163 lines
19 KiB
XML
163 lines
19 KiB
XML
<document id="97579D4D37C1B8DF5FBEBDCB67032FFA" ID-DOI="10.11646/zootaxa.1535.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5088515" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625853817661" checkinUser="felipe" docAuthor="DE SOUZA AMORIM, DALTON & RINDAL, EIRIK" docDate="2007" docId="039487A25B08FF9D4AB89AACFBADD237" docLanguage="en" docName="zt01535p092.pdf.imf" docOrigin="Zootaxa 1535 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.1535.1.1" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Eratomyia Published 2007, n. gen." docType="treatment" docVersion="4" lastPageNumber="14" masterDocId="FFADFFDA5B05FF904A2F9B03FF84D612" masterDocTitle="Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="14" updateTime="1698999120255" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="416E02E68D4BC32BDE107C37422BBB5D">Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha)</mods:title>
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<mods:namePart id="64CE5A94A015BAE438000ABDE5AF44B2">DE SOUZA AMORIM, DALTON</mods:namePart>
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<mods:namePart id="B8E3F64277350911E4D4177BBCFFAC88">RINDAL, EIRIK</mods:namePart>
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<mods:typeOfResource id="8C2FA4DD203F320EDEA6F58E76EC238B">text</mods:typeOfResource>
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<mods:title id="131554CFF10932889FC1E30DDB9C8571">Zootaxa</mods:title>
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<mods:part id="EFD8BE485102B650DFD503465CC96D10">
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<mods:date id="2D63AF9176E99B2E3936485D0F0C06F6">2007</mods:date>
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<mods:number id="508573A3C5E98FF8DAC1CFE36B2015D0">2007-07-30</mods:number>
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<mods:number id="2A8A2E81CB5E32FE61BB047B8F986FC7">1535</mods:number>
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<mods:url id="E84C65941194B62B63008E1E4944C14D">http://dx.doi.org/10.11646/zootaxa.1535.1.1</mods:url>
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<mods:classification id="E71528D22F4BFD62327791E41F8FE328">journal article</mods:classification>
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<mods:identifier id="640B3DC27B8D8BC5A8EC59AAF2F9FE91" type="DOI">10.11646/zootaxa.1535.1.1</mods:identifier>
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<treatment id="039487A25B08FF9D4AB89AACFBADD237" ID-DOI="http://doi.org/10.5281/zenodo.5088460" ID-Zenodo-Dep="5088460" LSID="urn:lsid:plazi:treatment:039487A25B08FF9D4AB89AACFBADD237" httpUri="http://treatment.plazi.org/id/039487A25B08FF9D4AB89AACFBADD237" lastPageNumber="14" pageId="13" pageNumber="14">
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<subSubSection id="C327653F5B08FF9D4AB89AACFEF2D7DB" box="[151,374,431,457]" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="8B8236B45B08FF9D4AB89AACFEF2D7DB" blockId="13.[151,374,431,492]" box="[151,374,431,457]" pageId="13" pageNumber="14">
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<heading id="D0CA81D85B08FF9D4AB89AACFEF2D7DB" bold="true" box="[151,374,431,457]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<emphasis id="B949EAA65B08FF9D4AB89AACFEF2D7DB" bold="true" box="[151,374,431,457]" pageId="13" pageNumber="14">
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<taxonomicName id="4C3D4D375B08FF9D4AB89AACFE92D7DB" ID-CoL="9CJJG" authority="Published, 2007" authorityName="Published" authorityYear="2007" box="[151,278,431,457]" class="Insecta" family="Rangomaramidae" genus="Eratomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B949EAA65B08FF9D4AB89AACFE92D7DB" bold="true" box="[151,278,431,457]" italics="true" pageId="13" pageNumber="14">Eratomyia</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27A57DD5B08FF9D4B0A9AACFEF2D7DB" box="[293,374,431,457]" pageId="13" pageNumber="14" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C327653F5B08FF9D4AB89AD1FEA2D7FE" box="[151,294,466,492]" pageId="13" pageNumber="14" type="description">
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<paragraph id="8B8236B45B08FF9D4AB89AD1FEA2D7FE" blockId="13.[151,374,431,492]" box="[151,294,466,492]" pageId="13" pageNumber="14">
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(
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<figureCitation id="13062A315B08FF9D4AB09AD1FE99D7FE" box="[159,285,466,492]" captionStart-0="FIGURES 7–10" captionStart-1="FIGURES 11–15" captionStartId-0="55.[151,269,1954,1978]" captionStartId-1="56.[151,269,1977,2001]" captionTargetBox-0="[201,1398,194,1934]" captionTargetBox-1="[215,1370,182,1963]" captionTargetId-0="figure-44@55.[201,1399,194,1940]" captionTargetId-1="figure-52@56.[214,1374,182,1963]" captionTargetPageId-0="55" captionTargetPageId-1="56" captionText-0="FIGURES 7–10. Eratomyia magnifica, sp. n. 7. Head (apical maxillary palpal segments not represented), lateral view. 8. Thorax. 9. Front tibial apex, inner view. 10. Wing." captionText-1="FIGURES 11–15. Eratomyia magnifica, sp. n. 11. Male terminalia, posterior view. 12. Male terminalia, lateral view. 13. Gonostyle, external view. 14. Aedeagus and gonocoxal bridge. 15. Tergites VIII–X and cerci, posterior view." figureDoi-0="http://doi.org/10.5281/zenodo.5088521" figureDoi-1="http://doi.org/10.5281/zenodo.5088523" httpUri-0="https://zenodo.org/record/5088521/files/figure.png" httpUri-1="https://zenodo.org/record/5088523/files/figure.png" pageId="13" pageNumber="14">Figs. 7–15</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C327653F5B08FF9D4AB89921FCC0D42E" box="[151,836,546,572]" pageId="13" pageNumber="14" type="type_taxon">
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<paragraph id="8B8236B45B08FF9D4AB89921FCC0D42E" blockId="13.[151,1437,546,1061]" box="[151,836,546,572]" pageId="13" pageNumber="14">
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<emphasis id="B949EAA65B08FF9D4AB89921FEBDD42E" bold="true" box="[151,313,546,572]" pageId="13" pageNumber="14">
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<typeStatus id="548688165B08FF9D4AB89921FF50D42E" box="[151,212,546,572]" pageId="13" pageNumber="14">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C3D4D375B08FF9D4B6F9921FDBCD429" ID-CoL="6GBVR" authority="Published, 2007" authorityName="Published" authorityYear="2007" box="[320,568,546,571]" class="Insecta" family="Rangomaramidae" genus="Eratomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="magnifica" status="sp. nov.">
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<emphasis id="B949EAA65B08FF9D4B6F9921FDBCD429" box="[320,568,546,571]" italics="true" pageId="13" pageNumber="14">Eratomyia magnifica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A57DD5B08FF9D486F9921FDFBD42E" box="[576,639,546,572]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
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, here designated
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</paragraph>
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</subSubSection>
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<subSubSection id="C327653F5B08FF9D4AE89949FF6DD4A5" pageId="13" pageNumber="14" type="diagnosis">
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<paragraph id="8B8236B45B08FF9D4AE89949FF6DD4A5" blockId="13.[151,1437,546,1061]" pageId="13" pageNumber="14">
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<emphasis id="B949EAA65B08FF9D4AE89949FEC3D476" bold="true" box="[199,327,586,612]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
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First sector of m-cu longer than in other genera of subfamily, m-cu reaching CuA more distally. M
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<subScript id="17B934F15B08FF9D4ADD9981FE8FD482" attach="left" box="[242,267,642,656]" fontSize="6" pageId="13" pageNumber="14">1+2</subScript>
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/M
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<subScript id="17B934F15B08FF9D4B029981FEC1D482" attach="left" box="[301,325,642,656]" fontSize="6" pageId="13" pageNumber="14">3+4</subScript>
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shorter than m-cu, r-m long. Tergites 1-4 and 6 shining brown, tergite 5 yellow. Midpleural pit absent.
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</paragraph>
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</subSubSection>
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<subSubSection id="C327653F5B08FF9D4AE899C6FAA0D4CD" box="[199,1316,709,735]" pageId="13" pageNumber="14" type="etymology">
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<paragraph id="8B8236B45B08FF9D4AE899C6FAA0D4CD" blockId="13.[151,1437,546,1061]" box="[199,1316,709,735]" pageId="13" pageNumber="14">
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<emphasis id="B949EAA65B08FF9D4AE899C6FED7D4CD" bold="true" box="[199,339,709,735]" pageId="13" pageNumber="14">Etymology.</emphasis>
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The species name is feminine, from the Greek
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<emphasis id="B949EAA65B08FF9D495199C5FC43D4CD" box="[894,967,710,735]" italics="true" pageId="13" pageNumber="14">eratos</emphasis>
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, lovely, meaning a lovely fly.
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</paragraph>
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</subSubSection>
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<subSubSection id="C327653F5B08FF9D4AE899EEFBADD237" pageId="13" pageNumber="14" type="discussion">
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<paragraph id="8B8236B45B08FF9D4AE899EEFBADD237" blockId="13.[151,1437,546,1061]" pageId="13" pageNumber="14">
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<emphasis id="B949EAA65B08FF9D4AE899EEFED1D515" bold="true" box="[199,341,749,775]" pageId="13" pageNumber="14">Comments.</emphasis>
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This genus, which is known only from the
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<typeStatus id="548688165B08FF9D497999EEFC0CD515" box="[854,904,749,775]" pageId="13" pageNumber="14">type</typeStatus>
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species, from
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<collectingCountry id="F32A76245B08FF9D4E1A99EEFB1CD515" box="[1077,1176,749,775]" name="Ecuador" pageId="13" pageNumber="14">Ecuador</collectingCountry>
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, appears in the phylogenetic analysis (below) as the sister group of
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<taxonomicName id="4C3D4D375B08FF9D48EB9815FCCDD53D" authorityName="Chandler" authorityYear="2002" box="[708,841,790,815]" class="Insecta" family="Rangomaramidae" genus="Chiletricha" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
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<emphasis id="B949EAA65B08FF9D48EB9815FCCDD53D" box="[708,841,790,815]" italics="true" pageId="13" pageNumber="14">Chiletricha</emphasis>
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</taxonomicName>
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, which is known from
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<collectingCountry id="F32A76245B08FF9D4E4D9816FB20D53D" box="[1122,1188,789,815]" name="Chile" pageId="13" pageNumber="14">Chile</collectingCountry>
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, southern
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<collectingCountry id="F32A76245B08FF9D4F0F9816FA1CD53D" box="[1312,1432,789,815]" name="Argentina" pageId="13" pageNumber="14">Argentina</collectingCountry>
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, and southern
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<collectingCountry id="F32A76245B08FF9D4B19983EFE05D545" box="[310,385,829,855]" name="Brazil" pageId="13" pageNumber="14">Brazil</collectingCountry>
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. It differs from the
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<taxonomicName id="4C3D4D375B08FF9D4848983DFCCDD545" box="[615,841,829,855]" class="Insecta" family="Rangomaramidae" genus="Chiletricha" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B949EAA65B08FF9D4848983DFD68D545" box="[615,748,830,855]" italics="true" pageId="13" pageNumber="14">Chiletricha</emphasis>
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species
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</taxonomicName>
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in wing features, with a much wider basal cell, R
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<subScript id="17B934F15B08FF9D4FBB984EFA18D549" attach="left" box="[1428,1436,845,859]" fontSize="6" pageId="13" pageNumber="14">5</subScript>
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separated from C, and M
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<subScript id="17B934F15B08FF9D4B97987BFE44D594" attach="left" box="[440,448,888,902]" fontSize="6" pageId="13" pageNumber="14">4</subScript>
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originating from CuA not as basally in the wing (
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<figureCitation id="13062A315B08FF9D4E2D986BFBE0D590" box="[1026,1124,872,898]" captionStart="FIGURES 7–10" captionStartId="55.[151,269,1954,1978]" captionTargetBox="[201,1398,194,1934]" captionTargetId="figure-44@55.[201,1399,194,1940]" captionTargetPageId="55" captionText="FIGURES 7–10. Eratomyia magnifica, sp. n. 7. Head (apical maxillary palpal segments not represented), lateral view. 8. Thorax. 9. Front tibial apex, inner view. 10. Wing." figureDoi="http://doi.org/10.5281/zenodo.5088521" httpUri="https://zenodo.org/record/5088521/files/figure.png" pageId="13" pageNumber="14">Figs. 10</figureCitation>
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,
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<figureCitation id="13062A315B08FF9D4E5F986BFB14D590" box="[1136,1168,872,898]" captionStart="FIGURES 16–19" captionStartId="57.[151,269,1915,1939]" captionTargetBox="[179,1398,194,1884]" captionTargetId="figure-44@57.[179,1398,194,1889]" captionTargetPageId="57" captionText="FIGURES 16–19. Chiletricha. 16, 17. Chiletricha seminuda (Freeman). 16. Wing. 17. Mouthparts, posterior view. 18, 19. Chiletricha marginata (Edwards). 18. Thorax. 19. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5088527" httpUri="https://zenodo.org/record/5088527/files/figure.png" pageId="13" pageNumber="14">16</figureCitation>
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). Mouthparts are fairly similar (
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<figureCitation id="13062A315B08FF9D4AD79890FEC8D5BF" box="[248,332,915,941]" captionStart="FIGURES 7–10" captionStartId="55.[151,269,1954,1978]" captionTargetBox="[201,1398,194,1934]" captionTargetId="figure-44@55.[201,1399,194,1940]" captionTargetPageId="55" captionText="FIGURES 7–10. Eratomyia magnifica, sp. n. 7. Head (apical maxillary palpal segments not represented), lateral view. 8. Thorax. 9. Front tibial apex, inner view. 10. Wing." figureDoi="http://doi.org/10.5281/zenodo.5088521" httpUri="https://zenodo.org/record/5088521/files/figure.png" pageId="13" pageNumber="14">Figs. 7</figureCitation>
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,
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<figureCitation id="13062A315B08FF9D4B779890FEFCD5BF" box="[344,376,915,941]" captionStart="FIGURES 16–19" captionStartId="57.[151,269,1915,1939]" captionTargetBox="[179,1398,194,1884]" captionTargetId="figure-44@57.[179,1398,194,1889]" captionTargetPageId="57" captionText="FIGURES 16–19. Chiletricha. 16, 17. Chiletricha seminuda (Freeman). 16. Wing. 17. Mouthparts, posterior view. 18, 19. Chiletricha marginata (Edwards). 18. Thorax. 19. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5088527" httpUri="https://zenodo.org/record/5088527/files/figure.png" pageId="13" pageNumber="14">17</figureCitation>
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). In the thorax, the mesepimeron of
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<taxonomicName id="4C3D4D375B08FF9D490D9890FC1AD5BE" authorityName="Published" authorityYear="2007" box="[802,926,915,940]" class="Insecta" family="Rangomaramidae" genus="Eratomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
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<emphasis id="B949EAA65B08FF9D490D9890FC1AD5BE" box="[802,926,915,940]" italics="true" pageId="13" pageNumber="14">Eratomyia</emphasis>
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</taxonomicName>
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is more slender on its ventral half, whereas the laterotergite is slightly larger (
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<figureCitation id="13062A315B08FF9D480898B8FDFED5C7" box="[551,634,955,981]" captionStart="FIGURES 7–10" captionStartId="55.[151,269,1954,1978]" captionTargetBox="[201,1398,194,1934]" captionTargetId="figure-44@55.[201,1399,194,1940]" captionTargetPageId="55" captionText="FIGURES 7–10. Eratomyia magnifica, sp. n. 7. Head (apical maxillary palpal segments not represented), lateral view. 8. Thorax. 9. Front tibial apex, inner view. 10. Wing." figureDoi="http://doi.org/10.5281/zenodo.5088521" httpUri="https://zenodo.org/record/5088521/files/figure.png" pageId="13" pageNumber="14">Figs. 8</figureCitation>
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,
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<figureCitation id="13062A315B08FF9D48A698B8FD2DD5C7" box="[649,681,955,981]" captionStart="FIGURES 16–19" captionStartId="57.[151,269,1915,1939]" captionTargetBox="[179,1398,194,1884]" captionTargetId="figure-44@57.[179,1398,194,1889]" captionTargetPageId="57" captionText="FIGURES 16–19. Chiletricha. 16, 17. Chiletricha seminuda (Freeman). 16. Wing. 17. Mouthparts, posterior view. 18, 19. Chiletricha marginata (Edwards). 18. Thorax. 19. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5088527" httpUri="https://zenodo.org/record/5088527/files/figure.png" pageId="13" pageNumber="14">18</figureCitation>
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). Tergite 9 has strong spines on a pair of distal projections (
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<figureCitation id="13062A315B08FF9D4F4298B8FF67D5EF" captionStart="FIGURES 11–15" captionStartId="56.[151,269,1977,2001]" captionTargetBox="[215,1370,182,1963]" captionTargetId="figure-52@56.[214,1374,182,1963]" captionTargetPageId="56" captionText="FIGURES 11–15. Eratomyia magnifica, sp. n. 11. Male terminalia, posterior view. 12. Male terminalia, lateral view. 13. Gonostyle, external view. 14. Aedeagus and gonocoxal bridge. 15. Tergites VIII–X and cerci, posterior view." figureDoi="http://doi.org/10.5281/zenodo.5088523" httpUri="https://zenodo.org/record/5088523/files/figure.png" pageId="13" pageNumber="14">Fig. 12–15</figureCitation>
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), as also seen in
|
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<taxonomicName id="4C3D4D375B08FF9D4B9398E0FDC5D5EE" authorityName="Chandler" authorityYear="2002" box="[444,577,995,1020]" class="Insecta" family="Rangomaramidae" genus="Chiletricha" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949EAA65B08FF9D4B9398E0FDC5D5EE" box="[444,577,995,1020]" italics="true" pageId="13" pageNumber="14">Chiletricha</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13062A315B08FF9D487A98E0FD34D5EF" box="[597,688,995,1021]" captionStart="FIGURES 16–19" captionStartId="57.[151,269,1915,1939]" captionTargetBox="[179,1398,194,1884]" captionTargetId="figure-44@57.[179,1398,194,1889]" captionTargetPageId="57" captionText="FIGURES 16–19. Chiletricha. 16, 17. Chiletricha seminuda (Freeman). 16. Wing. 17. Mouthparts, posterior view. 18, 19. Chiletricha marginata (Edwards). 18. Thorax. 19. Male terminalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5088527" httpUri="https://zenodo.org/record/5088527/files/figure.png" pageId="13" pageNumber="14">Fig. 19</figureCitation>
|
||
),
|
||
<taxonomicName id="4C3D4D375B08FF9D48E598E0FC44D5EE" box="[714,960,995,1020]" class="Insecta" family="Rangomaramidae" genus="Rhynchoheterotricha" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949EAA65B08FF9D48E598E0FC44D5EE" box="[714,960,995,1020]" italics="true" pageId="13" pageNumber="14">Rhynchoheterotricha</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3D4D375B08FF9D4E2798E0FB13D5EE" authorityName="Jaschhof" authorityYear="2004" box="[1032,1175,995,1020]" class="Insecta" family="Rangomaramidae" genus="Insulatricha" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949EAA65B08FF9D4E2798E0FB13D5EE" box="[1032,1175,995,1020]" italics="true" pageId="13" pageNumber="14">Insulatricha</emphasis>
|
||
</taxonomicName>
|
||
, and the gonocoxites have a pair of unique digitiform projections bearing setae and a pair of spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |