440 lines
57 KiB
XML
440 lines
57 KiB
XML
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<mods:title id="493509F343F416EBA76530AE7B7FC9AA">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<mods:namePart id="D12A49D4FA270A3D2E734C7478EDFBE4">Harzhauser, Mathias</mods:namePart>
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<mods:namePart id="41460FBB60E74E66D47D2582F40A9EAA">Landau, Bernard</mods:namePart>
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<mods:namePart id="777F21C44C3BAF1F0D2608FA16770352">Janssen, Ronald</mods:namePart>
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<mods:nameIdentifier id="C69F4D4093DF419D44E64CE47C72C817" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:title id="C4F6A3E455C82AA83378A06B8ECB24C6">Zootaxa</mods:title>
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<mods:part id="B19704BEB530E2414A2D780A1A1BEFE8">
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<mods:date id="9276B57470583994872657DA9284FC98">2022</mods:date>
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<mods:number id="A5F983CFF74A055EB306D86197C45630">2022-03-30</mods:number>
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<treatment id="039487D1FF0CFF27FFBAFF3C6BD3FF15" LSID="urn:lsid:plazi:treatment:039487D1FF0CFF27FFBAFF3C6BD3FF15" httpUri="http://treatment.plazi.org/id/039487D1FF0CFF27FFBAFF3C6BD3FF15" lastPageId="143" lastPageNumber="144" pageId="142" pageNumber="143">
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<subSubSection id="C327654CFF0CFF26FFBAFF3C6A77FF71" box="[151,517,151,178]" pageId="142" pageNumber="143" type="reference_group">
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<paragraph id="8B8236C7FF0CFF26FFBAFF3C6A77FF71" blockId="142.[151,517,151,178]" box="[151,517,151,178]" pageId="142" pageNumber="143">
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<emphasis id="B949EAD5FF0CFF26FFBAFF3C6A77FF71" bold="true" box="[151,517,151,178]" pageId="142" pageNumber="143">
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Genus
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<taxonomicName id="4C3D4D44FF0CFF26FFC3FF3C6A77FF71" ID-CoL="7PJLT" authority="Wenz, 1943" authorityName="Wenz" authorityYear="1943" box="[238,517,151,178]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
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<emphasis id="B949EAD5FF0CFF26FFC3FF3C691CFF72" bold="true" box="[238,366,151,178]" italics="true" pageId="142" pageNumber="143">Tomellana</emphasis>
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<bibRefCitation id="EFAC4B36FF0CFF26FE58FF3C6A77FF71" author="Wenz, W." box="[373,517,151,177]" pageId="142" pageNumber="143" pagination="1201 - 1506" refId="ref146470" refString="Wenz, W. (1943) Gastropoda. Teil 1: Allgemeiner Teil und Prosobranchia. xii + 1639 pp. In: Schindewolf, O. H. (Ed.) Handbuch der Palaozoologie. Band 6. Lfg. 8. Borntrager, Berlin, pp. 1201 - 1506." type="book chapter" year="1943">Wenz, 1943</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</paragraph>
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<subSubSection id="C327654CFF0CFF26FFBAFF746D7BFEA5" pageId="142" pageNumber="143" type="type_taxon">
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<paragraph id="8B8236C7FF0CFF26FFBAFF746D22FF39" blockId="142.[151,1437,223,2014]" box="[151,1360,223,250]" pageId="142" pageNumber="143">
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<materialsCitation id="3B553C9AFF0CFF26FFBAFF746D3EFF39" box="[151,1356,223,250]" location="West Africa" pageId="142" pageNumber="143" specimenCount="1" typeStatus="holotype">
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<emphasis id="B949EAD5FF0CFF26FFBAFF746940FF39" bold="true" box="[151,306,223,249]" pageId="142" pageNumber="143">
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<typeStatus id="54868865FF0CFF26FFBAFF7468A0FF39" box="[151,210,223,249]" pageId="142" pageNumber="143">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C3D4D44FF0CFF26FE14FF746AC2FF39" ID-CoL="VV47" authority="Lamarck, 1816" authorityName="Lamarck" authorityYear="1816" box="[313,688,223,249]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="lineata">
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<emphasis id="B949EAD5FF0CFF26FE14FF74698FFF39" box="[313,509,223,249]" italics="true" pageId="142" pageNumber="143">Clavatula lineata</emphasis>
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<bibRefCitation id="EFAC4B36FF0CFF26FD2EFF746AC2FF39" author="Lamarck, J. B. P. A. de" box="[515,688,223,249]" pageId="142" pageNumber="143" pagination="1 - 16" refId="ref137407" refString="Lamarck, J. B. P. A. de (1816) Liste des objets representes dans les planches de cette livraison. In: Lamarck, J. B. P. A. de (Ed.), Tableau Encyclopedique et Methodique des Trois Regnes de la Nature. Mollusques et Polypes Divers. Veuve Agasse, Imprimeur, Paris, pp. 1 - 16, pls. 391 - 488." type="book chapter" year="1816">Lamarck, 1816</bibRefCitation>
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</taxonomicName>
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, by typification of replaced name; present-day,
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<location id="8EE2601CFF0CFF26FBE9FF4B6D3EFF39" LSID="urn:lsid:plazi:treatment:039487D1FF0CFF27FFBAFF3C6BD3FF15:8EE2601CFF0CFF26FBE9FF4B6D3EFF39" box="[1220,1356,223,250]" name="West Africa" pageId="142" pageNumber="143">West Africa</location>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B8236C7FF0CFF26FFEAFEA86D7BFEA5" blockId="142.[151,1437,223,2014]" pageId="142" pageNumber="143">
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<emphasis id="B949EAD5FF0CFF26FFEAFEA86974FEDD" bold="true" box="[199,262,259,285]" pageId="142" pageNumber="143">Note.</emphasis>
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This genus was established as
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<taxonomicName id="4C3D4D44FF0CFF26FD7AFEA86AC2FEDD" ID-CoL="10101725-1d56-411e-accc-0a99384eccb2" box="[599,688,259,285]" class="Gastropoda" family="Clavatulidae" genus="Tomella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
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<emphasis id="B949EAD5FF0CFF26FD7AFEA86AC2FEDD" box="[599,688,259,285]" italics="true" pageId="142" pageNumber="143">Tomella</emphasis>
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</taxonomicName>
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by
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<collectingRegion id="49F9F825FF0CFF26FDFAFEA86B35FEDD" box="[727,839,259,285]" country="American Samoa" name="Swains Island" pageId="142" pageNumber="143">Swainson</collectingRegion>
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, (1840: 314) (
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<typeStatus id="54868865FF0CFF26FCF3FEA86C62FEDD" box="[990,1040,259,285]" pageId="142" pageNumber="143">type</typeStatus>
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species
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||
<taxonomicName id="4C3D4D44FF0CFF26FB46FEA868BDFE81" ID-CoL="VV47" authority="Lamarck, 1816" authorityName="Lamarck" authorityYear="1816" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="lineata">
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<emphasis id="B949EAD5FF0CFF26FB46FEA86D5FFEDD" box="[1131,1325,259,285]" italics="true" pageId="142" pageNumber="143">Clavatula lineata</emphasis>
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<bibRefCitation id="EFAC4B36FF0CFF26FA1FFEA868BDFE81" author="Lamarck, J. B. P. A. de" pageId="142" pageNumber="143" pagination="1 - 16" refId="ref137407" refString="Lamarck, J. B. P. A. de (1816) Liste des objets representes dans les planches de cette livraison. In: Lamarck, J. B. P. A. de (Ed.), Tableau Encyclopedique et Methodique des Trois Regnes de la Nature. Mollusques et Polypes Divers. Veuve Agasse, Imprimeur, Paris, pp. 1 - 16, pls. 391 - 488." type="book chapter" year="1816">Lamarck, 1816</bibRefCitation>
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</taxonomicName>
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by subsequent designation by Herrmannsen (1849: 579), which was a junior homonym of
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<taxonomicName id="4C3D4D44FF0CFF26FBEAFE8C6D52FE81" ID-CoL="10101725-1d56-411e-accc-0a99384eccb2" box="[1223,1312,295,321]" class="Gastropoda" family="Clavatulidae" genus="Tomella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
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<emphasis id="B949EAD5FF0CFF26FBEAFE8C6D52FE81" box="[1223,1312,295,321]" italics="true" pageId="142" pageNumber="143">Tomella</emphasis>
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</taxonomicName>
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Robineau- Desvoidy, 1830 [
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<taxonomicName id="4C3D4D44FF0CFF26FE79FEE169C3FEA5" box="[340,433,330,357]" class="Gastropoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="142" pageNumber="143" phylum="Mollusca" rank="order">Diptera</taxonomicName>
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]. Therefore,
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<bibRefCitation id="EFAC4B36FF0CFF26FD13FEE76B67FEA5" author="Wenz, W." box="[574,789,331,358]" pageId="142" pageNumber="143" pagination="1201 - 1506" refId="ref146470" refString="Wenz, W. (1943) Gastropoda. Teil 1: Allgemeiner Teil und Prosobranchia. xii + 1639 pp. In: Schindewolf, O. H. (Ed.) Handbuch der Palaozoologie. Band 6. Lfg. 8. Borntrager, Berlin, pp. 1201 - 1506." type="book chapter" year="1943">Wenz (1943: 1383)</bibRefCitation>
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introduced
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<taxonomicName id="4C3D4D44FF0CFF26FCB6FEE06C62FEA5" ID-CoL="7PJLT" authorityName="Wenz" authorityYear="1943" box="[923,1040,331,357]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
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<emphasis id="B949EAD5FF0CFF26FCB6FEE06C62FEA5" box="[923,1040,331,357]" italics="true" pageId="142" pageNumber="143">Tomellana</emphasis>
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</taxonomicName>
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as replacement name.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C327654CFF0CFF26FFEAFEC46C09FCD5" pageId="142" pageNumber="143" type="diagnosis">
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||
<paragraph id="8B8236C7FF0CFF26FFEAFEC46C90FE36" blockId="142.[151,1437,223,2014]" pageId="142" pageNumber="143">
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||
<emphasis id="B949EAD5FF0CFF26FFEAFEC46930FE49" bold="true" box="[199,322,367,393]" pageId="142" pageNumber="143">Diagnosis.</emphasis>
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‘
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<emphasis id="B949EAD5FF0CFF26FE7FFEC46C7BFE36" italics="true" pageId="142" pageNumber="143">
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||
differs from
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<taxonomicName id="4C3D4D44FF0CFF26FEF6FEDB6A45FE49" box="[475,567,368,393]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">Perrona</taxonomicName>
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in its short, concavely outlined spire, massive peripheral fold, lack of a basal angulation, very heavy parietal callus pad, and deep slit-like anal sinus. Protoconch small, papillate of 1.5 smooth whorls, followed by a whorl of sigmoid brephic axials. … The anal fasciole does not show on the spire whorls, being immersed by each succeeding whorl. Parietal callus pad white, very massive.”
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</emphasis>
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(
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<bibRefCitation id="EFAC4B36FF0CFF26FB35FE706CA4FE36" author="Powell, A. W. B." box="[1048,1238,475,502]" pageId="142" pageNumber="143" pagination="1 - 184" refId="ref141714" refString="Powell, A. W. B. (1966) The molluscan families Speightiidae and Turridae: an evaluation of the valid taxa, both recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Museum, 5, 1 - 184. [https: // www. biodiversitylibrary. org / item / 278249 # page / 4 / mode / 1 up]" type="journal article" year="1966">Powell 1966: 56</bibRefCitation>
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||
).
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||
</paragraph>
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||
<paragraph id="8B8236C7FF0CFF26FFEAFE546ABFFDFE" blockId="142.[151,1437,223,2014]" pageId="142" pageNumber="143">
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||
<emphasis id="B949EAD5FF0CFF26FFEAFE5469E6FDD9" bold="true" box="[199,404,511,537]" pageId="142" pageNumber="143">Included species.</emphasis>
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Central Paratethys Sea:
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<taxonomicName id="4C3D4D44FF0CFF26FD9EFE546DEAFDD9" authority="Schaffer, 1912" authorityName="Schaffer" authorityYear="1912" box="[691,1432,511,538]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="subSpecies" species="semimarginata" subGenus="Perrona" subSpecies="praecursor">
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||
<emphasis id="B949EAD5FF0CFF26FD9EFE546B44FDD9" box="[691,822,511,537]" italics="true" pageId="142" pageNumber="143">Pleurotoma</emphasis>
|
||
(
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<emphasis id="B949EAD5FF0CFF26FC64FDAB6BD7FDD9" box="[841,933,512,537]" italics="true" pageId="142" pageNumber="143">Perrona</emphasis>
|
||
)
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||
<emphasis id="B949EAD5FF0CFF26FC95FDAB6C97FDD9" box="[952,1253,512,537]" italics="true" pageId="142" pageNumber="143">semimarginata praecursor</emphasis>
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Schaffer, 1912
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</taxonomicName>
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,
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<taxonomicName id="4C3D4D44FF0CFF26FFBAFD88693CFDFD" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[151,334,547,573]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="aueri" status="sp. nov.">
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<emphasis id="B949EAD5FF0CFF26FFBAFD88693CFDFD" box="[151,334,547,573]" italics="true" pageId="142" pageNumber="143">Tomellana aueri</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A27A57AEFF0CFF26FE78FD8F69DFFDFE" box="[341,429,548,574]" pageId="142" pageNumber="143" rank="species">nov. sp.</taxonomicNameLabel>
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,
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<taxonomicName id="4C3D4D44FF0CFF26FE97FD886A02FDFD" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[442,624,547,573]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="dulai" status="sp. nov.">
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<emphasis id="B949EAD5FF0CFF26FE97FD886A02FDFD" box="[442,624,547,573]" italics="true" pageId="142" pageNumber="143">Tomellana dulai</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A27A57AEFF0CFF26FD5BFD8F6ABFFDFE" box="[630,717,548,574]" pageId="142" pageNumber="143" rank="species">nov. sp.</taxonomicNameLabel>
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||
</paragraph>
|
||
<paragraph id="8B8236C7FF0CFF26FFEAFDEC6CF6FD45" blockId="142.[151,1437,223,2014]" pageId="142" pageNumber="143">
|
||
Proto-Mediterraean Sea:?
|
||
<taxonomicName id="4C3D4D44FF0CFF26FED8FDEC6CD5FDA1" authority="Sacco, 1890" authorityName="Sacco" authorityYear="1890" box="[501,1191,583,610]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="variety" species="semimarginata" subGenus="Perrona" variety="servata">
|
||
<emphasis id="B949EAD5FF0CFF26FED8FDEC6A11FDA1" box="[501,611,583,609]" italics="true" pageId="142" pageNumber="143">Clavatula</emphasis>
|
||
(
|
||
<emphasis id="B949EAD5FF0CFF26FD58FDE36AA3FDA1" box="[629,721,584,609]" italics="true" pageId="142" pageNumber="143">Perrona</emphasis>
|
||
)
|
||
<emphasis id="B949EAD5FF0CFF26FDCFFDE36BFBFDA1" box="[738,905,584,609]" italics="true" pageId="142" pageNumber="143">semimarginata</emphasis>
|
||
var.
|
||
<emphasis id="B949EAD5FF0CFF26FCECFDE36C61FDA1" box="[961,1043,584,609]" italics="true" pageId="142" pageNumber="143">servata</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FB37FDEC6CD5FDA1" author="Sacco, F." box="[1050,1191,583,609]" pageId="142" pageNumber="143" pagination="189 - 340" refId="ref143023" refString="Sacco, F. (1890) Catalogo paleontologico del bacino terziaro del Piemonte. Bollettino della Societa Geologica Italiana, 9, 189 - 340. [https: // www. biodiversitylibrary. org / item / 210076 # page / 207 / mode / 1 up]" type="journal article" year="1890">Sacco, 1890</bibRefCitation>
|
||
</taxonomicName>
|
||
, an additional species from the Pliocene of Estepona (
|
||
<collectingCountry id="F32A7657FF0CFF26FED4FDC06A4EFD46" box="[505,572,619,646]" name="Spain" pageId="142" pageNumber="143">Spain</collectingCountry>
|
||
) will be described by
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FC19FDC06CF2FD45" author="Landau, B. & Harzhauser, M." box="[820,1152,619,645]" pageId="142" pageNumber="143" refId="ref137533" refString="Landau, B. & Harzhauser, M. (2022) The Pliocene Gastropoda (Mollusca) of Estepona, southern Spain. Part 14: Clavatulidae (Gastropoda, Conoidea). Cainozoic Research. [in press]" type="book" year="2022">Landau & Harzhauser (2022)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF0CFF26FFEAFD3B6D4FFD31" blockId="142.[151,1437,223,2014]" pageId="142" pageNumber="143">
|
||
Norteastern Atlantic:
|
||
<taxonomicName id="4C3D4D44FF0CFF26FE91FD246BD2FD69" authority="Des Moulins, 1842" authorityName="Des Moulins" authorityYear="1842" box="[444,928,655,682]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="jouannetii">
|
||
<emphasis id="B949EAD5FF0CFF26FE91FD246AC9FD69" box="[444,699,655,681]" italics="true" pageId="142" pageNumber="143">Pleurotoma jouannetii</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FDEEFD3B6BD2FD69" author="Des Moulins, M. Ch." box="[707,928,655,682]" pageId="142" pageNumber="143" pagination="109 - 185" refId="ref131318" refString="Des Moulins, M. Ch. (1842) Revision de quelques especes de Pleurotomes. Actes de la Societe linneenne de Bordeaux, 12, 109 - 185. [https: // www. biodiversitylibrary. org / item / 101660 # page / 164 / mode / 1 up]" type="journal article" year="1842">Des Moulins, 1842</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FF0CFF26FC80FD246DEAFD69" authority="Lamarck, 1822" authorityName="Lamarck" authorityYear="1822" box="[941,1432,655,681]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="semimarginata">
|
||
<emphasis id="B949EAD5FF0CFF26FC80FD246C93FD69" box="[941,1249,655,681]" italics="true" pageId="142" pageNumber="143">Pleurotoma semimarginata</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FBC7FD246DEAFD69" author="Lamarck, J. - B. M. de" box="[1258,1432,655,681]" pageId="142" pageNumber="143" refId="ref137485" refString="Lamarck, J. - B. M. de. (1822) Histoire naturelle des animaux sans vertebres. Tome Septieme. Published by the Author, Paris, 711 pp. [http: // www. biodiversitylibrary. org / item / 47433]" type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FF0CFF26FFBAFD1F691BFD0D" box="[151,361,692,717]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="jouanneti">
|
||
<emphasis id="B949EAD5FF0CFF26FFBAFD1F691BFD0D" box="[151,361,692,717]" italics="true" pageId="142" pageNumber="143">Perrona jouanneti</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<emphasis id="B949EAD5FF0CFF26FE53FD1F69EFFD0D" box="[382,413,692,717]" italics="true" pageId="142" pageNumber="143">sic</emphasis>
|
||
]
|
||
<emphasis id="B949EAD5FF0CFF26FE9FFD1F6982FD0D" box="[434,496,692,717]" italics="true" pageId="142" pageNumber="143">sensu</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FED1FD186A90FD0D" author="Landau, B. & Da Silva, C. M. & Mayoral, E." box="[508,738,691,718]" pageId="142" pageNumber="143" pagination="1 - 155" refId="ref137574" refString="Landau, B., Da Silva, C. M. & Mayoral, E. (2011) The Lower Pliocene gastropods of the Huelva Sands Formation, Guadalquivir Basin, Southwestern Spain. Palaeofocus, 4, 1 - 155." type="journal article" year="2011">
|
||
Landau
|
||
<emphasis id="B949EAD5FF0CFF26FD71FD1F6AE0FD0D" box="[604,658,691,717]" italics="true" pageId="142" pageNumber="143">et al</emphasis>
|
||
., 2011
|
||
</bibRefCitation>
|
||
and
|
||
<emphasis id="B949EAD5FF0CFF26FC0FFD1F6B12FD0D" box="[802,864,692,717]" italics="true" pageId="142" pageNumber="143">sensu</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FC46FD186C4EFD0D" author="Vera-Pelaez, J. L. & Lozano-Francisco, M. C." box="[875,1084,691,717]" pageId="142" pageNumber="143" pagination="1 - 67" refId="ref146020" refString="Vera-Pelaez, J. L. & Lozano-Francisco, M. C. (2001) Revision de la subfamilia Clavatulinae (Gastropoda, Prosobranchia, Turridae) con ocho especies nuevas del Plioceno andaluz (S. Espana). Pliocenica, Publicaciones del Museo Municipal Paleontologico de Estepona, 1, 1 - 67." type="journal article" year="2001">Vera-Peláez, 2001</bibRefCitation>
|
||
(
|
||
<emphasis id="B949EAD5FF0CFF26FB7CFD1F6C09FD0D" box="[1105,1147,692,717]" italics="true" pageId="142" pageNumber="143">non</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FBABFD1F6D18FD0D" author="Des Moulins, M. Ch." box="[1158,1386,691,718]" pageId="142" pageNumber="143" pagination="109 - 185" refId="ref131318" refString="Des Moulins, M. Ch. (1842) Revision de quelques especes de Pleurotomes. Actes de la Societe linneenne de Bordeaux, 12, 109 - 185. [https: // www. biodiversitylibrary. org / item / 101660 # page / 164 / mode / 1 up]" type="journal article" year="1842">Des Moulins, 1842</bibRefCitation>
|
||
), an additional species from the Pliocene of Huleva (
|
||
<collectingCountry id="F32A7657FF0CFF26FD9FFD7C6A87FD32" box="[690,757,727,754]" name="Spain" pageId="142" pageNumber="143">Spain</collectingCountry>
|
||
) will be described by
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FCC0FD7C6D4BFD31" author="Landau, B. & Harzhauser, M." box="[1005,1337,727,753]" pageId="142" pageNumber="143" refId="ref137533" refString="Landau, B. & Harzhauser, M. (2022) The Pliocene Gastropoda (Mollusca) of Estepona, southern Spain. Part 14: Clavatulidae (Gastropoda, Conoidea). Cainozoic Research. [in press]" type="book" year="2022">Landau & Harzhauser (2022)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF0CFF26FFEAFD576C09FCD5" blockId="142.[151,1437,223,2014]" box="[199,1147,763,790]" pageId="142" pageNumber="143">
|
||
West Africa (Recent):
|
||
<taxonomicName id="4C3D4D44FF0CFF26FEEFFD506B60FCD5" authority="Strebel, 1912" authorityName="Strebel" authorityYear="1912" box="[450,786,763,789]" class="Gastropoda" family="Clavatulidae" genus="Tomella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="hupferi">
|
||
<emphasis id="B949EAD5FF0CFF26FEEFFD506A01FCD5" box="[450,627,763,789]" italics="true" pageId="142" pageNumber="143">Tomella hupferi</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FD57FD506B60FCD5" author="Strebel, H." box="[634,786,763,789]" pageId="142" pageNumber="143" pagination="1 - 24" refId="ref145028" refString="Strebel, H. (1912) Bemerkungen zu den Clavatula - Gruppen Perrona und Tomella. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 29, 1 - 24, 1 pl. https: // doi. org / 10.5962 / bhl. title. 10744" type="journal article" year="1912">Strebel, 1912</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FF0CFF26FC30FD506C05FCD5" authority="Strebel, 1912" authorityName="Strebel" authorityYear="1912" box="[797,1143,763,789]" class="Gastropoda" family="Clavatulidae" genus="Tomella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="leschkei">
|
||
<emphasis id="B949EAD5FF0CFF26FC30FD506BA5FCD5" box="[797,983,763,789]" italics="true" pageId="142" pageNumber="143">Tomella leschkei</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FCF3FD506C05FCD5" author="Strebel, H." box="[990,1143,763,789]" pageId="142" pageNumber="143" pagination="1 - 24" refId="ref145028" refString="Strebel, H. (1912) Bemerkungen zu den Clavatula - Gruppen Perrona und Tomella. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 29, 1 - 24, 1 pl. https: // doi. org / 10.5962 / bhl. title. 10744" type="journal article" year="1912">Strebel, 1912</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFF0CFF26FFEAFCB46D4FFC09" pageId="142" pageNumber="143" type="reference_group">
|
||
<paragraph id="8B8236C7FF0CFF26FFEAFCB46A30FCF9" blockId="142.[151,1437,223,2014]" box="[199,578,799,825]" pageId="142" pageNumber="143">
|
||
<taxonomicName id="4C3D4D44FF0CFF26FFEAFCB46A4CFCF9" authority="Lamarck, 1816" authorityName="Lamarck" authorityYear="1816" box="[199,574,799,825]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="lineata">
|
||
<emphasis id="B949EAD5FF0CFF26FFEAFCB469F9FCF9" box="[199,395,799,825]" italics="true" pageId="142" pageNumber="143">Clavatula lineata</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FEBCFCB46A4CFCF9" author="Lamarck, J. B. P. A. de" box="[401,574,799,825]" pageId="142" pageNumber="143" pagination="1 - 16" refId="ref137407" refString="Lamarck, J. B. P. A. de (1816) Liste des objets representes dans les planches de cette livraison. In: Lamarck, J. B. P. A. de (Ed.), Tableau Encyclopedique et Methodique des Trois Regnes de la Nature. Mollusques et Polypes Divers. Veuve Agasse, Imprimeur, Paris, pp. 1 - 16, pls. 391 - 488." type="book chapter" year="1816">Lamarck, 1816</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF0CFF26FFEAFCE86D4FFC09" blockId="142.[151,1437,223,2014]" pageId="142" pageNumber="143">
|
||
<emphasis id="B949EAD5FF0CFF26FFEAFCE86AF4FC9D" bold="true" box="[199,646,835,861]" pageId="142" pageNumber="143">Stratigraphic and geographic range.</emphasis>
|
||
Central Paratethys Sea: early to middle Miocene (Eggenburgian to Badenian, late Burdigalian to Langhian),
|
||
<collectingCountry id="F32A7657FF0CFF26FDA1FCCC6A93FC41" box="[652,737,871,897]" name="Austria" pageId="142" pageNumber="143">Austria</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32A7657FF0CFF26FDD9FCCC6B2FFC41" box="[756,861,871,897]" name="Germany" pageId="142" pageNumber="143">Germany</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32A7657FF0CFF26FC5DFCC36BA1FC42" box="[880,979,872,898]" name="Hungary" pageId="142" pageNumber="143">Hungary</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32A7657FF0CFF26FCCBFCCC6C3DFC41" box="[998,1103,871,897]" name="Romania" pageId="142" pageNumber="143">Romania</collectingCountry>
|
||
; (Proto-)Mediterranean Sea:? late Miocene (Tortonian),
|
||
<collectingCountry id="F32A7657FF0CFF26FEF3FC206A61FC65" box="[478,531,907,933]" name="Italy" pageId="142" pageNumber="143">Italy</collectingCountry>
|
||
. Pliocene:
|
||
<collectingCountry id="F32A7657FF0CFF26FDBEFC206AA1FC65" box="[659,723,907,933]" name="Spain" pageId="142" pageNumber="143">Spain</collectingCountry>
|
||
(Landau
|
||
<emphasis id="B949EAD5FF0CFF26FC68FC276BF3FC65" box="[837,897,907,933]" italics="true" pageId="142" pageNumber="143">et al.</emphasis>
|
||
in prep); northeastern Atlantic: early Miocene (Burdigalian):
|
||
<collectingCountry id="F32A7657FF0CFF26FE10FC1B69FFFC0A" box="[317,397,944,970]" name="France" pageId="142" pageNumber="143">France</collectingCountry>
|
||
; present-day: West Africa, upper Pliocene, southwestern
|
||
<collectingCountry id="F32A7657FF0CFF26FB26FC046C39FC09" box="[1035,1099,943,969]" name="Spain" pageId="142" pageNumber="143">Spain</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FB74FC046D43FC09" author="Landau, B. & Da Silva, C. M. & Mayoral, E." box="[1113,1329,943,969]" pageId="142" pageNumber="143" pagination="1 - 155" refId="ref137574" refString="Landau, B., Da Silva, C. M. & Mayoral, E. (2011) The Lower Pliocene gastropods of the Huelva Sands Formation, Guadalquivir Basin, Southwestern Spain. Palaeofocus, 4, 1 - 155." type="journal article" year="2011">
|
||
Landau
|
||
<emphasis id="B949EAD5FF0CFF26FB9BFC1B6C9CFC09" box="[1206,1262,943,969]" italics="true" pageId="142" pageNumber="143">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFF0CFF26FFEAFC786A4DF8B1" pageId="142" pageNumber="143" type="discussion">
|
||
<paragraph id="8B8236C7FF0CFF26FFEAFC786D36FAF1" blockId="142.[151,1437,223,2014]" pageId="142" pageNumber="143">
|
||
<emphasis id="B949EAD5FF0CFF26FFEAFC78693EFC2D" bold="true" box="[199,332,979,1005]" pageId="142" pageNumber="143">Discussion.</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FE7CFC78699AFC2D" author="Powell, A. W. B." box="[337,488,979,1005]" pageId="142" pageNumber="143" pagination="1 - 184" refId="ref141714" refString="Powell, A. W. B. (1966) The molluscan families Speightiidae and Turridae: an evaluation of the valid taxa, both recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Museum, 5, 1 - 184. [https: // www. biodiversitylibrary. org / item / 278249 # page / 4 / mode / 1 up]" type="journal article" year="1966">Powell (1966</bibRefCitation>
|
||
, p. 56) considered
|
||
<taxonomicName id="4C3D4D44FF0CFF26FD9FFC786BDCFC2D" authority="Wenz, 1943" authorityName="Wenz" authorityYear="1943" box="[690,942,979,1006]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF0CFF26FD9FFC786B55FC2D" box="[690,807,979,1005]" italics="true" pageId="142" pageNumber="143">Tomellana</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FC01FC7F6BDCFC2D" author="Wenz, W." box="[812,942,979,1006]" pageId="142" pageNumber="143" pagination="1201 - 1506" refId="ref146470" refString="Wenz, W. (1943) Gastropoda. Teil 1: Allgemeiner Teil und Prosobranchia. xii + 1639 pp. In: Schindewolf, O. H. (Ed.) Handbuch der Palaozoologie. Band 6. Lfg. 8. Borntrager, Berlin, pp. 1201 - 1506." type="book chapter" year="1943">Wenz, 1943</bibRefCitation>
|
||
</taxonomicName>
|
||
monotypic but MolluscaBase (Eds.) (2021d) includes also
|
||
<taxonomicName id="4C3D4D44FF0CFF26FE1BFC5C6ACBFBD1" authority="(Strebel, 1912)" baseAuthorityName="Strebel" baseAuthorityYear="1912" box="[310,697,1015,1041]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="hupferi">
|
||
<emphasis id="B949EAD5FF0CFF26FE1BFC5C6A75FBD1" box="[310,519,1015,1041]" italics="true" pageId="142" pageNumber="143">Tomellana hupferi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FD38FC5C6AC2FBD1" author="Strebel, H." box="[533,688,1015,1041]" pageId="142" pageNumber="143" pagination="1 - 24" refId="ref145028" refString="Strebel, H. (1912) Bemerkungen zu den Clavatula - Gruppen Perrona und Tomella. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 29, 1 - 24, 1 pl. https: // doi. org / 10.5962 / bhl. title. 10744" type="journal article" year="1912">Strebel, 1912</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3D4D44FF0CFF26FDDBFC5C6CF3FBD1" authority="(Strebel, 1912)" baseAuthorityName="Strebel" baseAuthorityYear="1912" box="[758,1153,1015,1041]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="leschkei">
|
||
<emphasis id="B949EAD5FF0CFF26FDDBFC5C6BBDFBD1" box="[758,975,1015,1041]" italics="true" pageId="142" pageNumber="143">Tomellana leschkei</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FCF3FC5C6C0BFBD1" author="Strebel, H." box="[990,1145,1015,1041]" pageId="142" pageNumber="143" pagination="1 - 24" refId="ref145028" refString="Strebel, H. (1912) Bemerkungen zu den Clavatula - Gruppen Perrona und Tomella. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 29, 1 - 24, 1 pl. https: // doi. org / 10.5962 / bhl. title. 10744" type="journal article" year="1912">Strebel, 1912</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in this genus. Including these species documents that the callus pad of the
|
||
<typeStatus id="54868865FF0CFF26FDFFFBB76B70FBF6" box="[722,770,1052,1078]" pageId="142" pageNumber="143">type</typeStatus>
|
||
species is extraordinarily thickened and that this feature is not indicative for the genus. If the early whorls are used to characterize the genus then
|
||
<taxonomicName id="4C3D4D44FF0CFF26FB5DFB946C97FB99" authorityName="Wenz" authorityYear="1943" box="[1136,1253,1087,1113]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF0CFF26FB5DFB946C97FB99" box="[1136,1253,1087,1113]" italics="true" pageId="142" pageNumber="143">Tomellana</emphasis>
|
||
</taxonomicName>
|
||
is characterized by “
|
||
<emphasis id="B949EAD5FF0CFF26FFEBFBC86BE8FB05" italics="true" pageId="142" pageNumber="143">Falten …, die der Anwuchsrichtung folgend, geschweift sind, anfangs sich über die ganze Windung erstrecken, dann oben obsolet werden und schliesslich in kurze schräge Falten oberhalb der Naht auslaufen, die aber je nach der Art nur zwischen der zweiten uns siebenten Windung auftreten</emphasis>
|
||
” [by sigmoidal folds, which initially follow the growth lines across the entire whorl but become adapically obsolete and grade into short, oblique folds above the suture, which depending on species, appear only between the second and seventh whorl]
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FB9BFB586D0AFACE" author="Strebel, H." box="[1206,1400,1267,1294]" pageId="142" pageNumber="143" pagination="1 - 24" refId="ref145028" refString="Strebel, H. (1912) Bemerkungen zu den Clavatula - Gruppen Perrona und Tomella. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 29, 1 - 24, 1 pl. https: // doi. org / 10.5962 / bhl. title. 10744" type="journal article" year="1912">Strebel (1912: 5)</bibRefCitation>
|
||
. In addition, we observe a distinct, but narrow subsutural spiral cord on early teleoconch whorls of
|
||
<emphasis id="B949EAD5FF0CFF26FBEAFABC6D36FAF1" box="[1223,1348,1303,1329]" italics="true" pageId="142" pageNumber="143">
|
||
<taxonomicName id="4C3D4D44FF0CFF26FBEAFABC6D32FAF1" authorityName="Wenz" authorityYear="1943" box="[1223,1344,1303,1329]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">Tomellana</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF0CFF26FFEAFA976CF1F991" blockId="142.[151,1437,223,2014]" pageId="142" pageNumber="143">
|
||
Later teleoconch whorls bear a weakly carinate subsutural cord some distance from the suture. This feature is typically developed in
|
||
<taxonomicName id="4C3D4D44FF0CFF26FE95FACB6A54FAB9" baseAuthorityName="Strebel" baseAuthorityYear="1912" box="[440,550,1375,1401]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="hupferi">
|
||
<emphasis id="B949EAD5FF0CFF26FE95FACB6A54FAB9" box="[440,550,1375,1401]" italics="true" pageId="142" pageNumber="143">T. hupferi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3D4D44FF0CFF26FD4DFACB6BB7FAB9" authority="(Strebel, 1912)" baseAuthorityName="Strebel" baseAuthorityYear="1912" box="[608,965,1375,1401]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="subSpecies" species="hupferi" subSpecies="fusca">
|
||
<emphasis id="B949EAD5FF0CFF26FD4DFACB6B60FAB9" box="[608,786,1375,1401]" italics="true" pageId="142" pageNumber="143">T. hupferi fusca</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FC0EFAF46BCEFAB9" author="Strebel, H." box="[803,956,1375,1401]" pageId="142" pageNumber="143" pagination="1 - 24" refId="ref145028" refString="Strebel, H. (1912) Bemerkungen zu den Clavatula - Gruppen Perrona und Tomella. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 29, 1 - 24, 1 pl. https: // doi. org / 10.5962 / bhl. title. 10744" type="journal article" year="1912">Strebel, 1912</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FB2EFAF46C0EFABA" author="Nolf, F." box="[1027,1148,1375,1402]" pageId="142" pageNumber="143" pagination="17 - 37" refId="ref140261" refString="Nolf, F. (2015) The genus Tomellana (Mollusca: Gastropoda: Clavatulidae) in West Africa: a comprehensive Survey and establishment of Fusiturris kribiensis Bozzetti, 2015 as a junior synonym. Neptunea, 13, 17 - 37. [https: // neptuneaorg. files. wordpress. com / 2019 / 02 / np 13 - 4 - tekst. pdf # 17]" type="journal article" year="2015">Nolf 2015</bibRefCitation>
|
||
: pl. 9), and in
|
||
<taxonomicName id="4C3D4D44FF0CFF26FA08FACB694DFA5D" authority="(Strebel, 1912)" baseAuthorityName="Strebel" baseAuthorityYear="1912" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="leschkei">
|
||
<emphasis id="B949EAD5FF0CFF26FA08FACB6DEEFAB9" box="[1317,1436,1375,1401]" italics="true" pageId="142" pageNumber="143">T. leschkei</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FFB2FA286945FA5D" author="Strebel, H." box="[159,311,1411,1437]" pageId="142" pageNumber="143" pagination="1 - 24" refId="ref145028" refString="Strebel, H. (1912) Bemerkungen zu den Clavatula - Gruppen Perrona und Tomella. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 29, 1 - 24, 1 pl. https: // doi. org / 10.5962 / bhl. title. 10744" type="journal article" year="1912">Strebel, 1912</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FE56FA286983FA5E" author="Nolf, F." box="[379,497,1411,1438]" pageId="142" pageNumber="143" pagination="17 - 37" refId="ref140261" refString="Nolf, F. (2015) The genus Tomellana (Mollusca: Gastropoda: Clavatulidae) in West Africa: a comprehensive Survey and establishment of Fusiturris kribiensis Bozzetti, 2015 as a junior synonym. Neptunea, 13, 17 - 37. [https: // neptuneaorg. files. wordpress. com / 2019 / 02 / np 13 - 4 - tekst. pdf # 17]" type="journal article" year="2015">Nolf 2015</bibRefCitation>
|
||
: pl. 12), but rarely also seen in
|
||
<taxonomicName id="4C3D4D44FF0CFF26FC7FFA2F6CBAFA5D" authority="(Strebel, 1912)" baseAuthorityName="Strebel" baseAuthorityYear="1912" box="[850,1224,1411,1437]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="subSpecies" species="lineata" subSpecies="gracilis">
|
||
<emphasis id="B949EAD5FF0CFF26FC7FFA2F6C6BFA5D" box="[850,1049,1411,1437]" italics="true" pageId="142" pageNumber="143">T. lineata gracilis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FB0AFA286CCDFA5D" author="Strebel, H." box="[1063,1215,1411,1437]" pageId="142" pageNumber="143" pagination="1 - 24" refId="ref145028" refString="Strebel, H. (1912) Bemerkungen zu den Clavatula - Gruppen Perrona und Tomella. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 29, 1 - 24, 1 pl. https: // doi. org / 10.5962 / bhl. title. 10744" type="journal article" year="1912">Strebel, 1912</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FBFBFA286D3DFA5E" author="Nolf, F." box="[1238,1359,1411,1438]" pageId="142" pageNumber="143" pagination="17 - 37" refId="ref140261" refString="Nolf, F. (2015) The genus Tomellana (Mollusca: Gastropoda: Clavatulidae) in West Africa: a comprehensive Survey and establishment of Fusiturris kribiensis Bozzetti, 2015 as a junior synonym. Neptunea, 13, 17 - 37. [https: // neptuneaorg. files. wordpress. com / 2019 / 02 / np 13 - 4 - tekst. pdf # 17]" type="journal article" year="2015">Nolf 2015</bibRefCitation>
|
||
: pl. 8).
|
||
<taxonomicName id="4C3D4D44FF0CFF26FFBAFA036881FA01" box="[151,243,1448,1473]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF0CFF26FFBAFA036881FA01" box="[151,243,1448,1473]" italics="true" pageId="142" pageNumber="143">Perrona</emphasis>
|
||
</taxonomicName>
|
||
, as understood herein, has opisthocline, comma-shaped axial riblets between two adsutural spiral cords (
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FFB2FA606941FA25" author="Strebel, H." box="[159,307,1483,1509]" pageId="142" pageNumber="143" pagination="1 - 24" refId="ref145028" refString="Strebel, H. (1912) Bemerkungen zu den Clavatula - Gruppen Perrona und Tomella. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 29, 1 - 24, 1 pl. https: // doi. org / 10.5962 / bhl. title. 10744" type="journal article" year="1912">Strebel 1912</bibRefCitation>
|
||
), whereas
|
||
<taxonomicName id="4C3D4D44FF0CFF26FE84FA606A6CFA25" authorityName="Wenz" authorityYear="1943" box="[425,542,1483,1509]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF0CFF26FE84FA606A6CFA25" box="[425,542,1483,1509]" italics="true" pageId="142" pageNumber="143">Tomellana</emphasis>
|
||
</taxonomicName>
|
||
has a beaded suprasutural spiral cord. The siphonal canal of
|
||
<taxonomicName id="4C3D4D44FF0CFF26FBE5FA606D4FFA25" authorityName="Wenz" authorityYear="1943" box="[1224,1341,1483,1509]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF0CFF26FBE5FA606D4FFA25" box="[1224,1341,1483,1509]" italics="true" pageId="142" pageNumber="143">Tomellana</emphasis>
|
||
</taxonomicName>
|
||
is much longer and its spire is conical as opposed to slightly gradate in most
|
||
<taxonomicName id="4C3D4D44FF0CFF26FCB4FA5B6C27F9C9" box="[921,1109,1519,1545]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B949EAD5FF0CFF26FCB4FA5B6B87F9C9" box="[921,1013,1520,1545]" italics="true" pageId="142" pageNumber="143">Perrona</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
The separation of
|
||
<taxonomicName id="4C3D4D44FF0CFF26FA0AFA446DEEF9C9" authorityName="Wenz" authorityYear="1943" box="[1319,1436,1519,1545]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF0CFF26FA0AFA446DEEF9C9" box="[1319,1436,1519,1545]" italics="true" pageId="142" pageNumber="143">Tomellana</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C3D4D44FF0CFF26FFFBF9BF6940F9ED" box="[214,306,1556,1581]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF0CFF26FFFBF9BF6940F9ED" box="[214,306,1556,1581]" italics="true" pageId="142" pageNumber="143">Perrona</emphasis>
|
||
</taxonomicName>
|
||
is also supported by molecular data (N. Puillandre, pers. comm.,
|
||
<date id="FF831007FF0CFF26FB0FF9B86CB3F9ED" box="[1058,1217,1555,1581]" pageId="142" pageNumber="143" value="2021-10">October 2021</date>
|
||
), corroborating our interpretation of the systematic value of early teleoconch whorl sculpture in
|
||
<taxonomicName id="4C3D4D44FF0CFF26FCC3F99C6C0DF991" authorityName="Gray" authorityYear="1853" box="[1006,1151,1591,1617]" class="Gastropoda" family="Clavatulidae" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="family">Clavatulidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF0CFF26FFEAF9F06A4DF8B1" blockId="142.[151,1437,223,2014]" pageId="142" pageNumber="143">
|
||
So far, this genus was only known from recent species from West Africa (
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FCD2F9F06CEAF9B5" author="Strebel, H." box="[1023,1176,1627,1653]" pageId="142" pageNumber="143" pagination="1 - 24" refId="ref145028" refString="Strebel, H. (1912) Bemerkungen zu den Clavatula - Gruppen Perrona und Tomella. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 29, 1 - 24, 1 pl. https: // doi. org / 10.5962 / bhl. title. 10744" type="journal article" year="1912">Strebel, 1912</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FB89F9F06D50F9B6" author="Nolf, F." box="[1188,1314,1627,1654]" pageId="142" pageNumber="143" pagination="17 - 37" refId="ref140261" refString="Nolf, F. (2015) The genus Tomellana (Mollusca: Gastropoda: Clavatulidae) in West Africa: a comprehensive Survey and establishment of Fusiturris kribiensis Bozzetti, 2015 as a junior synonym. Neptunea, 13, 17 - 37. [https: // neptuneaorg. files. wordpress. com / 2019 / 02 / np 13 - 4 - tekst. pdf # 17]" type="journal article" year="2015">Nolf, 2015</bibRefCitation>
|
||
). Based on the early teleoconch morphology and overall shape, we assume that Miocene representatives of
|
||
<taxonomicName id="4C3D4D44FF0CFF26FBF0F9D46D20F959" authorityName="Wenz" authorityYear="1943" box="[1245,1362,1663,1689]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF0CFF26FBF0F9D46D20F959" box="[1245,1362,1663,1689]" italics="true" pageId="142" pageNumber="143">Tomellana</emphasis>
|
||
</taxonomicName>
|
||
might have been overlooked so far by placing the species in
|
||
<emphasis id="B949EAD5FF0CFF26FDD3F90F6B10F97D" box="[766,866,1700,1725]" italics="true" pageId="142" pageNumber="143">
|
||
<taxonomicName id="4C3D4D44FF0CFF26FDD3F90F6B2CF97D" box="[766,862,1700,1725]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">Perrona</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
An emblematic example is
|
||
<taxonomicName id="4C3D4D44FF0CFF26FB8DF908690AF921" authority="Des Moulins, 1842" authorityName="Des Moulins" authorityYear="1842" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="jouannetii">
|
||
<emphasis id="B949EAD5FF0CFF26FB8DF9086DEFF97D" box="[1184,1437,1699,1725]" italics="true" pageId="142" pageNumber="143">Pleurotoma jouannetii</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FFBAF963690AF921" author="Des Moulins, M. Ch." box="[151,376,1735,1762]" pageId="142" pageNumber="143" pagination="109 - 185" refId="ref131318" refString="Des Moulins, M. Ch. (1842) Revision de quelques especes de Pleurotomes. Actes de la Societe linneenne de Bordeaux, 12, 109 - 185. [https: // www. biodiversitylibrary. org / item / 101660 # page / 164 / mode / 1 up]" type="journal article" year="1842">Des Moulins, 1842</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the Burdigalian of Mérignac in
|
||
<collectingCountry id="F32A7657FF0CFF26FC10F9636BFEF922" box="[829,908,1736,1762]" name="France" pageId="142" pageNumber="143">France</collectingCountry>
|
||
, which was placed in
|
||
<taxonomicName id="4C3D4D44FF0CFF26FBB8F9636C83F921" box="[1173,1265,1736,1761]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF0CFF26FBB8F9636C83F921" box="[1173,1265,1736,1761]" italics="true" pageId="142" pageNumber="143">Perrona</emphasis>
|
||
</taxonomicName>
|
||
by all authors since
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FFF4F947690AF8C5" author="Peyrot, A." box="[217,376,1771,1798]" pageId="142" pageNumber="143" pagination="5 - 288" refId="ref141284" refString="Peyrot, A. (1931 & 1933) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 83, 5 - 116 (1931), 84, 5 - 288 (1933). [https: // www. biodiversitylibrary. org / item / 185325 # page / 11 / mode / 1 up]" type="journal article" year="1931">Peyrot (1931)</bibRefCitation>
|
||
. We did not study topotypic specimens of this species from Mérignac, but the early whorls of specimens from the Burdigalian of Léognan (
|
||
<collectingCountry id="F32A7657FF0CFF26FDA2F8A46A92F8E9" box="[655,736,1807,1833]" name="France" pageId="142" pageNumber="143">France</collectingCountry>
|
||
, NHMW 1858/0018/0104) (
|
||
<figureCitation id="13062A42FF0CFF26FB37F8A46C1FF8E9" box="[1050,1133,1807,1833]" captionStart="FIGURE 43" captionStartId="148.[151,250,1846,1870]" captionTargetBox="[215,1372,885,1823]" captionTargetId="figure-317@148.[215,1372,885,1823]" captionTargetPageId="148" captionText="FIGURE 43 A1–A3. Tomellana jouannetii (Des Moulins, 1842), NHMW 1858/0018/0104, Mérignac (France), Burdigalian." figureDoi="http://doi.org/10.5281/zenodo.6399927" httpUri="https://zenodo.org/record/6399927/files/figure.png" pageId="142" pageNumber="143">Fig. 43</figureCitation>
|
||
), differ greatly from that of typical
|
||
<taxonomicName id="4C3D4D44FF0CFF26FFC7F89F6934F88D" box="[234,326,1844,1869]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF0CFF26FFC7F89F6934F88D" box="[234,326,1844,1869]" italics="true" pageId="142" pageNumber="143">Perrona</emphasis>
|
||
</taxonomicName>
|
||
, which has short, comma-shaped riblets between two narrow adsutural spiral cords, but agrees well with that of extant
|
||
<taxonomicName id="4C3D4D44FF0CFF26FE46F8FC6A4DF8B1" box="[363,575,1879,1905]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B949EAD5FF0CFF26FE46F8FC6992F8B1" box="[363,480,1879,1905]" italics="true" pageId="142" pageNumber="143">Tomellana</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFF0CFF27FFEAF8D06BD3FF15" lastPageId="143" lastPageNumber="144" pageId="142" pageNumber="143" type="reference_group">
|
||
<paragraph id="8B8236C7FF0CFF26FFEAF8D06C37F81E" blockId="142.[151,1437,223,2014]" pageId="142" pageNumber="143">
|
||
<taxonomicName id="4C3D4D44FF0CFF26FFEAF8D069CDF855" authority="Gray, 1847" authorityName=", Gray" authorityYear="1847" box="[199,447,1915,1941]" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF0CFF26FFEAF8D0694DF855" box="[199,319,1915,1941]" italics="true" pageId="142" pageNumber="143">Pusionella</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FE69F8D069CDF855" author="Gray, J. E." box="[324,447,1915,1941]" pageId="142" pageNumber="143" pagination="129 - 219" refId="ref133560" refString="Gray, J. E. (1847) A list of the genera of recent mollusca, their synonyma and types. Proceedings of the Zoological Society of London, 15, 129 - 219. [http: // biodiversitylibrary. org / page / 12862913]" type="journal article" year="1847">Gray, 1847</bibRefCitation>
|
||
</taxonomicName>
|
||
differs distinctly in its wide and very shallow anal canal. The extant
|
||
<taxonomicName id="4C3D4D44FF0CFF26FB83F8D06941F879" authority="Strebel, 1914" authorityName="Strebel" authorityYear="1914" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="species" species="kraepelini">
|
||
<emphasis id="B949EAD5FF0CFF26FB83F8D06DEFF855" box="[1198,1437,1915,1941]" italics="true" pageId="142" pageNumber="143">Pusionella kraepelini</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FFBAF8346941F879" author="Strebel, H." box="[151,307,1951,1977]" pageId="142" pageNumber="143" pagination="87 - 125" refId="ref145074" refString="Strebel, H. (1914) Mollusca I, Gen. Pusionella. In: Michaelsen, W. (Ed.), Kenntnis der Meeresfauna Westafrikas. Band 1. Friedrichsen & Co., Hamburg, 87 - 125. https: // www. biodiversitylibrary. org / page / 7078288 # page / 101 / mode / 1 up" type="book chapter" year="1914">Strebel, 1914</bibRefCitation>
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="F32A7657FF0CFF26FEACF83469A1F879" box="[385,467,1951,1977]" name="Angola" pageId="142" pageNumber="143">Angola</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32A7657FF0CFF26FD3FF8346A2DF879" box="[530,607,1951,1977]" name="Gabon" pageId="142" pageNumber="143">Gabon</collectingCountry>
|
||
, is reminiscent in general shape and bears beads at the abapical suture (
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FFB2F8686940F81D" author="Strebel, H." box="[159,306,1987,2013]" pageId="142" pageNumber="143" pagination="87 - 125" refId="ref145074" refString="Strebel, H. (1914) Mollusca I, Gen. Pusionella. In: Michaelsen, W. (Ed.), Kenntnis der Meeresfauna Westafrikas. Band 1. Friedrichsen & Co., Hamburg, 87 - 125. https: // www. biodiversitylibrary. org / page / 7078288 # page / 101 / mode / 1 up" type="book chapter" year="1914">Strebel 1914</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAC4B36FF0CFF26FE13F86869A9F81D" author="Bernard, P. A." box="[318,475,1987,2013]" pageId="142" pageNumber="143" refId="ref129362" refString="Bernard, P. A. (1984) Coquillages du Gabon [Shells of Gabon]. Pierre A. Libreville, Gabon, Bernard, 140 pp., 75 pls." type="book" year="1984">Bernard 1984</bibRefCitation>
|
||
), but has a shallow anal sinus as typical for
|
||
<taxonomicName id="4C3D4D44FF0CFF26FCEBF8686C4CF81D" authorityName=", Gray" authorityYear="1847" box="[966,1086,1987,2013]" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="142" pageNumber="143" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF0CFF26FCEBF8686C4CF81D" box="[966,1086,1987,2013]" italics="true" pageId="142" pageNumber="143">Pusionella</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF0DFF27FFEAFF3D6BD3FF15" blockId="143.[151,1436,150,213]" pageId="143" pageNumber="144">
|
||
<emphasis id="B949EAD5FF0DFF27FFEAFF3D69DAFF70" bold="true" box="[199,424,150,176]" pageId="143" pageNumber="144">Paleoenvironment.</emphasis>
|
||
The fossil species derive from inner neritic environments. Present-day species are reported from soft bottom environments in
|
||
<quantity id="4CC59B22FF0DFF27FD35FF106AF1FF15" box="[536,643,186,213]" metricMagnitude="1" metricUnit="m" metricValue="2.85" metricValueMax="5.0" metricValueMin="0.7" pageId="143" pageNumber="144" unit="m" value="28.5" valueMax="50.0" valueMin="7.0">7 to 50 m</quantity>
|
||
water depth (
|
||
<bibRefCitation id="EFAC4B36FF0DFF27FC31FF116BE7FF15" author="Nolf, F." box="[796,917,186,213]" pageId="143" pageNumber="144" pagination="17 - 37" refId="ref140261" refString="Nolf, F. (2015) The genus Tomellana (Mollusca: Gastropoda: Clavatulidae) in West Africa: a comprehensive Survey and establishment of Fusiturris kribiensis Bozzetti, 2015 as a junior synonym. Neptunea, 13, 17 - 37. [https: // neptuneaorg. files. wordpress. com / 2019 / 02 / np 13 - 4 - tekst. pdf # 17]" type="journal article" year="2015">Nolf 2015</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |