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<document id="F7DDF3405435715EEB1A1E0FBEB3DED4" ID-DOI="10.5281/zenodo.5393630" ID-ISSN="1638-9387" ID-Zenodo-Dep="5393630" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630319175263" checkinUser="marcus" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; Spiegel, Didier Van Den" docDate="2007" docId="B36487D8FFD9D521D2D5FD42FC5AF95A" docLanguage="en" docName="Zoosystema.2007.29.3.417-456.pdf" docOrigin="Zoosystema 29 (3)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Glyphiulus percostulifer Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel 2007, n. sp." docType="treatment" docVersion="4" lastPageNumber="442" masterDocId="4F5DFFA0FFC1D538D20EFFF0FFA5FC3D" masterDocTitle="Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group" masterLastPageNumber="456" masterPageNumber="417" pageNumber="441" updateTime="1699225420998" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="F15B02A1987FA82779F83447032DB2FB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FEC7E1741F4D7FAB865BE7F4BB01A49D">
<mods:title id="23EAE0E954E865243DEDBD081D17A391">Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group</mods:title>
</mods:titleInfo>
<mods:name id="9843B36E38DF0EAAFB6C9527BCA8D710" type="personal">
<mods:role id="3DB71670013FBCCDF7F9A9DA63979F2D">
<mods:roleTerm id="3B52123057D95D01C32B9BAB7EB727E2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D41DD4981E600E33D81DCE98C3F47BC5">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="D0542D2AAE888C4787C521AB068CFB9B">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
<mods:nameIdentifier id="956C99535B5E32535B434C9CC1D6BF72" type="email">sgol@orc.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="57BFA1D3917F96D30304D3CCBF3BACE8" type="personal">
<mods:role id="A6D3030253F74037E1DE8FFE6056FDC3">
<mods:roleTerm id="F37FBB15FBD3937745813C9A827C8921">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9576A2B410CDE99567EA5C68A50660B6">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="43F7A58DC98EAC3C67B10A04F8203B04">Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="EB9F09A8426174DF7F54F67E3E0AB270" type="email">geoffroy@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="49A875A2C16F6D08E5E4EDEDF50098A3" type="personal">
<mods:role id="BB148F316FCD99C1EDBDB6DA142F7CA5">
<mods:roleTerm id="3C7B1C6EC6EB4F3B281806D4C27D0988">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AD968BE8924AC5218EAE9399D20412B3">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="09ADA875CD6EA744F6BEFD8E777BFA07">Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="6C810732D67789825B236D85A8FC9B75" type="email">collmill@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="69646F3F65E5591FDD58450FCBE010CC" type="personal">
<mods:role id="708E141C7C6140E7B7852DC6DD916FC5">
<mods:roleTerm id="BCE3FCB6FF8FF8D7BC0629530F9E1B99">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8ABC5A880F54252869000131E36A62B4">Spiegel, Didier Van Den</mods:namePart>
<mods:affiliation id="2665E56135063BED08ED9A502E3AE838">Musée royal de l’Afrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be</mods:affiliation>
</mods:name>
<mods:typeOfResource id="B75E89B16E6D63B4D3413E2A1289C879">text</mods:typeOfResource>
<mods:relatedItem id="8B9BD870BE9AE0BD8286DFFDE7DB7E9F" type="host">
<mods:titleInfo id="0C47B1015EEFF1D35F97C2143E7A510F">
<mods:title id="73CD8EA0E2E470D053D3A475A8D9714E">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="D323BC3FA15F264F2E77D359B8840B88">
<mods:date id="F2A272EA7EA98328EB740128A8D638DB">2007</mods:date>
<mods:detail id="20EA9008C2635D9E7469D50985E5271A" type="volume">
<mods:number id="108A2212FCF5637C06A47C1E3A55CFB4">29</mods:number>
</mods:detail>
<mods:detail id="7B8183A620632D72234F5907D30946D8" type="issue">
<mods:number id="61C91E00D882436BB118225173D80318">3</mods:number>
</mods:detail>
<mods:extent id="5E753539DEF1ECC9C11A5DCEECCC7D20" unit="page">
<mods:start id="BD2B51F07F605A9F3AC02C1E9B832E3E">417</mods:start>
<mods:end id="AFE5F529C0F3956C8CE1C4C948BF43C7">456</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="435C447DB1E19AE86470D8C4FE4054B4">journal article</mods:classification>
<mods:identifier id="76834600229143E153554E3D656C05E8" type="DOI">10.5281/zenodo.5393630</mods:identifier>
<mods:identifier id="27E01470B5CDDD4644696DFC1B69D0F6" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="61C62D848BE8DAD2939FA7CB8BA31077" type="Zenodo-Dep">5393630</mods:identifier>
</mods:mods>
<treatment id="B36487D8FFD9D521D2D5FD42FC5AF95A" ID-DOI="http://doi.org/10.5281/zenodo.5469542" ID-Zenodo-Dep="5469542" LSID="urn:lsid:plazi:treatment:B36487D8FFD9D521D2D5FD42FC5AF95A" httpUri="http://treatment.plazi.org/id/B36487D8FFD9D521D2D5FD42FC5AF95A" lastPageId="25" lastPageNumber="442" pageId="24" pageNumber="441">
<subSubSection id="73D76545FFD9D520D2D5FD42FDB8FEF3" box="[219,541,690,718]" pageId="24" pageNumber="441" type="nomenclature">
<paragraph id="3B7236CEFFD9D520D2D5FD42FDB8FEF3" blockId="24.[219,541,690,749]" box="[219,541,690,718]" pageId="24" pageNumber="441">
<heading id="603A81A2FFD9D520D2D5FD42FDB8FEF3" box="[219,541,690,718]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="441" reason="2">
<taxonomicName id="FCCD4D4DFFD9D520D2D5FD42FE7EFEF0" ID-CoL="3GGKN" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[219,475,690,717]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="441" phylum="Arthropoda" rank="species" species="percostulifer" status="sp. nov.">
<emphasis id="09B9EADCFFD9D520D2D5FD42FE7EFEF0" bold="true" box="[219,475,690,717]" italics="true" pageId="24" pageNumber="441">Glyphiulus percostulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFD9D520D3ECFD44FDB8FEF3" box="[482,541,692,718]" pageId="24" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="73D76545FFD9D520D339FD23FE65FED0" box="[311,448,723,749]" pageId="24" pageNumber="441" type="description">
<paragraph id="3B7236CEFFD9D520D339FD23FE65FED0" blockId="24.[219,541,690,749]" box="[311,448,723,749]" pageId="24" pageNumber="441">
(
<figureCitation id="A3F62A4BFFD9D520D330FD23FE34FED0" box="[318,401,723,749]" captionStart="FIG" captionStartId="26.[113,124,1497,1514]" captionTargetBox="[118,1205,215,1453]" captionTargetId="figure-126@26.[615,868,661,844]" captionTargetPageId="26" captionText="FIG. 20. — Glyphiulus percostulifer n. sp., paratype ♀: A, anterior part of body, lateral view; B, midbody segment section; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393676" httpUri="https://zenodo.org/record/5393676/files/figure.png" pageId="24" pageNumber="441">Figs 20</figureCitation>
;
<figureCitation id="A3F62A4BFFD9D520D392FD23FE1CFED0" box="[412,441,723,749]" captionStart="FIG" captionStartId="27.[113,124,1196,1213]" captionTargetBox="[128,1201,226,1152]" captionTargetId="figure-19@27.[110,1214,200,1174]" captionTargetPageId="27" captionText="FIG. 21. — Glyphiulus percostulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="24" pageNumber="441">21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="73D76545FFD9D520D27FFCFCFEB0FF95" pageId="24" pageNumber="441" type="materials_examined">
<paragraph id="3B7236CEFFD9D520D27FFCFCFEB0FF95" blockId="24.[113,647,780,936]" pageId="24" pageNumber="441">
<materialsCitation id="8BA53C93FFD9D520D27FFCFCFD26FF4E" collectingDate="1998-02-11" collectionCode="TYPE, MATERIAL, MNHN" collectorName="A. Bedos &amp; L. Deharveng" country="Laos" latitude="17.9" location="Tham The" longLatPrecision="766" longitude="104.49" municipality="Ban Nakok" pageId="24" pageNumber="441" specimenCode="GA 051" specimenCount="1" specimenCount-male="1" stateProvince="Khammouan" typeStatus="holotype">
<collectionCode id="5DDCAE0BFFD9D520D27FFCFCFEB7FF1F" box="[113,274,780,803]" pageId="24" pageNumber="441">TYPE MATERIAL</collectionCode>
. —
<emphasis id="09B9EADCFFD9D520D34BFCFCFED8FF19" bold="true" box="[325,381,780,804]" pageId="24" pageNumber="441">
<collectingCountry id="43DA765EFFD9D520D34BFCFCFEDCFF19" box="[325,377,780,804]" name="Laos" pageId="24" pageNumber="441">Laos</collectingCountry>
.
</emphasis>
<collectingRegion id="F909F82CFFD9D520D389FCFCFDF7FF1E" box="[391,594,780,804]" country="Laos" name="Khammouan" pageId="24" pageNumber="441">Khammouan Prov.</collectingRegion>
,
<collectingMunicipality id="DB16ACB4FFD9D520D050FCFCFF10FF03" pageId="24" pageNumber="441">Ban Nakok</collectingMunicipality>
(= Nakhok),
<location id="3E126015FFD9D520D34AFCD6FE0AFF03" LSID="urn:lsid:plazi:treatment:B36487D8FFD9D521D2D5FD42FC5AF95A:3E126015FFD9D520D34AFCD6FE0AFF03" box="[324,431,806,830]" country="Laos" latitude="17.9" longLatPrecision="766" longitude="104.49" municipality="Ban Nakok" name="Tham The" pageId="24" pageNumber="441" stateProvince="Khammouan">Tham Thê</location>
,
<geoCoordinate id="5EF95009FFD9D520D3B5FCD6FDB9FF03" box="[443,540,806,830]" degrees="104.49" direction="east" orientation="longitude" pageId="24" pageNumber="441" precision="555" value="104.49">104.49°E</geoCoordinate>
,
<geoCoordinate id="5EF95009FFD9D520D026FCD6FD27FF03" box="[552,642,806,830]" degrees="17.90" direction="north" orientation="latitude" pageId="24" pageNumber="441" precision="555" value="17.9">17.90°N</geoCoordinate>
, cave, hand collection,
<date id="4F73100EFFD9D520D35FFCB1FE18FF64" box="[337,445,833,857]" pageId="24" pageNumber="441" value="1998-02-11">
<collectingDate id="5F37E9E6FFD9D520D35FFCB1FE18FF64" box="[337,445,833,857]" pageId="24" pageNumber="441" value="1998-02-11">11.II.1998</collectingDate>
</date>
, leg.
<collectorName id="96385318FFD9D520D3FAFCB1FDE8FF64" box="[500,589,833,857]" pageId="24" pageNumber="441">A. Bedos</collectorName>
&amp;
<collectorName id="96385318FFD9D520D07CFCB1FF7EFF4E" pageId="24" pageNumber="441">L. Deharveng</collectorName>
(LAO-016),
<typeStatus id="E476886CFFD9D520D354FCABFE0AFF4E" box="[346,431,859,883]" pageId="24" pageNumber="441" type="holotype">holotype</typeStatus>
<specimenCount id="2DCBFD47FFD9D520D3BAFC91FE6DFF4E" box="[436,456,865,883]" pageId="24" pageNumber="441" type="male">♂</specimenCount>
(
<collectionCode id="5DDCAE0BFFD9D520D3DDFCABFD82FF4E" box="[467,551,859,883]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="441">MNHN</collectionCode>
<specimenCode id="6B6B9EB5FFD9D520D025FCABFDDFFF4E" box="[555,634,859,883]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="441">GA 051</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8BA53C93FFD9D520D27FFC86FDFCFFB0" box="[113,601,885,910]" collectingDate="1998-02-11" collectionCode="MNHN" collectorName="A. Bedos &amp; L. Deharveng" country="Laos" isTailEnumeration="true" latitude="17.9" location="Tham The" longLatPrecision="766" longitude="104.49" municipality="Ban Nakok" pageId="24" pageNumber="441" specimenCode="GA 051" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Khammouan" typeStatus="paratype">
<typeStatus id="E476886CFFD9D520D27FFC86FF68FFB3" box="[113,205,886,910]" pageId="24" pageNumber="441" type="paratype">paratypes</typeStatus>
<specimenCount id="2DCBFD47FFD9D520D2DCFC86FF5DFFB0" box="[210,248,886,910]" pageId="24" pageNumber="441" type="male">1 ♂</specimenCount>
,
<specimenCount id="2DCBFD47FFD9D520D30DFC86FE83FFB0" box="[259,294,886,910]" pageId="24" pageNumber="441" type="female">1 ♀</specimenCount>
,
<specimenCount id="2DCBFD47FFD9D520D33FFC86FE3EFFB0" box="[305,411,885,910]" pageId="24" pageNumber="441" type="male">1 subad. ♂</specimenCount>
(
<collectionCode id="5DDCAE0BFFD9D520D3A6FC86FE59FFB3" box="[424,508,886,910]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="441">MNHN</collectionCode>
<specimenCode id="6B6B9EB5FFD9D520D00FFC86FDF4FFB0" box="[513,593,885,910]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="441">GA 051</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8BA53C93FFD9D520D06CFC86FEB4FF95" collectionCode="ZMUM, SEM" country="Laos" pageId="24" pageNumber="441" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="2DCBFD47FFD9D520D06CFC86FD23FFB0" box="[610,646,886,910]" pageId="24" pageNumber="441" type="female">1 ♀</specimenCount>
(
<collectionCode id="5DDCAE0BFFD9D520D276FC60FF69FF95" box="[120,204,912,936]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="24" pageNumber="441">ZMUM</collectionCode>
/
<collectionCode id="5DDCAE0BFFD9D520D2DAFC60FEAFFF95" box="[212,266,912,936]" pageId="24" pageNumber="441">SEM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="73D76545FFD9D520D27FFC30FF4FF831" pageId="24" pageNumber="441" type="etymology">
<paragraph id="3B7236CEFFD9D520D27FFC30FF4FF831" blockId="24.[113,646,960,1036]" pageId="24" pageNumber="441">
ETYMOLOGY. — To emphasize the very close resemblance to
<taxonomicName id="FCCD4D4DFFD9D520D285FC2AFF5EFFCF" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[139,251,986,1010]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="441" phylum="Arthropoda" rank="species" species="costulifer" status="sp. nov.">
<emphasis id="09B9EADCFFD9D520D285FC2AFF5EFFCF" box="[139,251,986,1010]" italics="true" pageId="24" pageNumber="441">G. costulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFD9D520D30EFC2BFE92FFCF" box="[256,311,987,1010]" pageId="24" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
, but with more strongly developed tergal crests.
</paragraph>
</subSubSection>
<subSubSection id="73D76545FFD9D520D27FFBD5FE9DF8B6" pageId="24" pageNumber="441" type="diagnosis">
<paragraph id="3B7236CEFFD9D520D27FFBD5FE9DF8B6" blockId="24.[113,647,1060,1164]" pageId="24" pageNumber="441">DIAGNOSIS. — Differs from congeners in the small size, the completely unpigmented tegument and ocelli, coupled with certain details of leg and gonopod structure (see also key below).</paragraph>
</subSubSection>
<subSubSection id="73D76545FFD9D521D27FFB5EFE53FA9E" lastPageId="25" lastPageNumber="442" pageId="24" pageNumber="441" type="description">
<paragraph id="3B7236CEFFD9D520D27FFB5EFEA7F8FB" blockId="24.[113,647,1198,1699]" box="[113,258,1198,1224]" pageId="24" pageNumber="441">DESCRIPTION</paragraph>
<paragraph id="3B7236CEFFD9D520D27FFB3DFE76F95A" blockId="24.[113,647,1198,1699]" pageId="24" pageNumber="441">
<typeStatus id="E476886CFFD9D520D27FFB3DFF72F8DA" box="[113,215,1229,1255]" pageId="24" pageNumber="441" type="holotype">Holotype</typeStatus>
with 34p+2a+T, about
<quantity id="FC359B2BFFD9D520D3DAFB3EFD87F8D5" box="[468,546,1230,1256]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="24" pageNumber="441" unit="mm" value="14.0">14 mm</quantity>
long and 1.0 mm wide; adult
<typeStatus id="E476886CFFD9D520D333FB1EFE38F935" box="[317,413,1262,1288]" pageId="24" pageNumber="441" type="paratype">paratypes</typeStatus>
with 36p+2a+T, about
<quantity id="FC359B2BFFD9D520D27FFAFDFF18F91A" box="[113,189,1293,1319]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="24" pageNumber="441" unit="mm" value="14.0">14 mm</quantity>
long and 1.0 mm wide (
<specimenCount id="2DCBFD47FFD9D520D3B6FAE3FE6BF91A" box="[440,462,1299,1319]" pageId="24" pageNumber="441" type="male">♂</specimenCount>
), with 40p+1a+T or 38p+1a-T, about
<quantity id="FC359B2BFFD9D520D348FADDFE36F97A" box="[326,403,1325,1351]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="24" pageNumber="441" unit="mm" value="17.0">17 mm</quantity>
long and 1.0 mm wide (
<specimenCount id="2DCBFD47FFD9D520D274FAA2FF28F95B" box="[122,141,1362,1382]" pageId="24" pageNumber="441" type="female">♀</specimenCount>
); subadult
<specimenCount id="2DCBFD47FFD9D520D306FAA2FEBBF95B" box="[264,286,1362,1382]" pageId="24" pageNumber="441" type="male">♂</specimenCount>
with 31p+2a+T.
</paragraph>
<paragraph id="3B7236CEFFD9D520D286FA9CFF4DF99B" blockId="24.[113,647,1198,1699]" pageId="24" pageNumber="441">
Coloration uniformly pallid, ocelli not visible (
<figureCitation id="A3F62A4BFFD9D520D277FA7CFF79F99B" box="[121,220,1420,1446]" captionStart="FIG" captionStartId="26.[113,124,1497,1514]" captionTargetBox="[118,1205,215,1453]" captionTargetId="figure-126@26.[615,868,661,844]" captionTargetPageId="26" captionText="FIG. 20. — Glyphiulus percostulifer n. sp., paratype ♀: A, anterior part of body, lateral view; B, midbody segment section; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393676" httpUri="https://zenodo.org/record/5393676/files/figure.png" pageId="24" pageNumber="441">Fig. 20A</figureCitation>
).
</paragraph>
<paragraph id="3B7236CEFFD9D521D286FA5BFE53FA9E" blockId="24.[113,647,1198,1699]" lastBlockId="25.[113,647,1166,1699]" lastPageId="25" lastPageNumber="442" pageId="24" pageNumber="441">
Postcollar constriction, dorsal crests and carinotaxy pattern on postcollar terga (
<figureCitation id="A3F62A4BFFD9D520D3F5FA3BFDEFF9D8" box="[507,586,1483,1509]" captionStart="FIG" captionStartId="26.[113,124,1497,1514]" captionTargetBox="[118,1205,215,1453]" captionTargetId="figure-126@26.[615,868,661,844]" captionTargetPageId="26" captionText="FIG. 20. — Glyphiulus percostulifer n. sp., paratype ♀: A, anterior part of body, lateral view; B, midbody segment section; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393676" httpUri="https://zenodo.org/record/5393676/files/figure.png" pageId="24" pageNumber="441">Fig. 20</figureCitation>
A-E), segment’s texture and cross-section (
<figureCitation id="A3F62A4BFFD9D520D3F7FA1BFDEDFA38" box="[505,584,1515,1541]" captionStart="FIG" captionStartId="26.[113,124,1497,1514]" captionTargetBox="[118,1205,215,1453]" captionTargetId="figure-126@26.[615,868,661,844]" captionTargetPageId="26" captionText="FIG. 20. — Glyphiulus percostulifer n. sp., paratype ♀: A, anterior part of body, lateral view; B, midbody segment section; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393676" httpUri="https://zenodo.org/record/5393676/files/figure.png" pageId="24" pageNumber="441">Fig. 20</figureCitation>
B-D), paraprocts and hypoproct (
<figureCitation id="A3F62A4BFFD9D520D39DF9FAFE56FA18" box="[403,499,1546,1573]" captionStart="FIG" captionStartId="26.[113,124,1497,1514]" captionTargetBox="[118,1205,215,1453]" captionTargetId="figure-126@26.[615,868,661,844]" captionTargetPageId="26" captionText="FIG. 20. — Glyphiulus percostulifer n. sp., paratype ♀: A, anterior part of body, lateral view; B, midbody segment section; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393676" httpUri="https://zenodo.org/record/5393676/files/figure.png" pageId="24" pageNumber="441">Fig. 20F</figureCitation>
), ventral flaps of male segment 7, male legs 2 and 3 (
<figureCitation id="A3F62A4BFFD9D520D3F4F9DAFDDCFA78" box="[506,633,1578,1605]" captionStart="FIG" captionStartId="27.[113,124,1196,1213]" captionTargetBox="[128,1201,226,1152]" captionTargetId="figure-19@27.[110,1214,200,1174]" captionTargetPageId="27" captionText="FIG. 21. — Glyphiulus percostulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="24" pageNumber="441">Fig. 21D, E</figureCitation>
), penes (
<figureCitation id="A3F62A4BFFD9D520D2B4F9BAFEBBFA59" box="[186,286,1610,1636]" captionStart="FIG" captionStartId="27.[113,124,1196,1213]" captionTargetBox="[128,1201,226,1152]" captionTargetId="figure-19@27.[110,1214,200,1174]" captionTargetPageId="27" captionText="FIG. 21. — Glyphiulus percostulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="24" pageNumber="441">Fig. 21D</figureCitation>
) generally as in
<taxonomicName id="FCCD4D4DFFD9D520D3CBF9BAFDE5FA59" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[453,576,1610,1636]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="441" phylum="Arthropoda" rank="species" species="costulifer" status="sp. nov.">
<emphasis id="09B9EADCFFD9D520D3CBF9BAFDE5FA59" box="[453,576,1610,1636]" italics="true" pageId="24" pageNumber="441">G. costulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFD9D520D04BF9BAFD27FA59" box="[581,642,1610,1636]" pageId="24" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
, but antennae elongate, antennomeres 5-7 richer in sensilla (
<figureCitation id="A3F62A4BFFD9D520D2E2F979FEF5FA9E" box="[236,336,1673,1699]" captionStart="FIG" captionStartId="27.[113,124,1196,1213]" captionTargetBox="[128,1201,226,1152]" captionTargetId="figure-19@27.[110,1214,200,1174]" captionTargetPageId="27" captionText="FIG. 21. — Glyphiulus percostulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="24" pageNumber="441">Fig. 21A</figureCitation>
), gnathochilarium (mentum divided, n = 2) (
<figureCitation id="A3F62A4BFFD8D521D315FB7EFED2F895" box="[283,375,1166,1192]" captionStart="FIG" captionStartId="27.[113,124,1196,1213]" captionTargetBox="[128,1201,226,1152]" captionTargetId="figure-19@27.[110,1214,200,1174]" captionTargetPageId="27" captionText="FIG. 21. — Glyphiulus percostulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="25" pageNumber="442">Fig. 21B</figureCitation>
) slightly even more oligotrichous; collum (
<figureCitation id="A3F62A4BFFD8D521D33AFB5EFE33F8F5" box="[308,406,1198,1224]" captionStart="FIG" captionStartId="26.[113,124,1497,1514]" captionTargetBox="[118,1205,215,1453]" captionTargetId="figure-126@26.[615,868,661,844]" captionTargetPageId="26" captionText="FIG. 20. — Glyphiulus percostulifer n. sp., paratype ♀: A, anterior part of body, lateral view; B, midbody segment section; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393676" httpUri="https://zenodo.org/record/5393676/files/figure.png" pageId="25" pageNumber="442">Fig. 20A</figureCitation>
) with a nearly typical carinotaxy formula (I-IV+5a+pc+ma+pc+5a+IV-I); all tergal crests (
<figureCitation id="A3F62A4BFFD8D521D32FFB1DFECBF93A" box="[289,366,1261,1287]" captionStart="FIG" captionStartId="26.[113,124,1497,1514]" captionTargetBox="[118,1205,215,1453]" captionTargetId="figure-126@26.[615,868,661,844]" captionTargetPageId="26" captionText="FIG. 20. — Glyphiulus percostulifer n. sp., paratype ♀: A, anterior part of body, lateral view; B, midbody segment section; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393676" httpUri="https://zenodo.org/record/5393676/files/figure.png" pageId="25" pageNumber="442">Fig. 20</figureCitation>
A-E) on subsequent body segments higher, subdivided (2/2+I/i+3/3+I/i+2/2), ozoporiferous cones about as high as wide; epiproct (
<figureCitation id="A3F62A4BFFD8D521D277FABCFF54F95A" box="[121,241,1356,1383]" captionStart="FIG" captionStartId="26.[113,124,1497,1514]" captionTargetBox="[118,1205,215,1453]" captionTargetId="figure-126@26.[615,868,661,844]" captionTargetPageId="26" captionText="FIG. 20. — Glyphiulus percostulifer n. sp., paratype ♀: A, anterior part of body, lateral view; B, midbody segment section; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393676" httpUri="https://zenodo.org/record/5393676/files/figure.png" pageId="25" pageNumber="442">Fig. 20E, F</figureCitation>
) upturned and broadly rounded in caudal part, with one median and two paramedian crests; legs slightly longer than body diameter (
<figureCitation id="A3F62A4BFFD8D521D016FA7CFDDCF99B" box="[536,633,1420,1446]" captionStart="FIG" captionStartId="26.[113,124,1497,1514]" captionTargetBox="[118,1205,215,1453]" captionTargetId="figure-126@26.[615,868,661,844]" captionTargetPageId="26" captionText="FIG. 20. — Glyphiulus percostulifer n. sp., paratype ♀: A, anterior part of body, lateral view; B, midbody segment section; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393676" httpUri="https://zenodo.org/record/5393676/files/figure.png" pageId="25" pageNumber="442">Fig. 20B</figureCitation>
), always with an accessory filament/spine at base of claw, which is usually slightly more than half as long as claw (
<figureCitation id="A3F62A4BFFD8D521D2F3FA1BFEE2FA38" box="[253,327,1515,1541]" captionStart="FIG" captionStartId="27.[113,124,1196,1213]" captionTargetBox="[128,1201,226,1152]" captionTargetId="figure-19@27.[110,1214,200,1174]" captionTargetPageId="27" captionText="FIG. 21. — Glyphiulus percostulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="25" pageNumber="442">Fig. 21</figureCitation>
D-F); male leg 1 with normal, 5-segmented telopodites and large, paramedian, coxal processes (
<figureCitation id="A3F62A4BFFD8D521D310F9DAFE24FA79" box="[286,385,1578,1604]" captionStart="FIG" captionStartId="27.[113,124,1196,1213]" captionTargetBox="[128,1201,226,1152]" captionTargetId="figure-19@27.[110,1214,200,1174]" captionTargetPageId="27" captionText="FIG. 21. — Glyphiulus percostulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="25" pageNumber="442">Fig. 21C</figureCitation>
); each anterior gonopod plate with a high and subsecuriform distomedial process (
<figureCitation id="A3F62A4BFFD8D521D2C7F999FE8DFAB9" box="[201,296,1641,1668]" captionStart="FIG" captionStartId="27.[113,124,1196,1213]" captionTargetBox="[128,1201,226,1152]" captionTargetId="figure-19@27.[110,1214,200,1174]" captionTargetPageId="27" captionText="FIG. 21. — Glyphiulus percostulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="25" pageNumber="442">Fig. 21G</figureCitation>
); posterior gonopods with distally slightly plumose flagella (
<figureCitation id="A3F62A4BFFD8D521D38CF979FE4CFA9E" box="[386,489,1673,1699]" captionStart="FIG" captionStartId="27.[113,124,1196,1213]" captionTargetBox="[128,1201,226,1152]" captionTargetId="figure-19@27.[110,1214,200,1174]" captionTargetPageId="27" captionText="FIG. 21. — Glyphiulus percostulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium, ventral view; C, leg 1, caudal view; D, leg 2, caudal view; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393678" httpUri="https://zenodo.org/record/5393678/files/figure.png" pageId="25" pageNumber="442">Fig. 21H</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="6FB26646FFD9D520D0ABF993FC16FA9C" ID-DOI="http://doi.org/10.5281/zenodo.5393670" ID-Zenodo-Dep="5393670" httpUri="https://zenodo.org/record/5393670/files/figure.png" pageId="24" pageNumber="441" startId="24.[677,688,1635,1652]" targetBox="[684,1203,216,1590]" targetPageId="24">
<paragraph id="3B7236CEFFD9D520D0ABF993FC16FA9C" blockId="24.[677,1210,1635,1697]" pageId="24" pageNumber="441">
FIG. 18. —
<taxonomicName id="FCCD4D4DFFD9D520D0F4F993FC18FA49" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[762,957,1635,1652]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="441" phylum="Arthropoda" rank="species" species="subcostulifer" status="sp. nov.">
<emphasis id="09B9EADCFFD9D520D0F4F993FC18FA49" box="[762,957,1635,1652]" italics="true" pageId="24" pageNumber="441">Glyphiulus subcostulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFD9D520D1CFF993FC57FA49" box="[961,1010,1635,1652]" pageId="24" pageNumber="441" rank="species">n.sp.</taxonomicNameLabel>
, paratype ♂:
<emphasis id="09B9EADCFFD9D520D655F993FBCDFA49" bold="true" box="[1115,1128,1635,1652]" pageId="24" pageNumber="441">A</emphasis>
, midbody segment section;
<emphasis id="09B9EADCFFD9D520D139F989FCE1FAB7" bold="true" box="[823,836,1657,1674]" pageId="24" pageNumber="441">B</emphasis>
, segment 7, frontolateral view;
<emphasis id="09B9EADCFFD9D520D64BF989FBF6FAB7" bold="true" box="[1093,1107,1657,1674]" pageId="24" pageNumber="441">C</emphasis>
, claw.Scale bars: A, B, 0.2 mm; C, 0.01 mm.
</paragraph>
</caption>
<caption id="6FB26646FFD8D521D27FFBDAFC22F86F" ID-DOI="http://doi.org/10.5281/zenodo.5393672" ID-Zenodo-Dep="5393672" httpUri="https://zenodo.org/record/5393672/files/figure.png" pageId="25" pageNumber="442" startId="25.[113,124,1066,1083]" targetBox="[113,1210,221,1106]" targetPageId="25">
<paragraph id="3B7236CEFFD8D521D27FFBDAFC22F86F" blockId="25.[113,1210,1066,1106]" pageId="25" pageNumber="442">
FIG. 19.—
<taxonomicName id="FCCD4D4DFFD8D521D2C6FBDAFE29F806" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[200,396,1066,1083]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="442" phylum="Arthropoda" rank="species" species="subcostulifer" status="sp. nov.">
<emphasis id="09B9EADCFFD8D521D2C6FBDAFE29F806" box="[200,396,1066,1083]" italics="true" pageId="25" pageNumber="442">Glyphiulus subcostulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFD8D521D39EFBDAFE64F806" box="[400,449,1066,1083]" pageId="25" pageNumber="442" rank="species">n. sp.</taxonomicNameLabel>
,paratype ♂:
<emphasis id="09B9EADCFFD8D521D025FBDAFD9DF806" bold="true" box="[555,568,1066,1083]" pageId="25" pageNumber="442">A</emphasis>
, antenna;
<emphasis id="09B9EADCFFD8D521D083FBDAFD3FF806" bold="true" box="[653,666,1066,1083]" pageId="25" pageNumber="442">B</emphasis>
, gnathochilarium;
<emphasis id="09B9EADCFFD8D521D13CFBDAFCE5F806" bold="true" box="[818,832,1066,1083]" pageId="25" pageNumber="442">C</emphasis>
, leg 1, frontal view;
<emphasis id="09B9EADCFFD8D521D1EEFBDAFC4BF806" bold="true" box="[992,1006,1066,1083]" pageId="25" pageNumber="442">D</emphasis>
, leg 2;
<emphasis id="09B9EADCFFD8D521D629FBDAFB96F806" bold="true" box="[1063,1075,1066,1083]" pageId="25" pageNumber="442">E</emphasis>
, leg 3;
<emphasis id="09B9EADCFFD8D521D662FBDAFBD2F806" bold="true" box="[1132,1143,1066,1083]" pageId="25" pageNumber="442">F</emphasis>
, leg 11;
<emphasis id="09B9EADCFFD8D521D27FFBB1FFDAF86F" bold="true" box="[113,127,1089,1106]" pageId="25" pageNumber="442">G</emphasis>
, right anterior gonopod, frontal view;
<emphasis id="09B9EADCFFD8D521D3B7FBB1FE62F86F" bold="true" box="[441,455,1089,1106]" pageId="25" pageNumber="442">H</emphasis>
, posterior gonopods, frontal view. Scale bar: 0.2 mm.
</paragraph>
</caption>
<subSubSection id="73D76545FFD8D521D0ABFB7FFC5AF95A" pageId="25" pageNumber="442" type="discussion">
<paragraph id="3B7236CEFFD8D521D0ABFB7FFCACF89A" blockId="25.[677,1210,1167,1383]" box="[677,777,1167,1193]" pageId="25" pageNumber="442">REMARKS</paragraph>
<paragraph id="3B7236CEFFD8D521D0ABFB5EFC5AF95A" blockId="25.[677,1210,1167,1383]" pageId="25" pageNumber="442">Based on several troglomorphic traits, such as the completely unpigmented body and ocelli, as well as the elongated antennae and legs, this species seems to be troglobitic, although the small size may also be evidence of its being an inhabitant of the MSS (“milieu souterrain superficiel”).</paragraph>
</subSubSection>
</treatment>
</document>
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