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172 lines
25 KiB
XML
<document id="3905C65B598727969FC15DA43EC8AD0F" ID-DOI="10.11646/zootaxa.5325.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8243389" ID-ZooBank="670D8B76-9F1F-48C5-9484-E2EFD8B09C24" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691941684947" checkinUser="plazi" docAuthor="Abstract, Júlio Cezar Mário Chaul" docDate="2023" docId="680E87F4624DFFE749ABBA57FD0AFDAC" docLanguage="en" docName="zootaxa.5325.3.1.pdf" docOrigin="Zootaxa 5325 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5325.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zigrasimecia tonsora Barden & Grimaldi 2013" docType="treatment" docVersion="2" lastPageNumber="326" masterDocId="9437FF8C625AFFFE493CBC44FFE6FFCD" masterDocTitle="A revision of the Cretaceous ant genus Zigrasimecia Barden & Grimaldi, 2013 (Hymenoptera: Formicidae: † Zigrasimeciinae)" masterLastPageNumber="341" masterPageNumber="301" pageNumber="324" updateTime="1691942332973" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9C00B118E21A08753095BABD5A1C2B33">A revision of the Cretaceous ant genus Zigrasimecia Barden & Grimaldi, 2013 (Hymenoptera: Formicidae: † Zigrasimeciinae)</mods:title>
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<mods:namePart id="0D87203FF5FBC5B0D7A98F20F54BCA3A">Abstract, Júlio Cezar Mário Chaul</mods:namePart>
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<mods:typeOfResource id="85FF9CB2D31A363A1EA9EF978DDFD625">text</mods:typeOfResource>
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<mods:title id="4C3132236D69A26D877AF14C2438AC93">Zootaxa</mods:title>
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<mods:part id="5F6D7B3AF769F5B466A54105DA6FEE9D">
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<mods:date id="B6330674BFCB31E0D4FA9E6B59025A0A">2023</mods:date>
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<mods:number id="32A1674D77DD50F871F9703F60A4BA3A">2023-08-09</mods:number>
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<mods:number id="AC29B4F8F2790D76E39040F734624CD0">5325</mods:number>
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<mods:classification id="AEC8201008FC121C23D11B40FAF6C73E">journal article</mods:classification>
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<mods:identifier id="2772386FEF75FC5D0712FA3AA3872332" type="DOI">10.11646/zootaxa.5325.3.1</mods:identifier>
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<subSubSection id="A8BD6569624DFFE949ABBA57FD07F9E1" box="[151,737,1554,1581]" pageId="23" pageNumber="324" type="nomenclature">
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<paragraph id="E01836E2624DFFE949ABBA57FD07F9E1" blockId="23.[151,737,1554,1581]" box="[151,737,1554,1581]" pageId="23" pageNumber="324">
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<heading id="BB50818E624DFFE949ABBA57FD07F9E1" bold="true" box="[151,737,1554,1581]" fontSize="11" level="1" pageId="23" pageNumber="324" reason="1">
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<taxonomicName id="27A74D61624DFFE949ABBA57FD07F9E1" authority="Barden & Grimaldi, 2013" authorityName="Barden & Grimaldi" authorityYear="2013" box="[151,737,1554,1581]" class="Insecta" family="Formicidae" genus="Zigrasimecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="tonsora">
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<emphasis id="D2D3EAF0624DFFE949ABBA57FD07F9E1" bold="true" box="[151,737,1554,1581]" pageId="23" pageNumber="324">
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<emphasis id="D2D3EAF0624DFFE949ABBA57FE7EF9E0" bold="true" box="[151,408,1555,1581]" italics="true" pageId="23" pageNumber="324">Zigrasimecia tonsora</emphasis>
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<bibRefCitation id="84364B13624DFFE948A3BA57FD07F9E1" author="Barden, P. & Grimaldi, D. A." box="[415,737,1554,1581]" pageId="23" pageNumber="324" pagination="405 - 412" refId="ref23029" refString="Barden, P. & Grimaldi, D. A. (2013) A new genus of highly specialized ants in Cretaceous Burmese amber (Hymenoptera: Formicidae). Zootaxa, 3681 (4), 405 - 412. https: // doi. org / 10.11646 / zootaxa. 3681.4.5" type="journal article" year="2013">Barden & Grimaldi, 2013</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A8BD6569624DFFE949ABBA1EFCFFF880" pageId="23" pageNumber="324" type="diagnosis">
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<paragraph id="E01836E2624DFFE949ABBA1EFCFFF880" blockId="23.[151,1437,1626,2013]" pageId="23" pageNumber="324">
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<emphasis id="D2D3EAF0624DFFE949ABBA1EFE8BF9B9" bold="true" box="[151,365,1626,1652]" pageId="23" pageNumber="324">Diagnosis (queen)</emphasis>
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. Frons with a shallow V-shaped groove; vertex with a pair of cuticular ridges or protuberances (see comment below). Mandible apical tooth strongly offset from mandible's outer margin. Compound eyes ovoid (in relation to
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<taxonomicName id="27A74D61624DFFE94832BAE7FE70F970" authorityName="Zhuang, Ran, Li, Feng & Liu" authorityYear="2022" box="[270,406,1699,1725]" class="Insecta" family="Formicidae" genus="Zigrasimecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="goldingot">
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<emphasis id="D2D3EAF0624DFFE94832BAE7FE70F970" box="[270,406,1699,1725]" italics="true" pageId="23" pageNumber="324">Z. goldingot</emphasis>
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</taxonomicName>
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). Pilosity on dorsum of head composed of abundant, thin, flexuous setae (
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<figureCitation id="789C2A67624DFFE94DF5BAE7FAECF970" box="[1225,1290,1699,1725]" captionStart="FIGURE 1" captionStartId="2.[151,250,1788,1812]" captionTargetBox="[157,1432,1033,1755]" captionTargetId="figure-447@2.[151,1435,1026,1761]" captionTargetPageId="2" captionText="FIGURE 1. Schematic drawings of most of the measurements taken from the specimens (presented in the species descriptions or in Appendix 1). See Materials and Methods or Appendix 1 for a detailed explanation of the measurements. Codes: a, head length; b, head length 2; c, head width; d, head width 2; e, head width 3; f, head maximum depth; g, mesosoma length (see also Fig.6); h, mesosoma height; i, propodeal spiracle length; j, petiole anterior face length; k, posttergite 3 length; o, posttergite 4 length. Measurements a, g, j, k, and o summed equal total length (TL). Gaster measurements l, m, n, and p are additional metasomal measurements that are useful if the measurements k and o are blocked (see Appendix 1)." figureDoi="http://doi.org/10.5281/zenodo.8243391" httpUri="https://zenodo.org/record/8243391/files/figure.png" pageId="23" pageNumber="324">Fig. 1</figureCitation>
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, E of
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<bibRefCitation id="84364B13624DFFE94C77BAE7FE86F92C" author="Barden, P. & Grimaldi, D. A." pageId="23" pageNumber="324" pagination="405 - 412" refId="ref23029" refString="Barden, P. & Grimaldi, D. A. (2013) A new genus of highly specialized ants in Cretaceous Burmese amber (Hymenoptera: Formicidae). Zootaxa, 3681 (4), 405 - 412. https: // doi. org / 10.11646 / zootaxa. 3681.4.5" type="journal article" year="2013">Barden & Grimaldi, 2013</bibRefCitation>
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, better observed in Antweb's high-resolution version than in the article's figure in the pdf), similar setae also present on dorsum of pronotum, mesothorax and propodeum. Dorsal propodeal surface relatively long (in relation to
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<taxonomicName id="27A74D61624DFFE9480BBB4BFE59F8E4" authorityName="Zhuang, Ran, Li, Feng & Liu" authorityYear="2022" box="[311,447,1807,1833]" class="Insecta" family="Formicidae" genus="Zigrasimecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="goldingot">
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<emphasis id="D2D3EAF0624DFFE9480BBB4BFE59F8E4" box="[311,447,1807,1833]" italics="true" pageId="23" pageNumber="324">Z. goldingot</emphasis>
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</taxonomicName>
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). Petiolar node forming a distinct, truncate dorsal surface, apex transversally apparently without lateral protuberances (or apparently not bilobed).
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</paragraph>
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</subSubSection>
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<subSubSection id="A8BD6569624DFFE749FBBB12FD0AFDAC" lastPageId="25" lastPageNumber="326" pageId="23" pageNumber="324" type="discussion">
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<paragraph id="E01836E2624DFFE949FBBB12FCD7F810" blockId="23.[151,1437,1626,2013]" pageId="23" pageNumber="324">
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<emphasis id="D2D3EAF0624DFFE949FBBB12FEAEF8BD" bold="true" box="[199,328,1878,1904]" pageId="23" pageNumber="324">Comments</emphasis>
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. The
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<typeStatus id="3F1C8840624DFFE948B4BB13FE0CF8BC" box="[392,490,1879,1905]" pageId="23" pageNumber="324">holotype</typeStatus>
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of
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<taxonomicName id="27A74D61624DFFE94B33BB13FD65F8BD" authorityName="Barden & Grimaldi" authorityYear="2013" box="[527,643,1879,1904]" class="Insecta" family="Formicidae" genus="Zigrasimecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="tonsora">
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<emphasis id="D2D3EAF0624DFFE94B33BB13FD65F8BD" box="[527,643,1879,1904]" italics="true" pageId="23" pageNumber="324">Z. tonsora</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="27A74D61624DFFE94BACBB13FCC2F8BD" authorityName="Abstract" authorityYear="2023" box="[656,804,1879,1904]" class="Insecta" family="Formicidae" genus="Zigrasimecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="genus">
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<emphasis id="D2D3EAF0624DFFE94BACBB13FCC2F8BD" box="[656,804,1879,1904]" italics="true" pageId="23" pageNumber="324">Zigrasimecia</emphasis>
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</taxonomicName>
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genotype, is not a perfectly preserved specimen and, for this reason, and for being a queen rather than a worker, creates a taxonomical challenge in ascertaining conspecificity between it and any worker specimen. The reasons why the new species described here do not belong to
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<taxonomicName id="27A74D61624DFFE94C15BBDBFA7BF875" authorityName="Barden & Grimaldi" authorityYear="2013" box="[1321,1437,1951,1976]" class="Insecta" family="Formicidae" genus="Zigrasimecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="tonsora">
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<emphasis id="D2D3EAF0624DFFE94C15BBDBFA7BF875" box="[1321,1437,1951,1976]" italics="true" pageId="23" pageNumber="324">Z. tonsora</emphasis>
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</taxonomicName>
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are presented above (see above in
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<taxonomicName id="27A74D61624DFFE94B2BBB87FCC0F810" authority="Comments" authorityName="Comments" box="[535,806,1987,2013]" class="Insecta" family="Formicidae" genus="Zigrasimecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="goldingot">
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<emphasis id="D2D3EAF0624DFFE94B2BBB87FD46F810" box="[535,672,1987,2013]" italics="true" pageId="23" pageNumber="324">Z. goldingot</emphasis>
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Comments
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</taxonomicName>
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).
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</paragraph>
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<caption id="B4D8666A6242FFE649ABBAC5FB03F8E7" ID-DOI="http://doi.org/10.5281/zenodo.8243413" ID-Zenodo-Dep="8243413" httpUri="https://zenodo.org/record/8243413/files/figure.png" pageId="24" pageNumber="325" startId="24.[151,250,1665,1689]" targetBox="[151,1436,181,1642]" targetPageId="24" targetType="figure">
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<paragraph id="E01836E26242FFE649ABBAC5FB03F8E7" blockId="24.[151,1437,1665,1834]" pageId="24" pageNumber="325">
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<emphasis id="D2D3EAF06242FFE649ABBAC5FEC5F954" bold="true" box="[151,291,1665,1689]" pageId="24" pageNumber="325">FIGURE 13.</emphasis>
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Holotype of
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<taxonomicName id="27A74D616242FFE64888BAC6FD90F957" authorityName="Abstract" authorityYear="2023" box="[436,630,1666,1690]" class="Insecta" family="Formicidae" genus="Zigrasimecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="thate">
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<emphasis id="D2D3EAF06242FFE64888BAC6FD90F957" box="[436,630,1666,1690]" italics="true" pageId="24" pageNumber="325">Zigrasimecia thate</emphasis>
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</taxonomicName>
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(ANTWEB1038912).
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<emphasis id="D2D3EAF06242FFE64A5ABAC5FC9EF954" bold="true" box="[870,888,1665,1689]" pageId="24" pageNumber="325">A</emphasis>
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, full-face view of the head, black triangles indicate mandible dorsal setae;
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<emphasis id="D2D3EAF06242FFE648B6BAE2FE7DF973" bold="true" box="[394,411,1702,1726]" pageId="24" pageNumber="325">B</emphasis>
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, profile view of left side of body;
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<emphasis id="D2D3EAF06242FFE64A2CBAE1FCC4F970" bold="true" box="[784,802,1701,1725]" pageId="24" pageNumber="325">C</emphasis>
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, anterior view of the head;
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<emphasis id="D2D3EAF06242FFE64D6ABAE2FB8EF973" bold="true" box="[1110,1128,1702,1726]" pageId="24" pageNumber="325">D</emphasis>
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, dorsal view of the body;
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<emphasis id="D2D3EAF06242FFE64CB9BAE2FA70F973" bold="true" box="[1413,1430,1702,1726]" pageId="24" pageNumber="325">E</emphasis>
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, zoom of flagelliform setae on frons and vertex (same view as C);
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<emphasis id="D2D3EAF06242FFE64A08BA8EFCA5F92F" bold="true" box="[820,835,1738,1762]" pageId="24" pageNumber="325">F</emphasis>
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, detail of mesosomal profile. Abbreviations:
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<emphasis id="D2D3EAF06242FFE64C2DBA8EFAB0F92F" bold="true" box="[1297,1366,1738,1762]" pageId="24" pageNumber="325">Lmspl</emphasis>
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, lower mesopleuron;
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<emphasis id="D2D3EAF06242FFE64816BAAAFE88F8CB" bold="true" box="[298,366,1774,1798]" pageId="24" pageNumber="325">Lmtpl</emphasis>
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, lower metapleuron;
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<emphasis id="D2D3EAF06242FFE64B73BAAAFD9BF8CB" bold="true" box="[591,637,1774,1798]" pageId="24" pageNumber="325">mgo</emphasis>
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, metapleural gland opening;
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<emphasis id="D2D3EAF06242FFE64A8EBAAAFC38F8CB" bold="true" box="[946,990,1774,1798]" pageId="24" pageNumber="325">Ppd</emphasis>
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, propodeum;
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<emphasis id="D2D3EAF06242FFE64D4DBAAAFB7CF8CB" bold="true" box="[1137,1178,1774,1798]" pageId="24" pageNumber="325">Prn</emphasis>
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, pronotum;
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<emphasis id="D2D3EAF06242FFE64C20BAAAFAD2F8CB" bold="true" box="[1308,1332,1774,1798]" pageId="24" pageNumber="325">sp</emphasis>
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, spiracles (metanotal and propodeal);
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<emphasis id="D2D3EAF06242FFE6488CBB56FE10F8E7" bold="true" box="[432,502,1810,1834]" pageId="24" pageNumber="325">Umspl</emphasis>
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, upper mesopleuron. Scale bars are 0.2 mm in B and D, and 0.1 mm in A.
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</paragraph>
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</caption>
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<paragraph id="E01836E26242FFE749FBBB1CFE94FDD4" blockId="24.[151,1436,1880,2014]" lastBlockId="25.[151,1436,151,609]" lastPageId="25" lastPageNumber="326" pageId="24" pageNumber="325">
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The
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<typeStatus id="3F1C88406242FFE649C4BB1CFEBCF8BF" box="[248,346,1880,1906]" pageId="24" pageNumber="325">holotype</typeStatus>
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head appears slightly desiccated and wrinkled; the "coronal rugosities", a pair of vertexal cuticular ridges, could be a preservational feature (
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<bibRefCitation id="84364B136242FFE64B60BB38FCFAF85B" author="Cao, H. & Boudinot, B. E. & Wang, Z. & Miao, X. & Shih, C. & Ren, D. & Gao, T." box="[604,796,1916,1942]" pageId="24" pageNumber="325" pagination="161 - 173" refId="ref23785" refString="Cao, H., Boudinot, B. E., Wang, Z., Miao, X., Shih, C., Ren, D. & Gao, T. (2020 a). Two new iron maiden ants from Burmese amber (Hymenoptera: Formicidae: † Zigrasimeciini). Myrmecological News, 30, 161 - 173. https: // doi. org / 10.25849 / myrmecol. news _ 030: 161" type="journal article" year="2020">
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Cao
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<emphasis id="D2D3EAF06242FFE64BACBB39FD20F85B" box="[656,710,1916,1942]" italics="true" pageId="24" pageNumber="325">et al.</emphasis>
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, 2020a
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</bibRefCitation>
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: note 7), although an anonymous reviewer called attention to the note by these authors claiming the paired structure is, in fact, symmetrical. By the illustration in the original publication the ridges are not perfectly mirrored, but considering how much compression and stretching forces the fossils are subjected to, that degree of displacement of one ridge in relation to the other is still acceptable to account for them as a symmetrical paired structure, if they are a natural trait of the specimen. Indeed, most burmite ants observed by the author are not free from at least a minimal amount of assymetry due to preservation in at least one part of the body. So, whether the vertexal ridges are a natural trait of
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<taxonomicName id="27A74D616243FFE74AF3BD47FBA5FED1" authorityName="Barden & Grimaldi" authorityYear="2013" box="[975,1091,259,284]" class="Insecta" family="Formicidae" genus="Zigrasimecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="tonsora">
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<emphasis id="D2D3EAF06243FFE74AF3BD47FBA5FED1" box="[975,1091,259,284]" italics="true" pageId="25" pageNumber="326">Z. tonsora</emphasis>
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</taxonomicName>
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is an open question that might be solved as more specimens are discovered. In addition, a drawing of the toruli (
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<figureCitation id="789C2A676243FFE74D13BD63FB92FE8C" box="[1071,1140,295,321]" captionStart="FIGURE 2" captionStartId="5.[151,250,1793,1817]" captionTargetBox="[151,1434,814,1770]" captionTargetId="figure-1048@5.[151,1435,814,1770]" captionTargetPageId="5" captionText="FIGURE 2. Masticatory margin of mandible in Gerontoformica and Zigrasimecia. A, Gerontoformica sp. (ANTWEB1032629); B, Z. ferox (ANTWEB1008126); C, Z. chuyangsui (ANTWEB1032660); D, Z. tonsora (ANTWEB11008098). Black triangles indicate incision between apical and basal teeth (or basal lamella). Image of Z. tonsora by Philip Barden and image of Z. ferox by Vincent Perrichot, courtesy of Antweb.org." figureDoi="http://doi.org/10.5281/zenodo.8243393" httpUri="https://zenodo.org/record/8243393/files/figure.png" pageId="25" pageNumber="326">Fig. 2</figureCitation>
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, C of
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<bibRefCitation id="84364B136243FFE74D86BD63FF37FEA8" author="Barden, P. & Grimaldi, D. A." pageId="25" pageNumber="326" pagination="405 - 412" refId="ref23029" refString="Barden, P. & Grimaldi, D. A. (2013) A new genus of highly specialized ants in Cretaceous Burmese amber (Hymenoptera: Formicidae). Zootaxa, 3681 (4), 405 - 412. https: // doi. org / 10.11646 / zootaxa. 3681.4.5" type="journal article" year="2013">Barden & Grimaldi, 2013</bibRefCitation>
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) is not in perfect accordance with the image presented (
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<figureCitation id="789C2A676243FFE74A7FBD0FFC60FEA8" box="[835,902,331,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1788,1812]" captionTargetBox="[157,1432,1033,1755]" captionTargetId="figure-447@2.[151,1435,1026,1761]" captionTargetPageId="2" captionText="FIGURE 1. Schematic drawings of most of the measurements taken from the specimens (presented in the species descriptions or in Appendix 1). See Materials and Methods or Appendix 1 for a detailed explanation of the measurements. Codes: a, head length; b, head length 2; c, head width; d, head width 2; e, head width 3; f, head maximum depth; g, mesosoma length (see also Fig.6); h, mesosoma height; i, propodeal spiracle length; j, petiole anterior face length; k, posttergite 3 length; o, posttergite 4 length. Measurements a, g, j, k, and o summed equal total length (TL). Gaster measurements l, m, n, and p are additional metasomal measurements that are useful if the measurements k and o are blocked (see Appendix 1)." figureDoi="http://doi.org/10.5281/zenodo.8243391" httpUri="https://zenodo.org/record/8243391/files/figure.png" pageId="25" pageNumber="326">Fig. 1</figureCitation>
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, C of
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<bibRefCitation id="84364B136243FFE74AF0BD0FFB16FEA8" author="Barden, P. & Grimaldi, D. A." box="[972,1264,331,357]" pageId="25" pageNumber="326" pagination="405 - 412" refId="ref23029" refString="Barden, P. & Grimaldi, D. A. (2013) A new genus of highly specialized ants in Cretaceous Burmese amber (Hymenoptera: Formicidae). Zootaxa, 3681 (4), 405 - 412. https: // doi. org / 10.11646 / zootaxa. 3681.4.5" type="journal article" year="2013">Barden & Grimaldi, 2013</bibRefCitation>
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), so the tubular shape of the illustration is here considered a misinterpretation. Instead, the toruli appear to have a regular ring-like shape, as in most ants, being, at most, slightly raised. The mesosoma is well-preserved. In the metasoma, only the petiole is preserved, the gaster being entirely covered by an internal fracture of the amber; the shape of the petiole, especially the flat dorsal surface of the node, is better seen in photo of
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<figureCitation id="789C2A676243FFE74AA3BD9FFC3DFE38" box="[927,987,475,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1788,1812]" captionTargetBox="[157,1432,1033,1755]" captionTargetId="figure-447@2.[151,1435,1026,1761]" captionTargetPageId="2" captionText="FIGURE 1. Schematic drawings of most of the measurements taken from the specimens (presented in the species descriptions or in Appendix 1). See Materials and Methods or Appendix 1 for a detailed explanation of the measurements. Codes: a, head length; b, head length 2; c, head width; d, head width 2; e, head width 3; f, head maximum depth; g, mesosoma length (see also Fig.6); h, mesosoma height; i, propodeal spiracle length; j, petiole anterior face length; k, posttergite 3 length; o, posttergite 4 length. Measurements a, g, j, k, and o summed equal total length (TL). Gaster measurements l, m, n, and p are additional metasomal measurements that are useful if the measurements k and o are blocked (see Appendix 1)." figureDoi="http://doi.org/10.5281/zenodo.8243391" httpUri="https://zenodo.org/record/8243391/files/figure.png" pageId="25" pageNumber="326">Fig.1</figureCitation>
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, B than in drawing of
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<figureCitation id="789C2A676243FFE74DECBD9FFAEBFE38" box="[1232,1293,475,501]" captionStart="FIGURE 2" captionStartId="5.[151,250,1793,1817]" captionTargetBox="[151,1434,814,1770]" captionTargetId="figure-1048@5.[151,1435,814,1770]" captionTargetPageId="5" captionText="FIGURE 2. Masticatory margin of mandible in Gerontoformica and Zigrasimecia. A, Gerontoformica sp. (ANTWEB1032629); B, Z. ferox (ANTWEB1008126); C, Z. chuyangsui (ANTWEB1032660); D, Z. tonsora (ANTWEB11008098). Black triangles indicate incision between apical and basal teeth (or basal lamella). Image of Z. tonsora by Philip Barden and image of Z. ferox by Vincent Perrichot, courtesy of Antweb.org." figureDoi="http://doi.org/10.5281/zenodo.8243393" httpUri="https://zenodo.org/record/8243393/files/figure.png" pageId="25" pageNumber="326">Fig.2</figureCitation>
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, A of
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<bibRefCitation id="84364B136243FFE74C77BD9FFE88FDD4" author="Barden, P. & Grimaldi, D. A." pageId="25" pageNumber="326" pagination="405 - 412" refId="ref23029" refString="Barden, P. & Grimaldi, D. A. (2013) A new genus of highly specialized ants in Cretaceous Burmese amber (Hymenoptera: Formicidae). Zootaxa, 3681 (4), 405 - 412. https: // doi. org / 10.11646 / zootaxa. 3681.4.5" type="journal article" year="2013">Barden & Grimaldi (2013)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="E01836E26243FFE749FBBE67FD0AFDAC" blockId="25.[151,1436,151,609]" pageId="25" pageNumber="326">
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All characters of the diagnosis, except the ones compared to
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<taxonomicName id="27A74D616243FFE74A5ABE67FC08FDF0" box="[870,1006,547,573]" class="Insecta" family="Formicidae" genus="Zigrasimecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="goldingot">
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<emphasis id="D2D3EAF06243FFE74A5ABE67FC08FDF0" box="[870,1006,547,573]" italics="true" pageId="25" pageNumber="326">Z. goldingot</emphasis>
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</taxonomicName>
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(eye shape and propodeum shape), are expected to be shared between the queen and worker.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |