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<mods:title id="6537EC64E16BAC3D5A021EC7DA68B0B3">The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae)</mods:title>
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<mods:namePart id="D07E29D74530E590D9E17B4B642CAC9E">Ng, Peter K. L.</mods:namePart>
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<treatment id="1AD6EEE48235CBB3A5F50F3FBE885B93" ID-GBIF-Taxon="144148304" LSID="urn:lsid:plazi:treatment:1AD6EEE48235CBB3A5F50F3FBE885B93" httpUri="http://treatment.plazi.org/id/1AD6EEE48235CBB3A5F50F3FBE885B93" lastPageNumber="89" pageId="0" pageNumber="89">
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<taxonomicName id="718A730222A65E0711ECCB9DFF5A8DF7" ID-CoL="5699Z" authority="Ng &amp; Stuebing, 1990" authorityName="Ng &amp; Stuebing" authorityYear="1990" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thelphusula dicerophilus" order="Decapoda" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="dicerophilus">Thelphusula dicerophilus Ng &amp; Stuebing, 1990</taxonomicName>
Figure 6A, B
</paragraph>
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<paragraph id="9DDCEF476DBE98DA294F2E69D72089D5" pageId="0" pageNumber="89">
<taxonomicName id="FE1103C061863356A45FF719BAC3EDFF" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thelphusula dicerophilus" order="Decapoda" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="dicerophilus">Thelphusula dicerophilus</taxonomicName>
Ng &amp; Stuebing, 1990: 46, fig. 1, pl. 1;
<bibRefCitation id="DF416F73F5619D1A1AD3751FF4ADA108" author="Tan, SH" journalOrPublisher="Journal of Crustacean Biology" pageId="0" pageNumber="89" pagination="808 - 822" title="Revision of the genus Thelphusula Bott, 1969 (Decapoda: Brachyura: Gecarcinucidae) with description of a new genus and three new species." url="https://doi.org/10.2307/1549155" volume="18" year="1998">Tan and Ng 1998</bibRefCitation>
: 813, fig. 6C;
<bibRefCitation id="F5A94DAF7A5F3B277BB719D765CB3615" author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology Supplement" pageId="0" pageNumber="89" pagination="1 - 286" title="Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world." volume="17" year="2008">Ng et al. 2008</bibRefCitation>
: 73.
</paragraph>
</subSubSection>
<subSubSection id="5AD4D6487A16786278C2620C682F17E4" pageId="0" pageNumber="89" type="materials_examined">
<paragraph id="403F2B7F28370A396272B7F6B1A69586" pageId="0" pageNumber="89">Material examined.</paragraph>
<paragraph id="7022EA5AEBE718F8DC826560833C80BA" pageId="0" pageNumber="89">
Holotype: male (14.0
<normalizedToken id="120365BD295866A28E4E1C3D10D82FBD" originalValue="×">x</normalizedToken>
12.0 mm) (ZRC 1989.3588), in pitfall trap, adjacent to pool of rhinoceros mud wallow, Danum Valley, Lahad Datu, Sabah, ca.
<geoCoordinate id="3C7B6648A4684B077C03A9FE5F339203" direction="north" orientation="latitude" precision="925" value="4.9166665">4°55'N</geoCoordinate>
117°46', coll. R. Stuebing, 4 March l988. Paratypes: 1 female (18.3
<normalizedToken id="F9827C8F8D4D3C2FE3905639E267EBF6" originalValue="×">x</normalizedToken>
15.0 mm), 2 juveniles (ZRC 1989.3592-3594), in pitfall trap, adjacent to pool of rhinoceros mud wallow, Danum Valley, Lahad Datu, Sabah, ca.
<geoCoordinate id="B352FDBAE9C47E956E1D4ADE51FD6BDF" direction="north" orientation="latitude" precision="925" value="4.9166665">4°55'N</geoCoordinate>
117°46', coll. R. Stuebing, 1 March 1988; 1 male (18.5
<normalizedToken id="33241D988FE0384A0AC2477480803F66" originalValue="×">x</normalizedToken>
15.4 mm) (ZRC 1989.3591), in mist net in rhinoceros mud wallow pool, Danum Valley, Lahad Datu, Sabah, ca.
<geoCoordinate id="0B9F4485ABB8AFF5FAC20270C677023B" direction="north" orientation="latitude" precision="925" value="4.9166665">4°55'N</geoCoordinate>
117°46', coll. R. Stuebing, 2 March l988; 1 male, 1 female (ZRC 1989.3589-3590), in rhinoceros mud wallow pool, Danum Valley, Lahad Datu, Sabah, ca.
<geoCoordinate id="0534BA2F2FCCD7EE89A12588E4E6E51C" direction="north" orientation="latitude" precision="925" value="4.9166665">4°55'N</geoCoordinate>
117°46', coll. R. Stuebing, 2 March l988. Others: 1 female (11.0
<normalizedToken id="56A4585A6A20D9BAD36BF3CE52E28228" originalValue="×">x</normalizedToken>
9.5 mm) (ZRC 1997.0138), in pitfall trap, Danum Valley Field Centre, Lahad Datu, Sabah, coll. C.
<normalizedToken id="31225BE8CB378D3122EF0D633DEA1713" originalValue="Colón">Colon</normalizedToken>
, 14 October 1996; 1 male (16.4
<normalizedToken id="180F1E3206F765D5F2330263AA688234" originalValue="×">x</normalizedToken>
13.5 mm) (ZRC 1997.0140), in pitfall trap, Danum Valley Field Centre, Lahad Datu, Sabah, coll. C.
<normalizedToken id="C50FE6B5AEC32ECE805F99018E6164B8" originalValue="Colón">Colon</normalizedToken>
, 10 October 1996; 1 male (20.0
<normalizedToken id="8BD426B87D4970EE0ABC521813840A59" originalValue="×">x</normalizedToken>
15.8 mm) (ZRC 1997.0141), in pitfall trap, Danum Valley Field Centre, Lahad Datu, Sabah, coll. C.
<normalizedToken id="B363BED845F83C53E4653F757A33CDDF" originalValue="Colón">Colon</normalizedToken>
, 17 October 1996; 1 male (22.0
<normalizedToken id="E880FC56EBD7E20E27A3466D0C6AEA2E" originalValue="×">x</normalizedToken>
17.6 mm) (ZRC 1997.0142), in pitfall trap, Danum Valley Field Centre, Lahad Datu, Sabah, coll. C.
<normalizedToken id="5A36C72700C8A6D2B1EA67ACE18EE0CE" originalValue="Colón">Colon</normalizedToken>
, 19 October 1996; 1 male (10.5
<normalizedToken id="5187B9D2EE20B7A15B7EE5517427DF02" originalValue="×">x</normalizedToken>
9.0 mm), 1 female (17.8
<normalizedToken id="55451C02C7B6E97F9BB660484A6DCA5B" originalValue="×">x</normalizedToken>
14.9 mm) (ZRC 2017.1272), in mud and leaves under wooden walk-way, Orchid Trail, Danum Valley Field Centre, Lahad Datu, Sabah, at night, coll. local rangers, 20 July 2017; 1 female (26.1
<normalizedToken id="2B02F8E92A430D2A0AE645EE32CDDDD0" originalValue="×">x</normalizedToken>
21.3 mm) (ZRC 2017.1047), in pool along wooden walkway at night, Orchid Trail, Danum Valley Field Centre, Lahad Datu, Sabah, at night, coll. local rangers, 20 July 2017; 1 female (24.0
<normalizedToken id="4634CA62C636DFD972EF26F4D53E0060" originalValue="×">x</normalizedToken>
17.5 mm) (ZRC 1997.0139), Kunak, Baturong, Binuang River, Lahad Datu, Sabah, coll. R. Stuebing, 20 March 1989.
</paragraph>
</subSubSection>
<subSubSection id="B48A8E2C77E50AD6D579124D8C52B67F" pageId="0" pageNumber="89" type="colour">
<paragraph id="F168F8BD953AE21D261B1606950634E9" pageId="0" pageNumber="89">Colour.</paragraph>
<paragraph id="5ED79542E7DA2F9A71E2400060678078" pageId="0" pageNumber="89">In life, the carapace is reddish brown with the ambulatory legs lighter in colour; the chelipeds are orangish red with the fingers pale-yellow (Fig. 6A, B).</paragraph>
</subSubSection>
<subSubSection id="1FF2EC44294A684FAB5B31367ADD87F4" pageId="0" pageNumber="89" type="remarks">
<paragraph id="F2E6435C8F91F249BAA96154A3402AF5" pageId="0" pageNumber="89">Remarks.</paragraph>
<paragraph id="135B8A8A340DCE489EDD514E2DC1FAEC" pageId="0" pageNumber="89">
The present series of specimens do not change the original description of this species in any way. The species does grow substantially larger than the type series, with the largest specimen here, a female measuring 26.1
<normalizedToken id="767171D977E1E1F65F70D0849A8E3E30" originalValue="×">x</normalizedToken>
21.3 mm (ZRC 2017.1047).
</paragraph>
<paragraph id="C7057A4522B63A6DD9F1C32D37C091B2" pageId="0" pageNumber="89">
The available collection data indicates
<taxonomicName id="654E9B50F2B996EA1871D031AD79E699" lsidName="T. dicerophilus" pageId="0" pageNumber="89" rank="species" species="dicerophilus">T. dicerophilus</taxonomicName>
is a semiterrestrial nocturnal species and forages on the forest floor, usually in wet, swampy areas, digging burrows in the soft substrate; they were often caught in pitfall traps set near these areas (see also
<bibRefCitation id="F1382C822B238E118E5F95E1BEFC522F" author="Ng, PKL" journalOrPublisher="Indo-Malayan Zoology" pageId="0" pageNumber="89" pagination="45 - 51" title="Thelphusuladicerophilus sp. nov., a new species of freshwater crab (Crustacea: Decapoda: Brachyura: Gecarcinucidae) found in mud wallows of the Sumatran rhinoceros from Sabah, Borneo." volume="6" year="1990">Ng and Stuebing 1990</bibRefCitation>
). This contrasts with the more aquatic habits of its congener in Danum Valley,
<taxonomicName id="977D99AB08A9BFE6F60AB4014CAA5879" lsidName="T. capillodigitus" pageId="0" pageNumber="89" rank="species" species="capillodigitus">T. capillodigitus</taxonomicName>
sp. n.
</paragraph>
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