536 lines
66 KiB
XML
536 lines
66 KiB
XML
<document id="96F69BDFE0A27B04AC54A76B58ACEE3B" ID-CLB-Dataset="20492" ID-DOI="10.5169/seals-787049" ID-GBIF-Dataset="ce0756e2-5184-4b74-8472-ab69fed7951c" ID-ISSN="2624-9170" ID-Zenodo-Dep="6285683" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637935326602" checkinUser="carolina" docAuthor="Huber, Charles & Schmidt, Joachim" docDate="2017" docId="03948799FFE1FF80FF41FCD2FE8BFA0E" docLanguage="en" docName="ContribNatHist.36.1-85.pdf.imf" docOrigin="Contributions to Natural History 36" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Nebria (Epinebriola) delicata Huber & Schmidt, 2017, sp. nov." docType="treatment" docVersion="12" lastPageNumber="64" masterDocId="FFADFFE1FFDBFFC0FF9CFFE9FFB0FFA7" masterDocTitle="Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)" masterLastPageNumber="85" masterPageNumber="1" pageNumber="59" updateTime="1699470525451" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods id="5EA3572EC688D4BD1E188FCE5FD7D318" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0D76CB99FD050DF0B313A62BDBF185A5">
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<mods:title id="033E350E4E291184D77FFE2B56C30F48">Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)</mods:title>
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<mods:name id="66C04DB82B4B746A96B942FD20D66AB6" type="personal">
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<mods:roleTerm id="B2CE1EE04C96CA4C286A11083487582D">Author</mods:roleTerm>
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<mods:namePart id="A1144F5923839AC57D9A58F8ECF0BAA7">Huber, Charles</mods:namePart>
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<mods:roleTerm id="C33CDE9B8BD148684C22F7129B42C28F">Author</mods:roleTerm>
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<mods:namePart id="E8D59A9C5EE2C80B5E73BD6D02893558">Schmidt, Joachim</mods:namePart>
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<mods:typeOfResource id="2BAAC31E68E206E1314184AEEAA61117">text</mods:typeOfResource>
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<mods:titleInfo id="926E897294C24DAE7975FA609E43CBFB">
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<mods:title id="8F516DE266EECD4B01C1728699A8EDAA">Contributions to Natural History</mods:title>
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<mods:part id="78071994F7FE66809540377B0D6C7642">
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<mods:date id="A5F37EBFEC4A2C7284BEFAEB872B76BE">2017</mods:date>
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<mods:number id="F9A3C2845388FD483FBDEFD76402D2E9">2017-07-17</mods:number>
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<mods:detail id="1B610C0FC6E9D943D08923092D5E326B" type="volume">
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<mods:number id="19FBA3187CD784817C2E9F872D014334">36</mods:number>
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<mods:classification id="FB4B6EF20A5E1DCB8A5EBCBD162EF9BB">journal article</mods:classification>
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<mods:identifier id="BA9B1BE7896CF7449C20FA9C3EB4E6B7" type="CLB-Dataset">20492</mods:identifier>
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<mods:identifier id="A133F15F4A638A810D84F2E31C6D3208" type="DOI">10.5169/seals-787049</mods:identifier>
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<mods:identifier id="80F7DE7B4AD585DF055F9815B8A6369D" type="ISSN">2624-9170</mods:identifier>
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<treatment id="03948799FFE1FF80FF41FCD2FE8BFA0E" ID-DOI="http://doi.org/10.5281/zenodo.6315677" ID-GBIF-Taxon="193820253" ID-Zenodo-Dep="6315677" LSID="urn:lsid:plazi:treatment:03948799FFE1FF80FF41FCD2FE8BFA0E" httpUri="http://treatment.plazi.org/id/03948799FFE1FF80FF41FCD2FE8BFA0E" lastPageId="64" lastPageNumber="64" pageId="58" pageNumber="59">
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<subSubSection id="C3276504FFE1FFFAFF41FCD2FD58FCF3" box="[221,744,827,852]" pageId="58" pageNumber="59" type="nomenclature">
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<paragraph id="8B82368FFFE1FFFAFF41FCD2FD58FCF3" blockId="58.[221,744,827,852]" box="[221,744,827,852]" pageId="58" pageNumber="59">
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<heading id="D0CA81E3FFE1FFFAFF41FCD2FD58FCF3" box="[221,744,827,852]" centered="true" fontSize="12" level="2" pageId="58" pageNumber="59" reason="2">
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<emphasis id="B949EA9DFFE1FFFAFF41FCD2FD34FCF3" bold="true" box="[221,644,827,852]" pageId="58" pageNumber="59">
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<taxonomicName id="4C3D4D0CFFE1FFFAFF41FCD2FD96FCF3" ID-CoL="8M6VZ" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[221,550,827,852]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov." subGenus="Epinebriola">
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<emphasis id="B949EA9DFFE1FFFAFF41FCD2FE99FCF3" bold="true" box="[221,297,827,852]" italics="true" pageId="58" pageNumber="59">Nebria</emphasis>
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(
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<emphasis id="B949EA9DFFE1FFFAFEA5FCD2FE0BFCF3" bold="true" box="[313,443,827,852]" italics="true" pageId="58" pageNumber="59">Epinebriola</emphasis>
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)
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<emphasis id="B949EA9DFFE1FFFAFE57FCD2FD96FCF3" bold="true" box="[459,550,827,852]" italics="true" pageId="58" pageNumber="59">delicata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A57E6FFE1FFFAFDB0FCD2FD34FCF3" box="[556,644,827,852]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="13062A0AFFE1FFFAFD0CFCD5FD6FFCF3" box="[656,735,828,852]" captionStart="Fig" captionStartId="60.[221,250,161,179]" captionTargetBox="[463,1106,153,1396]" captionTargetId="figure-64@60.[463,1106,150,1396]" captionTargetPageId="60" captionText="Fig. 30: Nebria (Epi- nebriola) delicata sp. nov. Holotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285810" httpUri="https://zenodo.org/record/6285810/files/figure.png" pageId="58" pageNumber="59">Fig. 30</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3276504FFE1FFFAFF41FC6FFC90FC4E" pageId="58" pageNumber="59" type="materials_examined">
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<paragraph id="8B82368FFFE1FFFAFF41FC6FFD73FC63" blockId="58.[221,1107,899,1636]" pageId="58" pageNumber="59">
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<materialsCitation id="3B553CD2FFE1FFFAFF41FC6FFD73FC63" ID-GBIF-Occurrence="3497686315" collectingDate="2011-06-04" collectionCode="cSCHM" collectorName="J. Schmidt" elevation="5310" latitude="29.566668" location="Mt. Mendju Zari" longLatPrecision="1226" longitude="91.21667" pageId="58" pageNumber="59" specimenCount="1" specimenCount-male="1" stateProvince="Tibet" typeStatus="holotype">
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<typeStatus id="5486882DFFE1FFFAFF41FC6FFEF6FC39" box="[221,326,902,926]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D3BFD06FFE1FFFAFECCFC6AFED4FC39" box="[336,356,899,926]" pageId="58" pageNumber="59" type="male">♂</specimenCount>
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: S
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<collectingRegion id="49F9F86DFFE1FFFAFE11FC6FFE7DFC39" box="[397,461,902,926]" country="China" name="Xizang" pageId="58" pageNumber="59">Tibet</collectingRegion>
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,
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<location id="8EE26054FFE1FFFAFE44FC6FFD28FC39" LSID="urn:lsid:plazi:treatment:03948799FFE1FF80FF41FCD2FE8BFA0E:8EE26054FFE1FFFAFE44FC6FFD28FC39" box="[472,664,902,926]" latitude="29.566668" longLatPrecision="1226" longitude="91.21667" name="Mt. Mendju Zari" pageId="58" pageNumber="59" stateProvince="Tibet">Mt. Mendju Zari</location>
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,
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<location id="8EE26054FFE1FFFAFD3FFC6FFCA2FC39" LSID="urn:lsid:plazi:treatment:03948799FFE1FF80FF41FCD2FE8BFA0E:8EE26054FFE1FFFAFD3FFC6FFCA2FC39" box="[675,786,902,926]" latitude="29.566668" longLatPrecision="1226" longitude="91.21667" name="SE Lhasa" pageId="58" pageNumber="59" stateProvince="Tibet">SE Lhasa</location>
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,
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<geoCoordinate id="EE095048FFE1FFFAFC82FC6FFC39FC39" box="[798,905,902,926]" degrees="29" direction="north" minutes="34" orientation="latitude" pageId="58" pageNumber="59" precision="925" value="29.566668">29°34' N</geoCoordinate>
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,
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<geoCoordinate id="EE095048FFE1FFFAFC0AFC6FFC4CFC39" box="[918,1020,902,926]" degrees="91" direction="east" minutes="13" orientation="longitude" pageId="58" pageNumber="59" precision="925" value="91.21667">91°13' E</geoCoordinate>
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,
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<quantity id="4CC59B6AFFE1FFFAFB95FC6FFE86FC63" metricMagnitude="3" metricUnit="m" metricValue="5.31" metricValueMax="5.42" metricValueMin="5.2" pageId="58" pageNumber="59" unit="m" value="5310.0" valueMax="5420.0" valueMin="5200.0">
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<elevation id="0010D1BCFFE1FFFAFB95FC6FFE86FC63" metricMagnitude="3" metricUnit="m" metricValue="5.31" metricValueMax="5.42" metricValueMin="5.2" pageId="58" pageNumber="59" unit="m" value="5310.0" valueMax="5420.0" valueMin="5200.0">5200– 5420 m</elevation>
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</quantity>
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,
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<date id="FF83104FFFE1FFFAFEA3FC45FE12FC63" box="[319,418,940,964]" pageId="58" pageNumber="59" value="2011-06-04">
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<collectingDate id="EFC7E9A7FFE1FFFAFEA3FC45FE12FC63" box="[319,418,940,964]" pageId="58" pageNumber="59" value="2011-06-04">4.6.2011</collectingDate>
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</date>
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, leg.
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<collectorName id="26C85359FFE1FFFAFE43FC45FDE0FC63" box="[479,592,940,964]" pageId="58" pageNumber="59">J. Schmidt</collectorName>
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(
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<collectionCode id="ED2CAE4AFFE1FFFAFDC0FC45FD09FC63" box="[604,697,940,964]" pageId="58" pageNumber="59">cSCHM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8B82368FFFE1FFFAFE9FFC38FC90FC4E" blockId="58.[221,1107,899,1636]" box="[259,800,974,1001]" pageId="58" pageNumber="59">
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<materialsCitation id="3B553CD2FFE1FFFAFE9FFC38FCACFC4E" ID-GBIF-Occurrence="3497686385" box="[259,796,974,1001]" collectingDate="2011-06-04" collectionCode="cSCHM" collectorName="Zari, SE & J. Schmidt" elevation="5310" latitude="29.566668" location="Mt. Mendju Zari" longLatPrecision="1226" longitude="91.21667" pageId="58" pageNumber="59" specimenCount="1" specimenCount-female="1" stateProvince="Xizang" typeStatus="paratype">
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<typeStatus id="5486882DFFE1FFFAFE9FFC38FEC0FC4E" box="[259,368,977,1001]" pageId="58" pageNumber="59" type="paratype">Paratype</typeStatus>
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:
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<specimenCount id="9D3BFD06FFE1FFFAFEE4FC38FE11FC4E" box="[376,417,974,1001]" pageId="58" pageNumber="59" type="female">1 ♀</specimenCount>
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, same data as holotype (
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<collectionCode id="ED2CAE4AFFE1FFFAFD26FC38FCA7FC4E" box="[698,791,977,1001]" pageId="58" pageNumber="59">cSCHM</collectionCode>
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)
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||
</materialsCitation>
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||
.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3276504FFE1FFFEFE9FFC1FFBFCFCDE" lastPageId="62" lastPageNumber="63" pageId="58" pageNumber="59" type="description">
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<paragraph id="8B82368FFFE1FFFAFE9FFC1FFD55FB93" blockId="58.[221,1107,899,1636]" pageId="58" pageNumber="59">
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Body length
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<quantity id="4CC59B6AFFE1FFFAFE0BFC1FFE6BFBA9" box="[407,475,1014,1038]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="58" pageNumber="59" unit="mm" value="9.0">9 mm</quantity>
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. Colour dark brown to black. Appendages of the head, antennae, tibiae and tarsi brownish lightened.
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</paragraph>
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<paragraph id="8B82368FFFE1FFFAFE9FFBA8FE6BFA69" blockId="58.[221,1107,899,1636]" pageId="58" pageNumber="59">
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Head small. Labrum with anterior margin straight, bearing eight setae. Clypeus with anterior margin straight. Frons transversally wrinkled. Eyes small, weakly prominent, temples long. One supraorbital seta. Head with a faint collar constriction. Vertex impunctate. Antennae long and slender extending at least to the middle of the elytra. Antennal scape slender, elongate, cylindrical, longer than the eye's diameter, basally narrowed, with one dorsal seta. 2
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<superScript id="7C489BC7FFE1FFFAFF70FAF7FEB4FA96" attach="left" box="[236,260,1310,1329]" fontSize="8" pageId="58" pageNumber="59">nd</superScript>
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antennomere with one seta ventroapically. Maxillar palpomeres short. Maxillary stipes laterally straight, with 5 setae. Penultimate labial palpomere trisetose. Ligula spiny, with two lateral setae. Mentum with bifid medial tooth. Submentum with a row of 4+5 setae. Vertex impunctate. Microreticulation of the head isodiametric.
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</paragraph>
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<paragraph id="8B82368FFFE1FFFBFE9FFA35FE67FD3E" blockId="58.[221,1107,899,1636]" lastBlockId="59.[184,1069,156,1636]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
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Pronotum slender, ratio width/length of the pronotum = 1.25. Lateral margin faintly convex, slightly narrowed basally (ratio apical width/basal width = 1.07), rounded to the anterior angles, faintly rounded to the posterior angles, weakly concave in front of the posterior angles (
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<figureCitation id="13062A0AFFE1FFFAFCBAF9A5FC20F9C3" box="[806,912,1612,1636]" captionStart="Fig" captionStartId="61.[184,213,1317,1335]" captionTargetBox="[185,1068,153,1276]" captionTargetId="figure-106@61.[551,733,151,716]" captionTargetPageId="61" captionText="Fig. 31: Left part of the pronotum of Epinebriola species. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) tuberculata sp. nov.; F: N. (E.) numburica sp. nov.; G: N. (E.) retingensis sp. nov.; H: N. (E.) delicata sp. nov.; I: N. (E.) incarinata sp. nov.; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6285816" httpUri="https://zenodo.org/record/6285816/files/figure.png" pageId="58" pageNumber="59">Fig. 31H</figureCitation>
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). Lateral groove very narrow, narrow evenly apically, broadened basally. Anterior angles faintly acuminate, slightly protruding. Posterior angles acute, slightly protruding backwards. Basal margin medially straight, in front of the posterior angles weakly sinuate. Pronotal disc weakly convex. Basal fovea deep, anterior transverse impression weak, posterior transverse impression distinct, median line shallow. Lateral groove and anterior transverse impression impunctate; basal fovea and posterior transverse impression sparsely punctate. Apical margination of the pronotum restricted to lateral one-fourths, merges with the apex of the anterior angle. Base of the pronotum not margined. Basolateral seta present. One midlateral seta in the apical half of the pronotum, inserted within the narrow lateral groove; outline of the lateral margin at the insertion site of the seta weakly disturbed. Pronotum impunctate on disc, microreticulation isodiametric. Proepisternum smooth. Prosternal process laterally margined, apex bulging and unmargined.
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</paragraph>
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<paragraph id="8B82368FFFE0FFFBFF41FD4FFD3FFB93" blockId="59.[184,1069,156,1636]" pageId="59" pageNumber="60">Elytral silhouette slender, elongate, convex with the lateral margin almost straight in the middle, widest slightly behind the middle. Ratio length/width of the elytra = 1.71. Distinctly wider than the pronotum, ratio elytral width/pronotal width = 1.65. Elytral apex acutely rounded, apical carina faint. Basal margination straight, at right angles with the elytral suture, regularly merged with the lateral margination. Humeral carina weakly present. Striae distinct, deep, reaching the apex, striae 1–2 faintly punctate, the remaining striae coarsely punctate; lateral striae obliterate towards the apex. Intervals slightly convex. Interval 3 with 2–4 setae adjoining stria 3, intervals 5 and 7 asetose. Scutellar seta absent. Microreticulation isodiametric, interspersed with transverse meshes. Mesepisterna sparsely punctate.</paragraph>
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<paragraph id="8B82368FFFE0FFFBFF41FBA8FD4DFB6E" blockId="59.[184,1069,156,1636]" pageId="59" pageNumber="60">Sternum 2 laterally faintly punctate. Sternum 3 medially asetose. Sterna 4–6 posteriorly each with 1–3 paramedial setae; anal sternum with two paramedial setae in the male. Metepisterna twice as long as wide, sparsely punctate. Metacoxa basally bisetose, apically unisetose.</paragraph>
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<paragraph id="8B82368FFFE0FFFBFF41FB3FFD01FA0E" blockId="59.[184,1069,156,1636]" pageId="59" pageNumber="60">Legs long and slender. Tarsi slender, tarsomeres 1 and 2 dorsally with a few short setae. Protarsus of the male with tarsomeres 1–3 broadened, ventrally with pads of adhesive setae. Ventroapical tooth of the metatarsomere 4 small and only little projecting. Meso- and metatarsomeres 2–4 ventroapically with long setae. Tarsomeres 5 ventrally with two rows of 3–4 short setae. Metatarsomere 5 as long as metatarsomeres 3+4.</paragraph>
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<paragraph id="8B82368FFFE0FFFBFF41FA5FFC19F9C3" blockId="59.[184,1069,156,1636]" pageId="59" pageNumber="60">
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Male genitalia: Edeagus (
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<figureCitation id="13062A0AFFE0FFFBFE6DFA5FFDE4FA69" box="[497,596,1462,1486]" captionStart="Fig" captionStartId="38.[221,250,1317,1335]" captionTargetBox="[221,1106,153,1296]" captionTargetId="figure-70@38.[221,704,627,867]" captionTargetPageId="38" captionText="Fig. 20: Male edeagi of new Nebria species of the subgenus Epinebriola K. DANIEL, 1904. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) numburica sp. nov., with parameres; F: N. (E.) retingensis sp. nov.; G: N. (E.) incarinata sp. nov.; H: N. (E.) delicata sp. nov.; I: N. (E.) incarinata sp. nov., apex in dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6285764" httpUri="https://zenodo.org/record/6285764/files/figure.png" pageId="59" pageNumber="60">Fig. 20G</figureCitation>
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): Base of the median lobe small, with small basolateral lobes; the base decreasing abruptly to the thin mid-shaft. Mid-shaft strongly curved at base on inner side as well as on outer side; moderately curved to the apex. Apex straight, faintly deflected to the left, in front of the long tip on the ventral side faintly convex. Mid-shaft of the endophallus asetose.
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</paragraph>
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<caption id="DF426607FFE7FFFCFF41FF48FECAFEA4" ID-DOI="http://doi.org/10.5281/zenodo.6285810" ID-Zenodo-Dep="6285810" httpUri="https://zenodo.org/record/6285810/files/figure.png" pageId="60" pageNumber="61" startId="60.[221,250,161,179]" targetBox="[463,1106,153,1396]" targetPageId="60">
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<paragraph id="8B82368FFFE7FFFCFF41FF48FECAFEA4" blockId="60.[221,407,161,259]" pageId="60" pageNumber="61">
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Fig. 30:
|
||
<taxonomicName id="4C3D4D0CFFE7FFFCFEBFFF48FE9EFF69" authority="(Epinebriola)" authorityName="LATREILLE" authorityYear="1802" baseAuthorityName="Epinebriola" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949EA9DFFE7FFFCFEBFFF48FEEDFF14" bold="true" box="[291,349,161,179]" italics="true" pageId="60" pageNumber="61">Nebria</emphasis>
|
||
(
|
||
<emphasis id="B949EA9DFFE7FFFCFEF5FF48FE9EFF69" bold="true" pageId="60" pageNumber="61">
|
||
<emphasis id="B949EA9DFFE7FFFCFEF5FF48FE95FF69" bold="true" italics="true" pageId="60" pageNumber="61">Epinebriola</emphasis>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B949EA9DFFE7FFFCFEAEFF55FECAFEA4" bold="true" pageId="60" pageNumber="61">
|
||
<taxonomicName id="4C3D4D0CFFE7FFFCFEAEFF55FEC8FF69" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[306,376,188,206]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="delicata" status="sp.">
|
||
<emphasis id="B949EA9DFFE7FFFCFEAEFF55FEC8FF69" bold="true" box="[306,376,188,206]" italics="true" pageId="60" pageNumber="61">delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE7FFFCFEE0FF55FE27FF69" box="[380,407,188,206]" pageId="60" pageNumber="61">sp.</taxonomicNameLabel>
|
||
nov.
|
||
<typeStatus id="5486882DFFE7FFFCFE9AFF3FFEE9FF4F" box="[262,345,214,232]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
|
||
. Scale bar =
|
||
<quantity id="4CC59B6AFFE7FFFCFED9FF18FEC6FEA4" box="[325,374,241,259]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="60" pageNumber="61" unit="mm" value="5.0">5 mm</quantity>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B82368FFFE7FFFCFE9FFA5FFE1DF9C3" blockId="60.[221,1106,1462,1636]" pageId="60" pageNumber="61">
|
||
Female genitalia: Gonocoxa (
|
||
<figureCitation id="13062A0AFFE7FFFCFDCFFA5FFD14FA69" box="[595,676,1462,1486]" captionStart="Fig" captionStartId="62.[221,250,160,178]" captionTargetBox="[463,1106,155,734]" captionTargetId="figure-330@62.[463,1106,153,737]" captionTargetPageId="62" captionText="Fig. 32: Gonocoxae of Nebria (Epinebriola) delicata sp. nov., in caudal view." figureDoi="http://doi.org/10.5281/zenodo.6285822" httpUri="https://zenodo.org/record/6285822/files/figure.png" pageId="60" pageNumber="61">Fig. 32</figureCitation>
|
||
): Gonocoxite 2 very long, as long or longer than gonocoxite 1, slender, curved dagger-like, broad at base, distinctly narrowing near base, parallel sided in median part. Apex acuminate, slightly arcuate dorsally. Ventral preapical insertion furrow small, short-oval, with two nematiform setae.
|
||
</paragraph>
|
||
<caption id="DF426607FFE6FFFDFF24FACCFDCBFA20" ID-DOI="http://doi.org/10.5281/zenodo.6285816" ID-Zenodo-Dep="6285816" httpUri="https://zenodo.org/record/6285816/files/figure.png" pageId="61" pageNumber="62" startId="61.[184,213,1317,1335]" targetBox="[185,1068,153,1276]" targetPageId="61">
|
||
<paragraph id="8B82368FFFE6FFFDFF24FACCFDCBFA20" blockId="61.[184,1069,1317,1415]" pageId="61" pageNumber="62">
|
||
Fig. 31: Left part of the pronotum of
|
||
<taxonomicName id="4C3D4D0CFFE6FFFDFE63FACCFDD3FA90" authorityName="K. DANIEL" authorityYear="1904" box="[511,611,1317,1335]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="79" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE6FFFDFE63FACCFDD3FA90" bold="true" box="[511,611,1317,1335]" italics="true" pageId="61" pageNumber="62">Epinebriola</emphasis>
|
||
</taxonomicName>
|
||
species. A:
|
||
<taxonomicName id="4C3D4D0CFFE6FFFDFD4CFACCFC6AFA90" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[720,986,1317,1335]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="triseriata" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE6FFFDFD4CFACCFCBAFA90" bold="true" box="[720,778,1317,1335]" italics="true" pageId="61" pageNumber="62">Nebria</emphasis>
|
||
(
|
||
<emphasis id="B949EA9DFFE6FFFDFC84FACCFCCCFA90" bold="true" box="[792,892,1317,1335]" italics="true" pageId="61" pageNumber="62">Epinebriola</emphasis>
|
||
)
|
||
<emphasis id="B949EA9DFFE6FFFDFC16FACCFC6AFA90" bold="true" box="[906,986,1317,1335]" italics="true" pageId="61" pageNumber="62">triseriata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE6FFFDFC7CFACCFB9AFA90" box="[992,1066,1317,1335]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; B:
|
||
<taxonomicName id="4C3D4D0CFFE6FFFDFF4DFAA9FEF7FAF5" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[209,327,1344,1362]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="2" phylum="Arthropoda" rank="species" species="rupina" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE6FFFDFF4DFAA9FF6EFAF5" bold="true" box="[209,222,1344,1362]" italics="true" pageId="61" pageNumber="62">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE6FFFDFF6DFAA9FF4CFAF5" bold="true" box="[241,252,1344,1362]" italics="true" pageId="61" pageNumber="62">E</emphasis>
|
||
.)
|
||
<emphasis id="B949EA9DFFE6FFFDFE93FAA9FEF7FAF5" bold="true" box="[271,327,1344,1362]" italics="true" pageId="61" pageNumber="62">rupina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE6FFFDFED2FAA9FE27FAF5" box="[334,407,1344,1362]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; C:
|
||
<taxonomicName id="4C3D4D0CFFE6FFFDFE24FAA9FDC2FAF5" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[440,626,1344,1362]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="2" phylum="Arthropoda" rank="species" species="montisanimae" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE6FFFDFE24FAA9FE75FAF5" bold="true" box="[440,453,1344,1362]" italics="true" pageId="61" pageNumber="62">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE6FFFDFE44FAA9FE53FAF5" bold="true" box="[472,483,1344,1362]" italics="true" pageId="61" pageNumber="62">E</emphasis>
|
||
.)
|
||
<emphasis id="B949EA9DFFE6FFFDFE6AFAA9FDC2FAF5" bold="true" box="[502,626,1344,1362]" italics="true" pageId="61" pageNumber="62">montisanimae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE6FFFDFDE4FAA9FD71FAF5" box="[632,705,1344,1362]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; D:
|
||
<taxonomicName id="4C3D4D0CFFE6FFFDFD78FAA9FC34FAF5" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[740,900,1344,1362]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="2" phylum="Arthropoda" rank="species" species="impunctata" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE6FFFDFD78FAA9FD41FAF5" bold="true" box="[740,753,1344,1362]" italics="true" pageId="61" pageNumber="62">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE6FFFDFC98FAA9FCBFFAF5" bold="true" box="[772,783,1344,1362]" italics="true" pageId="61" pageNumber="62">E</emphasis>
|
||
.)
|
||
<emphasis id="B949EA9DFFE6FFFDFCBEFAA9FC34FAF5" bold="true" box="[802,900,1344,1362]" italics="true" pageId="61" pageNumber="62">impunctata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE6FFFDFC16FAA9FC63FAF5" box="[906,979,1344,1362]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; E:
|
||
<taxonomicName id="4C3D4D0CFFE6FFFDFC68FAA9FEACFACA" authorityName="C.Huber & J.Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="2" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE6FFFDFC68FAA9FBB1FAF5" bold="true" box="[1012,1025,1344,1362]" italics="true" pageId="61" pageNumber="62">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE6FFFDFB88FAA9FBAFFAF5" bold="true" box="[1044,1055,1344,1362]" italics="true" pageId="61" pageNumber="62">E</emphasis>
|
||
.)
|
||
<emphasis id="B949EA9DFFE6FFFDFF24FAB2FEACFACA" bold="true" box="[184,284,1371,1389]" italics="true" pageId="61" pageNumber="62">tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE6FFFDFEBCFAB2FED7FACA" box="[288,359,1371,1389]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; F:
|
||
<taxonomicName id="4C3D4D0CFFE6FFFDFE1EFAB2FDA9FACA" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[386,537,1371,1389]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="2" phylum="Arthropoda" rank="species" species="numburica" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE6FFFDFE1EFAB2FE3FFACA" bold="true" box="[386,399,1371,1389]" italics="true" pageId="61" pageNumber="62">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE6FFFDFE3DFAB2FE1CFACA" bold="true" box="[417,428,1371,1389]" italics="true" pageId="61" pageNumber="62">E</emphasis>
|
||
.)
|
||
<emphasis id="B949EA9DFFE6FFFDFE21FAB2FDA9FACA" bold="true" box="[445,537,1371,1389]" italics="true" pageId="61" pageNumber="62">numburica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE6FFFDFD82FAB2FDD5FACA" box="[542,613,1371,1389]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; G:
|
||
<taxonomicName id="4C3D4D0CFFE6FFFDFD19FAB2FC92FACA" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[645,802,1371,1389]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="2" phylum="Arthropoda" rank="species" species="retingensis" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE6FFFDFD19FAB2FD22FACA" bold="true" box="[645,658,1371,1389]" italics="true" pageId="61" pageNumber="62">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE6FFFDFD3FFAB2FD1EFACA" bold="true" box="[675,686,1371,1389]" italics="true" pageId="61" pageNumber="62">E</emphasis>
|
||
.)
|
||
<emphasis id="B949EA9DFFE6FFFDFD5CFAB2FC92FACA" bold="true" box="[704,802,1371,1389]" italics="true" pageId="61" pageNumber="62">retingensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE6FFFDFCBAFAB2FCDDFACA" box="[806,877,1371,1389]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; H:
|
||
<taxonomicName id="4C3D4D0CFFE6FFFDFC11FAB2FBBEFACA" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[909,1038,1371,1389]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="2" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE6FFFDFC11FAB2FC2AFACA" bold="true" box="[909,922,1371,1389]" italics="true" pageId="61" pageNumber="62">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE6FFFDFC37FAB2FC06FACA" bold="true" box="[939,950,1371,1389]" italics="true" pageId="61" pageNumber="62">E</emphasis>
|
||
.)
|
||
<emphasis id="B949EA9DFFE6FFFDFC54FAB2FBBEFACA" bold="true" box="[968,1038,1371,1389]" italics="true" pageId="61" pageNumber="62">delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE6FFFDFB8EFAB2FF50FA20" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; I:
|
||
<taxonomicName id="4C3D4D0CFFE6FFFDFF65FA9CFE3CFA20" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[249,396,1397,1415]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="2" phylum="Arthropoda" rank="species" species="incarinata" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE6FFFDFF65FA9CFEB6FA20" bold="true" box="[249,262,1397,1415]" italics="true" pageId="61" pageNumber="62">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE6FFFDFE84FA9CFE93FA20" bold="true" box="[280,291,1397,1415]" italics="true" pageId="61" pageNumber="62">E</emphasis>
|
||
.)
|
||
<emphasis id="B949EA9DFFE6FFFDFEA8FA9CFE3CFA20" bold="true" box="[308,396,1397,1415]" italics="true" pageId="61" pageNumber="62">incarinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE6FFFDFE0CFA9CFE68FA20" box="[400,472,1397,1415]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF426607FFE5FFFEFF41FF49FEFAFEA5" ID-DOI="http://doi.org/10.5281/zenodo.6285822" ID-Zenodo-Dep="6285822" httpUri="https://zenodo.org/record/6285822/files/figure.png" pageId="62" pageNumber="63" startId="62.[221,250,160,178]" targetBox="[463,1106,155,734]" targetPageId="62">
|
||
<paragraph id="8B82368FFFE5FFFEFF41FF49FEFAFEA5" blockId="62.[221,409,160,258]" pageId="62" pageNumber="63">
|
||
<emphasis id="B949EA9DFFE5FFFEFF41FF49FE29FF15" bold="true" box="[221,409,160,178]" pageId="62" pageNumber="63">Fig. 32: Gonocoxae of</emphasis>
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFF41FF52FE3FFF6A" authority="" authorityName="K. DANIEL" authorityYear="1904" box="[221,399,187,205]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE5FFFEFF41FF52FEA7FF6A" bold="true" box="[221,279,187,205]" italics="true" pageId="62" pageNumber="63">Nebria</emphasis>
|
||
(
|
||
<emphasis id="B949EA9DFFE5FFFEFEBFFF52FE37FF6A" bold="true" box="[291,391,187,205]" italics="true" pageId="62" pageNumber="63">Epinebriola</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B949EA9DFFE5FFFEFF41FF3FFEFAFEA5" bold="true" pageId="62" pageNumber="63">
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFF41FF3FFE93FF4F" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[221,291,214,232]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE5FFFEFF41FF3FFE93FF4F" bold="true" box="[221,291,214,232]" italics="true" pageId="62" pageNumber="63">delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE5FFFEFEB4FF3FFEDEFF4F" box="[296,366,214,232]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, in caudal view.
|
||
</emphasis>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B82368FFFE5FFFEFE9FFCD5FBFCFCDE" blockId="62.[221,1106,828,1598]" pageId="62" pageNumber="63">Body ratios: hea.w/fro.w = 1.38; prm.w/hea.w = 1.16; prm.w/prp.w = 1.47; pra.w/prp.w = 1.07; ely.w/prm.w = 1.71; prm.w/prm.l = 1.21; ely.l/ely.w = 1.69.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFE5FFFEFE9FFC6FFCEEFC63" pageId="62" pageNumber="63" type="etymology">
|
||
<paragraph id="8B82368FFFE5FFFEFE9FFC6FFCEEFC63" blockId="62.[221,1106,828,1598]" pageId="62" pageNumber="63">
|
||
Etymology: The specific epithet refers to the habitus and to the finely margined pronotum (
|
||
<emphasis id="B949EA9DFFE5FFFEFE3EFC45FDB9FC63" box="[418,521,940,964]" italics="true" pageId="62" pageNumber="63">delicatus</emphasis>
|
||
, latin = delicate, dainty, fine).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFE5FFFEFE9FFC38FCE5FA9E" pageId="62" pageNumber="63" type="diagnosis">
|
||
<paragraph id="8B82368FFFE5FFFEFE9FFC38FCE5FA9E" blockId="62.[221,1106,828,1598]" pageId="62" pageNumber="63">
|
||
Diagnosis:
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFE1DFC38FE4AFC4E" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[385,506,977,1001]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE5FFFEFE1DFC38FE4AFC4E" box="[385,506,977,1001]" italics="true" pageId="62" pageNumber="63">N. delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE5FFFEFD9CFC38FDE9FC4E" box="[512,601,977,1001]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is recognizable within the subgenus
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFC67FC38FEA1FBA9" authorityName="K. DANIEL" authorityYear="1904" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE5FFFEFC67FC38FEA1FBA9" italics="true" pageId="62" pageNumber="63">Epinebriola</emphasis>
|
||
</taxonomicName>
|
||
by its dainty habitus and by the anterior margin of the labrum with a row of eight setae instead of six. A labrum of eight setae is also present in
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFB9AFBF5FE93FBFE" authorityName="ANDREWES" authorityYear="1925" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="pindarica">
|
||
<emphasis id="B949EA9DFFE5FFFEFB9AFBF5FE93FBFE" italics="true" pageId="62" pageNumber="63">N. pindarica</emphasis>
|
||
</taxonomicName>
|
||
– distributed in
|
||
<collectingRegion id="49F9F86DFFE5FFFEFE42FBA8FDC1FBFE" box="[478,625,1089,1113]" country="India" name="Uttarakhand" pageId="62" pageNumber="63">Uttarakhand</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32A761FFFE5FFFEFDE7FBA8FD03FBFE" box="[635,691,1089,1113]" name="India" pageId="62" pageNumber="63">India</collectingCountry>
|
||
–, a large species of
|
||
<quantity id="4CC59B6AFFE5FFFEFC38FBA8FBAAFBFE" box="[932,1050,1089,1113]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="62" pageNumber="63" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 mm</quantity>
|
||
with the lateral margin of the pronotum distinctly concavely narrowed in front of the posterior angle and with hanging shoulders, whereas in the small (
|
||
<quantity id="4CC59B6AFFE5FFFEFB95FB65FBFBFB03" box="[1033,1099,1164,1188]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="62" pageNumber="63" unit="mm" value="9.0">9 mm</quantity>
|
||
) species
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFEA7FB58FE06FB6E" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[315,438,1201,1225]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE5FFFEFEA7FB58FE06FB6E" box="[315,438,1201,1225]" italics="true" pageId="62" pageNumber="63">N. delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE5FFFEFE22FB58FDA8FB6E" box="[446,536,1201,1225]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
|
||
the lateral margin of the pronotum is only faintly concave in front of the posterior angle, and the shoulders are pronounced.
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFBA5FB3FFE87FAB3" authorityName="C.Huber & J.Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE5FFFEFBA5FB3FFE87FAB3" italics="true" pageId="62" pageNumber="63">N. delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE5FFFEFEA3FB15FE28FAB3" box="[319,408,1276,1300]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is closely related to
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFD1FFB15FC91FAB3" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[643,801,1276,1300]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="retingensis" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE5FFFEFD1FFB15FC91FAB3" box="[643,801,1276,1300]" italics="true" pageId="62" pageNumber="63">N. retingensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE5FFFEFCB4FB15FC31FAB3" box="[808,897,1276,1300]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
|
||
due to the resemblance in the shape of the median lobes (
|
||
<figureCitation id="13062A0AFFE5FFFEFD35FAC8FCF8FA9E" box="[681,840,1313,1337]" captionStart="Fig" captionStartId="38.[221,250,1317,1335]" captionTargetBox="[221,1106,153,1296]" captionTargetId="figure-70@38.[221,704,627,867]" captionTargetPageId="38" captionText="Fig. 20: Male edeagi of new Nebria species of the subgenus Epinebriola K. DANIEL, 1904. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) numburica sp. nov., with parameres; F: N. (E.) retingensis sp. nov.; G: N. (E.) incarinata sp. nov.; H: N. (E.) delicata sp. nov.; I: N. (E.) incarinata sp. nov., apex in dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6285764" httpUri="https://zenodo.org/record/6285764/files/figure.png" pageId="62" pageNumber="63">Figs 20F, 20H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFE5FFFEFE9FFAAFFD67F999" pageId="62" pageNumber="63" type="discussion">
|
||
<paragraph id="8B82368FFFE5FFFEFE9FFAAFFD67F999" blockId="62.[221,1106,828,1598]" pageId="62" pageNumber="63">
|
||
Remark: Generally the species of the genus
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFD72FAAFFC88FAF9" authorityName="LATREILLE" authorityYear="1802" box="[750,824,1350,1374]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949EA9DFFE5FFFEFD72FAAFFC88FAF9" box="[750,824,1350,1374]" italics="true" pageId="62" pageNumber="63">Nebria</emphasis>
|
||
</taxonomicName>
|
||
have the anterior margin of the labrum with a row of six apical setae with the exception of
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFC58FA85FECBFA0E" authorityName="ANDREWES" authorityYear="1925" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="pindarica" subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE5FFFEFC58FA85FC65FA23" box="[964,981,1388,1412]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE5FFFEFC71FA85FEB0FA0E" italics="true" pageId="62" pageNumber="63">Epinebriola</emphasis>
|
||
)
|
||
<emphasis id="B949EA9DFFE5FFFEFE8EFA78FECBFA0E" box="[274,379,1425,1449]" italics="true" pageId="62" pageNumber="63">pindarica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFE16FA78FCF7FA0E" authority="K. Daniel, 1910" authorityName="K. Daniel" authorityYear="1910" box="[394,839,1425,1449]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="eugeniae" subGenus="Tyrrhenia">
|
||
<emphasis id="B949EA9DFFE5FFFEFE16FA78FE2BFA0E" box="[394,411,1425,1449]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE5FFFEFE2EFA78FDABFA0E" box="[434,539,1425,1449]" italics="true" pageId="62" pageNumber="63">Tyrrhenia</emphasis>
|
||
)
|
||
<emphasis id="B949EA9DFFE5FFFEFDB0FA78FD24FA0E" box="[556,660,1425,1449]" italics="true" pageId="62" pageNumber="63">eugeniae</emphasis>
|
||
K. Daniel, 1910
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFC15FA78FE51FA69" authority="Peyerimhoff, 1910" authorityName="Peyerimhoff" authorityYear="1910" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="exul" subGenus="Alpaeonebria">
|
||
<emphasis id="B949EA9DFFE5FFFEFC15FA78FC2AFA0E" box="[905,922,1425,1449]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE5FFFEFC2FFA78FBF9FA0E" box="[947,1097,1425,1449]" italics="true" pageId="62" pageNumber="63">Alpaeonebria</emphasis>
|
||
)
|
||
<emphasis id="B949EA9DFFE5FFFEFF41FA5FFEBCFA69" box="[221,268,1462,1486]" italics="true" pageId="62" pageNumber="63">exul</emphasis>
|
||
Peyerimhoff, 1910
|
||
</taxonomicName>
|
||
which all have eight setae, and of
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFCF3FA5FFE7DFA53" authority="Ledoux & Roux, 1998" authorityName="Ledoux & Roux" authorityYear="1998" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="lucifer" subGenus="Eonebria">
|
||
<emphasis id="B949EA9DFFE5FFFEFCF3FA5FFC30FA69" box="[879,896,1462,1486]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE5FFFEFC04FA5FFC4BFA69" box="[920,1019,1462,1486]" italics="true" pageId="62" pageNumber="63">Eonebria</emphasis>
|
||
)
|
||
<emphasis id="B949EA9DFFE5FFFEFB91FA5FFBE2FA69" box="[1037,1106,1462,1486]" italics="true" pageId="62" pageNumber="63">lucifer</emphasis>
|
||
Ledoux & Roux, 1998
|
||
</taxonomicName>
|
||
with ten setae (
|
||
<bibRefCitation id="EFAC4B7EFFE5FFFEFDE2FA35FCDAFA53" author="Ledoux, G. & Roux, P." box="[638,874,1500,1524]" pageId="62" pageNumber="63" refId="ref33536" refString="Ledoux, G. & Roux, P. (2005): Nebria (Coleoptera, Nebriidae) Faune mondiale. - 976 pp., Saint-Just-la-Pendue." type="book" year="2005">Ledoux & Roux 2005</bibRefCitation>
|
||
). Obviously the evolution to eight or more labral setae happened convergently several times in different subgenera within the genus
|
||
<taxonomicName id="4C3D4D0CFFE5FFFEFD18F9CFFD60F999" authorityName="LATREILLE" authorityYear="1802" box="[644,720,1574,1598]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949EA9DFFE5FFFEFD18F9CFFD60F999" box="[644,720,1574,1598]" italics="true" pageId="62" pageNumber="63">Nebria</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF426607FFE4FFFFFF24FCA9FD64FC20" ID-DOI="http://doi.org/10.5281/zenodo.6285826" ID-Zenodo-Dep="6285826" httpUri="https://zenodo.org/record/6285826/files/figure.png" pageId="63" pageNumber="64" startId="63.[184,213,832,850]" targetBox="[184,1068,153,811]" targetPageId="63">
|
||
<paragraph id="8B82368FFFE4FFFFFF24FCA9FD64FC20" blockId="63.[184,1068,832,903]" pageId="63" pageNumber="64">
|
||
Fig. 33: Habitat of
|
||
<taxonomicName id="4C3D4D0CFFE4FFFFFEC3FCA9FDEFFCF5" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[351,607,832,850]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE4FFFFFEC3FCA9FE29FCF5" bold="true" box="[351,409,832,850]" italics="true" pageId="63" pageNumber="64">Nebria</emphasis>
|
||
(
|
||
<emphasis id="B949EA9DFFE4FFFFFE3BFCA9FDBBFCF5" bold="true" box="[423,523,832,850]" italics="true" pageId="63" pageNumber="64">Epinebriola</emphasis>
|
||
)
|
||
<emphasis id="B949EA9DFFE4FFFFFD85FCA9FDEFFCF5" bold="true" box="[537,607,832,850]" italics="true" pageId="63" pageNumber="64">delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE4FFFFFDF9FCA9FD1EFCF5" box="[613,686,832,850]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
: Humid slope near mountain crest, 5400 m a.s.l., below big stones (white arrow) on the Mendju Zari Mountain SE of the Lhasa Valley in the central Transhimalaya (Gangdise Shan). Photo: Joachim Schmidt.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3276504FFE4FFFFFF41FC38FB94FBA9" pageId="63" pageNumber="64" type="distribution">
|
||
<paragraph id="8B82368FFFE4FFFFFF41FC38FB94FBA9" blockId="63.[184,1070,977,1636]" pageId="63" pageNumber="64">
|
||
Distribution (
|
||
<figureCitation id="13062A0AFFE4FFFFFEEFFC38FE72FC4E" box="[371,450,977,1001]" captionStart="Fig" captionStartId="76.[221,250,160,178]" captionTargetId="figure-8@75.[184,1068,152,1645]" captionTargetPageId="75" captionText="Fig. 41: Locations with occurrences of Nebria (Epinebriola) species from the Central and Eastern Himalaya and from South Tibet." figureDoi="http://doi.org/10.5281/zenodo.6285892" httpUri="https://zenodo.org/record/6285892/files/figure.png" pageId="63" pageNumber="64">Fig. 41</figureCitation>
|
||
): Known only from the
|
||
<typeStatus id="5486882DFFE4FFFFFD4BFC38FCBAFC4E" box="[727,778,977,1001]" pageId="63" pageNumber="64">type</typeStatus>
|
||
locality, the Mendju Zari Mountain SE of the Lhasa Valley in the central Transhimalaya (Gangdise Shan).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFE4FFFFFF41FBF5FDE0FBFE" pageId="63" pageNumber="64" type="biology_ecology">
|
||
<paragraph id="8B82368FFFE4FFFFFF41FBF5FDE0FBFE" blockId="63.[184,1070,977,1636]" pageId="63" pageNumber="64">
|
||
Habitat (
|
||
<figureCitation id="13062A0AFFE4FFFFFEDDFBF5FE22FB93" box="[321,402,1052,1076]" captionStart="Fig" captionStartId="63.[184,213,832,850]" captionTargetBox="[184,1068,153,811]" captionTargetId="figure-260@63.[184,1068,151,811]" captionTargetPageId="63" captionText="Fig. 33: Habitat of Nebria (Epinebriola) delicata sp. nov.: Humid slope near mountain crest, 5400 m a.s.l., below big stones (white arrow) on the Mendju Zari Mountain SE of the Lhasa Valley in the central Transhimalaya (Gangdise Shan). Photo: Joachim Schmidt." figureDoi="http://doi.org/10.5281/zenodo.6285826" httpUri="https://zenodo.org/record/6285826/files/figure.png" pageId="63" pageNumber="64">Fig. 33</figureCitation>
|
||
): Both specimens of the
|
||
<typeStatus id="5486882DFFE4FFFFFD21FBF5FD40FB93" box="[701,752,1052,1076]" pageId="63" pageNumber="64">type</typeStatus>
|
||
series were collected on a humid slope in the high alpine zone.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFE4FF80FF41FB8FFE8BFA0E" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64" type="discussion">
|
||
<paragraph id="8B82368FFFE4FF80FF41FB8FFE8BFA0E" blockId="63.[184,1070,977,1636]" lastBlockId="64.[221,1107,1164,1449]" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">
|
||
Remark (see
|
||
<figureCitation id="13062A0AFFE4FFFFFEE5FB8FFE79FBD9" box="[377,457,1126,1150]" captionStart="Fig" captionStartId="64.[221,251,1018,1036]" captionTargetBox="[221,992,153,958]" captionTargetId="figure-110@64.[423,633,150,988]" captionTargetPageId="64" captionText="Fig. 34: Right gonocoxa, in ventral view. A: Nebria (Epinebriola) retingensis sp. nov. (= Fig. 10J); B: N. (Eonebria) kurentzovi LAFER, 1989; C: N. (Eonebria) gemina LEDOUx, ROUx & WRASE, 1996; D: N. (Eonebria) amabilis LEDOUx, ROUx & SAWADA, 1991. Scale bar = 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285830" httpUri="https://zenodo.org/record/6285830/files/figure.png" pageId="63" pageNumber="64">Fig. 34</figureCitation>
|
||
): The gonocoxites 2 of
|
||
<taxonomicName id="4C3D4D0CFFE4FFFFFD7BFB8FFB9CFBD9" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[743,1068,1126,1150]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="retingensis" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE4FFFFFD7BFB8FFD48FBD9" box="[743,760,1126,1150]" italics="true" pageId="63" pageNumber="64">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE4FFFFFC8EFB8FFC27FBD9" box="[786,919,1126,1150]" italics="true" pageId="63" pageNumber="64">Epinebriola</emphasis>
|
||
)
|
||
<emphasis id="B949EA9DFFE4FFFFFC37FB8FFB9CFBD9" box="[939,1068,1126,1150]" italics="true" pageId="63" pageNumber="64">retingensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE4FFFFFF24FB65FEA0FB03" box="[184,272,1164,1188]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13062A0AFFE4FFFFFE82FB65FEC7FB03" box="[286,375,1164,1188]" captionStart="Fig" captionStartId="20.[221,250,1430,1448]" captionTargetBox="[228,1106,153,1365]" captionTargetId="figure-16@20.[574,757,751,1260]" captionTargetPageId="20" captionText="Fig. 10: Right gonocoxae of Epinebriola species, in ventral view. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) tuberculata sp. nov.; F: N. (E.) numburica sp. nov.; G: N. (E.) numburica sp. nov. in lateral view with the distale gonocoxite 2 dorsally grooved (arrow); H: N. (E.) christinae HUBER & SCHMIDT, 2007; I: N. (E.) pseudorestias sp. nov.; J: N. (E.) retingensis sp. nov.; Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285726" httpUri="https://zenodo.org/record/6285726/files/figure.png" pageId="63" pageNumber="64">Fig. 10J</figureCitation>
|
||
,
|
||
<figureCitation id="13062A0AFFE4FFFFFE1DFB65FE01FB03" box="[385,433,1164,1188]" captionStart="Fig" captionStartId="64.[221,251,1018,1036]" captionTargetBox="[221,992,153,958]" captionTargetId="figure-110@64.[423,633,150,988]" captionTargetPageId="64" captionText="Fig. 34: Right gonocoxa, in ventral view. A: Nebria (Epinebriola) retingensis sp. nov. (= Fig. 10J); B: N. (Eonebria) kurentzovi LAFER, 1989; C: N. (Eonebria) gemina LEDOUx, ROUx & WRASE, 1996; D: N. (Eonebria) amabilis LEDOUx, ROUx & SAWADA, 1991. Scale bar = 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285830" httpUri="https://zenodo.org/record/6285830/files/figure.png" pageId="63" pageNumber="64">34A</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C3D4D0CFFE4FFFFFE6CFB65FCB5FB03" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[496,773,1164,1188]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE4FFFFFE6CFB65FDB1FB03" box="[496,513,1164,1188]" italics="true" pageId="63" pageNumber="64">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE4FFFFFD85FB65FD2BFB03" box="[537,667,1164,1188]" italics="true" pageId="63" pageNumber="64">Epinebriola</emphasis>
|
||
)
|
||
<emphasis id="B949EA9DFFE4FFFFFD37FB65FCB5FB03" box="[683,773,1164,1188]" italics="true" pageId="63" pageNumber="64">delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE4FFFFFC90FB65FCD5FB03" box="[780,869,1164,1188]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13062A0AFFE4FFFFFCEEFB65FC70FB03" box="[882,960,1164,1188]" captionStart="Fig" captionStartId="62.[221,250,160,178]" captionTargetBox="[463,1106,155,734]" captionTargetId="figure-330@62.[463,1106,153,737]" captionTargetPageId="62" captionText="Fig. 32: Gonocoxae of Nebria (Epinebriola) delicata sp. nov., in caudal view." figureDoi="http://doi.org/10.5281/zenodo.6285822" httpUri="https://zenodo.org/record/6285822/files/figure.png" pageId="63" pageNumber="64">Fig. 32</figureCitation>
|
||
) are conspicuous in length and shape. Such long and dagger-shaped gonocoxites 2 are neither described nor mentioned in relevant
|
||
<taxonomicName id="4C3D4D0CFFE4FFFFFD71FB3FFC89FB49" authorityName="LATREILLE" authorityYear="1802" box="[749,825,1238,1262]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949EA9DFFE4FFFFFD71FB3FFC89FB49" box="[749,825,1238,1262]" italics="true" pageId="63" pageNumber="64">Nebria</emphasis>
|
||
</taxonomicName>
|
||
literature (
|
||
<bibRefCitation id="EFAC4B7EFFE4FFFFFC26FB3FFE85FAB3" author="Ledoux, G. & Roux, P." pageId="63" pageNumber="64" refId="ref33536" refString="Ledoux, G. & Roux, P. (2005): Nebria (Coleoptera, Nebriidae) Faune mondiale. - 976 pp., Saint-Just-la-Pendue." type="book" year="2005">Ledoux & Roux 2005</bibRefCitation>
|
||
). In the present case the gonocoxite 2 of
|
||
<taxonomicName id="4C3D4D0CFFE4FFFFFCBAFB15FC78FAB3" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[806,968,1276,1300]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="retingensis" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFE4FFFFFCBAFB15FC78FAB3" box="[806,968,1276,1300]" italics="true" pageId="63" pageNumber="64">N. retingensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE4FFFFFC4DFB15FB9DFAB3" box="[977,1069,1276,1300]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13062A0AFFE4FFFFFF23FAC8FE91FA9E" box="[191,289,1313,1337]" captionStart="Fig" captionStartId="64.[221,251,1018,1036]" captionTargetBox="[221,992,153,958]" captionTargetId="figure-110@64.[423,633,150,988]" captionTargetPageId="64" captionText="Fig. 34: Right gonocoxa, in ventral view. A: Nebria (Epinebriola) retingensis sp. nov. (= Fig. 10J); B: N. (Eonebria) kurentzovi LAFER, 1989; C: N. (Eonebria) gemina LEDOUx, ROUx & WRASE, 1996; D: N. (Eonebria) amabilis LEDOUx, ROUx & SAWADA, 1991. Scale bar = 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285830" httpUri="https://zenodo.org/record/6285830/files/figure.png" pageId="63" pageNumber="64">Fig. 34A</figureCitation>
|
||
) is compared with those of some species of the subgenus
|
||
<taxonomicName id="4C3D4D0CFFE4FFFFFC5BFAC8FE5DFAF9" authority="SEMENOV & ZNOJKO, 1928" authorityName="SEMENOV & ZNOJKO" authorityYear="1928" class="Insecta" family="Carabidae" genus="Eonebria" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949EA9DFFE4FFFFFC5BFAC8FB9CFA9E" box="[967,1068,1313,1337]" italics="true" pageId="63" pageNumber="64">Eonebria</emphasis>
|
||
SEMENOV & ZNOJKO, 1928
|
||
</taxonomicName>
|
||
. The gonocoxites 2 of
|
||
<taxonomicName id="4C3D4D0CFFE4FFFFFC99FAAFFEF4FA23" authority="LAFER, 1989" authorityName="LAFER" authorityYear="1989" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="kurentzovi" subGenus="Eonebria">
|
||
<emphasis id="B949EA9DFFE4FFFFFC99FAAFFCA6FAF9" box="[773,790,1350,1374]" italics="true" pageId="63" pageNumber="64">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE4FFFFFCAEFAAFFC2BFAF9" box="[818,923,1350,1374]" italics="true" pageId="63" pageNumber="64">Eonebria</emphasis>
|
||
)
|
||
<emphasis id="B949EA9DFFE4FFFFFC33FAAFFB9CFAF9" box="[943,1068,1350,1374]" italics="true" pageId="63" pageNumber="64">kurentzovi</emphasis>
|
||
LAFER, 1989
|
||
</taxonomicName>
|
||
(Anisimovka, Primorsk. Kr. Litovka, Far East
|
||
<collectingCountry id="F32A761FFFE4FFFFFCD0FA85FC2CFA23" box="[844,924,1388,1412]" name="Russia" pageId="63" pageNumber="64">Russia</collectingCountry>
|
||
; coll. NMBE.
|
||
<figureCitation id="13062A0AFFE4FFFFFF24FA78FEA5FA0E" box="[184,277,1425,1449]" captionStart="Fig" captionStartId="64.[221,251,1018,1036]" captionTargetBox="[221,992,153,958]" captionTargetId="figure-110@64.[423,633,150,988]" captionTargetPageId="64" captionText="Fig. 34: Right gonocoxa, in ventral view. A: Nebria (Epinebriola) retingensis sp. nov. (= Fig. 10J); B: N. (Eonebria) kurentzovi LAFER, 1989; C: N. (Eonebria) gemina LEDOUx, ROUx & WRASE, 1996; D: N. (Eonebria) amabilis LEDOUx, ROUx & SAWADA, 1991. Scale bar = 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285830" httpUri="https://zenodo.org/record/6285830/files/figure.png" pageId="63" pageNumber="64">Fig. 34B</figureCitation>
|
||
) and of
|
||
<taxonomicName id="4C3D4D0CFFE4FFFFFEEFFA78FDD3FA0E" authorityName="Ledoux & Roux&Wrase" authorityYear="1996" box="[371,611,1425,1449]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="gemina" subGenus="Eonebria">
|
||
<emphasis id="B949EA9DFFE4FFFFFEEFFA78FE34FA0E" box="[371,388,1425,1449]" italics="true" pageId="63" pageNumber="64">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE4FFFFFE07FA78FE4EFA0E" box="[411,510,1425,1449]" italics="true" pageId="63" pageNumber="64">Eonebria</emphasis>
|
||
)
|
||
<emphasis id="B949EA9DFFE4FFFFFD93FA78FDD3FA0E" box="[527,611,1425,1449]" italics="true" pageId="63" pageNumber="64">gemina</emphasis>
|
||
</taxonomicName>
|
||
LEDOUx, ROUx & WRASE, 1996 (Qin Ling Shan, Tabai Shan,
|
||
<collectingRegion id="49F9F86DFFE4FFFFFE17FA5FFE58FA69" box="[395,488,1462,1486]" country="China" name="Shaanxi" pageId="63" pageNumber="64">Shaanxi</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32A761FFFE4FFFFFE6FFA5FFD88FA69" box="[499,568,1462,1486]" name="China" pageId="63" pageNumber="64">China</collectingCountry>
|
||
; coll. NMBE.
|
||
<figureCitation id="13062A0AFFE4FFFFFD51FA5FFC99FA69" box="[717,809,1462,1486]" captionStart="Fig" captionStartId="54.[221,250,832,850]" captionTargetBox="[221,1105,153,808]" captionTargetId="figure-185@54.[221,1105,153,812]" captionTargetPageId="54" captionText="Fig. 27: Habitat of Nebria (Epinebriola) numburica sp. nov.: Headwater of the Dudhkund Khola, 4100 m a.s.l., subalpine zone. Specimens were collected alongside the stream. Photo: Joachim Schmidt." figureDoi="http://doi.org/10.5281/zenodo.6285800" httpUri="https://zenodo.org/record/6285800/files/figure.png" pageId="63" pageNumber="64">Fig. 27C</figureCitation>
|
||
) look similar to that of
|
||
<taxonomicName id="4C3D4D0CFFE4FFFFFF24FA35FE5AFA53" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[184,490,1500,1524]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="retingensis" status="sp. nov." subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFFE4FFFFFF24FA35FF79FA53" box="[184,201,1500,1524]" italics="true" pageId="63" pageNumber="64">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE4FFFFFF43FA35FEEEFA53" box="[223,350,1500,1524]" italics="true" pageId="63" pageNumber="64">Epinebriola</emphasis>
|
||
)
|
||
<emphasis id="B949EA9DFFE4FFFFFEF2FA35FE5AFA53" box="[366,490,1500,1524]" italics="true" pageId="63" pageNumber="64">retingensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFE4FFFFFE73FA35FDFAFA53" box="[495,586,1500,1524]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, whereas in
|
||
<taxonomicName id="4C3D4D0CFFE4FFFFFD4DFA35FC7AFA53" authorityName="Ledoux; Roux & Sawada" authorityYear="1991" box="[721,970,1500,1524]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="amabilis" subGenus="Eonebria">
|
||
<emphasis id="B949EA9DFFE4FFFFFD4DFA35FD52FA53" box="[721,738,1500,1524]" italics="true" pageId="63" pageNumber="64">N</emphasis>
|
||
. (
|
||
<emphasis id="B949EA9DFFE4FFFFFD64FA35FCEBFA53" box="[760,859,1500,1524]" italics="true" pageId="63" pageNumber="64">Eonebria</emphasis>
|
||
)
|
||
<emphasis id="B949EA9DFFE4FFFFFCF7FA35FC7AFA53" box="[875,970,1500,1524]" italics="true" pageId="63" pageNumber="64">amabilis</emphasis>
|
||
</taxonomicName>
|
||
LEDOUx, ROUx & SAWADA, 1991 (Heka,
|
||
<collectingRegion id="49F9F86DFFE4FFFFFD8DF9E8FDDBF9BE" box="[529,619,1537,1561]" country="China" name="Qinghai" pageId="63" pageNumber="64">Qinghai</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32A761FFFE4FFFFFDEAF9E8FD0BF9BE" box="[630,699,1537,1561]" name="China" pageId="63" pageNumber="64">China</collectingCountry>
|
||
; coll. NMBE.
|
||
<figureCitation id="13062A0AFFE4FFFFFCD2F9E8FC1DF9BE" box="[846,941,1537,1561]" captionStart="Fig" captionStartId="64.[221,251,1018,1036]" captionTargetBox="[221,992,153,958]" captionTargetId="figure-110@64.[423,633,150,988]" captionTargetPageId="64" captionText="Fig. 34: Right gonocoxa, in ventral view. A: Nebria (Epinebriola) retingensis sp. nov. (= Fig. 10J); B: N. (Eonebria) kurentzovi LAFER, 1989; C: N. (Eonebria) gemina LEDOUx, ROUx & WRASE, 1996; D: N. (Eonebria) amabilis LEDOUx, ROUx & SAWADA, 1991. Scale bar = 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285830" httpUri="https://zenodo.org/record/6285830/files/figure.png" pageId="63" pageNumber="64">Fig. 34D</figureCitation>
|
||
) the gonocoxite 2 is shaped in a rather "normal", not dagger-like way. The subgenus
|
||
<taxonomicName id="4C3D4D0CFFE4FFFFFF24F9A5FEADF9C3" authorityName="SEMENOV & ZNOJKO" authorityYear="1928" box="[184,285,1612,1636]" class="Insecta" family="Carabidae" genus="Eonebria" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949EA9DFFE4FFFFFF24F9A5FEADF9C3" box="[184,285,1612,1636]" italics="true" pageId="63" pageNumber="64">Eonebria</emphasis>
|
||
</taxonomicName>
|
||
currently consists of 74 taxa, and the selected three taxa may not be representative for the whole subgenus. But the gonocoxites 2 of the selected
|
||
<taxonomicName id="4C3D4D0CFF9BFF80FF41FB58FE2FFB6E" box="[221,415,1201,1225]" class="Insecta" family="Carabidae" genus="Eonebria" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B949EA9DFF9BFF80FF41FB58FEF2FB6E" box="[221,322,1201,1225]" italics="true" pageId="64" pageNumber="65">Eonebria</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
may point to a certain variability of the gonocoxite shape as in the subgenus
|
||
<taxonomicName id="4C3D4D0CFF9BFF80FE05FB3FFDABFB49" authorityName="K. DANIEL" authorityYear="1904" box="[409,539,1238,1262]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFF9BFF80FE05FB3FFDABFB49" box="[409,539,1238,1262]" italics="true" pageId="64" pageNumber="65">Epinebriola</emphasis>
|
||
</taxonomicName>
|
||
. A re-evaluation of the subgenus
|
||
<taxonomicName id="4C3D4D0CFF9BFF80FC05FB3FFC4EFB49" authorityName="SEMENOV & ZNOJKO" authorityYear="1928" box="[921,1022,1238,1262]" class="Insecta" family="Carabidae" genus="Eonebria" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949EA9DFF9BFF80FC05FB3FFC4EFB49" box="[921,1022,1238,1262]" italics="true" pageId="64" pageNumber="65">Eonebria</emphasis>
|
||
</taxonomicName>
|
||
including the disregarded gonocoxal characters may be required, but such an analysis was not the objective of the present study. For now we leave
|
||
<taxonomicName id="4C3D4D0CFF9BFF80FC13FAC8FB9AFA9E" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[911,1066,1313,1337]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="retingensis" status="sp. nov.">
|
||
<emphasis id="B949EA9DFF9BFF80FC13FAC8FB9AFA9E" box="[911,1066,1313,1337]" italics="true" pageId="64" pageNumber="65">N. retingensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FF9BFF80FBADFAC8FEBDFAF9" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C3D4D0CFF9BFF80FED8FAAFFE0CFAF9" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[324,444,1350,1374]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="delicata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFF9BFF80FED8FAAFFE0CFAF9" box="[324,444,1350,1374]" italics="true" pageId="64" pageNumber="65">N. delicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FF9BFF80FE5FFAAFFDAAFAF9" box="[451,538,1350,1374]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
within the subgenus
|
||
<taxonomicName id="4C3D4D0CFF9BFF80FC90FAAFFC3BFAF9" authorityName="K. DANIEL" authorityYear="1904" box="[780,907,1350,1374]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
|
||
<emphasis id="B949EA9DFF9BFF80FC90FAAFFC3BFAF9" box="[780,907,1350,1374]" italics="true" pageId="64" pageNumber="65">Epinebriola</emphasis>
|
||
</taxonomicName>
|
||
, and we interpret the dagger-like gonocoxite 2 as being convergent to those of some
|
||
<taxonomicName id="4C3D4D0CFF9BFF80FC71FA85FE8BFA0E" class="Insecta" family="Carabidae" genus="Eonebria" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B949EA9DFF9BFF80FC71FA85FBE2FA23" box="[1005,1106,1388,1412]" italics="true" pageId="64" pageNumber="65">Eonebria</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |