195 lines
22 KiB
XML
195 lines
22 KiB
XML
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<mods:title id="5366EE8EA3FDF64B9509DF6956838D8F">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<mods:namePart id="6A6F09F691846D3CBA453EB117DA590B">Harzhauser, Mathias</mods:namePart>
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<mods:nameIdentifier id="201BB51D7F48008E8A54B88E6A9EBC0B" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:namePart id="E2B7C0802215C79155EDE4CA890DE3C3">Landau, Bernard</mods:namePart>
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<mods:nameIdentifier id="D7092BA7B0F4EFB4FF1BE42893432958" type="email">bernardmlandau@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0B2BA029758C26193617D61AC766BAA0">Janssen, Ronald</mods:namePart>
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<mods:nameIdentifier id="885A2951C14DC641ADEB90B6C70C25E7" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:title id="4CEF59DFE38B0EC7A8C55F3D54A711DB">Zootaxa</mods:title>
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<mods:part id="DE5E372750C4735B5F40A70FF684CFD6">
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<mods:date id="127FBA0D5DA6360888AFA11AA0E86D37">2022</mods:date>
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<mods:number id="55FC210ACD91687342B5809D858A13AC">2022-03-30</mods:number>
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<subSubSection id="C327654CFFB2FF98FFBAFA0D6BE2FA00" box="[151,912,1446,1473]" pageId="48" pageNumber="49" type="nomenclature">
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<paragraph id="8B8236C7FFB2FF98FFBAFA0D6BE2FA00" blockId="48.[151,912,1446,1508]" box="[151,912,1446,1473]" pageId="48" pageNumber="49">
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<heading id="D0CA81ABFFB2FF98FFBAFA0D6BE2FA00" bold="true" box="[151,912,1446,1473]" fontSize="11" level="2" pageId="48" pageNumber="49" reason="4">
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<taxonomicName id="4C3D4D44FFB2FF98FFBAFA0D6AD3FA00" authority="Jekelius, 1944" authorityName="Jekelius" authorityYear="1944" box="[151,673,1446,1473]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="carasiensis">
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<emphasis id="B949EAD5FFB2FF98FFBAFA0D6999FA00" bold="true" box="[151,491,1446,1473]" italics="true" pageId="48" pageNumber="49">Granulatocincta carasiensis</emphasis>
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<bibRefCitation id="EFAC4B36FFB2FF98FEDFFA0D6AD3FA00" author="Jekelius, E." box="[498,673,1446,1472]" pageId="48" pageNumber="49" pagination="1 - 167" refId="ref135955" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului Geolologic al Romaniei, 5, 1 - 167, 65 pls." type="journal article" year="1944">Jekelius, 1944</bibRefCitation>
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</taxonomicName>
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||
species inquirenda
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||
</emphasis>
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||
</heading>
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</paragraph>
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<subSubSection id="C327654CFFB2FF98FFBAFA61688AFA24" box="[151,248,1482,1508]" pageId="48" pageNumber="49" type="description">
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<paragraph id="8B8236C7FFB2FF98FFBAFA61688AFA24" blockId="48.[151,912,1446,1508]" box="[151,248,1482,1508]" pageId="48" pageNumber="49">
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<figureCitation id="13062A42FFB2FF98FFBAFA61688AFA24" box="[151,248,1482,1508]" captionStart="FIGURE 17" captionStartId="52.[151,250,1723,1747]" captionTargetBox="[248,1339,181,1698]" captionTargetId="figure-26@52.[248,1339,181,1698]" captionTargetPageId="52" captionText="FIGURE 17 A1–A3. Granulatocincta callim nov. sp., holotype, NHMW 1866/0001/0549, Wiesen (Austria). B1–B3. Granulatocincta callim nov.sp.,paralectotype,NHMW 1846/0037/0351b, Wiesen (Austria).C1–C3. Granulatocincta callim nov. sp., paralectotype, NHMW 1846/0037/0351, Wiesen (Austria). D1–D2. Granulatocincta callim nov. sp., paralectotype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). E. Granulatocincta carasiensis (Jekelius, 1944), MNG P– 4965, Soceni (Romania). F1–F2. Granulatocincta contorta (Švagrovský, 1958), holotype, Z 3076, Kuzmice (Slovakia). G1–G2. Granulatocincta contorta (Švagrovský, 1958), paratype, SNMB Z 3074, Kuzmice (Slovakia). H1–H3. Granulatocincta pelliscrocodili nov. sp., holotype, RGM 776 907, Gödet Creek near Tilkikaya (Turkey). I1–I2. Granulatocincta pelliscrocodili nov. sp., paratype, RGM 776 906, southwest bank of Gödet Creek near Tilkikaya (Turkey). J1–J3. Granulatocincta nataliae (Hoernes & Auinger, 1891), holotype, NHMW 1855/0035/0021, Belgrade/Rakovica (Serbia)." figureDoi="http://doi.org/10.5281/zenodo.6399871" httpUri="https://zenodo.org/record/6399871/files/figure.png" pageId="48" pageNumber="49">Fig. 17E</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C327654CFFB2FF98FFBAF9A46B21F9E6" box="[151,851,1550,1575]" pageId="48" pageNumber="49" type="reference_group">
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<paragraph id="8B8236C7FFB2FF98FFBAF9A46B21F9E6" blockId="48.[151,851,1550,1575]" box="[151,851,1550,1575]" pageId="48" pageNumber="49">
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<treatmentCitationGroup id="AB2D11E9FFB2FF98FFBAF9A46B21F9E6" box="[151,851,1550,1575]" pageId="48" pageNumber="49">
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<emphasis id="B949EAD5FFB2FF98FFBAF9A469F4F9E6" box="[151,390,1551,1575]" italics="true" pageId="48" pageNumber="49">
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*
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<taxonomicName id="4C3D4D44FFB2FF98FF87F9A469F4F9E6" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[170,390,1551,1575]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="carasiensis" status="sp. nov.">Clavatula carasiensis</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A27A57AEFFB2FF98FEA0F9A469B2F9E7" box="[397,448,1551,1575]" pageId="48" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
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—
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<treatmentCitation id="0A9C10D6FFB2FF98FEF7F9A46AFDF9E7" author="Jekelius, E." box="[474,655,1551,1575]" page="87" pageId="48" pageNumber="49" year="1944">
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<bibRefCitation id="EFAC4B36FFB2FF98FEF7F9A46AFDF9E7" author="Jekelius, E." box="[474,655,1551,1575]" pageId="48" pageNumber="49" pagination="1 - 167" refId="ref135955" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului Geolologic al Romaniei, 5, 1 - 167, 65 pls." type="journal article" year="1944">Jekelius 1944: 87</bibRefCitation>
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</treatmentCitation>
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, pl. 23, figs 13–15.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C327654CFFB2FF98FFBAF9FE69EDF873" pageId="48" pageNumber="49" type="materials_examined">
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<paragraph id="8B8236C7FFB2FF98FFBAF9FE6C02F91C" blockId="48.[151,1437,1621,2044]" pageId="48" pageNumber="49">
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<emphasis id="B949EAD5FFB2FF98FFBAF9FE693AF9AF" bold="true" box="[151,328,1621,1647]" pageId="48" pageNumber="49">Type material.</emphasis>
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<typeStatus id="54868865FFB2FF98FE7CF9FE69CCF9AF" box="[337,446,1621,1647]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="ED2CAE02FFB2FF98FEE6F9FE6A7EF9AF" box="[459,524,1621,1647]" country="0" httpUri="http://grbio.org/cool/9drb-g62q" name="Sammlung Eisfeld des Museums der Natur Gotha" pageId="48" pageNumber="49">MNG</collectionCode>
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P– 4965, SL:
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<quantity id="4CC59B22FFB2FF98FD94F9FE6B66F9AF" box="[697,788,1621,1648]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="48" pageNumber="49" unit="mm" value="3.5">3.5 mm</quantity>
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,
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<collectionCode id="ED2CAE02FFB2FF98FC0FF9FE6B21F9AF" box="[802,851,1621,1647]" country="Germany" name="Museum Donaueschingen" pageId="48" pageNumber="49">MD</collectionCode>
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:
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<quantity id="4CC59B22FFB2FF98FC4CF9FE6BCEF9AF" box="[865,956,1621,1647]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="48" pageNumber="49" unit="mm" value="1.6">1.6 mm</quantity>
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, Soceni (
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<collectingCountry id="F32A7657FFB2FF98FB05F9FE6CE1F9B0" box="[1064,1171,1621,1648]" name="Romania" pageId="48" pageNumber="49">Romania</collectingCountry>
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), illustrated in
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<bibRefCitation id="EFAC4B36FFB2FF98FA6FF9FE68A9F953" author="Jekelius, E." pageId="48" pageNumber="49" pagination="1 - 167" refId="ref135955" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului Geolologic al Romaniei, 5, 1 - 167, 65 pls." type="journal article" year="1944">Jekelius (1944</bibRefCitation>
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, pl. 23, fig. 15), stored in the National Museum of Geology of
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<collectingCountry id="F32A7657FFB2FF98FC94F9D26C6CF953" box="[953,1054,1657,1683]" name="Romania" pageId="48" pageNumber="49">Romania</collectingCountry>
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(
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<collectingRegion id="49F9F825FFB2FF98FB02F9D26CD7F954" box="[1071,1189,1657,1684]" country="Romania" name="Bucuresti" pageId="48" pageNumber="49">Bucharest</collectingRegion>
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), fig. 16E.
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<typeStatus id="54868865FFB2FF98FA05F9D26DEBF953" box="[1320,1433,1657,1683]" pageId="48" pageNumber="49" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="ED2CAE02FFB2FF98FFBAF93668AAF977" box="[151,216,1693,1719]" country="0" httpUri="http://grbio.org/cool/9drb-g62q" name="Sammlung Eisfeld des Museums der Natur Gotha" pageId="48" pageNumber="49">MNG</collectionCode>
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P– 4965, Soceni (
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<collectingCountry id="F32A7657FFB2FF98FE87F9366A67F978" box="[426,533,1693,1720]" name="Romania" pageId="48" pageNumber="49">Romania</collectingCountry>
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),
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<typeStatus id="54868865FFB2FF98FD07F9366AFEF977" box="[554,652,1693,1719]" pageId="48" pageNumber="49" type="syntype">syntypes</typeStatus>
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of
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<taxonomicName id="4C3D4D44FFB2FF98FD99F9366C3DF977" authority="Jekelius 1944" authorityName="Jekelius" authorityYear="1944" box="[692,1103,1693,1719]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="carasiensis">
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<emphasis id="B949EAD5FFB2FF98FD99F9366BDBF977" box="[692,937,1693,1719]" italics="true" pageId="48" pageNumber="49">Clavatula carasiensis</emphasis>
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<bibRefCitation id="EFAC4B36FFB2FF98FC9CF9366C3DF977" author="Jekelius, E." box="[945,1103,1693,1719]" pageId="48" pageNumber="49" pagination="1 - 167" refId="ref135955" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului Geolologic al Romaniei, 5, 1 - 167, 65 pls." type="journal article" year="1944">Jekelius 1944</bibRefCitation>
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||
</taxonomicName>
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, illustrated in
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||
<bibRefCitation id="EFAC4B36FFB2FF98FBDCF9366DEAF977" author="Jekelius, E." box="[1265,1432,1693,1719]" pageId="48" pageNumber="49" pagination="1 - 167" refId="ref135955" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului Geolologic al Romaniei, 5, 1 - 167, 65 pls." type="journal article" year="1944">Jekelius (1944</bibRefCitation>
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, pl. 23, figs 14–15), stored in the National Museum of Geology of
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||
<collectingCountry id="F32A7657FFB2FF98FC54F96A6BACF91B" box="[889,990,1729,1755]" name="Romania" pageId="48" pageNumber="49">Romania</collectingCountry>
|
||
(
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<collectingRegion id="49F9F825FFB2FF98FCC0F96A6C16F91C" box="[1005,1124,1729,1756]" country="Romania" name="Bucuresti" pageId="48" pageNumber="49">Bucharest</collectingRegion>
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).
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</paragraph>
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<paragraph id="8B8236C7FFB2FF98FFEAF94E69EDF873" blockId="48.[151,1437,1621,2044]" pageId="48" pageNumber="49">
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<emphasis id="B949EAD5FFB2FF98FFEAF94E69C9F93F" bold="true" box="[199,443,1765,1791]" pageId="48" pageNumber="49">Revised description.</emphasis>
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Only tiny, juvenile shells comprising three teleoconch whorls available. Small slender fusiform with high spire; apical angle ~30°. Protoconch high, turreted, paucispiral with strongly convex initial part and weakly convex last whorl. First teleoconch whorl weakly convex with indistinct subsutural spiral cord. Commashaped riblets crossed by weak spiral cords on later teleoconch whorls resulting in beaded appearance. Whorl profile slightly angulated at mid-whorl. Last preserved whorl with slightly protruding weakly beaded shoulder. Base low, moderately constricted.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C327654CFFB2FF99FFEAF8166BD5FEDC" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="discussion">
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<paragraph id="8B8236C7FFB2FF99FFEAF8166B74FF39" blockId="48.[151,1437,1621,2044]" lastBlockId="49.[151,1437,151,357]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
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<emphasis id="B949EAD5FFB2FF98FFEAF816693EF817" bold="true" box="[199,332,1981,2007]" pageId="48" pageNumber="49">Discussion.</emphasis>
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The status of this species is unclear as only juvenile specimens are known. Its early whorl sculpture differs from other Sarmatian species such as
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<taxonomicName id="4C3D4D44FFB2FF98FDA3F84A6C75F83C" authority="(Hornes, 1854)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hornes" baseAuthorityYear="1854" box="[654,1031,2017,2044]" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="doderleini">
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||
<emphasis id="B949EAD5FFB2FF98FDA3F84A6B26F83B" box="[654,852,2017,2043]" italics="true" pageId="48" pageNumber="49">Olegia doderleini</emphasis>
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||
(
|
||
<bibRefCitation id="EFAC4B36FFB2FF98FC4EF84A6B8DF83C" author="Hornes, M." box="[867,1023,2017,2044]" pageId="48" pageNumber="49" pagination="297 - 382" refId="ref135553" refString="Hornes, M. (1854) Die fossilen Mollusken des Tertiar-Beckens von Wien. Abhandlungen der K. K. Geologischen Reichsanstalt, 3, 297 - 382, pls. XXXIII - XL (1854). [https: // opac. geologie. ac. at / ais 312 / dokumente / Abh- 3 - GRA-gesamt. pdf]" type="journal article" year="1854">Hörnes, 1854</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FFB2FF98FB3EF84A6CFDF83B" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[1043,1167,2017,2043]" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="mandici" status="sp. nov.">
|
||
<emphasis id="B949EAD5FFB2FF98FB3EF84A6CFDF83B" box="[1043,1167,2017,2043]" italics="true" pageId="48" pageNumber="49">O. mandici</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFFB2FF98FBBBF84A6C9FF83B" box="[1174,1261,2017,2043]" pageId="48" pageNumber="49" rank="species">nov. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C3D4D44FFB2FF99FA0EF84A699AFF71" authority="(Simionescu & Barbu, 1940)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Simionescu & Barbu" baseAuthorityYear="1940" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" lastPageId="49" lastPageNumber="50" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rumana">
|
||
<emphasis id="B949EAD5FFB2FF98FA0EF84A6DEEF83B" box="[1315,1436,2017,2043]" italics="true" pageId="48" pageNumber="49">O. rumana</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FFB3FF99FFB2FF3C6992FF71" author="Simionescu, I. & Barbu, I. Z." box="[159,480,151,177]" pageId="49" pageNumber="50" pagination="1 - 194" refId="ref144291" refString="Simionescu, I. & Barbu, I. Z. (1940) La faune sarmatienne de Roumanie. Memoriile Institutului Geologic al Romaniei, 3, 1 - 194, 11 pls." type="journal article" year="1940">Simionescu & Barbu, 1940</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The beads on the axial riblets on early teleoconch whorls are reminiscent of the Sarmatian
|
||
<taxonomicName id="4C3D4D44FFB3FF99FE22FF106A67FF15" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[271,533,187,213]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="callim" status="sp. nov.">
|
||
<emphasis id="B949EAD5FFB3FF99FE22FF106A67FF15" box="[271,533,187,213]" italics="true" pageId="49" pageNumber="50">Granulatocincta callim</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFFB3FF99FD36FF106A01FF15" box="[539,627,187,213]" pageId="49" pageNumber="50" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, but the latter develops well-defined beads at the abapical suture at same growth stage and a central spiral cord of delicate beads.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFB3FF99FFEAFEA96BD5FEDC" blockId="49.[151,1437,151,357]" box="[199,935,258,285]" pageId="49" pageNumber="50">
|
||
<emphasis id="B949EAD5FFB3FF99FFEAFEA969DAFEDC" bold="true" box="[199,424,258,284]" pageId="49" pageNumber="50">Paleoenvironment.</emphasis>
|
||
Coastal inner neritic (
|
||
<bibRefCitation id="EFAC4B36FFB3FF99FD8DFEA86BEAFEDD" author="Lukeneder, S. & Zuschin, M. & Harzhauser, M. & Mandic, O." box="[672,920,259,285]" pageId="49" pageNumber="50" pagination="767 - 784" refId="ref138028" refString="Lukeneder, S., Zuschin, M., Harzhauser, M. & Mandic, O. (2011) Spatiotemporal signals and palaeoenvironments of endemic molluscan assemblages in the marine system of the Sarmatian Paratethys. Acta Palaeontologica Polonica, 56, 767 - 784. [https: // www. app. pan. pl / archive / published / app 56 / app 20100046. pdf] https: // doi. org / 10.4202 / app. 2010.0046" type="journal article" year="2011">
|
||
Lukeneder
|
||
<emphasis id="B949EAD5FFB3FF99FC0DFEA86B2BFEDD" box="[800,857,259,285]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFB3FF99FFEAFE8D6938FEA5" pageId="49" pageNumber="50" type="distribution">
|
||
<paragraph id="8B8236C7FFB3FF99FFEAFE8D6938FEA5" blockId="49.[151,1437,151,357]" pageId="49" pageNumber="50">
|
||
<emphasis id="B949EAD5FFB3FF99FFEAFE8D6A0EFE81" bold="true" box="[199,636,294,321]" pageId="49" pageNumber="50">Distribution in Central Paratethys.</emphasis>
|
||
Sarmatian (middle Miocene):
|
||
<emphasis id="B949EAD5FFB3FF99FCC0FE8C6CCFFE80" box="[1005,1213,295,320]" italics="true" pageId="49" pageNumber="50">Pannonian Basin:</emphasis>
|
||
Soceni (
|
||
<collectingCountry id="F32A7657FFB3FF99FA01FE8C6DE7FE82" box="[1324,1429,295,322]" name="Romania" pageId="49" pageNumber="50">Romania</collectingCountry>
|
||
) (
|
||
<bibRefCitation id="EFAC4B36FFB3FF99FFB2FEE0694CFEA5" author="Jekelius, E." box="[159,318,331,357]" pageId="49" pageNumber="50" pagination="1 - 167" refId="ref135955" refString="Jekelius, E. (1944) Sarmat und Pont von Soceni (Banat). Memoriile Institutului Geolologic al Romaniei, 5, 1 - 167, 65 pls." type="journal article" year="1944">Jekelius 1944</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |