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<document id="7F87A56BBCCC948EFA20E34BE703F1EC" ID-DOI="10.5281/zenodo.5393630" ID-ISSN="1638-9387" ID-Zenodo-Dep="5393630" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630319175263" checkinUser="marcus" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; Spiegel, Didier Van Den" docDate="2007" docId="B36487D8FFE0D51BD31FFF28FBCBFDEB" docLanguage="en" docName="Zoosystema.2007.29.3.417-456.pdf" docOrigin="Zoosystema 29 (3)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Glyphiulus Gervais 1847" docType="key" docVersion="4" lastPageNumber="452" masterDocId="4F5DFFA0FFC1D538D20EFFF0FFA5FC3D" masterDocTitle="Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group" masterLastPageNumber="456" masterPageNumber="417" pageNumber="450" updateTime="1699225420998" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E3CB64C0C536BB9EBC4EBF3B8EE2495F">Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group</mods:title>
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<mods:namePart id="AEEBA474D9FA4389B7B58E21A319A548">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="1CD25738DA0A7658A377C55D96B8588C">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
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<mods:namePart id="E8FFB6BEAC9846367FBA3E762B236CE7">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="09B740019D39EAD56DF1DC6D56BD3A95">Muséum national dHistoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
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<mods:namePart id="7D5B168409D2C7E54EB3494A037CF460">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="510A4695EAEB71CC6EF7F8792E18D1C8">Muséum national dHistoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
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<mods:namePart id="2A6AE9D392FE57026CBB5AFF5722C9BE">Spiegel, Didier Van Den</mods:namePart>
<mods:affiliation id="25EE278ED0DE92197EE2B2B98233067F">Musée royal de lAfrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be</mods:affiliation>
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<mods:date id="D44A1F2BF89C9DAAE70C5ABE4C1C8A03">2007</mods:date>
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KEY TO SPECIES OF
<taxonomicName id="FCCD4D4DFFE0D519D3EEFF28FCFFFCCD" ID-CoL="7P3VR" authority="GERVAIS, 1847 OF THE" authorityName="Gervais" authorityYear="1847" box="[480,858,216,243]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="33" pageNumber="450" phylum="Arthropoda" rank="genus">
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GERVAIS, 1847 OF THE
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GROUP
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1. At least some crests on collum more or less obliterated (
<figureCitation id="A3F62A4BFFE0D519D125FEEBFC3EFD0E" box="[811,923,283,308]" captionStart="FIG" captionStartId="4.[113,124,1635,1652]" captionTargetBox="[117,1206,215,1509]" captionTargetId="figure-135@4.[633,720,627,1075]" captionTargetPageId="4" captionText="FIG. 1. — Glyphiulus obliteratus n. sp., paratype ♀: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, dorsal view; D, midbody segments, lateral view; E, midbody segment section; F, claw of midbody leg. Scale bars: A, D, 0.5 mm; B, C, E, 0.2 mm; F, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.5393632" httpUri="https://zenodo.org/record/5393632/files/figure.png" pageId="33" pageNumber="450">Figs 1A, B</figureCitation>
;
<figureCitation id="A3F62A4BFFE0D519D1A9FEECFC40FD0E" box="[935,997,284,308]" captionStart="FIG" captionStartId="8.[113,124,1459,1476]" captionTargetBox="[117,1206,215,1418]" captionTargetId="figure-124@8.[395,674,761,927]" captionTargetPageId="8" captionText="FIG. 3.— Glyphiulus subobliteratus n.sp., paratype ♀: A, anterior part of body,lateral view; B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393636" httpUri="https://zenodo.org/record/5393636/files/figure.png" pageId="33" pageNumber="450">3A, B</figureCitation>
;
<figureCitation id="A3F62A4BFFE0D519D1FCFEEBFB95FD0E" box="[1010,1072,283,307]" captionStart="FIG" captionStartId="11.[113,124,1635,1652]" captionTargetBox="[117,1205,215,1535]" captionTargetId="figure-109@11.[654,745,613,1104]" captionTargetPageId="11" captionText="FIG. 6. — Glyphiulus mediobliteratus n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, telson, ventral view. Scale bars: A, C, D, 0.5 mm; B, E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393642" httpUri="https://zenodo.org/record/5393642/files/figure.png" pageId="33" pageNumber="450">6A, B</figureCitation>
;
<figureCitation id="A3F62A4BFFE0D519D632FEECFF59FD6D" captionStart="FIG" captionStartId="15.[113,124,1077,1094]" captionTargetBox="[118,1205,215,1041]" captionTargetId="figure-46@15.[389,662,581,765]" captionTargetPageId="15" captionText="FIG. 10. — Glyphiulus obliteratoides n. sp., paratype ♀: A, telson, ventral view; B, midbody segment section; C, claw of midbody leg; D, claw. Scale bars: A, 0.2 mm; B, 0.5 mm; C, 0.05 mm; D, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.5393650" httpUri="https://zenodo.org/record/5393650/files/figure.png" pageId="33" pageNumber="450">10A, B</figureCitation>
;
<figureCitation id="A3F62A4BFFE0D519D306FEC8FE90FD6D" box="[264,309,312,336]" captionStart="FIG" captionStartId="17.[113,124,910,927]" captionTargetBox="[139,1184,203,875]" captionTargetId="figure-19@17.[139,1184,203,875]" captionTargetPageId="17" captionText="FIG. 12.— Glyphiulus parobliteratus n. sp.,holotype ♂: A, schematic presentation of carinotaxy pattern of collum;B, midbody segment section; C, midbody segments, lateral view. Scale bar: A, not to scale; B, C, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5393656" httpUri="https://zenodo.org/record/5393656/files/figure.png" pageId="33" pageNumber="450">12A</figureCitation>
) ...................................................................................................................... 2
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— Crests on collum fully developed, no significant parts of collum smooth (
<figureCitation id="A3F62A4BFFE0D519D1E4FEA5FBCCFD51" box="[1002,1129,341,365]" captionStart="FIG" captionStartId="20.[113,124,1657,1674]" captionTargetBox="[117,1206,216,1545]" captionTargetId="figure-120@20.[641,729,688,1087]" captionTargetPageId="20" captionText="FIG. 14. — Glyphiulus costulifer n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, E, F, 0.2 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393660" httpUri="https://zenodo.org/record/5393660/files/figure.png" pageId="33" pageNumber="450">Figs 14A, B</figureCitation>
;
<figureCitation id="A3F62A4BFFE0D519D2E4FE82FEB2FDB7" box="[234,279,370,394]" captionStart="FIG" captionStartId="23.[113,124,1426,1443]" captionTargetBox="[117,1206,215,1382]" captionTargetId="figure-58@23.[389,662,753,934]" captionTargetPageId="23" captionText="FIG. 17.— Glyphiulus subcostulifer n. sp., paratype ♂: A, ocellarium;B, collum,dorsal view;C, midbody segments,dorsal view;D, same, lateral view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, 0.1 mm; B, C, F, 0.2 mm; D, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393668" httpUri="https://zenodo.org/record/5393668/files/figure.png" pageId="33" pageNumber="450">17B</figureCitation>
;
<figureCitation id="A3F62A4BFFE0D519D32CFE82FEF5FDB7" box="[290,336,370,394]" captionStart="FIG" captionStartId="26.[113,124,1497,1514]" captionTargetBox="[118,1205,215,1453]" captionTargetId="figure-126@26.[615,868,661,844]" captionTargetPageId="26" captionText="FIG. 20. — Glyphiulus percostulifer n. sp., paratype ♀: A, anterior part of body, lateral view; B, midbody segment section; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393676" httpUri="https://zenodo.org/record/5393676/files/figure.png" pageId="33" pageNumber="450">20A</figureCitation>
;
<figureCitation id="A3F62A4BFFE0D519D355FE82FE03FDB4" box="[347,422,370,394]" captionStart="FIG" captionStartId="28.[113,124,1657,1674]" captionTargetBox="[116,1205,215,1587]" captionTargetId="figure-146@28.[628,876,845,1029]" captionTargetPageId="28" captionText="FIG. 22.— Glyphiulus paracostulifer n.sp., paratype ♀: A, anterior part of body,lateral view;B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393680" httpUri="https://zenodo.org/record/5393680/files/figure.png" pageId="33" pageNumber="450">22A, B</figureCitation>
;
<figureCitation id="A3F62A4BFFE0D519D3BFFE81FE5EFDB7" box="[433,507,369,394]" captionStart="FIG" captionStartId="31.[113,124,1520,1537]" captionTargetBox="[118,1205,215,1480]" captionTargetId="figure-97@31.[647,736,596,974]" captionTargetPageId="31" captionText="FIG. 25.— Glyphiulus intermedius n. sp., paratype ♀: A, anterior part of body,lateral view;B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, midbody segment section. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393686" httpUri="https://zenodo.org/record/5393686/files/figure.png" pageId="33" pageNumber="450">25A, B</figureCitation>
) ........................................................................................ 8
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2. Pleurosternal (= lowest) crests on segments 2-4 very prominent, arcuate (
<figureCitation id="A3F62A4BFFE0D519D1ECFE55FB91FD80" box="[994,1076,421,445]" captionStart="FIG" captionStartId="4.[113,124,1635,1652]" captionTargetBox="[117,1206,215,1509]" captionTargetId="figure-135@4.[633,720,627,1075]" captionTargetPageId="4" captionText="FIG. 1. — Glyphiulus obliteratus n. sp., paratype ♀: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, dorsal view; D, midbody segments, lateral view; E, midbody segment section; F, claw of midbody leg. Scale bars: A, D, 0.5 mm; B, C, E, 0.2 mm; F, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.5393632" httpUri="https://zenodo.org/record/5393632/files/figure.png" pageId="33" pageNumber="450">Figs 1A</figureCitation>
;
<figureCitation id="A3F62A4BFFE0D519D64FFE55FBC7FD80" box="[1089,1122,421,445]" captionStart="FIG" captionStartId="8.[113,124,1459,1476]" captionTargetBox="[117,1206,215,1418]" captionTargetId="figure-124@8.[395,674,761,927]" captionTargetPageId="8" captionText="FIG. 3.— Glyphiulus subobliteratus n.sp., paratype ♀: A, anterior part of body,lateral view; B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393636" httpUri="https://zenodo.org/record/5393636/files/figure.png" pageId="33" pageNumber="450">3A</figureCitation>
); flagella of posterior gonopods missing; southern
<collectingCountry id="43DA765EFFE0D519D0D3FE32FCB8FDE7" box="[733,797,450,474]" name="China" pageId="33" pageNumber="450">China</collectingCountry>
.............................................. 3
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— Pleurosternal crests on segments 2-4 not particularly prominent, as on subsequent segments (
<figureCitation id="A3F62A4BFFE0D519D2FFFE0BFEE6FE2E" box="[241,323,507,531]" captionStart="FIG" captionStartId="11.[113,124,1635,1652]" captionTargetBox="[117,1205,215,1535]" captionTargetId="figure-109@11.[654,745,613,1104]" captionTargetPageId="11" captionText="FIG. 6. — Glyphiulus mediobliteratus n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, telson, ventral view. Scale bars: A, C, D, 0.5 mm; B, E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393642" httpUri="https://zenodo.org/record/5393642/files/figure.png" pageId="33" pageNumber="450">Figs 6A</figureCitation>
;
<figureCitation id="A3F62A4BFFE0D519D340FE0BFED8FE2E" box="[334,381,507,531]" captionStart="FIG" captionStartId="15.[113,124,1077,1094]" captionTargetBox="[118,1205,215,1041]" captionTargetId="figure-46@15.[389,662,581,765]" captionTargetPageId="15" captionText="FIG. 10. — Glyphiulus obliteratoides n. sp., paratype ♀: A, telson, ventral view; B, midbody segment section; C, claw of midbody leg; D, claw. Scale bars: A, 0.2 mm; B, 0.5 mm; C, 0.05 mm; D, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.5393650" httpUri="https://zenodo.org/record/5393650/files/figure.png" pageId="33" pageNumber="450">10A</figureCitation>
); flagella of posterior gonopods present or absent ................................. 5
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3. Body of adults
<quantity id="FC359B2BFFE0D519D374FDDFFE4AFE7B" box="[378,495,559,583]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="33" pageNumber="450" unit="mm" value="1.25" valueMax="1.4" valueMin="1.1">1.1-1.4 mm</quantity>
wide; carinotaxy formula of midbody segments 1/1+I/i+3+I/i+1/1 (
<figureCitation id="A3F62A4BFFE0D519D2FCFDBBFE8DFE5E" box="[242,296,587,611]" captionStart="FIG" captionStartId="4.[113,124,1635,1652]" captionTargetBox="[117,1206,215,1509]" captionTargetId="figure-135@4.[633,720,627,1075]" captionTargetPageId="4" captionText="FIG. 1. — Glyphiulus obliteratus n. sp., paratype ♀: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, dorsal view; D, midbody segments, lateral view; E, midbody segment section; F, claw of midbody leg. Scale bars: A, D, 0.5 mm; B, C, E, 0.2 mm; F, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.5393632" httpUri="https://zenodo.org/record/5393632/files/figure.png" pageId="33" pageNumber="450">Fig. 1</figureCitation>
A-E); male telopodites 1 slightly reduced, 4-segmented (
<figureCitation id="A3F62A4BFFE0D519D163FDBBFC1BFE5E" box="[877,958,587,611]" captionStart="FIG" captionStartId="6.[113,124,1635,1652]" captionTargetBox="[120,1189,215,1584]" captionTargetId="figure-24@6.[99,1225,189,1626]" captionTargetPageId="6" captionText="FIG. 2. — Glyphiulus obliteratus n. sp., paratype ♂: A, antenna; B, gnathochilarium; C, leg 1, frontal view; D, leg 2; E, leg 3; F, leg 9; G, anterior gonopods,caudal view;H, posterior gonopods,frontal view;I, same,caudal view.Abbreviations:c, caudal piece; d, digitiform process; f, frontal piece; l, setose lobe; s, subsecuriform process; t, (?)telopodite. Scale bar: A-F, 0.2 mm; G-I, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.5393634" httpUri="https://zenodo.org/record/5393634/files/figure.png" pageId="33" pageNumber="450">Fig. 2C</figureCitation>
); cave in
<collectingRegion id="F909F82CFFE0D519D610FDBCFEE3FEBD" country="China" name="Yunnan" pageId="33" pageNumber="450">Yunnan Province</collectingRegion>
,
<collectingCountry id="43DA765EFFE0D519D35EFD98FE35FEBD" box="[336,400,616,640]" name="China" pageId="33" pageNumber="450">China</collectingCountry>
..............................................................................
<taxonomicName id="FCCD4D4DFFE0D519D1A7FD99FB95FEBD" ID-CoL="3GGKG" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[937,1072,616,640]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="33" pageNumber="450" phylum="Arthropoda" rank="species" species="obliteratus" status="sp. nov.">
<emphasis id="09B9EADCFFE0D519D1A7FD99FB95FEBD" box="[937,1072,616,640]" italics="true" pageId="33" pageNumber="450">G. obliteratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFE0D519D638FD99FBCBFEBD" box="[1078,1134,617,640]" pageId="33" pageNumber="450" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
<paragraph id="3B7236CEFFE0D519D2B3FD75FBCBFE87" blockId="33.[189,1135,559,698]" pageId="33" pageNumber="450">
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— Body of adults
<quantity id="FC359B2BFFE0D519D381FD75FDABFEA1" box="[399,526,645,669]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="33" pageNumber="450" unit="mm" value="1.9" valueMax="2.2" valueMin="1.6">1.6-2.2 mm</quantity>
wide; two divided crests below ozoporiferous cone; male telopodites 1 slightly reduced in size but 5-segmented ................................................ 4
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<paragraph id="3B7236CEFFE0D519D2B3FD25FBCBFF34" blockId="33.[189,1134,725,835]" pageId="33" pageNumber="450">
<keyLead id="803C95FBFFE0D519D2B3FD25FBCBFF34" pageId="33" pageNumber="450">
4. Carinotaxy formula of midbody segments 2/2+I/i+3+I/i+2/2; body unpigmented, only ocelli black; cave in
<collectingRegion id="F909F82CFFE0D519D3B6FD02FDD3FF37" box="[440,630,754,778]" country="China" name="Guizhou" pageId="33" pageNumber="450">Guizhou Province</collectingRegion>
,
<collectingCountry id="43DA765EFFE0D519D08EFD02FD65FF37" box="[640,704,754,778]" name="China" pageId="33" pageNumber="450">China</collectingCountry>
..............................................
<taxonomicName id="FCCD4D4DFFE0D519D606FD02FBCBFF34" ID-CoL="3GGLF" authorityName="Mauries &amp; Nguyen Duy-Jacquemin" authorityYear="1997" box="[1032,1134,754,777]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="33" pageNumber="450" phylum="Arthropoda" rank="species" species="zorzini">
<emphasis id="09B9EADCFFE0D519D606FD02FBCBFF34" box="[1032,1134,754,777]" italics="true" pageId="33" pageNumber="450">G. zorzini</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE0D519D2B3FCFFFBCBFF7E" blockId="33.[189,1134,725,835]" pageId="33" pageNumber="450">
<keyLead id="803C95FBFFE0D519D2B3FCFFFBCBFF7E" pageId="33" pageNumber="450">
— Carinotaxy formula of midbody segments 2/2+I/i+3/3+I/i+2/2 (
<figureCitation id="A3F62A4BFFE0D519D189FCFEFC5DFF1A" box="[903,1016,782,807]" captionStart="FIG" captionStartId="8.[113,124,1459,1476]" captionTargetBox="[117,1206,215,1418]" captionTargetId="figure-124@8.[395,674,761,927]" captionTargetPageId="8" captionText="FIG. 3.— Glyphiulus subobliteratus n.sp., paratype ♀: A, anterior part of body,lateral view; B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393636" httpUri="https://zenodo.org/record/5393636/files/figure.png" pageId="33" pageNumber="450">Fig. 3A, B</figureCitation>
); body and ocelli pigmented, at least partly dark; cave in
<collectingRegion id="F909F82CFFE0D519D0BBFCDCFCC6FF7E" box="[693,867,811,835]" country="China" name="Yunnan" pageId="33" pageNumber="450">Yunnan Province</collectingRegion>
....
<taxonomicName id="FCCD4D4DFFE0D519D185FCDCFB95FF7E" ID-CoL="3GGL8" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[907,1072,811,835]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="33" pageNumber="450" phylum="Arthropoda" rank="species" species="subobliteratus" status="sp. nov.">
<emphasis id="09B9EADCFFE0D519D185FCDCFB95FF7E" box="[907,1072,811,835]" italics="true" pageId="33" pageNumber="450">G. subobliteratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFE0D519D639FCDCFBCBFF7E" box="[1079,1134,812,835]" pageId="33" pageNumber="450" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="80392E6BFFE0D519D2B3FCAEFBCBF81E" pageId="33" pageNumber="450">
<paragraph id="3B7236CEFFE0D519D2B3FCAEFBCBFFF1" blockId="33.[189,1134,862,1059]" pageId="33" pageNumber="450">
<keyLead id="803C95FBFFE0D519D2B3FCAEFBCBFFF1" pageId="33" pageNumber="450">
5. Axial crest on midbody segments strongly reduced, carinotaxy formula 2/2+I/i+1/1+m+1/ 1+I/i+2/2 (
<figureCitation id="A3F62A4BFFE0D519D353FC8BFE3EFFAE" box="[349,411,891,915]" captionStart="FIG" captionStartId="11.[113,124,1635,1652]" captionTargetBox="[117,1205,215,1535]" captionTargetId="figure-109@11.[654,745,613,1104]" captionTargetPageId="11" captionText="FIG. 6. — Glyphiulus mediobliteratus n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, telson, ventral view. Scale bars: A, C, D, 0.5 mm; B, E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393642" httpUri="https://zenodo.org/record/5393642/files/figure.png" pageId="33" pageNumber="450">Fig. 6</figureCitation>
); legs slightly shorter than body diameter (
<figureCitation id="A3F62A4BFFE0D519D156FC8BFC03FFAE" box="[856,934,891,915]" captionStart="FIG" captionStartId="11.[113,124,1635,1652]" captionTargetBox="[117,1205,215,1535]" captionTargetId="figure-109@11.[654,745,613,1104]" captionTargetPageId="11" captionText="FIG. 6. — Glyphiulus mediobliteratus n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, telson, ventral view. Scale bars: A, C, D, 0.5 mm; B, E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393642" httpUri="https://zenodo.org/record/5393642/files/figure.png" pageId="33" pageNumber="450">Fig. 6F</figureCitation>
); anterior gonopod coxosternum with particularly high median outgrowths, devoid of a notch at distal margin (
<figureCitation id="A3F62A4BFFE0D519D2FCFC45FEE6FFF0" box="[242,323,949,973]" captionStart="FIG" captionStartId="13.[113,124,1198,1215]" captionTargetBox="[94,1236,194,1174]" captionTargetId="figure-19@13.[94,1236,194,1174]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, antenna;B, gnathochilarium; C, leg 1, frontal view;D, leg 2; E, leg 3; F, leg 9; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393646" httpUri="https://zenodo.org/record/5393646/files/figure.png" pageId="33" pageNumber="450">Fig. 8G</figureCitation>
); cave in North
<collectingCountry id="43DA765EFFE0D519D3E5FC45FDE1FFF0" box="[491,580,949,973]" name="Vietnam" pageId="33" pageNumber="450">Vietnam</collectingCountry>
...........................................
<taxonomicName id="FCCD4D4DFFE0D519D177FC45FB95FFF0" ID-CoL="3GGK9" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[889,1072,949,973]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="33" pageNumber="450" phylum="Arthropoda" rank="species" species="mediobliteratus" status="sp. nov.">
<emphasis id="09B9EADCFFE0D519D177FC45FB95FFF0" box="[889,1072,949,973]" italics="true" pageId="33" pageNumber="450">G. mediobliteratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFE0D519D638FC45FBCBFFF1" box="[1078,1134,949,972]" pageId="33" pageNumber="450" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE0D519D2B3FC22FBCBF81E" blockId="33.[189,1134,862,1059]" pageId="33" pageNumber="450">
<keyLead id="803C95FBFFE0D519D2B3FC22FBCBF81E" pageId="33" pageNumber="450">
— Axial crest on midbody segments not so strikingly reduced; legs subequal to or even longer than body diameter; anterior gonopod coxosternum deeply notched at distal margin; caves in
<collectingRegion id="F909F82CFFE0D519D308FBFBFE66F81E" box="[262,451,1035,1059]" country="China" name="Guizhou" pageId="33" pageNumber="450">Guizhou Province</collectingRegion>
,
<collectingCountry id="43DA765EFFE0D519D3C3FBFBFDA8F81E" box="[461,525,1035,1059]" name="China" pageId="33" pageNumber="450">China</collectingCountry>
....................................................................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="80392E6BFFE0D519D2B3FBCEFBCBF8F4" pageId="33" pageNumber="450">
<paragraph id="3B7236CEFFE0D519D2B3FBCEFBCBF8B2" blockId="33.[189,1134,1086,1225]" pageId="33" pageNumber="450">
<keyLead id="803C95FBFFE0D519D2B3FBCEFBCBF8B2" pageId="33" pageNumber="450">
6. Carinotaxy formula of midbody segments 1/0+1+I/i+3/3+I/i+1+1/0 (
<figureCitation id="A3F62A4BFFE0D519D1A1FBCEFB8CF86B" box="[943,1065,1086,1110]" captionStart="FIG" captionStartId="17.[113,124,910,927]" captionTargetBox="[139,1184,203,875]" captionTargetId="figure-19@17.[139,1184,203,875]" captionTargetPageId="17" captionText="FIG. 12.— Glyphiulus parobliteratus n. sp.,holotype ♂: A, schematic presentation of carinotaxy pattern of collum;B, midbody segment section; C, midbody segments, lateral view. Scale bar: A, not to scale; B, C, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5393656" httpUri="https://zenodo.org/record/5393656/files/figure.png" pageId="33" pageNumber="450">Fig. 12B, C</figureCitation>
); male telopodites 1 normal, 5-segmented (
<figureCitation id="A3F62A4BFFE0D519D060FBABFD75F84E" box="[622,720,1115,1139]" captionStart="FIG" captionStartId="18.[113,124,1657,1674]" captionTargetBox="[167,1180,227,1580]" captionTargetId="figure-24@18.[94,1206,201,1668]" captionTargetPageId="18" captionText="FIG. 13. — Glyphiulus parobliteratus n. sp., holotype ♂: A, antenna;B, gnathochilarium;C, leg 1, frontal view;D, leg 2; E, leg 3;F, leg 13; G, anterior gonopods, caudal view; H, posterior gonopods caudal view, respectively. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393658" httpUri="https://zenodo.org/record/5393658/files/figure.png" pageId="33" pageNumber="450">Fig. 13C</figureCitation>
); anterior gonopods as in
<figureCitation id="A3F62A4BFFE0D519D1E5FBABFBCDF84E" box="[1003,1128,1115,1139]" captionStart="FIG" captionStartId="18.[113,124,1657,1674]" captionTargetBox="[167,1180,227,1580]" captionTargetId="figure-24@18.[94,1206,201,1668]" captionTargetPageId="18" captionText="FIG. 13. — Glyphiulus parobliteratus n. sp., holotype ♂: A, antenna;B, gnathochilarium;C, leg 1, frontal view;D, leg 2; E, leg 3;F, leg 13; G, anterior gonopods, caudal view; H, posterior gonopods caudal view, respectively. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393658" httpUri="https://zenodo.org/record/5393658/files/figure.png" pageId="33" pageNumber="450">Figure 13G</figureCitation>
, posterior gonopods as in
<figureCitation id="A3F62A4BFFE0D519D3E3FB88FDC0F8AD" box="[493,613,1144,1168]" captionStart="FIG" captionStartId="18.[113,124,1657,1674]" captionTargetBox="[167,1180,227,1580]" captionTargetId="figure-24@18.[94,1206,201,1668]" captionTargetPageId="18" captionText="FIG. 13. — Glyphiulus parobliteratus n. sp., holotype ♂: A, antenna;B, gnathochilarium;C, leg 1, frontal view;D, leg 2; E, leg 3;F, leg 13; G, anterior gonopods, caudal view; H, posterior gonopods caudal view, respectively. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393658" httpUri="https://zenodo.org/record/5393658/files/figure.png" pageId="33" pageNumber="450">Figure 13H</figureCitation>
.........................................
<taxonomicName id="FCCD4D4DFFE0D519D187FB88FB95F8AD" ID-CoL="3GGKM" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[905,1072,1144,1168]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="33" pageNumber="450" phylum="Arthropoda" rank="species" species="parobliteratus" status="sp. nov.">
<emphasis id="09B9EADCFFE0D519D187FB88FB95F8AD" box="[905,1072,1144,1168]" italics="true" pageId="33" pageNumber="450">G. parobliteratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFE0D519D638FB88FBCBF8B2" box="[1078,1134,1144,1167]" pageId="33" pageNumber="450" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE0D519D2B3FB65FBCBF8F4" blockId="33.[189,1134,1086,1225]" pageId="33" pageNumber="450">
<keyLead id="803C95FBFFE0D519D2B3FB65FBCBF8F4" pageId="33" pageNumber="450">
— Carinotaxy formula of midbody segments different; male telopodites 1 reduced in size but 5-segmented (
<figureCitation id="A3F62A4BFFE0D519D374FB41FE72F8F4" box="[378,471,1201,1225]" captionStart="FIG" captionStartId="16.[113,124,1182,1199]" captionTargetBox="[97,1218,203,1159]" captionTargetId="figure-24@16.[97,1218,203,1159]" captionTargetPageId="16" captionText="FIG. 11. — Glyphiulus obliteratoides n. sp., paratype ♂: A, antenna; B, gnathochilarium; C, leg 1, frontal view; D, leg 2, caudal view; E, leg 3; F, leg 9; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393652" httpUri="https://zenodo.org/record/5393652/files/figure.png" pageId="33" pageNumber="450">Fig. 11C</figureCitation>
); anterior gonopods and posterior gonopods not as above ...... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="80392E6BFFE0D519D2B3FB15FBCAF952" pageId="33" pageNumber="450">
<paragraph id="3B7236CEFFE0D519D2B3FB15FBCBF90B" blockId="33.[189,1135,1253,1391]" pageId="33" pageNumber="450">
<keyLead id="803C95FBFFE0D519D2B3FB15FBCBF90B" pageId="33" pageNumber="450">
7. Carinotaxy formula of midbody segments 1/0+1+I/i+3+I/i+1+1/0 (
<figureCitation id="A3F62A4BFFE0D519D198FB15FC6AF8C0" box="[918,975,1253,1277]" captionStart="FIG" captionStartId="14.[113,124,1459,1476]" captionTargetBox="[117,1206,215,1417]" captionTargetId="figure-126@14.[587,853,634,821]" captionTargetPageId="14" captionText="FIG. 9. — Glyphiulus obliteratoides n. sp., paratype ♀: A, anterior part of body,lateral view;B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, dorsal view. Scale bars: A-E, 0.5 mm; F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393648" httpUri="https://zenodo.org/record/5393648/files/figure.png" pageId="33" pageNumber="450">Fig. 9</figureCitation>
A-E); body and ocelli pigmented; medial outgrowth of anterior gonopod coxosternum folded (
<figureCitation id="A3F62A4BFFE0D519D607FAF1FBC3F924" box="[1033,1126,1281,1305]" captionStart="FIG" captionStartId="16.[113,124,1182,1199]" captionTargetBox="[97,1218,203,1159]" captionTargetId="figure-24@16.[97,1218,203,1159]" captionTargetPageId="16" captionText="FIG. 11. — Glyphiulus obliteratoides n. sp., paratype ♂: A, antenna; B, gnathochilarium; C, leg 1, frontal view; D, leg 2, caudal view; E, leg 3; F, leg 9; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393652" httpUri="https://zenodo.org/record/5393652/files/figure.png" pageId="33" pageNumber="450">Fig. 11G</figureCitation>
) ...................................................................................................
<taxonomicName id="FCCD4D4DFFE0D519D181FAEEFB95F90B" ID-CoL="3GGKF" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[911,1072,1310,1334]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="33" pageNumber="450" phylum="Arthropoda" rank="species" species="obliteratoides" status="sp. nov.">
<emphasis id="09B9EADCFFE0D519D181FAEEFB95F90B" box="[911,1072,1310,1334]" italics="true" pageId="33" pageNumber="450">G. obliteratoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFE0D519D638FAEFFBCBF90B" box="[1078,1134,1311,1334]" pageId="33" pageNumber="450" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
<paragraph id="3B7236CEFFE0D519D2B3FACBFBCAF952" blockId="33.[189,1135,1253,1391]" pageId="33" pageNumber="450">
<keyLead id="803C95FBFFE0D519D2B3FACBFBCAF952" pageId="33" pageNumber="450">
— Carinotaxy formula of midbody segments 2/2+I/i+3/3+I/i+2/2; body and ocelli unpigmented medial outgrowth of anterior gonopod coxosternum not folded ....
<taxonomicName id="FCCD4D4DFFE0D519D1AAFAA8FBA9F952" ID-CoL="3GGKZ" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" baseAuthorityName="Meng &amp; Zhang" baseAuthorityYear="1993" box="[932,1036,1368,1391]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="33" pageNumber="450" phylum="Arthropoda" rank="species" species="sinensis" status="comb. nov.">
<emphasis id="09B9EADCFFE0D519D1AAFAA8FBA9F952" box="[932,1036,1368,1391]" italics="true" pageId="33" pageNumber="450">G. sinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFE0D519D61CFAA8FBCAF952" box="[1042,1135,1367,1391]" pageId="33" pageNumber="450" rank="species">n. comb.</taxonomicNameLabel>
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</paragraph>
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<keyStep id="80392E6BFFE0D519D2B3FA7BFBCBFA0E" pageId="33" pageNumber="450">
<paragraph id="3B7236CEFFE0D519D2B3FA7BFBCBF9C4" blockId="33.[189,1135,1419,1587]" pageId="33" pageNumber="450">
<keyLead id="803C95FBFFE0D519D2B3FA7BFBCBF9C4" pageId="33" pageNumber="450">
8. Carinotaxy formula of collum typical for the
<taxonomicName id="FCCD4D4DFFE0D519D095FA7BFD5BF99E" ID-CoL="3GGJZ" baseAuthorityName="Gervais" baseAuthorityYear="1847" box="[667,766,1419,1443]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="33" pageNumber="450" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="09B9EADCFFE0D519D095FA7BFD5BF99E" box="[667,766,1419,1443]" italics="true" pageId="33" pageNumber="450">granulatus</emphasis>
</taxonomicName>
-group, I-VI+7a+pc+ma+pc+7a+VI-I, that of midbody segments 2+I/i+3/3+I/i+2 (
<figureCitation id="A3F62A4BFFE0D519D0AAFA58FD4CF9FD" box="[676,745,1448,1472]" captionStart="FIG" captionStartId="20.[113,124,1657,1674]" captionTargetBox="[117,1206,216,1545]" captionTargetId="figure-120@20.[641,729,688,1087]" captionTargetPageId="20" captionText="FIG. 14. — Glyphiulus costulifer n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, E, F, 0.2 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393660" httpUri="https://zenodo.org/record/5393660/files/figure.png" pageId="33" pageNumber="450">Fig. 14</figureCitation>
A-E), texture of both lateralmost crests unusually micropunctate (
<figureCitation id="A3F62A4BFFE0D519D3F6FA34FDF0F9E1" box="[504,597,1476,1500]" captionStart="FIG" captionStartId="21.[113,124,1635,1652]" captionTargetBox="[259,1063,215,1592]" captionTargetId="figure-148@21.[629,890,854,1037]" captionTargetPageId="21" captionText="FIG. 15. — Glyphiulus costulifer n. sp., paratype ♂: A, midbody segment section; B, midbody metazonite enlarged, lateral view; C, segment 7, frontolateral view;D, anterior gonopod,caudal view;E, posterior gonopod,caudal view; F, posterior gonopod,enlarged caudal view. Scale bars: A, C, 0.2 mm; B, D, E, 0.1 mm; F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393664" httpUri="https://zenodo.org/record/5393664/files/figure.png" pageId="33" pageNumber="450">Fig. 15B</figureCitation>
); male legs 1 and gonopods as in
<figureCitation id="A3F62A4BFFE0D519D1A0FA34FBCCF9E1" box="[942,1129,1476,1500]" captionStart="FIG" captionStartId="22.[113,124,1149,1166]" captionTargetBox="[116,1207,225,1110]" captionTargetId="figure-24@22.[94,1229,215,1123]" captionTargetPageId="22" captionText="FIG. 16. — Glyphiulus costulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium; C, leg 1, frontal view; D, leg 2; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, posterior gonopods, caudal view. Abbreviations: d, digitiform process; f, flagellum. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393666" httpUri="https://zenodo.org/record/5393666/files/figure.png" pageId="33" pageNumber="450">Figure 16C, G, H</figureCitation>
; cave in
<collectingCountry id="43DA765EFFE0D519D336FA12FEC2F9C4" box="[312,359,1506,1529]" name="Laos" pageId="33" pageNumber="450">Laos</collectingCountry>
......................................................................................
<taxonomicName id="FCCD4D4DFFE0D519D1B7FA11FB95F9C4" ID-CoL="3GGJN" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[953,1072,1505,1529]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="33" pageNumber="450" phylum="Arthropoda" rank="species" species="costulifer" status="sp. nov.">
<emphasis id="09B9EADCFFE0D519D1B7FA11FB95F9C4" box="[953,1072,1505,1529]" italics="true" pageId="33" pageNumber="450">G. costulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFE0D519D638FA12FBCBF9C4" box="[1078,1134,1506,1529]" pageId="33" pageNumber="450" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE0D519D2B3FA0EFBCBFA0E" blockId="33.[189,1135,1419,1587]" pageId="33" pageNumber="450">
<keyLead id="803C95FBFFE0D519D2B3FA0EFBCBFA0E" pageId="33" pageNumber="450">— Carinotaxy formula of collum not as above, both lateralmost crests usual, smooth; male legs 1 and gonopods not as above ............................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="80392E6BFFE0D51AD2B3F9BEFBCBFDBD" lastPageId="34" lastPageNumber="451" pageId="33" pageNumber="450">
<paragraph id="3B7236CEFFE0D51AD2B3F9BEFBCBFD30" blockId="33.[189,1134,1614,1695]" lastBlockId="34.[189,1134,216,384]" lastPageId="34" lastPageNumber="451" pageId="33" pageNumber="450">
<keyLead id="803C95FBFFE0D51AD2B3F9BEFBCBFD30" lastPageId="34" lastPageNumber="451" pageId="33" pageNumber="450">
9. Carinotaxy formula of collum I-III+P+M+P+IV-I (i.e. all crests complete), that of midbody segments 2/2+I/i+3/3+I/i+2/2; anterior gonopod coxosternum without a deep notch at distal margin (unknown in
<taxonomicName id="FCCD4D4DFFE0D519D066F978FD7BFAA2" ID-CoL="3GGJX" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[616,734,1671,1695]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="33" pageNumber="450" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="09B9EADCFFE0D519D066F978FD7BFAA2" box="[616,734,1671,1695]" italics="true" pageId="33" pageNumber="450">G. formosus</emphasis>
</taxonomicName>
); male telopodites 1 strongly reduced, 1-3-segmented (unknown in
<taxonomicName id="FCCD4D4DFFE3D51AD01CFF28FD23FCCD" ID-CoL="3GGJX" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[530,646,216,240]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="451" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="09B9EADCFFE3D51AD01CFF28FD23FCCD" box="[530,646,216,240]" italics="true" pageId="34" pageNumber="451">G. formosus</emphasis>
</taxonomicName>
); flagella of posterior gonopods plumose distally (unknown in
<taxonomicName id="FCCD4D4DFFE3D51AD376FF05FE48FD30" ID-CoL="3GGJX" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[376,493,245,269]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="451" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="09B9EADCFFE3D51AD376FF05FE48FD30" box="[376,493,245,269]" italics="true" pageId="34" pageNumber="451">G. formosus</emphasis>
</taxonomicName>
) ........................................................................................ 10
</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE3D51AD2B3FEE2FBCBFDBD" blockId="34.[189,1134,216,384]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FEE2FBCBFDBD" pageId="34" pageNumber="451">— Carinotaxy formula of collum not as above, at least some of the crests being abbreviated; carinotaxy formula of midbody segments as above or different; anterior gonopod coxosternum often deeply notched at distal margin; male telopodites 1 usually less reduced; flagella of posterior gonopods, when present, bare or plumose ................................................. 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="80392E6BFFE3D51AD2B3FE6CFBCBFDD3" pageId="34" pageNumber="451">
<paragraph id="3B7236CEFFE3D51AD2B3FE6CFBCBFDEC" blockId="34.[189,1134,412,494]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FE6CFBCBFDEC" pageId="34" pageNumber="451">
10. Adult body about
<quantity id="FC359B2BFFE3D51AD3BAFE6CFDAFFD89" box="[436,522,412,436]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="451" unit="mm" value="1.5">1.5 mm</quantity>
wide; male femora 6 and 7 with a glandular adenostyle distoventrally;
<collectingRegion id="F909F82CFFE3D51AD38CFE49FDE5FDEC" box="[386,576,441,465]" country="China" name="Zhejiang" pageId="34" pageNumber="451">Zhejiang Province</collectingRegion>
,
<collectingCountry id="43DA765EFFE3D51AD044FE49FD2FFDEC" box="[586,650,441,465]" name="China" pageId="34" pageNumber="451">China</collectingCountry>
..................................................
<taxonomicName id="FCCD4D4DFFE3D51AD1E2FE49FBCBFDEC" ID-CoL="6KLWH" authorityName="Zhang &amp; Li" authorityYear="1982" box="[1004,1134,441,465]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="451" phylum="Arthropoda" rank="species" species="recticullus">
<emphasis id="09B9EADCFFE3D51AD1E2FE49FBCBFDEC" box="[1004,1134,441,465]" italics="true" pageId="34" pageNumber="451">G. recticullus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE3D51AD2B3FE26FBCBFDD3" blockId="34.[189,1134,412,494]" box="[189,1134,470,494]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FE26FBCBFDD3" box="[189,1134,470,494]" pageId="34" pageNumber="451">
— Body width at least
<quantity id="FC359B2BFFE3D51AD3B9FE26FDA9FDD0" box="[439,524,470,494]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="34" pageNumber="451" unit="mm" value="1.8">1.8 mm</quantity>
; male femora without adenostyles ................................. 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="80392E6BFFE3D51AD2B3FDFBFBCBFE44" pageId="34" pageNumber="451">
<paragraph id="3B7236CEFFE3D51AD2B3FDFBFBCBFE02" blockId="34.[189,1134,522,633]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FDFBFBCBFE02" pageId="34" pageNumber="451">
11. Coloration light yellow-brown; only 12 ocelli in a subtriangular patch, black; adult body
<quantity id="FC359B2BFFE3D51AD2E4FDD7FE99FE02" box="[234,316,551,575]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="34" pageNumber="451" unit="mm" value="1.8">1.8 mm</quantity>
wide; cave in
<collectingRegion id="F909F82CFFE3D51AD3C0FDD7FD2CFE02" box="[462,649,551,575]" country="China" name="Guangxi" pageId="34" pageNumber="451">Guangxi Province</collectingRegion>
,
<collectingCountry id="43DA765EFFE3D51AD09DFDD7FD76FE02" box="[659,723,551,575]" name="China" pageId="34" pageNumber="451">China</collectingCountry>
...........................................
<taxonomicName id="FCCD4D4DFFE3D51AD608FDD8FBCBFE02" ID-CoL="7XJKZ" authorityName="Loksa" authorityYear="1960" box="[1030,1134,551,575]" class="Diplopoda" family="Cambalopsidae" genus="Octoglyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="451" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="09B9EADCFFE3D51AD608FDD8FBCBFE02" box="[1030,1134,551,575]" italics="true" pageId="34" pageNumber="451">G. pulcher</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE3D51AD2B3FDB4FBCBFE44" blockId="34.[189,1134,522,633]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FDB4FBCBFE44" pageId="34" pageNumber="451">
— Coloration darker, epigean species; fewer ocelli per eye patch; body about
<quantity id="FC359B2BFFE3D51AD1FFFDB4FBCBFE66" box="[1009,1134,580,604]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="34" pageNumber="451" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.5-2.6 mm</quantity>
wide ......................................................................................................................... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="80392E6BFFE3D51AD2B3FD65FBCBFF39" pageId="34" pageNumber="451">
<paragraph id="3B7236CEFFE3D51AD2B3FD65FBCBFE90" blockId="34.[189,1134,661,772]" box="[189,1134,661,686]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FD65FBCBFE90" box="[189,1134,661,686]" pageId="34" pageNumber="451">
12. Nine ocelli in two rows per patch;
<materialsCitation id="8BA53C93FFE3D51AD041FD66FBCBFE90" box="[591,1134,661,686]" country="Hong Kong" location="Hong Kong" pageId="34" pageNumber="451" specimenCount="1" specimenCount-female="1">
<collectingCountry id="43DA765EFFE3D51AD041FD66FD6CFE93" box="[591,713,662,686]" name="Hong Kong" pageId="34" pageNumber="451">Hong Kong</collectingCountry>
(
<specimenCount id="2DCBFD47FFE3D51AD0D7FD6AFD4EFE90" box="[729,747,666,685]" pageId="34" pageNumber="451" type="female"></specimenCount>
) ....................................
<taxonomicName id="FCCD4D4DFFE3D51AD1F7FD66FBCBFE90" ID-CoL="3GGJX" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[1017,1134,661,685]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="451" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="09B9EADCFFE3D51AD1F7FD66FBCBFE90" box="[1017,1134,661,685]" italics="true" pageId="34" pageNumber="451">G. formosus</emphasis>
</taxonomicName>
</materialsCitation>
</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE3D51AD2B3FD43FBCBFF39" blockId="34.[189,1134,661,772]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FD43FBCBFF39" pageId="34" pageNumber="451">
— Seven ocelli in three rows per patch; male telopodites reduced, 3-segmented; half-anterior gonopod with a subsecuriform coxosternum; Mt. Bana,
<collectingRegion id="F909F82CFFE3D51AD144FD3FFBA3FEDA" box="[842,1030,719,743]" country="Vietnam" name="Da Nang" pageId="34" pageNumber="451">Danang Province</collectingRegion>
,
<collectingCountry id="43DA765EFFE3D51AD61CFD3FFBCBFEDA" box="[1042,1134,719,743]" name="Vietnam" pageId="34" pageNumber="451">Vietnam</collectingCountry>
...................................................................................................................
<taxonomicName id="FCCD4D4DFFE3D51AD1F8FD1CFBCBFF39" ID-CoL="3GGK8" authorityName="Attems" authorityYear="1938" box="[1014,1134,748,772]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="451" phylum="Arthropoda" rank="species" species="mediator">
<emphasis id="09B9EADCFFE3D51AD1F8FD1CFBCBFF39" box="[1014,1134,748,772]" italics="true" pageId="34" pageNumber="451">G. mediator</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="80392E6BFFE3D51AD2B3FCD0FBCBFF96" pageId="34" pageNumber="451">
<paragraph id="3B7236CEFFE3D51AD2B3FCD0FBCBFF4C" blockId="34.[189,1134,800,939]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FCD0FBCBFF4C" pageId="34" pageNumber="451">
13. Carinotaxy formula of collum I-V+P+ma+P+V-I, that of midbody segments 3/3+I/i+3+I/ i+3/3; not only male coxa 1 but also femur 1 with a median outgrowth; northern
<collectingCountry id="43DA765EFFE3D51AD61DFCCDFBC8FF68" box="[1043,1133,829,853]" name="Thailand" pageId="34" pageNumber="451">Thailand</collectingCountry>
..................................................................................................................
<taxonomicName id="FCCD4D4DFFE3D51AD1FAFCAAFBCBFF4C" ID-CoL="3GGKY" authorityName="Mauries" authorityYear="1983" box="[1012,1134,858,881]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="451" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="09B9EADCFFE3D51AD1FAFCAAFBCBFF4C" box="[1012,1134,858,881]" italics="true" pageId="34" pageNumber="451">G. siamensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE3D51AD2B3FC87FBCBFF96" blockId="34.[189,1134,800,939]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FC87FBCBFF96" pageId="34" pageNumber="451">— Carinotaxy formulae not as above; only male coxae 1 with a pair of paramedian processes ................................................................................................................................. 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="80392E6BFFE3D51AD2B3FC38FBCBFFC0" pageId="34" pageNumber="451">
<paragraph id="3B7236CEFFE3D51AD2B3FC38FBCBFFDD" blockId="34.[189,1134,968,1021]" box="[189,1134,968,992]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FC38FBCBFFDD" box="[189,1134,968,992]" pageId="34" pageNumber="451">14. Carinotaxy formula of midbody segments 2+I/i+3+I/i+2 .......................................... 15</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE3D51AD2B3FC15FBCBFFC0" blockId="34.[189,1134,968,1021]" box="[189,1134,997,1021]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FC15FBCBFFC0" box="[189,1134,997,1021]" pageId="34" pageNumber="451">— Carinotaxy formula of midbody segments 2/2+I/i+3/3+I/i+2/2 ................................ 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="80392E6BFFE3D51AD2B3FBE9FBCBF8E3" pageId="34" pageNumber="451">
<paragraph id="3B7236CEFFE3D51AD2B3FBE9FBCBF8BA" blockId="34.[189,1134,1049,1246]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FBE9FBCBF8BA" pageId="34" pageNumber="451">
15. Carinotaxy formula of collum I-III+4c+5a+pc+ma+pc+5a+4c+III-I (
<figureCitation id="A3F62A4BFFE3D51AD1D5FBE9FBFAF80F" box="[987,1119,1049,1074]" captionStart="FIG" captionStartId="28.[113,124,1657,1674]" captionTargetBox="[116,1205,215,1587]" captionTargetId="figure-146@28.[628,876,845,1029]" captionTargetPageId="28" captionText="FIG. 22.— Glyphiulus paracostulifer n.sp., paratype ♀: A, anterior part of body,lateral view;B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393680" httpUri="https://zenodo.org/record/5393680/files/figure.png" pageId="34" pageNumber="451">Fig. 22A, B</figureCitation>
); male telopodites 1 reduced in size (
<figureCitation id="A3F62A4BFFE3D51AD057FBC6FD12F873" box="[601,695,1078,1102]" captionStart="FIG" captionStartId="30.[113,124,1286,1303]" captionTargetBox="[115,1207,204,1257]" captionTargetId="figure-23@30.[115,1207,204,1257]" captionTargetPageId="30" captionText="FIG. 24. — Glyphiulus paracostulifer n. sp., paratype ♂: A, antenna; B, leg 1, frontal view; C, leg 2, caudal view; D, leg 3; E, leg 11; F, anterior gonopods, caudal view; G, posterior gonopods, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393684" httpUri="https://zenodo.org/record/5393684/files/figure.png" pageId="34" pageNumber="451">Fig. 24B</figureCitation>
); half-anterior gonopod coxosternum narrowly notched at distal margin (
<figureCitation id="A3F62A4BFFE3D51AD03DFBA3FD2BF856" box="[563,654,1107,1131]" captionStart="FIG" captionStartId="30.[113,124,1286,1303]" captionTargetBox="[115,1207,204,1257]" captionTargetId="figure-23@30.[115,1207,204,1257]" captionTargetPageId="30" captionText="FIG. 24. — Glyphiulus paracostulifer n. sp., paratype ♂: A, antenna; B, leg 1, frontal view; C, leg 2, caudal view; D, leg 3; E, leg 11; F, anterior gonopods, caudal view; G, posterior gonopods, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393684" httpUri="https://zenodo.org/record/5393684/files/figure.png" pageId="34" pageNumber="451">Fig. 24F</figureCitation>
); cave in
<collectingRegion id="F909F82CFFE3D51AD0E0FBA3FC0EF856" box="[750,939,1107,1131]" country="China" name="Guizhou" pageId="34" pageNumber="451">Guizhou Province</collectingRegion>
,
<collectingCountry id="43DA765EFFE3D51AD1BBFBA3FC50F856" box="[949,1013,1107,1131]" name="China" pageId="34" pageNumber="451">China</collectingCountry>
................. ...................................................................................................
<taxonomicName id="FCCD4D4DFFE3D51AD183FB80FB95F8B5" ID-CoL="3GGKH" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[909,1072,1136,1160]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="451" phylum="Arthropoda" rank="species" species="paracostulifer" status="sp. nov.">
<emphasis id="09B9EADCFFE3D51AD183FB80FB95F8B5" box="[909,1072,1136,1160]" italics="true" pageId="34" pageNumber="451">G. paracostulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFE3D51AD638FB80FBCBF8BA" box="[1078,1134,1136,1159]" pageId="34" pageNumber="451" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE3D51AD2B3FB7DFBCBF8E3" blockId="34.[189,1134,1049,1246]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FB7DFBCBF8E3" pageId="34" pageNumber="451">
— Carinotaxy formula of collum I-IV+P+ma+P+IV-I; male telopodites 1 not reduced in size; half-anterior gonopod coxosternum broadly and barely emarginate at distal margin; cave in northern
<collectingCountry id="43DA765EFFE3D51AD394FB36FE56F8E3" box="[410,499,1222,1246]" name="Vietnam" pageId="34" pageNumber="451">Vietnam</collectingCountry>
......................................................................
<taxonomicName id="FCCD4D4DFFE3D51AD1D7FB36FBCBF8E3" ID-CoL="7WC45" authorityName=", Mauries" authorityYear="1977" box="[985,1134,1222,1246]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="451" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="09B9EADCFFE3D51AD1D7FB36FBCBF8E3" box="[985,1134,1222,1246]" italics="true" pageId="34" pageNumber="451">G. vietnamicus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="80392E6BFFE3D51AD2B3FB0AFBCBF9BB" pageId="34" pageNumber="451">
<paragraph id="3B7236CEFFE3D51AD2B3FB0AFBCBF971" blockId="34.[189,1134,1274,1414]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FB0AFBCBF971" pageId="34" pageNumber="451">
16. Coloration entirely pallid, ocelli not visible; adult body up to about 1.0 mm wide; paramedian outgrowths of anterior gonopod half-coxosternum especially high and large (e.g.,
<figureCitation id="A3F62A4BFFE3D51AD32AFAC4FE24F971" box="[292,385,1332,1356]" captionStart="FIG" captionStartId="27.[113,124,1196,1213]" captionTargetBox="[128,1201,226,1152]" captionTargetId="figure-19@27.[110,1214,200,1174]" captionTargetPageId="27" captionText="FIG. 21. — Glyphiulus percostulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="34" pageNumber="451">Fig. 21G</figureCitation>
) ......................................................................................................... 17
</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE3D51AD2B3FAA1FBCBF9BB" blockId="34.[189,1134,1274,1414]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FAA1FBCBF9BB" pageId="34" pageNumber="451">
— Coloration usually darker, ocelli always dark and discernible; body at least
<quantity id="FC359B2BFFE3D51AD1EEFAA1FB94F955" box="[992,1073,1361,1385]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="34" pageNumber="451" unit="mm" value="1.3">1.3 mm</quantity>
wide; paramedian outgrowths of anterior gonopod half-coxosternum less conspicuous ...... 18
</keyLead>
</paragraph>
</keyStep>
<keyStep id="80392E6BFFE3D51AD2B3FA52FBCBFA9D" pageId="34" pageNumber="451">
<paragraph id="3B7236CEFFE3D51AD2B3FA52FBCBFA2D" blockId="34.[189,1135,1442,1696]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3FA52FBCBFA2D" pageId="34" pageNumber="451">
17. Ozoporiferous cones much wider than high; epiproct normal, smooth dorsally; male telopodites 1 reduced, 2-segmented; half-anterior gonopod with a particularly deep notch, such that both distal outgrowths are extremely prominent; posterior gonopods without flagella;
<collectingRegion id="F909F82CFFE3D51AD331FA09FECEFA2D" box="[319,363,1529,1552]" country="Indonesia" name="Jawa Timur" pageId="34" pageNumber="451">Java</collectingRegion>
,
<collectingCountry id="43DA765EFFE3D51AD37BFA08FE7CFA2D" box="[373,473,1528,1552]" name="Indonesia" pageId="34" pageNumber="451">Indonesia</collectingCountry>
(introduction from continental Southeast Asia?) ...
<taxonomicName id="FCCD4D4DFFE3D51AD1FCFA09FBCBFA2D" ID-CoL="3GGK3" authorityName="Carl" authorityYear="1911" box="[1010,1134,1529,1552]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="451" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="09B9EADCFFE3D51AD1FCFA09FBCBFA2D" box="[1010,1134,1529,1552]" italics="true" pageId="34" pageNumber="451">G. javanicus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE3D51AD2B3F9E6FBCBFA9D" blockId="34.[189,1135,1442,1696]" pageId="34" pageNumber="451">
<keyLead id="803C95FBFFE3D51AD2B3F9E6FBCBFA9D" pageId="34" pageNumber="451">
— Ozoporiferous cones about as high as wide (
<figureCitation id="A3F62A4BFFE3D51AD0B4F9E5FCA1FA10" box="[698,772,1557,1581]" captionStart="FIG" captionStartId="26.[113,124,1497,1514]" captionTargetBox="[118,1205,215,1453]" captionTargetId="figure-126@26.[615,868,661,844]" captionTargetPageId="26" captionText="FIG. 20. — Glyphiulus percostulifer n. sp., paratype ♀: A, anterior part of body, lateral view; B, midbody segment section; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393676" httpUri="https://zenodo.org/record/5393676/files/figure.png" pageId="34" pageNumber="451">Fig. 20</figureCitation>
A-E); epiproct unusual in showing two paramedian crests (
<figureCitation id="A3F62A4BFFE3D51AD3EEF9C2FD98FA77" box="[480,573,1586,1610]" captionStart="FIG" captionStartId="26.[113,124,1497,1514]" captionTargetBox="[118,1205,215,1453]" captionTargetId="figure-126@26.[615,868,661,844]" captionTargetPageId="26" captionText="FIG. 20. — Glyphiulus percostulifer n. sp., paratype ♀: A, anterior part of body, lateral view; B, midbody segment section; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393676" httpUri="https://zenodo.org/record/5393676/files/figure.png" pageId="34" pageNumber="451">Fig. 20E</figureCitation>
); male telopodites 1 normal, 5-segmented (
<figureCitation id="A3F62A4BFFE3D51AD60CF9C2FBC4FA77" box="[1026,1121,1586,1610]" captionStart="FIG" captionStartId="27.[113,124,1196,1213]" captionTargetBox="[128,1201,226,1152]" captionTargetId="figure-19@27.[110,1214,200,1174]" captionTargetPageId="27" captionText="FIG. 21. — Glyphiulus percostulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="34" pageNumber="451">Fig. 21C</figureCitation>
); half-anterior gonopod weakly emarginated at distal margin, only paramedian outgrowths prominent (
<figureCitation id="A3F62A4BFFE3D51AD368F99BFE63FABE" box="[358,454,1643,1667]" captionStart="FIG" captionStartId="27.[113,124,1196,1213]" captionTargetBox="[128,1201,226,1152]" captionTargetId="figure-19@27.[110,1214,200,1174]" captionTargetPageId="27" captionText="FIG. 21. — Glyphiulus percostulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="34" pageNumber="451">Fig. 21G</figureCitation>
); posterior gonopods with flagella (
<figureCitation id="A3F62A4BFFE3D51AD13FF99BFC34FABE" box="[817,913,1643,1667]" captionStart="FIG" captionStartId="27.[113,124,1196,1213]" captionTargetBox="[128,1201,226,1152]" captionTargetId="figure-19@27.[110,1214,200,1174]" captionTargetPageId="27" captionText="FIG. 21. — Glyphiulus percostulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="34" pageNumber="451">Fig. 21H</figureCitation>
); cave in
<collectingCountry id="43DA765EFFE3D51AD1FDF99CFB87FABE" box="[1011,1058,1644,1667]" name="Laos" pageId="34" pageNumber="451">Laos</collectingCountry>
.......... .....................................................................................................
<taxonomicName id="FCCD4D4DFFE3D51AD195F978FB95FA9D" ID-CoL="3GGKN" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[923,1072,1672,1696]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="34" pageNumber="451" phylum="Arthropoda" rank="species" species="percostulifer" status="sp. nov.">
<emphasis id="09B9EADCFFE3D51AD195F978FB95FA9D" box="[923,1072,1672,1696]" italics="true" pageId="34" pageNumber="451">G. percostulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFE3D51AD638F979FBCBFA9D" box="[1078,1134,1673,1696]" pageId="34" pageNumber="451" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="80392E6BFFE2D51BD2B3FF28FBCBFDEB" pageId="35" pageNumber="452">
<paragraph id="3B7236CEFFE2D51BD2B3FF28FBCBFD5F" blockId="35.[189,1134,216,470]" pageId="35" pageNumber="452">
<keyLead id="803C95FBFFE2D51BD2B3FF28FBCBFD5F" pageId="35" pageNumber="452">
18. Carinotaxy formula of collum particularly complex, I-II+3c+4a+5c+VI+7c+8a+pc+ma +pc+8a+7c+VI+5c+4a+3c+II-I (
<figureCitation id="A3F62A4BFFE2D51BD038FF05FD30FD30" box="[566,661,245,269]" captionStart="FIG" captionStartId="23.[113,124,1426,1443]" captionTargetBox="[117,1206,215,1382]" captionTargetId="figure-58@23.[389,662,753,934]" captionTargetPageId="23" captionText="FIG. 17.— Glyphiulus subcostulifer n. sp., paratype ♂: A, ocellarium;B, collum,dorsal view;C, midbody segments,dorsal view;D, same, lateral view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, 0.1 mm; B, C, F, 0.2 mm; D, E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393668" httpUri="https://zenodo.org/record/5393668/files/figure.png" pageId="35" pageNumber="452">Fig. 17B</figureCitation>
); male telopodites 1 reduced in size, but still 5-segmented (
<figureCitation id="A3F62A4BFFE2D51BD375FEE1FE7FFD14" box="[379,474,273,297]" captionStart="FIG" captionStartId="25.[113,124,1066,1083]" captionTargetBox="[113,1210,221,1106]" captionTargetId="figure-19@25.[79,1229,184,1072]" captionTargetPageId="25" captionText="FIG. 19.— Glyphiulus subcostulifer n. sp.,paratype ♂: A, antenna;B, gnathochilarium;C, leg 1, frontal view; D, leg 2; E, leg 3; F, leg 11; G, right anterior gonopod, frontal view; H, posterior gonopods, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393672" httpUri="https://zenodo.org/record/5393672/files/figure.png" pageId="35" pageNumber="452">Fig. 19C</figureCitation>
); half-anterior gonopod coxosternum subsecuriform (
<figureCitation id="A3F62A4BFFE2D51BD60FFEE1FBC4FD14" box="[1025,1121,273,297]" captionStart="FIG" captionStartId="25.[113,124,1066,1083]" captionTargetBox="[113,1210,221,1106]" captionTargetId="figure-19@25.[79,1229,184,1072]" captionTargetPageId="25" captionText="FIG. 19.— Glyphiulus subcostulifer n. sp.,paratype ♂: A, antenna;B, gnathochilarium;C, leg 1, frontal view; D, leg 2; E, leg 3; F, leg 11; G, right anterior gonopod, frontal view; H, posterior gonopods, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393672" httpUri="https://zenodo.org/record/5393672/files/figure.png" pageId="35" pageNumber="452">Fig. 19G</figureCitation>
); flagella of posterior gonopods plumose distally (
<figureCitation id="A3F62A4BFFE2D51BD0DBFEDEFC93FD7B" box="[725,822,302,326]" captionStart="FIG" captionStartId="25.[113,124,1066,1083]" captionTargetBox="[113,1210,221,1106]" captionTargetId="figure-19@25.[79,1229,184,1072]" captionTargetPageId="25" captionText="FIG. 19.— Glyphiulus subcostulifer n. sp.,paratype ♂: A, antenna;B, gnathochilarium;C, leg 1, frontal view; D, leg 2; E, leg 3; F, leg 11; G, right anterior gonopod, frontal view; H, posterior gonopods, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393672" httpUri="https://zenodo.org/record/5393672/files/figure.png" pageId="35" pageNumber="452">Fig. 19H</figureCitation>
); cave in
<collectingCountry id="43DA765EFFE2D51BD199FEDFFC63FD7B" box="[919,966,303,326]" name="Laos" pageId="35" pageNumber="452">Laos</collectingCountry>
........................ .....................................................................................................
<taxonomicName id="FCCD4D4DFFE2D51BD196FEBBFB95FD5E" ID-CoL="3GGL4" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[920,1072,331,355]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="35" pageNumber="452" phylum="Arthropoda" rank="species" species="subcostulifer" status="sp. nov.">
<emphasis id="09B9EADCFFE2D51BD196FEBBFB95FD5E" box="[920,1072,331,355]" italics="true" pageId="35" pageNumber="452">G. subcostulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFE2D51BD638FEBBFBCBFD5F" box="[1078,1134,331,354]" pageId="35" pageNumber="452" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="3B7236CEFFE2D51BD2B3FE98FBCBFDEB" blockId="35.[189,1134,216,470]" pageId="35" pageNumber="452">
<keyLead id="803C95FBFFE2D51BD2B3FE98FBCBFDEB" pageId="35" pageNumber="452">
— Carinotaxy formula of collum I+2c+III-IV+5c+6a+pc+ma+pc+6a+5c+IV-III+2c+I (
<figureCitation id="A3F62A4BFFE2D51BD2FCFE74FECFFDA1" box="[242,362,388,412]" captionStart="FIG" captionStartId="31.[113,124,1520,1537]" captionTargetBox="[118,1205,215,1480]" captionTargetId="figure-97@31.[647,736,596,974]" captionTargetPageId="31" captionText="FIG. 25.— Glyphiulus intermedius n. sp., paratype ♀: A, anterior part of body,lateral view;B, same,dorsal view;C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, midbody segment section. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393686" httpUri="https://zenodo.org/record/5393686/files/figure.png" pageId="35" pageNumber="452">Fig. 25A, B</figureCitation>
); male telopodites 1 normal, 5-segmented (
<figureCitation id="A3F62A4BFFE2D51BD110FE74FCD9FDA1" box="[798,892,388,412]" captionStart="FIG" captionStartId="32.[113,124,1116,1133]" captionTargetBox="[137,1184,222,1056]" captionTargetId="figure-24@32.[99,1242,189,1097]" captionTargetPageId="32" captionText="FIG. 26. — Glyphiulus intermedius n. sp., ♂ paratype: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 11; G, anterior gonopods, caudal view; H, posterior gonopods, caudal view; I, left posterior gonopod, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393688" httpUri="https://zenodo.org/record/5393688/files/figure.png" pageId="35" pageNumber="452">Fig. 26C</figureCitation>
); half-anterior gonopod coxosternum as in
<figureCitation id="A3F62A4BFFE2D51BD3A2FE51FD83FD84" box="[428,550,417,441]" captionStart="FIG" captionStartId="32.[113,124,1116,1133]" captionTargetBox="[137,1184,222,1056]" captionTargetId="figure-24@32.[99,1242,189,1097]" captionTargetPageId="32" captionText="FIG. 26. — Glyphiulus intermedius n. sp., ♂ paratype: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 11; G, anterior gonopods, caudal view; H, posterior gonopods, caudal view; I, left posterior gonopod, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393688" httpUri="https://zenodo.org/record/5393688/files/figure.png" pageId="35" pageNumber="452">Figure 26G</figureCitation>
; posterior gonopods as in
<figureCitation id="A3F62A4BFFE2D51BD13BFE51FC15FD84" box="[821,944,417,441]" captionStart="FIG" captionStartId="32.[113,124,1116,1133]" captionTargetBox="[137,1184,222,1056]" captionTargetId="figure-24@32.[99,1242,189,1097]" captionTargetPageId="32" captionText="FIG. 26. — Glyphiulus intermedius n. sp., ♂ paratype: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 11; G, anterior gonopods, caudal view; H, posterior gonopods, caudal view; I, left posterior gonopod, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393688" httpUri="https://zenodo.org/record/5393688/files/figure.png" pageId="35" pageNumber="452">Figure 26H</figureCitation>
, flagella bare; cave in
<collectingRegion id="F909F82CFFE2D51BD308FE4EFE1EFDEB" box="[262,443,446,470]" country="China" name="Sichuan" pageId="35" pageNumber="452">Sichuan Province</collectingRegion>
,
<collectingCountry id="43DA765EFFE2D51BD3CBFE4EFDA0FDEB" box="[453,517,446,470]" name="China" pageId="35" pageNumber="452">China</collectingCountry>
...................................................................
<taxonomicName id="FCCD4D4DFFE2D51BD1D4FE4EFBCBFDEB" ID-CoL="3GGK2" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[986,1134,446,470]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="35" pageNumber="452" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="09B9EADCFFE2D51BD1D4FE4EFBCBFDEB" box="[986,1134,446,470]" italics="true" pageId="35" pageNumber="452">G. intermedius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>