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<document id="FE6BE1C84D067C09C44F0FB9377E94BF" ID-DOI="10.37828/em.2023.67.10" ID-ISSN="2336-9744" ID-Zenodo-Dep="13246270" ID-ZooBank="urn:lsid:zoobank.org:pub:41D35BDE-CCED-4E32-A3CC-4808CD36FE1D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722971496884" checkinUser="felipe" docAuthor="Volynkin, Anton V., Titov, Sergey V. &amp; Saldaitis, Aidas" docDate="2023" docId="039487A6565CFFC8C1A0F9C7FC32FBB6" docLanguage="en" docName="EcolMontenegrina.67.83-90.pdf" docOrigin="Ecologica Montenegrina 67" docSource="http://dx.doi.org/10.37828/em.2023.67.10" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Apamea ingloria subsp. postnikovi Volynkin, Titov &amp; Saldaitis, 2023, ssp. n." docType="treatment" docUuid="0BE2A0EE-E8CC-45CB-9A65-AD037BC50948" docUuidSource="ZooBank" docVersion="3" lastPageNumber="89" masterDocId="FFADFFDE565DFFCEC137FFCDFFE0FFBE" masterDocTitle="A new subspecies of Apamea ingloria (A. Bang-Haas, 1912) from northern Tien Shan Mountains (Lepidoptera: Noctuidae: Noctuinae: Xylenini: Apameina)" masterLastPageNumber="90" masterPageNumber="83" pageNumber="84" updateTime="1722979797270" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5E26FD5785325B2CAF941BCB6E209D64">A new subspecies of Apamea ingloria (A. Bang-Haas, 1912) from northern Tien Shan Mountains (Lepidoptera: Noctuidae: Noctuinae: Xylenini: Apameina)</mods:title>
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<mods:namePart id="7F76F83C8160B3D68F615732960FA61D">Volynkin, Anton V.</mods:namePart>
<mods:nameIdentifier id="65BE33D986F88E2872EE2213D09FFCCE" type="ORCID">0000-0001-9447-4925</mods:nameIdentifier>
<mods:affiliation id="DAC264CA6411BB3D5576348D16FA8E23">Altai State University, Lenina Avenue, 61, RF- 656049, Barnaul, Russia. https: // orcid. org / 0000 - 0001 - 9447 - 4925</mods:affiliation>
</mods:name>
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<mods:roleTerm id="1BFDF2BCCAD01B6ACC5BD7F5FFC90413">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7776C8689B2EC3C6DA028A8F43DBD1CD">Titov, Sergey V.</mods:namePart>
<mods:affiliation id="A302E8B2F53F271456EB99B677E3F6A1">Institute of Zoology, Entomology laboratory, Al-Farabi Avenue, 93, KZ- 050060, Almaty, Kazakhstan. E-mail: sandibicumar @ gmail. com; https: // orcid. org / 0000 - 0003 - 2196 - 280 X &amp; Scientific Center &quot; Bioarchaeology &quot;, Institute of Archaeological Research, Margulan University, Olzhabay Batyr str. 60, KZ- 140002, Pavlodar, Kazakhstan &amp; Department of Biology and Ecology; the Research Centre for Environmental &quot; Monitoring &quot;, Toraighyrov University, Lomova str. 64, KZ- 140008, Pavlodar, Kazakhstan</mods:affiliation>
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<mods:namePart id="6D7DA133AAAD6CCD257C913BA7D7FF03">Saldaitis, Aidas</mods:namePart>
<mods:nameIdentifier id="0FABA3D66E16F62E83CEC5456FADC1A6" type="ORCID">0000-0003-0999-3996</mods:nameIdentifier>
<mods:affiliation id="3D646B198D2287C510F8E2851D53657B">Nature Research Centre, Akademijos str., 2, LT- 08412, Vilnius- 21, Lithuania. E-mail: saldrasa @ gmail. com; https: // orcid. org / 0000 - 0003 - 0999 - 3996 * Corresponding author. E-mail: monstruncusarctia @ gmail. com</mods:affiliation>
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<mods:title id="2CEA17EE4F9C6B411F01B4A3811D0F0F">Ecologica Montenegrina</mods:title>
</mods:titleInfo>
<mods:part id="7271E4B087BD1FD50C69CB9591B3BB68">
<mods:date id="76A6A49878BFCA8DA631858DDC572B74">2023</mods:date>
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<mods:number id="132AB89F740029026CD64221E6AD8EBE">2023-10-17</mods:number>
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<mods:number id="9EA03E320CB25ECFC4F921A10BA969E2">67</mods:number>
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<treatment id="039487A6565CFFC8C1A0F9C7FC32FBB6" ID-DOI="http://doi.org/10.5281/zenodo.13246958" ID-Zenodo-Dep="13246958" LSID="urn:lsid:zoobank.org:act:0BE2A0EE-E8CC-45CB-9A65-AD037BC50948" httpUri="http://treatment.plazi.org/id/039487A6565CFFC8C1A0F9C7FC32FBB6" lastPageId="6" lastPageNumber="89" pageId="1" pageNumber="84">
<subSubSection id="C327653B565CFFCFC1A0F9C7FB69F9FC" pageId="1" pageNumber="84" type="nomenclature">
<paragraph id="8B8236B0565CFFCFC1A0F9C7FDDAF99A" blockId="1.[151,1161,1546,1636]" box="[151,570,1546,1572]" pageId="1" pageNumber="84">
<heading id="D0CA81DC565CFFCFC1A0F9C7FDDAF99A" box="[151,570,1546,1572]" fontSize="11" level="2" pageId="1" pageNumber="84" reason="2">
<emphasis id="B949EAA2565CFFCFC1A0F9C7FDDAF99A" bold="true" box="[151,570,1546,1572]" pageId="1" pageNumber="84">
<taxonomicName id="4C3D4D33565CFFCFC1A0F9C7FE06F99A" authorityName="Volynkin &amp; Titov &amp; Saldaitis" authorityYear="2023" box="[151,486,1546,1572]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="subSpecies" species="ingloria" status="subsp. nov." subSpecies="postnikovi">
<emphasis id="B949EAA2565CFFCFC1A0F9C7FE06F99A" bold="true" box="[151,486,1546,1572]" italics="true" pageId="1" pageNumber="84">Apamea ingloria postnikovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57D9565CFFCFC0DAF9C7FDDAF99A" box="[493,570,1546,1572]" pageId="1" pageNumber="84" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B0565CFFCFC1A0F9E6FB69F9FC" blockId="1.[151,1161,1546,1636]" box="[151,1161,1579,1602]" pageId="1" pageNumber="84">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="C327653B565CFFCFC1A0F987FE43F9DA" box="[151,419,1610,1636]" pageId="1" pageNumber="84" type="description">
<paragraph id="8B8236B0565CFFCFC1A0F987FE43F9DA" blockId="1.[151,1161,1546,1636]" box="[151,419,1610,1636]" pageId="1" pageNumber="84">
(
<figureCitation id="13062A35565CFFCFC1A8F987FEE3F9DA" box="[159,259,1610,1636]" captionStart="Figures 1–8" captionStartId="2.[151,237,1980,2003]" captionTargetBox="[207,1373,227,1952]" captionTargetId="figure-10@2.[151,1436,223,1962]" captionTargetPageId="2" captionText="Figures 1–8. Apamea spp.: adults. Depositories of the specimens: 1 in WIGJ; 2–8 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246272" httpUri="https://zenodo.org/record/13246272/files/figure.png" pageId="1" pageNumber="84">Figs 1-3</figureCitation>
,
<figureCitation id="13062A35565CFFCFC027F987FEA9F9DA" box="[272,329,1610,1636]" captionStart="Figures 9–12" captionStartId="3.[151,237,1980,2003]" captionTargetBox="[182,1436,249,1943]" captionTargetId="figure-11@3.[151,1436,223,1962]" captionTargetPageId="3" captionText="Figures 9–12. Apamea ingloria sspp.: male genitalia. Depositories of the specimens dissected: 9 in WIGJ; 10–12 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246274" httpUri="https://zenodo.org/record/13246274/files/figure.png" pageId="1" pageNumber="84">9-10</figureCitation>
,
<figureCitation id="13062A35565CFFCFC062F987FE7AF9DA" box="[341,410,1610,1636]" captionStart="Figures 13–16" captionStartId="4.[151,237,1977,2000]" captionTargetBox="[151,1436,220,1940]" captionTargetId="figure-10@4.[151,1436,220,1959]" captionTargetPageId="4" captionText="Figures 13–16. Apamea ingloria sspp.: sternite A1+2. Depositories of the specimens dissected: 13 in WIGJ; 14–16 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246278" httpUri="https://zenodo.org/record/13246278/files/figure.png" pageId="1" pageNumber="84">13-14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C327653B565CFFCFC1A0F940FC23F8B2" pageId="1" pageNumber="84" type="materials_examined">
<paragraph id="8B8236B0565CFFCFC1A0F940FACBF955" blockId="1.[151,1437,1677,1804]" pageId="1" pageNumber="84">
<emphasis id="B949EAA2565CFFCFC1A0F940FEADF919" bold="true" box="[151,333,1677,1703]" pageId="1" pageNumber="84">Type material</emphasis>
.
<materialsCitation id="3B553CED565CFFCFC055F940FACBF955" collectingDate="2022-07-21" collectingDateMax="2022-07-22" collectingDateMin="2022-07-21" collectionCode="V, WIGJ" collectorName="A. Volynkin" country="Kazakhstan" elevation="3050" latitude="43.327587" location="Figs" longLatPrecision="1" longitude="80.31928" pageId="1" pageNumber="84" specimenCount="1" specimenCount-male="1" stateProvince="Almaty Oblysy" typeStatus="holotype">
<emphasis id="B949EAA2565CFFCFC055F940FE34F919" bold="true" box="[354,468,1677,1703]" pageId="1" pageNumber="84">
<typeStatus id="54868812565CFFCFC055F940FE34F919" box="[354,468,1677,1703]" pageId="1" pageNumber="84" type="holotype">Holotype</typeStatus>
</emphasis>
(
<location id="8EE2606B565CFFCFC0DEF940FDC2F919" LSID="urn:lsid:plazi:treatment:039487A6565CFFC8C1A0F9C7FC32FBB6:8EE2606B565CFFCFC0DEF940FDC2F919" box="[489,546,1677,1703]" country="Kazakhstan" latitude="43.327587" longLatPrecision="1" longitude="80.31928" name="Figs" pageId="1" pageNumber="84" stateProvince="Almaty Oblysy">Figs</location>
): male, “
<date id="FF831070565CFFCFC3A3F940FCB3F919" box="[660,851,1677,1703]" pageId="1" pageNumber="84" value="2022-07-21" valueMax="2022-07-22" valueMin="2022-07-21">
<collectingDate id="EFC7E998565CFFCFC3A3F940FCB3F919" box="[660,851,1677,1703]" pageId="1" pageNumber="84" value="2022-07-21" valueMax="2022-07-22" valueMin="2022-07-21">21–22.VII.2022</collectingDate>
</date>
, | SE
<collectingCountry id="F32A7620565CFFCFC291F940FBD2F919" box="[934,1074,1677,1703]" name="Kazakhstan" pageId="1" pageNumber="84">Kazakhstan</collectingCountry>
,
<collectingRegion id="49F9F852565CFFCFC573F940FAE0F919" box="[1092,1280,1677,1703]" country="Kazakhstan" name="Almaty Oblysy" pageId="1" pageNumber="84">Almaty Region</collectingRegion>
, | Raiymbek District, E Ketmen | Ridge, Ketmen Pass, |
<geoCoordinate id="EE095077565CFFCFC39FF962FCB7F977" box="[680,855,1711,1737]" degrees="43" direction="north" minutes="19" orientation="latitude" pageId="1" pageNumber="84" precision="1" seconds="39.32" value="43.327587">43°19'39.32&quot;N</geoCoordinate>
<geoCoordinate id="EE095077565CFFCFC256F962FBE0F977" box="[865,1024,1711,1737]" degrees="80" direction="east" minutes="19" orientation="longitude" pageId="1" pageNumber="84" precision="1" seconds="9.41" value="80.31928">80°19'9.41&quot;E</geoCoordinate>
, |
<quantity id="4CC59B55565CFFCFC528F962FB5CF977" box="[1055,1212,1711,1737]" metricMagnitude="3" metricUnit="m" metricValue="3.05" metricValueMax="3.1" metricValueMin="3.0" pageId="1" pageNumber="84" unit="m" value="3050.0" valueMax="3100.0" valueMin="3000.0">
<elevation id="0010D183565CFFCFC528F962FB5CF977" box="[1055,1212,1711,1737]" metricMagnitude="3" metricUnit="m" metricValue="3.05" metricValueMax="3.1" metricValueMin="3.0" pageId="1" pageNumber="84" unit="m" value="3050.0" valueMax="3100.0" valueMin="3000.0">3000–3100m</elevation>
</quantity>
, alpine meadows | with rock outcrops, | Volynkin A.
<collectionCode id="ED2CAE75565CFFCFC315F91CFDDAF955" box="[546,570,1745,1771]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="84" type="Museum">V</collectionCode>
. &amp; Titov S.
<collectionCode id="ED2CAE75565CFFCFC3F7F91CFD3AF955" box="[704,730,1745,1771]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="84" type="Museum">V</collectionCode>
. leg.” / “Slide | AV7424
<specimenCount id="9D3BFD39565CFFCFC2CCF91CFBEEF955" box="[1019,1038,1745,1771]" count="1" pageId="1" pageNumber="84" type="male">♂</specimenCount>
|
<collectorName id="26C85366565CFFCFC514F91CFB58F955" box="[1059,1208,1745,1771]" pageId="1" pageNumber="84">A. Volynkin</collectorName>
” (
<collectionCode id="ED2CAE75565CFFCFC5E5F91CFAFFF955" box="[1234,1311,1745,1771]" pageId="1" pageNumber="84">WIGJ</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B8236B0565CFFCFC1C2F93FFC23F8B2" blockId="1.[151,1437,1677,1804]" box="[245,963,1778,1804]" pageId="1" pageNumber="84">
<materialsCitation id="3B553CED565CFFCFC1C2F93FFC5EF8B2" box="[245,958,1778,1804]" collectionCode="CAV" pageId="1" pageNumber="84" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="B949EAA2565CFFCFC1C2F93FFE93F8B2" bold="true" box="[245,371,1778,1804]" pageId="1" pageNumber="84">
<typeStatus id="54868812565CFFCFC1C2F93FFE93F8B2" box="[245,371,1778,1804]" pageId="1" pageNumber="84" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D3BFD39565CFFCFC0B4F93FFE00F8B2" box="[387,480,1778,1804]" count="2" pageId="1" pageNumber="84" type="male">2 males</specimenCount>
, the same data as in the
<typeStatus id="54868812565CFFCFC3CBF93FFC82F8B2" box="[764,866,1778,1804]" pageId="1" pageNumber="84" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED2CAE75565CFFCFC245F93FFC56F8B2" box="[882,950,1778,1804]" pageId="1" pageNumber="84">CAV</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C327653B565CFFCBC1A0F8FBFF3AF9DC" lastPageId="5" lastPageNumber="88" pageId="1" pageNumber="84" type="diagnosis">
<paragraph id="8B8236B0565CFFCBC1A0F8FBFF3AF9DC" blockId="1.[151,1437,1846,2040]" lastBlockId="5.[151,1437,1237,1634]" lastPageId="5" lastPageNumber="88" pageId="1" pageNumber="84">
<emphasis id="B949EAA2565CFFCFC1A0F8FBFEF0F8EE" bold="true" box="[151,272,1846,1872]" pageId="1" pageNumber="84">Diagnosis</emphasis>
.
<taxonomicName id="4C3D4D33565CFFCFC028F8FBFD87F8EE" authorityName="Volynkin &amp; Titov &amp; Saldaitis" authorityYear="2023" box="[287,615,1846,1872]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="subSpecies" species="ingloria" status="subsp. nov." subSpecies="postnikovi">
<emphasis id="B949EAA2565CFFCFC028F8FBFD87F8EE" box="[287,615,1846,1872]" italics="true" pageId="1" pageNumber="84">Apamea ingloria postnikovi</emphasis>
</taxonomicName>
<emphasis id="B949EAA2565CFFCFC358F8FBFD5DF8EE" bold="true" box="[623,701,1846,1872]" pageId="1" pageNumber="84">
<taxonomicNameLabel id="A27A57D9565CFFCFC358F8FBFD5DF8EE" box="[623,701,1846,1872]" pageId="1" pageNumber="84" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13062A35565CFFCFC3FAF8FBFCD6F8EE" box="[717,822,1846,1872]" captionStart="Figures 1–8" captionStartId="2.[151,237,1980,2003]" captionTargetBox="[207,1373,227,1952]" captionTargetId="figure-10@2.[151,1436,223,1962]" captionTargetPageId="2" captionText="Figures 1–8. Apamea spp.: adults. Depositories of the specimens: 1 in WIGJ; 2–8 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246272" httpUri="https://zenodo.org/record/13246272/files/figure.png" pageId="1" pageNumber="84">Figs 1–3</figureCitation>
) differs from the nominate subspecies (
<figureCitation id="13062A35565CFFCFC43BF8FBFA95F8EE" box="[1292,1397,1846,1872]" captionStart="Figures 1–8" captionStartId="2.[151,237,1980,2003]" captionTargetBox="[207,1373,227,1952]" captionTargetId="figure-10@2.[151,1436,223,1962]" captionTargetPageId="2" captionText="Figures 1–8. Apamea spp.: adults. Depositories of the specimens: 1 in WIGJ; 2–8 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246272" httpUri="https://zenodo.org/record/13246272/files/figure.png" pageId="1" pageNumber="84">Figs 4–6</figureCitation>
) in the darker, brownish-grey body and forewing ground colour lacking the bronze suffusion which is typical of
<emphasis id="B949EAA2565CFFCFC1A0F8B4FED9F82D" box="[151,313,1913,1939]" italics="true" pageId="1" pageNumber="84">
A. i.
<taxonomicName id="4C3D4D33565CFFCFC1EDF8B4FED9F82D" baseAuthorityName="A. Bang-Haas" baseAuthorityYear="1912" box="[218,313,1913,1939]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="ingloria">ingloria</taxonomicName>
</emphasis>
, the more diffuse forewing pattern, and the considerably darker, brownish-grey hindwing. Compared to
<taxonomicName id="4C3D4D33565CFFCFC00EF856FE53F80B" authorityName="Varga" authorityYear="1982" box="[313,435,1947,1973]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="B949EAA2565CFFCFC00EF856FE53F80B" box="[313,435,1947,1973]" italics="true" pageId="1" pageNumber="84">A. kaszabi</emphasis>
</taxonomicName>
(
<figureCitation id="13062A35565CFFCFC0F3F856FDCFF80B" box="[452,559,1947,1973]" captionStart="Figures 1–8" captionStartId="2.[151,237,1980,2003]" captionTargetBox="[207,1373,227,1952]" captionTargetId="figure-10@2.[151,1436,223,1962]" captionTargetPageId="2" captionText="Figures 1–8. Apamea spp.: adults. Depositories of the specimens: 1 in WIGJ; 2–8 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246272" httpUri="https://zenodo.org/record/13246272/files/figure.png" pageId="1" pageNumber="84">Figs 7, 8</figureCitation>
), the new subspecies has a somewhat shorter and broader forewing, more contrast dark forewing markings, a broader whitish edge of the reniform stigma, and a markedly darker hindwing. The male trifine brush organ of the new subspecies (
<figureCitation id="13062A35565CFFCFC24BF813FBE1F846" box="[892,1025,2014,2040]" captionStart="Figures 13–16" captionStartId="4.[151,237,1977,2000]" captionTargetBox="[151,1436,220,1940]" captionTargetId="figure-10@4.[151,1436,220,1959]" captionTargetPageId="4" captionText="Figures 13–16. Apamea ingloria sspp.: sternite A1+2. Depositories of the specimens dissected: 13 in WIGJ; 14–16 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246278" httpUri="https://zenodo.org/record/13246278/files/figure.png" pageId="1" pageNumber="84">Figs 13, 14</figureCitation>
) has a longer, distally broader and apically rounded flat sclerotisation corresponding to a remnant of a lever whereas it is apically pointed in the nominate subspecies (
<figureCitation id="13062A355658FFCBC0ACFB3AFDC6FAAF" box="[411,550,1271,1297]" captionStart="Figures 13–16" captionStartId="4.[151,237,1977,2000]" captionTargetBox="[151,1436,220,1940]" captionTargetId="figure-10@4.[151,1436,220,1959]" captionTargetPageId="4" captionText="Figures 13–16. Apamea ingloria sspp.: sternite A1+2. Depositories of the specimens dissected: 13 in WIGJ; 14–16 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246278" httpUri="https://zenodo.org/record/13246278/files/figure.png" pageId="5" pageNumber="88">Figs 15, 16</figureCitation>
). Unlike
<emphasis id="B949EAA25658FFCBC3A2FB3AFCA8FAAF" box="[661,840,1271,1297]" italics="true" pageId="5" pageNumber="88">A. i. postnikovi</emphasis>
<emphasis id="B949EAA25658FFCBC267FB3AFC7FFAAF" bold="true" box="[848,927,1271,1297]" pageId="5" pageNumber="88">ssp. n.</emphasis>
, the trifine brush organ of
<taxonomicName id="4C3D4D335658FFCBC5D5FB3AFABBFAAF" authorityName="Varga" authorityYear="1982" box="[1250,1371,1271,1297]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="B949EAA25658FFCBC5D5FB3AFABBFAAF" box="[1250,1371,1271,1297]" italics="true" pageId="5" pageNumber="88">A. kaszabi</emphasis>
</taxonomicName>
(
<figureCitation id="13062A355658FFCBC45BFB3AFF55FA8D" captionStart="Figures 17–18" captionStartId="5.[151,237,1074,1097]" captionTargetBox="[177,1353,189,1043]" captionTargetId="figure-11@5.[151,1436,189,1056]" captionTargetPageId="5" captionText="Figures 17–18. Apamea kaszabi: male genitalia (17) and sternite A1+2 (18). The specimen dissected is deposited in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246282" httpUri="https://zenodo.org/record/13246282/files/figure.png" pageId="5" pageNumber="88">Fig. 18</figureCitation>
) is reduced to a basal plate. Additionally, the sternum A1+2 has the antero-lateral (outer) apodemes in both the subspecies of
<emphasis id="B949EAA25658FFCBC093FAF6FDDBFAEB" box="[420,571,1339,1365]" italics="true" pageId="5" pageNumber="88">
A. i.
<taxonomicName id="4C3D4D335658FFCBC0EBFAF6FDDBFAEB" baseAuthorityName="A. Bang-Haas" baseAuthorityYear="1912" box="[476,571,1339,1365]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="ingloria">ingloria</taxonomicName>
</emphasis>
(
<figureCitation id="13062A355658FFCBC37CFAF6FD31FAEB" box="[587,721,1339,1365]" captionStart="Figures 13–16" captionStartId="4.[151,237,1977,2000]" captionTargetBox="[151,1436,220,1940]" captionTargetId="figure-10@4.[151,1436,220,1959]" captionTargetPageId="4" captionText="Figures 13–16. Apamea ingloria sspp.: sternite A1+2. Depositories of the specimens dissected: 13 in WIGJ; 14–16 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246278" httpUri="https://zenodo.org/record/13246278/files/figure.png" pageId="5" pageNumber="88">Figs 13–16</figureCitation>
) whereas they are absent in
<taxonomicName id="4C3D4D335658FFCBC516FAF6FB7AFAEB" authorityName="Varga" authorityYear="1982" box="[1057,1178,1339,1365]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="B949EAA25658FFCBC516FAF6FB7AFAEB" box="[1057,1178,1339,1365]" italics="true" pageId="5" pageNumber="88">A. kaszabi</emphasis>
</taxonomicName>
(
<figureCitation id="13062A355658FFCBC59DFAF6FAE1FAEB" box="[1194,1281,1339,1365]" captionStart="Figures 17–18" captionStartId="5.[151,237,1074,1097]" captionTargetBox="[177,1353,189,1043]" captionTargetId="figure-11@5.[151,1436,189,1056]" captionTargetPageId="5" captionText="Figures 17–18. Apamea kaszabi: male genitalia (17) and sternite A1+2 (18). The specimen dissected is deposited in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246282" httpUri="https://zenodo.org/record/13246282/files/figure.png" pageId="5" pageNumber="88">Fig. 18</figureCitation>
). In the male genitalia,
<emphasis id="B949EAA25658FFCBC038FA90FE28FAC9" box="[271,456,1373,1399]" italics="true" pageId="5" pageNumber="88">A. i. postnikovi</emphasis>
<emphasis id="B949EAA25658FFCBC0E3FA90FDC6FAC9" bold="true" box="[468,550,1373,1399]" pageId="5" pageNumber="88">ssp. n.</emphasis>
(
<figureCitation id="13062A355658FFCBC30DFA90FD5AFAC9" box="[570,698,1373,1399]" captionStart="Figures 9–12" captionStartId="3.[151,237,1980,2003]" captionTargetBox="[182,1436,249,1943]" captionTargetId="figure-11@3.[151,1436,223,1962]" captionTargetPageId="3" captionText="Figures 9–12. Apamea ingloria sspp.: male genitalia. Depositories of the specimens dissected: 9 in WIGJ; 10–12 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246274" httpUri="https://zenodo.org/record/13246274/files/figure.png" pageId="5" pageNumber="88">Figs 9, 10</figureCitation>
) differs from
<emphasis id="B949EAA25658FFCBC25BFA90FBECFAC9" box="[876,1036,1373,1399]" italics="true" pageId="5" pageNumber="88">
A. i.
<taxonomicName id="4C3D4D335658FFCBC29AFA90FBECFAC9" baseAuthorityName="A. Bang-Haas" baseAuthorityYear="1912" box="[941,1036,1373,1399]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="ingloria">ingloria</taxonomicName>
</emphasis>
(
<figureCitation id="13062A355658FFCBC528FA90FB4DFAC9" box="[1055,1197,1373,1399]" captionStart="Figures 9–12" captionStartId="3.[151,237,1980,2003]" captionTargetBox="[182,1436,249,1943]" captionTargetId="figure-11@3.[151,1436,223,1962]" captionTargetPageId="3" captionText="Figures 9–12. Apamea ingloria sspp.: male genitalia. Depositories of the specimens dissected: 9 in WIGJ; 10–12 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246274" httpUri="https://zenodo.org/record/13246274/files/figure.png" pageId="5" pageNumber="88">Figs 11, 12</figureCitation>
) in the shorter and narrower setose field of the cucullus, the shorter ampulla, the somewhat broader clavus, and the slightly longer base of the distal cornutus in the phallus vesica. Compared to
<taxonomicName id="4C3D4D335658FFCBC2F3FA6DFBDDFA04" authorityName="Varga" authorityYear="1982" box="[964,1085,1440,1466]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="B949EAA25658FFCBC2F3FA6DFBDDFA04" box="[964,1085,1440,1466]" italics="true" pageId="5" pageNumber="88">A. kaszabi</emphasis>
</taxonomicName>
(
<figureCitation id="13062A355658FFCBC579FA6DFB44FA04" box="[1102,1188,1440,1466]" captionStart="Figures 17–18" captionStartId="5.[151,237,1074,1097]" captionTargetBox="[177,1353,189,1043]" captionTargetId="figure-11@5.[151,1436,189,1056]" captionTargetPageId="5" captionText="Figures 17–18. Apamea kaszabi: male genitalia (17) and sternite A1+2 (18). The specimen dissected is deposited in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246282" httpUri="https://zenodo.org/record/13246282/files/figure.png" pageId="5" pageNumber="88">Fig. 17</figureCitation>
), the new subspecies has a slightly broader cucullus with a shorter setose field, a broader clavus, and a somewhat longer base of the distal cornutus. The male genitalia of the externally vaguely similar
<taxonomicName id="4C3D4D335658FFCBC2C0FA2EFB8CFA43" box="[1015,1132,1507,1533]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="fervida">
<emphasis id="B949EAA25658FFCBC2C0FA2EFB8CFA43" box="[1015,1132,1507,1533]" italics="true" pageId="5" pageNumber="88">A. fervida</emphasis>
</taxonomicName>
(illustrated by
<bibRefCitation id="EFAC4B415658FFCBC412FA2EFECCF9A1" author="Zilli, A. &amp; Varga, Z. &amp; Ronkay, G. &amp; Ronkay, L." pageId="5" pageNumber="88" refId="ref2802" refString="Zilli, A., Varga, Z. Ronkay, G. &amp; Ronkay, L. (2009) A Taxonomic Atlas of the Eurasian and North African Noctoidea. Volume 3. Apameini 1. Heterocera Press, Budapest, 393 pp." type="book" year="2009">
Zilli
<emphasis id="B949EAA25658FFCBC457FA2EFA7CFA43" box="[1376,1436,1507,1533]" italics="true" pageId="5" pageNumber="88">et al.</emphasis>
(2009: 283))
</bibRefCitation>
from
<collectingRegion id="49F9F8525658FFCBC042F9C8FE34F9A1" box="[373,468,1541,1567]" country="China" name="Qinghai" pageId="5" pageNumber="88">Qinghai</collectingRegion>
differ from
<emphasis id="B949EAA25658FFCBC350F9C8FCFAF9A1" box="[615,794,1541,1567]" italics="true" pageId="5" pageNumber="88">A. i. postnikovi</emphasis>
<emphasis id="B949EAA25658FFCBC215F9C8FC90F9A1" bold="true" box="[802,880,1541,1567]" pageId="5" pageNumber="88">ssp. n.</emphasis>
in the shorter cucullus with a markedly longer setose field stretching towards the pollex, the broader digitus, and the longer latero-posterior processes of the juxta.
</paragraph>
</subSubSection>
<caption id="DF426638565FFFCCC1A0F871FB84F86D" ID-DOI="http://doi.org/10.5281/zenodo.13246272" ID-Zenodo-Dep="13246272" box="[151,1124,1980,2003]" httpUri="https://zenodo.org/record/13246272/files/figure.png" pageId="2" pageNumber="85" startId="2.[151,237,1980,2003]" targetBox="[207,1373,227,1952]" targetPageId="2" targetType="figure">
<paragraph id="8B8236B0565FFFCCC1A0F871FB84F86D" blockId="2.[151,1124,1980,2003]" box="[151,1124,1980,2003]" pageId="2" pageNumber="85">
<emphasis id="B949EAA2565FFFCCC1A0F871FEFBF86D" bold="true" box="[151,283,1980,2003]" pageId="2" pageNumber="85">Figures 1–8</emphasis>
.
<taxonomicName id="4C3D4D33565FFFCCC01EF871FE57F86D" box="[297,439,1980,2003]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B949EAA2565FFFCCC01EF871FE60F86D" box="[297,384,1980,2003]" italics="true" pageId="2" pageNumber="85">Apamea</emphasis>
spp.
</taxonomicName>
: adults. Depositories of the specimens: 1 in WIGJ; 2–8 in CAV.
</paragraph>
</caption>
<caption id="DF426638565EFFCDC1A0F871FF37F84C" ID-DOI="http://doi.org/10.5281/zenodo.13246274" ID-Zenodo-Dep="13246274" httpUri="https://zenodo.org/record/13246274/files/figure.png" pageId="3" pageNumber="86" startId="3.[151,237,1980,2003]" targetBox="[182,1436,249,1943]" targetPageId="3" targetType="figure">
<paragraph id="8B8236B0565EFFCDC1A0F871FF37F84C" blockId="3.[151,1436,1980,2034]" pageId="3" pageNumber="86">
<emphasis id="B949EAA2565EFFCDC1A0F871FECCF86D" bold="true" box="[151,300,1980,2003]" pageId="3" pageNumber="86">Figures 9–12</emphasis>
.
<taxonomicName id="4C3D4D33565EFFCDC00BF871FE13F86D" baseAuthorityName="A. Bang-Haas" baseAuthorityYear="1912" box="[316,499,1980,2003]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="ingloria">
<emphasis id="B949EAA2565EFFCDC00BF871FE13F86D" box="[316,499,1980,2003]" italics="true" pageId="3" pageNumber="86">Apamea ingloria</emphasis>
</taxonomicName>
sspp.: male genitalia. Depositories of the specimens dissected: 9 in WIGJ; 10–12 in CAV.
</paragraph>
</caption>
<caption id="DF4266385659FFCAC1A0F874FF37F851" ID-DOI="http://doi.org/10.5281/zenodo.13246278" ID-Zenodo-Dep="13246278" httpUri="https://zenodo.org/record/13246278/files/figure.png" pageId="4" pageNumber="87" startId="4.[151,237,1977,2000]" targetBox="[151,1436,220,1940]" targetPageId="4" targetType="figure">
<paragraph id="8B8236B05659FFCAC1A0F874FF37F851" blockId="4.[151,1436,1977,2031]" pageId="4" pageNumber="87">
<emphasis id="B949EAA25659FFCAC1A0F874FED8F86E" bold="true" box="[151,312,1977,2000]" pageId="4" pageNumber="87">Figures 13–16</emphasis>
.
<taxonomicName id="4C3D4D335659FFCAC070F874FE1CF86E" baseAuthorityName="A. Bang-Haas" baseAuthorityYear="1912" box="[327,508,1977,2000]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="ingloria">
<emphasis id="B949EAA25659FFCAC070F874FE1CF86E" box="[327,508,1977,2000]" italics="true" pageId="4" pageNumber="87">Apamea ingloria</emphasis>
</taxonomicName>
sspp.: sternite A1+2. Depositories of the specimens dissected: 13 in WIGJ; 14–16 in CAV.
</paragraph>
</caption>
<caption id="DF4266385658FFCBC1A0FBFFFF37FBD6" ID-DOI="http://doi.org/10.5281/zenodo.13246282" ID-Zenodo-Dep="13246282" httpUri="https://zenodo.org/record/13246282/files/figure.png" pageId="5" pageNumber="88" startId="5.[151,237,1074,1097]" targetBox="[177,1353,189,1043]" targetPageId="5" targetType="figure">
<paragraph id="8B8236B05658FFCBC1A0FBFFFF37FBD6" blockId="5.[151,1436,1074,1128]" pageId="5" pageNumber="88">
<emphasis id="B949EAA25658FFCBC1A0FBFFFED9FBF7" bold="true" box="[151,313,1074,1097]" pageId="5" pageNumber="88">Figures 17–18</emphasis>
.
<taxonomicName id="4C3D4D335658FFCBC07DFBFFFE1BFBF7" authorityName="Varga" authorityYear="1982" box="[330,507,1074,1097]" class="Insecta" family="Noctuidae" genus="Apamea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="B949EAA25658FFCBC07DFBFFFE1BFBF7" box="[330,507,1074,1097]" italics="true" pageId="5" pageNumber="88">Apamea kaszabi</emphasis>
</taxonomicName>
: male genitalia (17) and sternite A1+2 (18). The specimen dissected is deposited in CAV.
</paragraph>
</caption>
<subSubSection id="C327653B5658FFC8C1A0F941FE8AFD44" lastPageId="6" lastPageNumber="89" pageId="5" pageNumber="88" type="description">
<paragraph id="8B8236B05658FFC8C1A0F941FC64FD09" blockId="5.[151,1437,1676,2039]" lastBlockId="6.[151,1437,196,695]" lastPageId="6" lastPageNumber="89" pageId="5" pageNumber="88">
<emphasis id="B949EAA25658FFCBC1A0F941FEC8F918" bold="true" box="[151,296,1676,1702]" pageId="5" pageNumber="88">Description</emphasis>
.
<emphasis id="B949EAA25658FFCBC00BF941FCE7F918" bold="true" box="[316,775,1676,1702]" pageId="5" pageNumber="88">External morphology of male adult</emphasis>
(
<figureCitation id="13062A355658FFCBC22CF941FC6BF918" box="[795,907,1676,1702]" captionStart="Figures 1–8" captionStartId="2.[151,237,1980,2003]" captionTargetBox="[207,1373,227,1952]" captionTargetId="figure-10@2.[151,1436,223,1962]" captionTargetPageId="2" captionText="Figures 1–8. Apamea spp.: adults. Depositories of the specimens: 1 in WIGJ; 2–8 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246272" httpUri="https://zenodo.org/record/13246272/files/figure.png" pageId="5" pageNumber="88">Figs 1–3</figureCitation>
). Forewing length 19.5–20.0 mm. Antenna filiform. Head and thorax brownish-grey with admixture of pale grey scales. Forewing ground colour brownish-grey, pattern slate black, diffuse. Basal dash short, with two short branches. Antemedial line sinuous, sloping outwards from costal to anal margins. Medial area intensely suffused with slate black in cell between orbicular and reniform stigmata, and between claviform stigma and postmedial line. Claviform stigma large, reaching middle of medial area. Orbicular stigma elliptical, oblique, pale grey with grey core suffusion. Reniform stigma large, constricted medially, pale grey with grey core suffusion. Postmedial line dentate on veins, curved inwards between vein R and costal margin, its outer margin edged with pale grey suffusion. Subterminal line indistinct, pale grey, irregularly wavy, edged with row of triangular diffuse slate black spots inwardly. Terminal line thin, interrupted on veins. Cilia brownish-grey with admixture of slate black scales. Hindwing dark brownish-grey, paler subbasally. Cilia as ground colour. Abdomen pale brownish-grey.
<emphasis id="B949EAA2565BFFC8C064FF09FDE8FF60" bold="true" box="[339,520,196,222]" pageId="6" pageNumber="89">Male genitalia</emphasis>
(
<figureCitation id="13062A35565BFFC8C32EFF09FD77FF60" box="[537,663,196,222]" captionStart="Figures 9–12" captionStartId="3.[151,237,1980,2003]" captionTargetBox="[182,1436,249,1943]" captionTargetId="figure-11@3.[151,1436,223,1962]" captionTargetPageId="3" captionText="Figures 9–12. Apamea ingloria sspp.: male genitalia. Depositories of the specimens dissected: 9 in WIGJ; 10–12 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13246274" httpUri="https://zenodo.org/record/13246274/files/figure.png" pageId="6" pageNumber="89">Figs 9, 10</figureCitation>
). Vinculum U-shaped with anteriorly dilated arms forming short saccus. Valva elongate and relatively narrow, somewhat upcurved subbasally and almost straight medially. Sacculus short with broad, semiglobular and serrulate clavus. Clasper narrow, stretching parallel to ventral margin of valva. Ampulla short and thin, stick-shaped, slightly downcurved. Costa with distal-inner trapezoidal protrusion bearing dense field of setae, and broad, laterally flattened and apically rounded digitus reaching outer margin of cucullus. Cucullus large, elongate dorsally, with broadly triangular and apically rounded pollex (ventral lobe), bearing corona stretching along its entire outer margin, and short and narrow setose field medially-basally. Transtilla long, band-shaped. Juxta shield-like with two short triangular latero-posterior processes, and one short medial anterior process. Phallus cylindrical, somewhat dilated and downcurved distally, with short and rounded coecum. Vesica with two short spike-like cornuti on broad plate-like bases subbasally and medially, broadly conical and apically rounded postero-ventral diverticulum, and upcurved and distally tapering distal section with short semiglobular apical diverticulum. Vesica ejaculatorius directed dorsally. Tegumen equal in length to vinculum, with broad trapezoidal peniculus. Tuba analis narrow, weakly sclerotised. Uncus elongate, slender, hook-like downcurved, apically pointed, distal half of its dorsal margin bearing dense cluster of hair-like setae.
</paragraph>
<paragraph id="8B8236B0565BFFC8C1A0FD2DFE8AFD44" blockId="6.[151,362,736,762]" box="[151,362,736,762]" pageId="6" pageNumber="89">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C327653B565BFFC8C1A0FCEEFCCDFC1C" pageId="6" pageNumber="89" type="distribution">
<paragraph id="8B8236B0565BFFC8C1A0FCEEFCCDFC1C" blockId="6.[151,1437,803,930]" pageId="6" pageNumber="89">
<emphasis id="B949EAA2565BFFC8C1A0FCEEFE18FC83" bold="true" box="[151,504,803,829]" pageId="6" pageNumber="89">Distribution and bionomics</emphasis>
. The new subspecies is currently known only from its
<typeStatus id="54868812565BFFC8C586FCEEFB04FC83" box="[1201,1252,803,829]" pageId="6" pageNumber="89">type</typeStatus>
locality in the central part of the Ketmen Ridge (Southeast
<collectingCountry id="F32A7620565BFFC8C38AFC88FCA9FCE1" box="[701,841,837,863]" name="Kazakhstan" pageId="6" pageNumber="89">Kazakhstan</collectingCountry>
,
<collectingRegion id="49F9F852565BFFC8C26FFC88FBF2FCE1" box="[856,1042,837,863]" country="Kazakhstan" name="Almaty Oblysy" pageId="6" pageNumber="89">Almaty Region</collectingRegion>
) which is part of Northern Tien Shan Massif (
<figureCitation id="13062A35565BFFC8C077FCAAFE7CFC3F" box="[320,412,871,897]" captionStart="Figure 19" captionStartId="6.[151,227,1922,1945]" captionTargetBox="[151,1436,1146,1903]" captionTargetId="figure-300@6.[151,1436,1146,1903]" captionTargetPageId="6" captionText="Figure 19. Distribution map of Apamea ingloria sspp." figureDoi="http://doi.org/10.5281/zenodo.13246284" httpUri="https://zenodo.org/record/13246284/files/figure.png" pageId="6" pageNumber="89">Fig. 19</figureCitation>
). The specimens were collected at light of mercury vapor bulb in the belt of alpine meadows at an altitude between
<quantity id="4CC59B55565BFFC8C322FC45FD57FC1C" box="[533,695,904,930]" metricMagnitude="3" metricUnit="m" metricValue="3.05" metricValueMax="3.1" metricValueMin="3.0" pageId="6" pageNumber="89" unit="m" value="3050.0" valueMax="3100.0" valueMin="3000.0">3000–3100 m</quantity>
(
<figureCitation id="13062A35565BFFC8C3F0FC45FCC1FC1C" box="[711,801,904,930]" captionStart="Figure 20" captionStartId="7.[151,227,1065,1088]" captionTargetBox="[151,1436,189,1046]" captionTargetId="figure-11@7.[151,1436,189,1046]" captionTargetPageId="7" captionText="Figure 20. Southeast Kazakhstan, Almaty Region, Ketmen Ridge, vicinities of Ketmen Pass, 43°19'39.32&quot;N 80°19'9.41&quot;E, ca. 3000m, the habitat of Apamea ingloria postnikovi spp. n. (photo by A. Volynkin)." figureDoi="http://doi.org/10.5281/zenodo.13246288" httpUri="https://zenodo.org/record/13246288/files/figure.png" pageId="6" pageNumber="89">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C327653B565BFFC8C1A0FC01FC32FBB6" pageId="6" pageNumber="89" type="etymology">
<paragraph id="8B8236B0565BFFC8C1A0FC01FC32FBB6" blockId="6.[151,1436,972,1032]" pageId="6" pageNumber="89">
<emphasis id="B949EAA2565BFFC8C1A0FC01FEFFFC58" bold="true" box="[151,287,972,998]" pageId="6" pageNumber="89">Etymology</emphasis>
. The new subspecies is named after Nikolay Postnikov (
<collectingRegion id="49F9F852565BFFC8C2C1FC01FBB2FC58" box="[1014,1106,972,998]" country="Kazakhstan" name="Almaty Oblysy" pageId="6" pageNumber="89">Almaty</collectingRegion>
,
<collectingCountry id="F32A7620565BFFC8C552FC01FB13FC58" box="[1125,1267,972,998]" name="Kazakhstan" pageId="6" pageNumber="89">Kazakhstan</collectingCountry>
), a renowned photographer and organiser of the collecting trip to the Ketmen Ridge.
</paragraph>
</subSubSection>
</treatment>
</document>
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