211 lines
18 KiB
XML
211 lines
18 KiB
XML
<document ID-DOI="10.1093/isd/ixab025" ID-ISSN="2399-3421" approvalRequired="23" approvalRequired_for_textStreams="22" approvalRequired_for_treatments="1" checkinTime="1665268061086" checkinUser="felipe" docAuthor="Ward, Philip S. & Branstetter, Michael G." docDate="2022" docId="0394878EFFF7FFDAB831FB3ABCD1FC3F" docLanguage="en" docName="InsectSystDivers.6.1.4.1-31.pdf" docOrigin="Insect Systematics and Diversity (AIFB) 6 (1)" docSource="http://dx.doi.org/10.1093/isd/ixab025" docStyle="DocumentStyle:20C34595BA01015AC07A3B955E96DF74.5:InsectSystDivers.2017-.journal_article" docStyleId="20C34595BA01015AC07A3B955E96DF74" docStyleName="InsectSystDivers.2017-.journal_article" docStyleVersion="5" docTitle="Pseudomyrmex ereptor Ward & Branstetter 2022, sp. nov." docType="treatment" docVersion="1" lastPageNumber="24" masterDocId="FFADFFF6FFE1FFCDBB03FFBDBE0EFF97" masterDocTitle="Species Paraphyly and Social Parasitism: Phylogenomics, Morphology, and Geography Clarify the Evolution of the Pseudomyrmex elongatulus Group (Hymenoptera:" masterLastPageNumber="31" masterPageNumber="1" pageNumber="23" updateTime="1665421853293" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Species Paraphyly and Social Parasitism: Phylogenomics, Morphology, and Geography Clarify the Evolution of the Pseudomyrmex elongatulus Group (Hymenoptera:</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ward, Philip S.</mods:namePart>
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<mods:affiliation>Department of Entomology and Nematology, University of California, Davis, CA 95616, USA,</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Branstetter, Michael G.</mods:namePart>
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<mods:affiliation>U. S. Department of Agriculture,</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Insect Systematics and Diversity</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="series">
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<mods:title>AIFB</mods:title>
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</mods:detail>
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<mods:detail type="pubDate">
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<mods:number>2022-01-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>6</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>31</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.1093/isd/ixab025</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1093/isd/ixab025</mods:identifier>
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<mods:identifier type="ISSN">2399-3421</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:0394878EFFF7FFDAB831FB3ABCD1FC3F" httpUri="http://treatment.plazi.org/id/0394878EFFF7FFDAB831FB3ABCD1FC3F" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<subSubSection pageId="22" pageNumber="23" type="nomenclature">
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<paragraph blockId="22.[818,1107,1159,1179]" box="[818,1107,1159,1179]" pageId="22" pageNumber="23">
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<heading box="[818,1107,1159,1179]" fontSize="8" level="2" pageId="22" pageNumber="23" reason="4">
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<emphasis bold="true" box="[818,1107,1159,1179]" pageId="22" pageNumber="23">
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<taxonomicName authority="Ward & Branstetter, 2022" authorityName="Ward & Branstetter" authorityYear="2022" box="[818,1032,1159,1178]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ereptor" status="sp. nov.">
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<emphasis bold="true" box="[818,1032,1159,1178]" italics="true" pageId="22" pageNumber="23">Pseudomyrmex ereptor</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1038,1107,1159,1179]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="22.[817,1443,1188,1501]" box="[818,960,1188,1208]" pageId="22" pageNumber="23">Figs. 12 and 24</paragraph>
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</subSubSection>
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<subSubSection lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
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<paragraph blockId="22.[817,1443,1188,1501]" pageId="22" pageNumber="23">
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Zoobank LSID:
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<uri pageId="22" pageNumber="23">
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urn:lsid:zoobank.org:act:
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<uuid pageId="22" pageNumber="23">3F4584C3-EB0C-4C74- B067-01CBEDC8F4EB</uuid>
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</uri>
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</paragraph>
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<paragraph blockId="22.[817,1443,1188,1501]" pageId="22" pageNumber="23">
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<materialsCitation accessionNumber="PSW07360" collectingDate="1985-03-21" collectionCode="UNAM" collectorName="P. S. Ward" elevation="500" latitude="18.583334" location="Los Tuxtlas" longLatPrecision="1275" longitude="-95.083336" pageId="22" pageNumber="23" specimenCode="CASENT0863524" specimenCount="1" stateProvince="MEXICO Veracruz" typeStatus="holotype">
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<typeStatus box="[850,937,1276,1296]" pageId="22" pageNumber="23">Holotype</typeStatus>
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alate queen,
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<collectingRegion box="[1064,1248,1276,1296]" isEnumeration="true" pageId="22" pageNumber="23">
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<collectingRegion box="[1064,1155,1276,1296]" country="Mexico" name="Mexico" pageId="22" pageNumber="23">MEXICO</collectingRegion>
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<collectingRegion box="[1163,1248,1277,1296]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="22" pageNumber="23">Veracruz</collectingRegion>
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</collectingRegion>
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:
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<location LSID="urn:lsid:plazi:treatment:0394878EFFF7FFDAB831FB3ABCD1FC3F:8EE26043FFF7FFDBBFE9FB40BB57FA87" box="[1258,1369,1276,1296]" latitude="18.583334" longLatPrecision="1275" longitude="-95.083336" name="Los Tuxtlas" pageId="22" pageNumber="23" stateProvince="MEXICO Veracruz">Los Tuxtlas</location>
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,
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<locationDeviation location="Sontecomapan" pageId="22" pageNumber="23">
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<quantity box="[1380,1442,1276,1296]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="23" unit="km" value="10.0">10 km</quantity>
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NNW Sontecomapan
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</locationDeviation>
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,
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<quantity box="[1029,1090,1305,1325]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="m" value="500.0">
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<elevation box="[1029,1090,1305,1325]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="m" value="500.0">500 m</elevation>
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</quantity>
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,
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<geoCoordinate box="[1101,1195,1305,1325]" degrees="18" direction="north" minutes="35" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="18.583334">
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18° 35
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<emphasis box="[1164,1170,1305,1325]" italics="true" pageId="22" pageNumber="23">′</emphasis>
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N
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</geoCoordinate>
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,
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<geoCoordinate box="[1206,1302,1305,1325]" degrees="95" direction="west" minutes="05" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="-95.083336">
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95° 05
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<emphasis box="[1269,1275,1305,1325]" italics="true" pageId="22" pageNumber="23">′</emphasis>
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W
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</geoCoordinate>
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,
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<date box="[1311,1438,1305,1325]" pageId="22" pageNumber="23" value="1985-03-21">
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<collectingDate box="[1311,1438,1305,1325]" pageId="22" pageNumber="23" value="1985-03-21">21 Mar 1985</collectingDate>
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</date>
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, ex dead twig of liana [in nest of
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<taxonomicName baseAuthorityName="Dalle Torre" baseAuthorityYear="1892" box="[1107,1234,1335,1355]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="elongatulus">
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<emphasis box="[1107,1234,1335,1355]" italics="true" pageId="22" pageNumber="23">P. elongatulus</emphasis>
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</taxonomicName>
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], rainforest,
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<collectorName box="[1351,1441,1335,1355]" pageId="22" pageNumber="23">P. S. Ward</collectorName>
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<accessionNumber box="[817,919,1364,1384]" httpUri="http://www.ncbi.nlm.nih.gov/protein/PSW07360" pageId="22" pageNumber="23">PSW07360</accessionNumber>
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A (
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<collectionCode box="[947,1019,1364,1384]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="22" pageNumber="23">UNAM</collectionCode>
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) (
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<specimenCode box="[1037,1212,1364,1384]" collectionCode="CASENT" pageId="22" pageNumber="23">CASENT0863524</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph blockId="22.[817,1443,1188,1501]" box="[850,1385,1393,1413]" pageId="22" pageNumber="23">
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Other material examined. Known only from the
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<typeStatus box="[1297,1381,1393,1413]" pageId="22" pageNumber="23">holotype</typeStatus>
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.
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</paragraph>
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<paragraph blockId="22.[817,1443,1188,1501]" pageId="22" pageNumber="23">
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Queen measurements (
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<emphasis box="[1063,1075,1424,1442]" italics="true" pageId="22" pageNumber="23">n</emphasis>
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= 1). HW 0.98, HL 1.25, MFC 0.058, LHT 0.76, CI 0.78, FCI 0.059, REL 0.44, REL2 0.56, FI 0.46, PLI 0.61, PWI 0.75, MSC 11.
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</paragraph>
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<paragraph blockId="22.[817,1443,1543,1944]" lastBlockId="23.[146,770,148,197]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<emphasis box="[818,986,1543,1562]" italics="true" pageId="22" pageNumber="23">Queen Diagnosis.</emphasis>
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Small species (see HW, HL, and LHT measurements), with moderately elongate head (CI 0.78); upper surface of mandible smooth and shiny, with scattered punctures; eyes of moderate length (REL 0.44, REL2 0.56); frontal carinae separated by basal scape width; petiole relatively short (PLI 0.61, PL/HL 0.52) and broad (PWI 0.75, DPW/LHT 0.64), with stout, recurved anteroventral tooth; in profile, petiole with convex anterodorsal face ascending gradually to summit in posterior quarter of node, then rounding into steep posterior face; postpetiole very broad, ovorectangular in dorsal view (PPW/LHT 1.01); profemur robust (FI 0.46); hind leg short (LHT/HL 0.61). Head sublucid, densely but finely punctulate, the punctures separated by one to several diameters, the interspaces smooth and shiny, or (towards vertex) finely coriarious. Standing pilosity scattered on head, mesosoma (MSC 11), and gaster, short and sparse on petiole, postpetiole, and fourth abdominal (first gastric) tergite. Uniformly light orange-brown.
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</paragraph>
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<paragraph blockId="23.[145,770,239,552]" pageId="23" pageNumber="24">
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<emphasis box="[146,252,239,258]" italics="true" pageId="23" pageNumber="24">Comments.</emphasis>
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The small size (HL 1.25, LHT 0.76), smooth and shiny mandibles, and very broad (ovorectangular) postpetiole (PPW/LHT 1.01) are distinctive features of
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<taxonomicName authorityName="Ward & Branstetter" authorityYear="2022" box="[443,532,298,316]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="ereptor">
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<emphasis box="[443,532,298,316]" italics="true" pageId="23" pageNumber="24">P. ereptor</emphasis>
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</taxonomicName>
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that set it apart from the queen of its apparent host species,
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<taxonomicName baseAuthorityName="Dalle Torre" baseAuthorityYear="1892" box="[481,611,327,347]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="elongatulus">
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<emphasis box="[481,611,327,347]" italics="true" pageId="23" pageNumber="24">P. elongatulus</emphasis>
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</taxonomicName>
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, and from other species in the
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<taxonomicName baseAuthorityName="Dalle Torre" baseAuthorityYear="1892" box="[284,414,356,376]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="elongatulus">
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<emphasis box="[284,414,356,376]" italics="true" pageId="23" pageNumber="24">P. elongatulus</emphasis>
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</taxonomicName>
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group. Queens of
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<taxonomicName baseAuthorityName="Dalle Torre" baseAuthorityYear="1892" box="[601,731,356,376]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="elongatulus">
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<emphasis box="[601,731,356,376]" italics="true" pageId="23" pageNumber="24">P. elongatulus</emphasis>
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</taxonomicName>
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are larger (HL 1.37–1.42, LHT 0.90–0.94), with striate mandibles, and a slender, pyriform postpetiole as seen in dorsal view (PPW/LHT 0.78–0.83). Our UCE phylogeny shows that
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<taxonomicName authorityName="Ward & Branstetter" authorityYear="2022" box="[552,639,445,463]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="ereptor">
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<emphasis box="[552,639,445,463]" italics="true" pageId="23" pageNumber="24">P. ereptor</emphasis>
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</taxonomicName>
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is sister to the entire assemblage of
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<taxonomicName baseAuthorityName="Dalle Torre" baseAuthorityYear="1892" box="[340,467,473,493]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="elongatulus">
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<emphasis box="[340,467,473,493]" italics="true" pageId="23" pageNumber="24">P. elongatulus</emphasis>
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</taxonomicName>
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samples (Fig. 1). It is well separated from an alate queen of
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<taxonomicName baseAuthorityName="Dalle Torre" baseAuthorityYear="1892" box="[411,538,503,523]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="elongatulus">
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<emphasis box="[411,538,503,523]" italics="true" pageId="23" pageNumber="24">P. elongatulus</emphasis>
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</taxonomicName>
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(D1985) collected in the same nest, thus confirming its status as a different species.
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</paragraph>
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<paragraph blockId="23.[145,771,593,936]" pageId="23" pageNumber="24">
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<emphasis box="[146,381,593,613]" italics="true" pageId="23" pageNumber="24">Distribution and Biology.</emphasis>
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The only known specimen of
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<taxonomicName authorityName="Ward & Branstetter" authorityYear="2022" box="[660,748,594,612]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="ereptor">
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<emphasis box="[660,748,594,612]" italics="true" pageId="23" pageNumber="24">P. ereptor</emphasis>
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</taxonomicName>
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— an alate queen—was collected in a nest of
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<taxonomicName baseAuthorityName="Dalle Torre" baseAuthorityYear="1892" box="[548,677,623,643]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="elongatulus">
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<emphasis box="[548,677,623,643]" italics="true" pageId="23" pageNumber="24">P. elongatulus</emphasis>
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</taxonomicName>
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in a dead twig attached to a liana, on Vereda Cima, Estación de Biología ‘Los Tuxtlas’,
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<collectingRegion box="[232,316,682,701]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="23" pageNumber="24">Veracruz</collectingRegion>
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. The nest (collection code
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<accessionNumber box="[563,667,681,701]" httpUri="http://www.ncbi.nlm.nih.gov/protein/PSW07360" pageId="23" pageNumber="24">PSW07360</accessionNumber>
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) contained numerous workers, alate queens, males, and brood of
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<taxonomicName baseAuthorityName="Dalle Torre" baseAuthorityYear="1892" box="[637,764,711,731]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="elongatulus">
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<emphasis box="[637,764,711,731]" italics="true" pageId="23" pageNumber="24">P. elongatulus</emphasis>
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</taxonomicName>
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.
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<taxonomicName authorityName="Ward & Branstetter" authorityYear="2022" box="[146,359,740,760]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="ereptor">
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<emphasis box="[146,359,740,760]" italics="true" pageId="23" pageNumber="24">Pseudomyrmex ereptor</emphasis>
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</taxonomicName>
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apparently represents a workerless, inquiline parasite of
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<taxonomicName baseAuthorityName="Dalle Torre" baseAuthorityYear="1892" box="[251,379,769,789]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="elongatulus">
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<emphasis box="[251,379,769,789]" italics="true" pageId="23" pageNumber="24">P. elongatulus</emphasis>
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</taxonomicName>
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. The fact that only a single alate queen of
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<taxonomicName authorityName="Ward & Branstetter" authorityYear="2022" box="[146,236,800,818]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="ereptor">
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<emphasis box="[146,236,800,818]" italics="true" pageId="23" pageNumber="24">P. ereptor</emphasis>
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</taxonomicName>
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was found in the nest suggests that she was a recent arrival. The absence of a dealate queen of
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<taxonomicName baseAuthorityName="Dalle Torre" baseAuthorityYear="1892" box="[531,661,828,848]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="elongatulus">
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<emphasis box="[531,661,828,848]" italics="true" pageId="23" pageNumber="24">P. elongatulus</emphasis>
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</taxonomicName>
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should not be over-interpreted; it could simply indicate that this was part of a larger polydomous colony.
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<taxonomicName box="[402,544,887,907]" class="Insecta" family="Formicidae" genus="Pseudomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
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<emphasis box="[402,544,887,907]" italics="true" pageId="23" pageNumber="24">Pseudomyrmex</emphasis>
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</taxonomicName>
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colonies are often comprised of disjunct nests occupying several dead twigs on a plant.
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</paragraph>
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</subSubSection>
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||
</treatment>
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||
</document> |