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<document id="73D79425BED285958580AC225F0A4698" ID-DOI="10.5281/zenodo.6464279" ID-ISBN="978-1-7397467-0-4" ID-ISSN="2754-9844" ID-ZooBank="FE033A11-ACCB-4C6D-B185-63420F8CA79B" IM.tables_requiresApprovalFor="GgImagineBatch" checkinTime="1699386246229" checkinUser="felipe" docAuthor="Hansson, Christer" docDate="2022" docId="03B38785651BB823FDE96F4B2190CD7D" docLanguage="en" docName="TaxMonogNeotropHymenopt.1.1-137.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 1 (1)" docSource="http://dx.doi.org/10.5281/zenodo.6464278" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Closterocerus byrrus Hansson 2022, sp.nov." docType="treatment" docVersion="2" lastPageNumber="33" masterDocId="FF8AFFFD653AB801FFC86C732579CE20" masterDocTitle="Eulophidae Of Costa Rica (Hymenoptera: Chalcidoidea), 4 The Genus Closterocerus Westwood S. Str." masterLastPageNumber="137" masterPageNumber="1" pageNumber="32" updateTime="1699839903426" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="5A17E8F8D74E43177FC29BACA8BFBEE5">Eulophidae Of Costa Rica (Hymenoptera: Chalcidoidea), 4 The Genus Closterocerus Westwood S. Str.</mods:title>
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<mods:namePart id="C78698F0334B6499B8D038387349CC4F">Hansson, Christer</mods:namePart>
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<mods:title id="68D1AC8308625E45CA405C3EFAC1FA91">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
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<mods:part id="4F6243FA12E8870A2311C17E7D957E1F">
<mods:date id="37A9A1F71F44578C136C6FD2A7F1D43F">2022</mods:date>
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<mods:title id="964E43171FFF97259F8DD6C087DCC5A7">Oxford, England</mods:title>
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<mods:number id="7553C8FDFAFB1F3A2081BD0E609675C1">2022-04-28</mods:number>
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<mods:identifier id="BD06E7E6847A1FA9D31E230BD8150278" type="DOI">10.5281/zenodo.6464279</mods:identifier>
<mods:identifier id="3B13C804C7E1C5DB31125F234FDDAEFA" type="ISBN">978-1-7397467-0-4</mods:identifier>
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<treatment id="03B38785651BB823FDE96F4B2190CD7D" LSID="urn:lsid:plazi:treatment:03B38785651BB823FDE96F4B2190CD7D" httpUri="http://treatment.plazi.org/id/03B38785651BB823FDE96F4B2190CD7D" lastPageId="34" lastPageNumber="33" pageId="33" pageNumber="32">
<subSubSection id="C3006518651BB820FDE96F4B2622CD70" box="[545,859,824,848]" pageId="33" pageNumber="32" type="nomenclature">
<paragraph id="8BA53693651BB820FDE96F4B2622CD70" blockId="33.[545,859,824,880]" box="[545,859,824,848]" pageId="33" pageNumber="32">
<heading id="D0ED81FF651BB820FDE96F4B2622CD70" bold="true" box="[545,859,824,848]" centered="true" fontSize="10" level="2" pageId="33" pageNumber="32" reason="2">
<emphasis id="B96EEA81651BB820FDE96F4B2622CD70" bold="true" box="[545,859,824,848]" pageId="33" pageNumber="32">
<taxonomicName id="4C1A4D10651BB820FDE96F4B267FCD70" authority="Hansson, 2022" authorityName="Hansson" authorityYear="2022" box="[545,774,824,848]" class="Insecta" family="Eulophidae" genus="Closterocerus" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="byrrus" status="sp.nov.">
<emphasis id="B96EEA81651BB820FDE96F4B267FCD70" bold="true" box="[545,774,824,848]" italics="true" pageId="33" pageNumber="32">Closterocerus byrrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D57FA651BB820FCC56F4B2622CD70" box="[781,859,824,848]" pageId="33" pageNumber="32" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C3006518651BB820FDA46F2B2669CD50" box="[620,784,856,880]" pageId="33" pageNumber="32" type="description">
<paragraph id="8BA53693651BB820FDA46F2B2669CD50" blockId="33.[545,859,824,880]" box="[620,784,856,880]" pageId="33" pageNumber="32">
(
<figureCitation id="13212A16651BB820FDBC6F2B2671CD50" box="[628,776,856,880]" captionStart="Figs 195–199" captionStartId="132.[213,260,1495,1519]" captionTargetBox="[213,1256,190,1486]" captionTargetId="figure-7@132.[213,1256,190,1486]" captionTargetPageId="132" captionText="Figs 195–199. Closterocerus spp. C. byrrus, holotype ♀ – (195) habitus dorsal view; (196) habitus lateral view. C. albicrus, holotype ♀ – (197) habitus dorsal view; (198) habitus lateral view. C. purpureus, holotype ♀ – (199) habitus dorsal view (excluding head). The image of C. purpureus (type no. 5.1299) are ©The Trustees of the Natural History Museum, London, and made available to you under Creative Commons License 4.0 (https://creativecommons.org/licenses/by/4.0/)." figureDoi="http://doi.org/10.5281/zenodo.10117052" httpUri="https://zenodo.org/record/10117052/files/figure.png" pageId="33" pageNumber="32">Figs 195, 196</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3006518651BB823FF7D6FEB24F0CC37" lastPageId="34" lastPageNumber="33" pageId="33" pageNumber="32" type="diagnosis">
<paragraph id="8BA53693651BB820FF7D6FEB27D3CAB0" blockId="33.[181,1224,920,1168]" pageId="33" pageNumber="32">
<emphasis id="B96EEA81651BB820FF7D6FEB2450CD90" bold="true" box="[181,297,920,944]" pageId="33" pageNumber="32">Diagnosis.</emphasis>
Antenna flattened, scape widest in apical part (
<figureCitation id="13212A16651BB820FCDB6FEB2616CD90" box="[787,879,920,944]" captionStart="Figs 195–199" captionStartId="132.[213,260,1495,1519]" captionTargetBox="[213,1256,190,1486]" captionTargetId="figure-7@132.[213,1256,190,1486]" captionTargetPageId="132" captionText="Figs 195–199. Closterocerus spp. C. byrrus, holotype ♀ – (195) habitus dorsal view; (196) habitus lateral view. C. albicrus, holotype ♀ – (197) habitus dorsal view; (198) habitus lateral view. C. purpureus, holotype ♀ – (199) habitus dorsal view (excluding head). The image of C. purpureus (type no. 5.1299) are ©The Trustees of the Natural History Museum, London, and made available to you under Creative Commons License 4.0 (https://creativecommons.org/licenses/by/4.0/)." figureDoi="http://doi.org/10.5281/zenodo.10117052" httpUri="https://zenodo.org/record/10117052/files/figure.png" pageId="33" pageNumber="32">Fig. 196</figureCitation>
); pronotal collar without a carina along anterior margin (
<figureCitation id="13212A16651BB820FE626FCB277ECDF0" box="[426,519,952,976]" captionStart="Figs 195–199" captionStartId="132.[213,260,1495,1519]" captionTargetBox="[213,1256,190,1486]" captionTargetId="figure-7@132.[213,1256,190,1486]" captionTargetPageId="132" captionText="Figs 195–199. Closterocerus spp. C. byrrus, holotype ♀ – (195) habitus dorsal view; (196) habitus lateral view. C. albicrus, holotype ♀ – (197) habitus dorsal view; (198) habitus lateral view. C. purpureus, holotype ♀ – (199) habitus dorsal view (excluding head). The image of C. purpureus (type no. 5.1299) are ©The Trustees of the Natural History Museum, London, and made available to you under Creative Commons License 4.0 (https://creativecommons.org/licenses/by/4.0/)." figureDoi="http://doi.org/10.5281/zenodo.10117052" httpUri="https://zenodo.org/record/10117052/files/figure.png" pageId="33" pageNumber="32">Fig. 195</figureCitation>
); midlobe of mesoscutum with strong reticulation in anterior one-half and weak reticulation in posterior one-half, and with two pairs of long and strong setae (
<figureCitation id="13212A16651BB820FB546FAB2599CA30" captionStart="Figs 195–199" captionStartId="132.[213,260,1495,1519]" captionTargetBox="[213,1256,190,1486]" captionTargetId="figure-7@132.[213,1256,190,1486]" captionTargetPageId="132" captionText="Figs 195–199. Closterocerus spp. C. byrrus, holotype ♀ – (195) habitus dorsal view; (196) habitus lateral view. C. albicrus, holotype ♀ – (197) habitus dorsal view; (198) habitus lateral view. C. purpureus, holotype ♀ – (199) habitus dorsal view (excluding head). The image of C. purpureus (type no. 5.1299) are ©The Trustees of the Natural History Museum, London, and made available to you under Creative Commons License 4.0 (https://creativecommons.org/licenses/by/4.0/)." figureDoi="http://doi.org/10.5281/zenodo.10117052" httpUri="https://zenodo.org/record/10117052/files/figure.png" pageId="33" pageNumber="32">Fig. 195</figureCitation>
); mesoscutellum golden-red with very weak reticulation, almost smooth (
<figureCitation id="13212A16651BB820FC386F8B213DCA30" box="[1008,1092,1016,1040]" captionStart="Figs 195–199" captionStartId="132.[213,260,1495,1519]" captionTargetBox="[213,1256,190,1486]" captionTargetId="figure-7@132.[213,1256,190,1486]" captionTargetPageId="132" captionText="Figs 195–199. Closterocerus spp. C. byrrus, holotype ♀ – (195) habitus dorsal view; (196) habitus lateral view. C. albicrus, holotype ♀ – (197) habitus dorsal view; (198) habitus lateral view. C. purpureus, holotype ♀ – (199) habitus dorsal view (excluding head). The image of C. purpureus (type no. 5.1299) are ©The Trustees of the Natural History Museum, London, and made available to you under Creative Commons License 4.0 (https://creativecommons.org/licenses/by/4.0/)." figureDoi="http://doi.org/10.5281/zenodo.10117052" httpUri="https://zenodo.org/record/10117052/files/figure.png" pageId="33" pageNumber="32">Fig.195</figureCitation>
); fore wing hyaline with a narrow infuscate band below stigmal vein, reaching to hind margin, and with apical margin weakly infuscate (
<figureCitation id="13212A16651BB820FE0C684B2720CA70" box="[452,601,1080,1104]" captionStart="Figs 195–199" captionStartId="132.[213,260,1495,1519]" captionTargetBox="[213,1256,190,1486]" captionTargetId="figure-7@132.[213,1256,190,1486]" captionTargetPageId="132" captionText="Figs 195–199. Closterocerus spp. C. byrrus, holotype ♀ – (195) habitus dorsal view; (196) habitus lateral view. C. albicrus, holotype ♀ – (197) habitus dorsal view; (198) habitus lateral view. C. purpureus, holotype ♀ – (199) habitus dorsal view (excluding head). The image of C. purpureus (type no. 5.1299) are ©The Trustees of the Natural History Museum, London, and made available to you under Creative Commons License 4.0 (https://creativecommons.org/licenses/by/4.0/)." figureDoi="http://doi.org/10.5281/zenodo.10117052" httpUri="https://zenodo.org/record/10117052/files/figure.png" pageId="33" pageNumber="32">Figs 195, 196</figureCitation>
); coxae pale brown, fore and mid femora white, hind femur dark brown (
<figureCitation id="13212A16651BB820FEF6682B24E5CA50" box="[318,412,1112,1136]" captionStart="Figs 195–199" captionStartId="132.[213,260,1495,1519]" captionTargetBox="[213,1256,190,1486]" captionTargetId="figure-7@132.[213,1256,190,1486]" captionTargetPageId="132" captionText="Figs 195–199. Closterocerus spp. C. byrrus, holotype ♀ – (195) habitus dorsal view; (196) habitus lateral view. C. albicrus, holotype ♀ – (197) habitus dorsal view; (198) habitus lateral view. C. purpureus, holotype ♀ – (199) habitus dorsal view (excluding head). The image of C. purpureus (type no. 5.1299) are ©The Trustees of the Natural History Museum, London, and made available to you under Creative Commons License 4.0 (https://creativecommons.org/licenses/by/4.0/)." figureDoi="http://doi.org/10.5281/zenodo.10117052" httpUri="https://zenodo.org/record/10117052/files/figure.png" pageId="33" pageNumber="32">Fig. 196</figureCitation>
); petiole about as long as wide and smooth; gaster elongate (
<figureCitation id="13212A16651BB820FBEF682B21FDCA50" box="[1063,1156,1112,1136]" captionStart="Figs 195–199" captionStartId="132.[213,260,1495,1519]" captionTargetBox="[213,1256,190,1486]" captionTargetId="figure-7@132.[213,1256,190,1486]" captionTargetPageId="132" captionText="Figs 195–199. Closterocerus spp. C. byrrus, holotype ♀ – (195) habitus dorsal view; (196) habitus lateral view. C. albicrus, holotype ♀ – (197) habitus dorsal view; (198) habitus lateral view. C. purpureus, holotype ♀ – (199) habitus dorsal view (excluding head). The image of C. purpureus (type no. 5.1299) are ©The Trustees of the Natural History Museum, London, and made available to you under Creative Commons License 4.0 (https://creativecommons.org/licenses/by/4.0/)." figureDoi="http://doi.org/10.5281/zenodo.10117052" httpUri="https://zenodo.org/record/10117052/files/figure.png" pageId="33" pageNumber="32">Fig. 195</figureCitation>
), 3.1× as long as wide; length of body female
<quantity id="4CE29B76651BB820FD90680B27DFCAB0" box="[600,678,1144,1168]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="33" pageNumber="32" unit="mm" value="1.4">1.4mm</quantity>
.
</paragraph>
<paragraph id="8BA53693651BB820FF7D68CB2713CAF0" blockId="33.[181,1224,1208,1712]" box="[181,618,1208,1232]" pageId="33" pageNumber="32">
Female
<typeStatus id="54A18831651BB820FEC268CB2415CAF0" box="[266,364,1208,1232]" pageId="33" pageNumber="32">holotype</typeStatus>
: length of body
<quantity id="4CE29B76651BB820FDDF68CB271CCAF0" box="[535,613,1208,1232]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="33" pageNumber="32" unit="mm" value="1.4">1.4mm</quantity>
.
</paragraph>
<paragraph id="8BA53693651BB820FF1268AA24CFCBF0" blockId="33.[181,1224,1208,1712]" pageId="33" pageNumber="32">Antenna dark brown. Frons golden-green. Vertex metallic bluish-green in anterior one-half, golden-green in posterior one-half. Mesoscutum with midlobe metallic bluish-green, sidelobes metallic bluish-purple; mesoscutellum golden-red; dorsellum golden-green. Propodeum dark brown with metallic tinges. Legs with coxae pale brown; fore and mid femora white, hind femur dark brown; tibiae white; tarsi infuscate. Fore wing hyaline with a narrow infuscate band below stigmal vein, reaching to hind margin, and with apical margin weakly infuscate, hind wing hyaline. Petiole dark brown. Gastral tergite 1 metallic bluish-green, remaining tergites dark brown with metallic purple and green tinges.</paragraph>
<paragraph id="8BA53693651BB820FF1269AA27D9C870" blockId="33.[181,1224,1208,1712]" pageId="33" pageNumber="32">Antenna flattened, scape widest in apical part. Frons with weak reticulation below frontofacial suture, with strong reticulation above suture. Vertex with strong reticulation in anterior one-half, with weak reticulation in posterior one-half. Frontofacial suture V-shaped. Subtorular sutures dark and difficult to see. Occipital margin rounded.</paragraph>
<paragraph id="8BA53693651BB820FF126A2B21BEC890" blockId="33.[181,1224,1208,1712]" pageId="33" pageNumber="32">Pronotal collar without a carina along anterior margin. Mesoscutum with midlobe with strong reticulation in anterior one-half and weak reticulation in posterior one-half, sidelobes with strong reticulation; notauli distinct in anterior one-half; midlobe of mesoscutum with two pairs of long</paragraph>
<paragraph id="8BA536936518B823FF1D6CCC2632CF37" blockId="34.[213,1256,191,471]" pageId="34" pageNumber="33">and strong setae. Mesoscutellum with very weak reticulation, almost smooth. Dorsellum convex and smooth. Propodeum smooth and shiny; callus with two setae. Fore wing with speculum closed; with an indistinct stigmal hairline and with radial cell bare.</paragraph>
<paragraph id="8BA536936518B823FF326D6C21F1CF17" blockId="34.[213,1256,191,471]" box="[250,1160,287,311]" pageId="34" pageNumber="33">Petiole about as long as wide and smooth. Gaster elongate; tergites smooth and shiny.</paragraph>
<paragraph id="8BA536936518B823FF326D4C211DCFF7" blockId="34.[213,1256,191,471]" pageId="34" pageNumber="33">Relative measurements: head length dorsal 13; head length frontal 18.5; head width 29; POL 6; OOL 2.5; lateral ocellus maximum width 2; eye length 14; malar space 5.5; mouth width 6.5; mesosoma length 36; mesosoma width 24; mesoscutellum length 14; mesoscutellum width 14; fore wing length 69; fore wing width 43; marginal vein length 32; postmarginal vein length 6; stigmal vein length 5.5; fore wing marginal fringe length 4; gaster length 61; gaster width 20.</paragraph>
<paragraph id="8BA536936518B823FF1D6D8C24F0CC37" blockId="34.[213,393,511,535]" box="[213,393,511,535]" pageId="34" pageNumber="33">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30065186518B823FF1D6E4C24EBCC77" box="[213,402,575,599]" pageId="34" pageNumber="33" type="biology_ecology">
<paragraph id="8BA536936518B823FF1D6E4C24EBCC77" blockId="34.[213,402,575,599]" box="[213,402,575,599]" pageId="34" pageNumber="33">
<emphasis id="B96EEA816518B823FF1D6E4C246CCC77" bold="true" box="[213,277,575,599]" pageId="34" pageNumber="33">Hosts</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30065186518B823FF1D6E0D2490CCB7" box="[213,489,638,663]" pageId="34" pageNumber="33" type="distribution">
<paragraph id="8BA536936518B823FF1D6E0D2490CCB7" blockId="34.[213,489,638,663]" box="[213,489,638,663]" pageId="34" pageNumber="33">
<emphasis id="B96EEA816518B823FF1D6E0D241ECCB6" bold="true" box="[213,359,638,662]" pageId="34" pageNumber="33">Distribution.</emphasis>
<collectingCountry id="F30D76036518B823FEA56E0C249CCCB7" box="[365,485,639,663]" name="Costa Rica" pageId="34" pageNumber="33">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30065186518B823FF1D6ECD2754CD30" pageId="34" pageNumber="33" type="materials_examined">
<paragraph id="8BA536936518B823FF1D6ECD24CACCF6" blockId="34.[213,1255,702,784]" box="[213,435,702,726]" pageId="34" pageNumber="33">
<emphasis id="B96EEA816518B823FF1D6ECD24CACCF6" bold="true" box="[213,435,702,726]" pageId="34" pageNumber="33">Material examined.</emphasis>
</paragraph>
<paragraph id="8BA536936518B823FF326EAD2754CD30" blockId="34.[213,1255,702,784]" pageId="34" pageNumber="33">
Type material.
<materialsCitation id="3B723CCE6518B823FE426EAE2754CD30" collectingDate="2003-09-20" collectingDateMax="2003-10-05" collectingDateMin="2003-09-20" collectionCode="NHMUK" collectorName="D. Rubi" country="Costa Rica" elevation="2460" location="Valle de Silencio" pageId="34" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Limon" typeStatus="holotype">
<typeStatus id="54A188316518B823FE426EAE249ACCD3" box="[394,483,733,755]" pageId="34" pageNumber="33">Holotype</typeStatus>
<specimenCount id="9D1CFD1A6518B823FE226EAD2485CCD3" box="[490,508,734,755]" pageId="34" pageNumber="33" type="female">♀</specimenCount>
“
<collectingCountry id="F30D76036518B823FDC26EAD27E2CCD3" box="[522,667,734,755]" name="Costa Rica" pageId="34" pageNumber="33">COSTA RICA</collectingCountry>
:
<collectingRegion id="49DEF8716518B823FD6C6EAE279FCCD3" box="[676,742,733,755]" country="Costa Rica" name="Limon" pageId="34" pageNumber="33">Limón</collectingRegion>
,
<location id="8EC560486518B823FD276EAE26EDCCD3" LSID="urn:lsid:plazi:treatment:03B38785651BB823FDE96F4B2190CD7D:8EC560486518B823FD276EAE26EDCCD3" box="[751,916,733,755]" country="Costa Rica" name="Valle de Silencio" pageId="34" pageNumber="33" stateProvince="Limon">Valle de Silencio</location>
,
<quantity id="4CE29B766518B823FC556EAE269BCCD3" box="[925,994,733,755]" metricMagnitude="3" metricUnit="m" metricValue="2.46" pageId="34" pageNumber="33" unit="m" value="2460.0">
<elevation id="0037D1A06518B823FC556EAE269BCCD3" box="[925,994,733,755]" metricMagnitude="3" metricUnit="m" metricValue="2.46" pageId="34" pageNumber="33" unit="m" value="2460.0">2460m</elevation>
</quantity>
,
<date id="FFA410536518B823FC246EAE2103CCD3" box="[1004,1146,733,755]" pageId="34" pageNumber="33" value="2003-09-20" valueMax="2003-10-05" valueMin="2003-09-20">
<collectingDate id="EFE0E9BB6518B823FC246EAE2103CCD3" box="[1004,1146,733,755]" pageId="34" pageNumber="33" value="2003-09-20" valueMax="2003-10-05" valueMin="2003-09-20">20.ix-5.x.2003</collectingDate>
</date>
,
<collectorName id="26EF53456518B823FB4C6EAD21A8CCD3" box="[1156,1233,733,755]" pageId="34" pageNumber="33">D. Rubí</collectorName>
et al., LS340258/577465” (
<collectionCode id="ED0BAE566518B823FE0A6E89275ACD30" box="[450,547,762,784]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="34" pageNumber="33" type="Museum">NHMUK</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30065186518B823FF1D6F362190CD7D" box="[213,1257,837,861]" pageId="34" pageNumber="33" type="etymology">
<paragraph id="8BA536936518B823FF1D6F362190CD7D" blockId="34.[213,1257,837,861]" box="[213,1257,837,861]" pageId="34" pageNumber="33">
<emphasis id="B96EEA816518B823FF1D6F36242CCD7D" bold="true" box="[213,341,837,861]" pageId="34" pageNumber="33">Etymology.</emphasis>
From the Latin
<taxonomicName id="4C1A4D106518B823FE356F36273ACD7D" authorityName="Hansson" authorityYear="2022" box="[509,579,837,861]" class="Insecta" family="Eulophidae" genus="Closterocerus" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="byrrus">
<emphasis id="B96EEA816518B823FE356F36273ACD7D" box="[509,579,837,861]" italics="true" pageId="34" pageNumber="33">byrrus</emphasis>
</taxonomicName>
= red, flame-coloured, referring to the colour of mesoscutellum.
</paragraph>
</subSubSection>
</treatment>
</document>
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