>
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</taxonomicName>
|
||
, fixed by subsequent designation by
|
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<treatmentCitation id="0004109AFFEF8E1189CAFE8FA902FEB7" author="Fenyes, J. S." box="[199,377,341,365]" page="26" pageId="15" pageNumber="42" year="1918">
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<bibRefCitation id="E5344B7AFFEF8E1189CAFE8FA902FEB7" author="Fenyes, J. S." box="[199,377,341,365]" pageId="15" pageNumber="42" pagination="1 - 110" refId="ref15394" refString="Fenyes, J. S. (1918) Coleoptera. Fam. Staphylinidae, Subfam. Aleocharinae. In: Wytsmam, P. A. (Ed.), Genera Insectorum. Vol. XXVII. M. Nijhoff, The Hague, pp. 1 - 110." type="book chapter" year="1918">Fenyes, 1918: 26</bibRefCitation>
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</treatmentCitation>
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);
|
||
<treatmentCitation id="0004109AFFEF8E118884FE8FAA4BFEB7" author="Ashe, J. S." box="[393,560,341,365]" page="371" pageId="15" pageNumber="42" year="1992">
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<bibRefCitation id="E5344B7AFFEF8E118884FE8FAA4BFEB7" author="Ashe, J. S." box="[393,560,341,365]" pageId="15" pageNumber="42" pagination="335 - 406" refId="ref14897" refString="Ashe, J. S. (1992) Phylogeny and revision of genera of the subtribe Bolitocharina (Coleoptera: Staphylinidae: Aleocharinae). University of Kansas science bulletin, 54 (10), 335 - 406. https: // doi. org / 10.5962 / bhl. part. 19642" type="journal article" year="1992">Ashe, 1992: 371</bibRefCitation>
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</treatmentCitation>
|
||
(as a synonym of
|
||
<taxonomicName id="46A54D08FFEF8E118BE6FE8FAB27FEB7" box="[747,860,341,365]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B3D1EA99FFEF8E118BE6FE8FAB27FEB7" box="[747,860,341,365]" italics="true" pageId="15" pageNumber="42">Phymatura</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
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<subSubSection id="C9BF6500FFEF8E11899AFE4CACBBFE0E" pageId="15" pageNumber="42" type="discussion">
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<paragraph id="811A368BFFEF8E11899AFE4CACBBFE0E" blockId="15.[151,1437,406,468]" pageId="15" pageNumber="42">
|
||
<emphasis id="B3D1EA99FFEF8E11899AFE4CA970FE6A" bold="true" box="[151,267,406,432]" pageId="15" pageNumber="42">Remarks.</emphasis>
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||
The genus
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||
<taxonomicName id="46A54D08FFEF8E118884FE4CAA7EFE6A" box="[393,517,406,432]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B3D1EA99FFEF8E118884FE4CAA7EFE6A" box="[393,517,406,432]" italics="true" pageId="15" pageNumber="42">Phymatura</emphasis>
|
||
</taxonomicName>
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||
can be distinguished from the genus
|
||
<taxonomicName id="46A54D08FFEF8E118A93FE4CAC53FE6A" authorityName="Cameron" authorityYear="1920" box="[926,1064,406,432]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="42" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B3D1EA99FFEF8E118A93FE4CAC53FE6A" box="[926,1064,406,432]" italics="true" pageId="15" pageNumber="42">Pseudatheta</emphasis>
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||
</taxonomicName>
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by the following character states: mesoventrite with a longitudinal carina; flabellum of hind wing with at least three setose lobes.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C9BF6500FFEF8E11899AFDC3AA9CFDE9" box="[151,743,536,563]" pageId="15" pageNumber="42" type="multiple">
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<paragraph id="811A368BFFEF8E11899AFDC3AA9CFDE9" blockId="15.[151,743,536,563]" box="[151,743,536,563]" pageId="15" pageNumber="42">
|
||
<heading id="DA5281E7FFEF8E11899AFDC3AA9CFDE9" bold="true" box="[151,743,536,563]" fontSize="11" level="1" pageId="15" pageNumber="42" reason="1">
|
||
<emphasis id="B3D1EA99FFEF8E11899AFDC3AA9CFDE9" bold="true" box="[151,743,536,563]" pageId="15" pageNumber="42">
|
||
Key to species of the genus
|
||
<taxonomicName id="46A54D08FFEF8E1188E1FDC2AA08FDE9" ID-CoL="8VRZ4" box="[492,627,536,563]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B3D1EA99FFEF8E1188E1FDC2AA08FDE9" bold="true" box="[492,627,536,563]" italics="true" pageId="15" pageNumber="42">Phymatura</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F9B2761BFFEF8E118B97FDC3AA9CFDE9" box="[666,743,537,563]" name="Japan" pageId="15" pageNumber="42">Japan</collectingCountry>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9BF6500FFEF8E11899AFD84ADE7FD4A" pageId="15" pageNumber="42" type="key">
|
||
<keyStep id="3A512E2EFFEF8E11899AFD84ADE7FD4A" pageId="15" pageNumber="42">
|
||
<paragraph id="811A368BFFEF8E11899AFD84ADE6FDA9" blockId="15.[151,1437,605,657]" box="[151,1437,605,627]" pageId="15" pageNumber="42">
|
||
<keyLead id="3A5495BEFFEF8E11899AFD84ADE6FDA9" box="[151,1437,605,627]" pageId="15" pageNumber="42">
|
||
1. Body nearly uniformly reddish; eyes smaller, length shorter than 40% length of head...................
|
||
<taxonomicName id="46A54D08FFEF8E118DF4FD87ADE6FDA9" authority="Assing" authorityName="Assing" authorityYear="2021" box="[1273,1437,605,627]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="42" phylum="Arthropoda" rank="species" species="russa">
|
||
<emphasis id="B3D1EA99FFEF8E118DF4FD87AD2FFDA9" box="[1273,1364,605,627]" italics="true" pageId="15" pageNumber="42">Ph. russa</emphasis>
|
||
Assing
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="811A368BFFEF8E11899AFDA1ADE7FD4A" blockId="15.[151,1437,605,657]" box="[151,1436,635,657]" pageId="15" pageNumber="42">
|
||
<keyLead id="3A5495BEFFEF8E11899AFDA1ADE7FD4A" box="[151,1436,635,657]" pageId="15" pageNumber="42">
|
||
- Body uniformly darker; eyes larger, length longer than 40% length of head...............
|
||
<taxonomicName id="46A54D08FFEF8E118D6AFDA1AD44FD4A" authority="Cameron" authorityName="Cameron" authorityYear="2023" box="[1127,1343,635,656]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="42" phylum="Arthropoda" rank="species" species="japonica" status="stat. rev.">
|
||
<emphasis id="B3D1EA99FFEF8E118D6AFDA1AC9AFD4A" box="[1127,1249,635,656]" italics="true" pageId="15" pageNumber="42">Ph. japonica</emphasis>
|
||
Cameron
|
||
</taxonomicName>
|
||
<emphasis id="B3D1EA99FFEF8E118C48FDA1ADE7FD4A" bold="true" box="[1349,1436,635,656]" pageId="15" pageNumber="42">
|
||
<taxonomicNameLabel id="A8E257E2FFEF8E118C48FDA1ADE7FD4A" box="[1349,1436,635,656]" pageId="15" pageNumber="42">stat. rev.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |