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<mods:title id="BF48C608848DBC2771971BB6EA9573BE">Two new species and a new record of Bactrocera Macquart (Diptera: Tephritidae: Dacinae: Dacini) from India with an updated key to species of subgenus Bactrocera</mods:title>
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<mods:namePart id="FAE9F3DE6BCDFF16D2C5868BA99A37C0">Abhishek, Venkateshaiah</mods:namePart>
<mods:affiliation id="4E5EECD6731165F3A329D9310CBD98D4">ICAR-National Bureau of Agricultural Insect Resources, Bengaluru- 560024, Karnataka, India. &amp; Keladi Shivappa Nayaka University of Agricultural and Horticultural Sciences, Shivamogga, Karnataka, India</mods:affiliation>
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<mods:namePart id="D151817B0387A6778EB36339F85DC199">David, Karamankodu Jacob</mods:namePart>
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<mods:affiliation id="FA74E6612D614636CB1299BF2D5425DF">ICAR-National Bureau of Agricultural Insect Resources, Bengaluru- 560024, Karnataka, India. &amp; Keladi Shivappa Nayaka University of Agricultural and Horticultural Sciences, Shivamogga, Karnataka, India &amp; ICAR-National Bureau of Agricultural Insect Resources, Bengaluru- 560024, Karnataka, India.</mods:affiliation>
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<taxonomicName id="4C3D4D52ED2AFF8A1C9EF969FD33F958" ID-CoL="8B6CV" authority="Drew" authorityName="Drew, Drew and Raghu" authorityYear="2002" box="[151,701,1698,1725]" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="290" phylum="Arthropoda" rank="species" species="paraosbeckiae" subGenus="Bactrocera">
<emphasis id="B949EAC3ED2AFF8A1C9EF969FE93F958" bold="true" box="[151,285,1699,1725]" italics="true" pageId="7" pageNumber="290">Bactrocera</emphasis>
(
<emphasis id="B949EAC3ED2AFF8A1D27F969FE3AF958" bold="true" box="[302,436,1699,1725]" italics="true" pageId="7" pageNumber="290">Bactrocera</emphasis>
)
<emphasis id="B949EAC3ED2AFF8A1DCCF968FDFCF958" bold="true" box="[453,626,1698,1725]" italics="true" pageId="7" pageNumber="290">paraosbeckiae</emphasis>
Drew
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(
<figureCitation id="13062A54ED2AFF8A1EC5F968FCF0F959" box="[716,894,1698,1724]" captionStart="FIGURES 1724" captionStartId="8.[151,261,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-24@8.[151,1436,181,1585]" captionTargetPageId="8" captionText="FIGURES 1724. Bactrocera (Bactrocera) paraosbeckiae Drew. 17, head; 18, scutum (dorsal view); 19, thorax (lateral view); 20, abdomen; 21, wing; 22, epandrium and surstyli (lateral view); 23, epandrium and surstyli (posterior view); 24, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209983" httpUri="https://zenodo.org/record/13209983/files/figure.png" pageId="7" pageNumber="290">Figures 1724</figureCitation>
)
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<treatmentCitation id="0A9C10C0ED2AFF8A1C9EF922FC1BF91A" author="Drew, R. A. I. &amp; Raghu, S." box="[151,917,1767,1791]" page="340" pageId="7" pageNumber="290" year="2002">
<taxonomicName id="4C3D4D52ED2AFF8A1C9EF922FC1BF91A" ID-CoL="8B6CV" authority="Drew, Drew and Raghu, 2002: 340" authorityName="Drew, Drew and Raghu" authorityPageNumber="340" authorityYear="2002" box="[151,917,1767,1791]" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="290" phylum="Arthropoda" rank="species" species="paraosbeckiae" subGenus="Bactrocera">
<emphasis id="B949EAC3ED2AFF8A1C9EF922FE86F91A" box="[151,264,1768,1791]" italics="true" pageId="7" pageNumber="290">Bactrocera</emphasis>
(
<emphasis id="B949EAC3ED2AFF8A1D1EF922FE06F91A" box="[279,392,1768,1791]" italics="true" pageId="7" pageNumber="290">Bactrocera</emphasis>
)
<emphasis id="B949EAC3ED2AFF8A1D9EF92DFDA5F91A" box="[407,555,1767,1791]" italics="true" pageId="7" pageNumber="290">paraosbeckiae</emphasis>
<materialsCitation id="3B553C8CED2AFF8A1E38F922FC1BF91A" box="[561,917,1767,1791]" country="United Kingdom" location="London" pageId="7" pageNumber="290" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EFAC4B20ED2AFF8A1E38F922FC1BF91A" author="Drew, R. A. I. &amp; Raghu, S." box="[561,917,1767,1791]" pageId="7" pageNumber="290" pagination="327 - 352" refId="ref12646" refString="Drew, R. A. I. &amp; Raghu, S. (2002) The fruit fly fauna (Diptera: Tephritidae: Dacinae) of the rainforest habitat of the Western Ghats, India. Raffles Bulletin of Zoology, 50 (2), 327 - 352." type="journal article" year="2002">Drew, Drew and Raghu, 2002: 340</bibRefCitation>
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.
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male in NHM,
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,
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.
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Material examined:
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,
<collectingCountry id="F32A7641ED2AFF8A1DA4F8E4FE71F8AD" box="[429,511,1838,1864]" name="India" pageId="7" pageNumber="290">INDIA</collectingCountry>
,
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,
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,
<collectionCode id="ED2CAE14ED2AFF8A1EDDF8E7FCA6F8A2" box="[724,808,1837,1863]" pageId="7" pageNumber="290">CPCRI</collectionCode>
,
<date id="FF831011ED2AFF8A1F3DF8E7FC26F8AD" box="[820,936,1837,1864]" pageId="7" pageNumber="290" value="2015-02-19">
<collectingDate id="EFC7E9F9ED2AFF8A1F3DF8E7FC26F8AD" box="[820,936,1837,1864]" pageId="7" pageNumber="290" value="2015-02-19">19.ii.2015</collectingDate>
</date>
,
<collectorName id="26C85307ED2AFF8A1FBCF8E7FBBDF8AD" box="[949,1075,1837,1864]" pageId="7" pageNumber="290">David, K.J</collectorName>
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.;
<materialsCitation id="3B553C8CED2AFF8A184CF8E7FE50F88E" collectingDate="2012-11-13" collectorName="Devaraj" country="India" location="Maroor" pageId="7" pageNumber="290" specimenCount="2" specimenCount-male="2" stateProvince="Kerala">
<specimenCount id="9D3BFD58ED2AFF8A184CF8E7FB0EF8A2" box="[1093,1152,1837,1863]" count="2" pageId="7" pageNumber="290" type="male">2♂♂</specimenCount>
,
<collectingCountry id="F32A7641ED2AFF8A1885F8E4FB50F8AD" box="[1164,1246,1838,1864]" name="India" pageId="7" pageNumber="290">INDIA</collectingCountry>
,
<collectingRegion id="49F9F833ED2AFF8A18E2F8E7FAB6F8A2" box="[1259,1336,1837,1863]" country="India" name="Kerala" pageId="7" pageNumber="290">Kerala</collectingRegion>
, Calicut,
<location id="8EE2600AED2AFF8A1C9EF898FF63F889" LSID="urn:lsid:plazi:treatment:039487C7ED2AFF841C9EF969FDF4FF30:8EE2600AED2AFF8A1C9EF898FF63F889" box="[151,237,1874,1900]" country="India" name="Maroor" pageId="7" pageNumber="290" stateProvince="Kerala">Maroor</location>
,
<date id="FF831011ED2AFF8A1CF3F89BFEFAF88E" box="[250,372,1873,1899]" pageId="7" pageNumber="290" value="2012-11-13">
<collectingDate id="EFC7E9F9ED2AFF8A1CF3F89BFEFAF88E" box="[250,372,1873,1899]" pageId="7" pageNumber="290" value="2012-11-13">13.xi.2012</collectingDate>
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,
<collectorName id="26C85307ED2AFF8A1D8BF89BFE50F88E" box="[386,478,1873,1899]" pageId="7" pageNumber="290">Devaraj</collectorName>
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;
<materialsCitation id="3B553C8CED2AFF8A1DE2F89BFB7FF88E" box="[491,1265,1873,1900]" collectingDate="2012-09-14" collectionCode="R" collectorName="Sawai" country="India" location="Konkan Region" pageId="7" pageNumber="290" specimenCount="1" specimenCount-male="1" stateProvince="Maharashtra">
<specimenCount id="9D3BFD58ED2AFF8A1DE2F89BFD81F88E" box="[491,527,1873,1899]" count="1" pageId="7" pageNumber="290" type="male">1♂</specimenCount>
,
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,
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,
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,
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<collectingDate id="EFC7E9F9ED2AFF8A1FD2F89BFBDBF88E" box="[987,1109,1873,1899]" pageId="7" pageNumber="290" value="2012-09-14">14.ix.2012</collectingDate>
</date>
, H.
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.
<collectorName id="26C85307ED2AFF8A18A0F89BFB7FF88E" box="[1193,1265,1873,1899]" pageId="7" pageNumber="290">Sawai</collectorName>
</materialsCitation>
;
<materialsCitation id="3B553C8CED2AFF8A18F7F89BFD6BF86A" collectingDate="2017-10-18" collectorName="Latha, S." country="India" location="Maroor" pageId="7" pageNumber="290" specimenCount="3" specimenCount-male="3" stateProvince="Kerala">
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,
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, Calicut,
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,
<date id="FF831011ED2AFF8A1DB9F8BFFDACF86A" box="[432,546,1909,1935]" pageId="7" pageNumber="290" value="2017-10-18">
<collectingDate id="EFC7E9F9ED2AFF8A1DB9F8BFFDACF86A" box="[432,546,1909,1935]" pageId="7" pageNumber="290" value="2017-10-18">18.x.2017</collectingDate>
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,
<collectorName id="26C85307ED2AFF8A1E27F8BFFD1FF86A" box="[558,657,1909,1935]" pageId="7" pageNumber="290">Latha, S.</collectorName>
(NIM)
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.
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<emphasis id="B949EAC3ED25FF851C9EF982FEDEF984" bold="true" box="[151,336,1608,1633]" pageId="8" pageNumber="291">FIGURES 1724.</emphasis>
<taxonomicName id="4C3D4D52ED25FF851D5DF983FCAEF985" authority="Drew." authorityName="Drew, Drew and Raghu" authorityYear="2002" box="[340,800,1609,1633]" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="291" phylum="Arthropoda" rank="species" species="paraosbeckiae" subGenus="Bactrocera">
<emphasis id="B949EAC3ED25FF851D5DF983FD50F984" box="[340,734,1609,1633]" italics="true" pageId="8" pageNumber="291">Bactrocera (Bactrocera) paraosbeckiae</emphasis>
Drew.
</taxonomicName>
17, head; 18, scutum (dorsal view); 19, thorax (lateral view); 20, abdomen; 21, wing; 22, epandrium and surstyli (lateral view); 23, epandrium and surstyli (posterior view); 24, glans of phallus.
</paragraph>
</caption>
<subSubSection id="C327655AED25FF841CCEF903FDF4FF30" lastPageId="9" lastPageNumber="292" pageId="8" pageNumber="291" type="diagnosis">
<paragraph id="8B8236D1ED25FF851CCEF903FAB5F859" blockId="8.[151,1437,1737,2016]" pageId="8" pageNumber="291">
<emphasis id="B949EAC3ED25FF851CCEF903FECEF906" bold="true" box="[199,320,1737,1763]" pageId="8" pageNumber="291">Diagnosis.</emphasis>
This species has been adequately described by
<bibRefCitation id="EFAC4B20ED25FF851F6BF900FBE8F901" author="Drew, R. A. I. &amp; Romig, M. C." box="[866,1126,1738,1764]" pageId="8" pageNumber="291" refId="ref12751" refString="Drew, R. A. I. &amp; Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia. CAB International, Wallingford, vii + 653 pp. https: // doi. org / 10.1079 / 9781780640358.0000" type="book" year="2013">Drew &amp; Romig (2013)</bibRefCitation>
except for some additional genitalia characters. Male genitalia (
<figureCitation id="13062A54ED25FF851E2FF924FD43F8ED" box="[550,717,1774,1800]" captionStart="FIGURES 1724" captionStartId="8.[151,261,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-24@8.[151,1436,181,1585]" captionTargetPageId="8" captionText="FIGURES 1724. Bactrocera (Bactrocera) paraosbeckiae Drew. 17, head; 18, scutum (dorsal view); 19, thorax (lateral view); 20, abdomen; 21, wing; 22, epandrium and surstyli (lateral view); 23, epandrium and surstyli (posterior view); 24, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209983" httpUri="https://zenodo.org/record/13209983/files/figure.png" pageId="8" pageNumber="291">Figs 22, 23, 24</figureCitation>
,
<figureCitation id="13062A54ED25FF851ED1F924FD78F8ED" box="[728,758,1774,1800]" captionStart="FIGURES4144" captionStartId="12.[151,264,1204,1228]" captionTargetBox="[151,1436,181,1180]" captionTargetId="figure-24@12.[151,1436,181,1180]" captionTargetPageId="12" captionText="FIGURES4144. Comparisonbetween theepandrium(lateral view).41, Bactrocera(Bactrocera) ettinabhujaAbhishek &amp;David, sp. nov.; 42, Bactrocera (Bactrocera) kyrdemkulai Abhishek &amp; David, sp. nov.; 43, Bactrocera (Bactrocera) paraosbeckiae Drew.; 44, Bactrocera (Bactrocera) rubigina (Wang and Zhao)." figureDoi="http://doi.org/10.5281/zenodo.13209993" httpUri="https://zenodo.org/record/13209993/files/figure.png" pageId="8" pageNumber="291">43</figureCitation>
,
<figureCitation id="13062A54ED25FF851F09F924FC90F8ED" box="[768,798,1774,1800]" captionStart="FIGURES 4548" captionStartId="13.[151,264,1761,1785]" captionTargetBox="[151,1436,182,1733]" captionTargetId="figure-17@13.[151,1436,181,1737]" captionTargetPageId="13" captionText="FIGURES 4548. Comparison between the epandrium (posterior view). 45, Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov.; 46, Bactrocera (Bactrocera) kyrdemkulai Abhishek &amp; David, sp. nov.; 47, Bactrocera (Bactrocera) paraosbeckiae Drew.; 48, Bactrocera (Bactrocera) rubigina (Wang and Zhao)." figureDoi="http://doi.org/10.5281/zenodo.13209995" httpUri="https://zenodo.org/record/13209995/files/figure.png" pageId="8" pageNumber="291">47</figureCitation>
,
<figureCitation id="13062A54ED25FF851F20F927FCC6F8ED" box="[809,840,1773,1800]" captionStart="FIGURES 4952" captionStartId="14.[151,264,994,1018]" captionTargetBox="[151,1437,181,970]" captionTargetId="figure-24@14.[151,1437,181,970]" captionTargetPageId="14" captionText="FIGURES 4952. Comparison between the glans of phallus. 49, Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov.; 50, Bactrocera (Bactrocera) kyrdemkulai Abhishek &amp; David, sp. nov.; 51, Bactrocera (Bactrocera) paraosbeckiae Drew; 52, Bactrocera (Bactrocera) rubigina (Wang and Zhao)." figureDoi="http://doi.org/10.5281/zenodo.13209999" httpUri="https://zenodo.org/record/13209999/files/figure.png" pageId="8" pageNumber="291">51</figureCitation>
): Epandrium and surstyli inverted U-shaped in outline (posterior view) epandrium shorter than lateral surstylus, lateral surstylus broad (
<quantity id="4CC59B34ED25FF851804F8D8FBE8F8C9" box="[1037,1126,1810,1836]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="291" unit="mm" value="0.1">0.1 mm</quantity>
), posterior lobe of surstylus, pointed, curved back, not longer than anterior lobe; ventral rim of lateral surstylus as long as the dorsal rim (
<quantity id="4CC59B34ED25FF851971F8FCFF49F891" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="291" unit="mm" value="0.2">0.2 mm</quantity>
). Proctiger membraneous, dome shaped, smaller than epandrium. Medial surstylus, longer than lateral surstylus tapering towards apex with a pair of thick prensisetae. Phallus elongate (1.82.0 mm) excluding glans (
<quantity id="4CC59B34ED25FF851924F8B4FA1BF87D" box="[1325,1429,1918,1944]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="291" unit="mm" value="0.4">0.40 mm</quantity>
) (
<figureCitation id="13062A54ED25FF851C96F868FF7BF859" box="[159,245,1954,1980]" captionStart="FIGURES 1724" captionStartId="8.[151,261,1608,1632]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-24@8.[151,1436,181,1585]" captionTargetPageId="8" captionText="FIGURES 1724. Bactrocera (Bactrocera) paraosbeckiae Drew. 17, head; 18, scutum (dorsal view); 19, thorax (lateral view); 20, abdomen; 21, wing; 22, epandrium and surstyli (lateral view); 23, epandrium and surstyli (posterior view); 24, glans of phallus." figureDoi="http://doi.org/10.5281/zenodo.13209983" httpUri="https://zenodo.org/record/13209983/files/figure.png" pageId="8" pageNumber="291">Figs 24</figureCitation>
,
<figureCitation id="13062A54ED25FF851D08F86BFEAEF859" box="[257,288,1953,1980]" captionStart="FIGURES 4952" captionStartId="14.[151,264,994,1018]" captionTargetBox="[151,1437,181,970]" captionTargetId="figure-24@14.[151,1437,181,970]" captionTargetPageId="14" captionText="FIGURES 4952. Comparison between the glans of phallus. 49, Bactrocera (Bactrocera) ettinabhuja Abhishek &amp; David, sp. nov.; 50, Bactrocera (Bactrocera) kyrdemkulai Abhishek &amp; David, sp. nov.; 51, Bactrocera (Bactrocera) paraosbeckiae Drew; 52, Bactrocera (Bactrocera) rubigina (Wang and Zhao)." figureDoi="http://doi.org/10.5281/zenodo.13209999" httpUri="https://zenodo.org/record/13209999/files/figure.png" pageId="8" pageNumber="291">51</figureCitation>
); 0.75 of glans sclerotised with unpatterned praeputium; subapical lobe and basal lobe present.
</paragraph>
<paragraph id="8B8236D1ED25FF851CCEF80CFE23F805" blockId="8.[151,1437,1737,2016]" box="[199,429,1990,2016]" pageId="8" pageNumber="291">Attractant: Cue lure.</paragraph>
<paragraph id="8B8236D1ED24FF841CCEFF5DFDAEFF54" blockId="9.[199,634,151,213]" box="[199,544,151,177]" pageId="9" pageNumber="292">Host plants: No known record.</paragraph>
<paragraph id="8B8236D1ED24FF841CCEFF71FDF4FF30" blockId="9.[199,634,151,213]" box="[199,634,187,213]" pageId="9" pageNumber="292">
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<typeStatus id="54868873ED24FF841CCEFF71FF71FF30" box="[199,255,187,213]" pageId="9" pageNumber="292">Type</typeStatus>
locality:
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<collectingCountry id="F32A7641ED24FF841DDCFF71FD81FF30" box="[469,527,187,213]" name="India" pageId="9" pageNumber="292">India</collectingCountry>
(
<collectingRegion id="49F9F833ED24FF841E14FF71FDE0FF30" box="[541,622,187,213]" country="India" name="Kerala" pageId="9" pageNumber="292">Kerala</collectingRegion>
)
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.
</paragraph>
</subSubSection>
</treatment>
</document>