treatments-xml/data/B3/64/87/B36487D8FFC8D535D0F2FA5BFB03FA9E.xml
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<document id="059525F555ACBAB5027E28508B7C9507" ID-DOI="10.5281/zenodo.5393630" ID-ISSN="1638-9387" ID-Zenodo-Dep="5393630" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630319175263" checkinUser="marcus" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; Spiegel, Didier Van Den" docDate="2007" docId="B36487D8FFC8D535D0F2FA5BFB03FA9E" docLanguage="en" docName="Zoosystema.2007.29.3.417-456.pdf" docOrigin="Zoosystema 29 (3)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Glyphiulus mediobliteratus Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel 2007, n. sp." docType="treatment" docVersion="3" lastPageNumber="430" masterDocId="4F5DFFA0FFC1D538D20EFFF0FFA5FC3D" masterDocTitle="Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group" masterLastPageNumber="456" masterPageNumber="417" pageNumber="426" updateTime="1699225420998" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="85076774591238981BF269432E227F57">Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group</mods:title>
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<mods:namePart id="25142430EECEE32FAA6BE9752E5B2F8F">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="BFF7B10E3567B741596BE82C96FAB8D7">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
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<mods:namePart id="CBE133C91220400400B657A700DBC87A">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="6704223C88878526B07A8FCAD5B35957">Muséum national dHistoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
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<mods:namePart id="EB85279B5B5AEC3533510295BC317939">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="B4D798186E0435B91DF90214F1589BA8">Muséum national dHistoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
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<mods:affiliation id="768BBF7F8A9592D23E141A8B53CF3D5F">Musée royal de lAfrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be</mods:affiliation>
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<emphasis id="09B9EADCFFC8D531D0F2FA5BFB84F9FB" bold="true" box="[764,1057,1451,1478]" italics="true" pageId="9" pageNumber="426">Glyphiulus mediobliteratus</emphasis>
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<figureCitation id="A3F62A4BFFC8D531D18CFA3BFC7FF9DB" box="[898,986,1483,1510]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="11.[113,124,1635,1652]" captionStartId-1="12.[113,124,1073,1090]" captionStartId-2="13.[113,124,1198,1215]" captionTargetBox-0="[117,1205,215,1535]" captionTargetBox-1="[117,1206,215,1027]" captionTargetBox-2="[94,1236,194,1174]" captionTargetId-0="figure-109@11.[654,745,613,1104]" captionTargetId-1="figure-51@12.[389,662,576,758]" captionTargetId-2="figure-19@13.[94,1236,194,1174]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="FIG. 6. — Glyphiulus mediobliteratus n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, telson, ventral view. Scale bars: A, C, D, 0.5 mm; B, E, F, 0.2 mm." captionText-1="FIG. 7. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, midbody segment section;B, both gonopods in situ, frontal view;C, anterior gonopod, frontal view; D, anterior gonopod, caudal view; E, posterior gonopod, subfrontal view; F, claw. Scale bars: A, 0.5 mm; B-D, 0.1 mm; E, F, 0.05 mm." captionText-2="FIG. 8. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, antenna;B, gnathochilarium; C, leg 1, frontal view;D, leg 2; E, leg 3; F, leg 9; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5393642" figureDoi-1="http://doi.org/10.5281/zenodo.5393644" figureDoi-2="http://doi.org/10.5281/zenodo.5393646" httpUri-0="https://zenodo.org/record/5393642/files/figure.png" httpUri-1="https://zenodo.org/record/5393644/files/figure.png" httpUri-2="https://zenodo.org/record/5393646/files/figure.png" pageId="9" pageNumber="426">Figs 6-8</figureCitation>
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20.
<collectionCode id="5DDCAE0BFFCDD534D2C3FB38FF4FF8DD" box="[205,234,1224,1248]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="12" pageNumber="429">VI</collectionCode>
.2003
</collectingDate>
</date>
, leg.
<collectorName id="96385318FFCDD534D35BFB39FE1DF8DD" box="[341,440,1224,1248]" pageId="12" pageNumber="429">P. Trontelj</collectorName>
&amp;
<collectorName id="96385318FFCDD534D3D6FB39FD84F8DD" box="[472,545,1224,1248]" pageId="12" pageNumber="429">B. Sket</collectorName>
,
<typeStatus id="E476886CFFCDD534D025FB39FD22F8DC" box="[555,647,1225,1249]" pageId="12" pageNumber="429" type="paratype">paratypes</typeStatus>
<specimenCount id="2DCBFD47FFCDD534D27FFB13FF3DF8C7" box="[113,152,1251,1275]" pageId="12" pageNumber="429" type="male">1 ♂</specimenCount>
(
<collectionCode id="5DDCAE0BFFCDD534D2A8FB13FF59F8C6" box="[166,252,1251,1275]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="12" pageNumber="429">ZMUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8BA53C93FFCDD534D303FB13FE56F8C7" box="[269,499,1250,1275]" collectionCode="MNHN" country="Vietnam" pageId="12" pageNumber="429" specimenCode="GA 046" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="2DCBFD47FFCDD534D303FB13FE94F8C7" box="[269,305,1251,1275]" pageId="12" pageNumber="429" type="female">1 ♀</specimenCount>
(
<collectionCode id="5DDCAE0BFFCDD534D331FB13FE36F8C6" box="[319,403,1251,1275]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="429">MNHN</collectionCode>
<specimenCode id="6B6B9EB5FFCDD534D397FB13FE49F8C7" box="[409,492,1250,1275]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="429">GA 046</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="6FB26646FFCBD532D27FF993FDAEFA9C" ID-DOI="http://doi.org/10.5281/zenodo.5393640" ID-Zenodo-Dep="5393640" httpUri="https://zenodo.org/record/5393640/files/figure.png" pageId="10" pageNumber="427" startId="10.[113,124,1635,1652]" targetBox="[120,1205,223,1547]" targetPageId="10">
<paragraph id="3B7236CEFFCBD532D27FF993FDAEFA9C" blockId="10.[113,1211,1635,1697]" pageId="10" pageNumber="427">
FIG. 5. —
<taxonomicName id="FCCD4D4DFFCBD532D2CDF993FE30FA49" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[195,405,1635,1652]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="427" phylum="Arthropoda" rank="species" species="subobliteratus" status="sp. nov.">
<emphasis id="09B9EADCFFCBD532D2CDF993FE30FA49" box="[195,405,1635,1652]" italics="true" pageId="10" pageNumber="427">Glyphiulus subobliteratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFCBD532D394F993FE68FA49" box="[410,461,1635,1652]" pageId="10" pageNumber="427" rank="species">n. sp.</taxonomicNameLabel>
, paratype ♂:
<emphasis id="09B9EADCFFCBD532D032F993FDECFA49" bold="true" box="[572,585,1635,1652]" pageId="10" pageNumber="427">A</emphasis>
, antenna;
<emphasis id="09B9EADCFFCBD532D0ACF993FD0AFA49" bold="true" box="[674,687,1635,1652]" pageId="10" pageNumber="427">B</emphasis>
, gnathochilarium;
<emphasis id="09B9EADCFFCBD532D144F993FCFDFA49" bold="true" box="[842,856,1635,1652]" pageId="10" pageNumber="427">C</emphasis>
, leg 1, frontal view;
<emphasis id="09B9EADCFFCBD532D60FF993FBAAFA49" bold="true" box="[1025,1039,1635,1652]" pageId="10" pageNumber="427">D</emphasis>
, same, caudal view;
<emphasis id="09B9EADCFFCBD532D27FF989FFD8FAB7" bold="true" box="[113,125,1657,1674]" pageId="10" pageNumber="427">E</emphasis>
, leg 2;
<emphasis id="09B9EADCFFCBD532D2B4F989FF60FAB7" bold="true" box="[186,197,1657,1674]" pageId="10" pageNumber="427">F</emphasis>
, leg 3;
<emphasis id="09B9EADCFFCBD532D30FF989FEAAFAB7" bold="true" box="[257,271,1657,1674]" pageId="10" pageNumber="427">G</emphasis>
, anterior gonopods, caudal view;
<emphasis id="09B9EADCFFCBD532D026F989FDEFFAB7" bold="true" box="[552,586,1657,1674]" pageId="10" pageNumber="427">H -K</emphasis>
, posterior gonopods, frontal, caudal, lateral, and mesal views, respectively. Scale bar: A, B, E, F, 0.3 mm; C, D, G-K, 0.2 mm.
</paragraph>
</caption>
<caption id="6FB26646FFCAD533D27FF993FF53FA9C" ID-DOI="http://doi.org/10.5281/zenodo.5393642" ID-Zenodo-Dep="5393642" httpUri="https://zenodo.org/record/5393642/files/figure.png" pageId="11" pageNumber="428" startId="11.[113,124,1635,1652]" targetBox="[117,1205,215,1535]" targetPageId="11">
<paragraph id="3B7236CEFFCAD533D27FF993FF53FA9C" blockId="11.[113,1211,1635,1697]" pageId="11" pageNumber="428">
FIG. 6. —
<taxonomicName id="FCCD4D4DFFCAD533D2C8F993FE02FA49" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[198,423,1635,1652]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="428" phylum="Arthropoda" rank="species" species="mediobliteratus" status="sp. nov.">
<emphasis id="09B9EADCFFCAD533D2C8F993FE02FA49" box="[198,423,1635,1652]" italics="true" pageId="11" pageNumber="428">Glyphiulus mediobliteratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFCAD533D3A3F993FE47FA49" box="[429,482,1635,1652]" pageId="11" pageNumber="428" rank="species">n. sp.</taxonomicNameLabel>
, paratype ♂:
<emphasis id="09B9EADCFFCAD533D058F993FDC6FA49" bold="true" box="[598,611,1635,1652]" pageId="11" pageNumber="428">A</emphasis>
, anterior part of body, lateral view;
<emphasis id="09B9EADCFFCAD533D198F993FC06FA49" bold="true" box="[918,931,1635,1652]" pageId="11" pageNumber="428">B</emphasis>
, same, dorsal view;
<emphasis id="09B9EADCFFCAD533D65BF993FBC6FA49" bold="true" box="[1109,1123,1635,1652]" pageId="11" pageNumber="428">C</emphasis>
, midbody segments, lateral view;
<emphasis id="09B9EADCFFCAD533D339F989FEE0FAB7" bold="true" box="[311,325,1657,1674]" pageId="11" pageNumber="428">D</emphasis>
, same, dorsal view;
<emphasis id="09B9EADCFFCAD533D3FFF989FE58FAB7" bold="true" box="[497,509,1657,1674]" pageId="11" pageNumber="428">E</emphasis>
, caudal body end, lateral view;
<emphasis id="09B9EADCFFCAD533D107F989FCB1FAB7" bold="true" box="[777,788,1657,1674]" pageId="11" pageNumber="428">F</emphasis>
, telson, ventral view. Scale bars: A, C, D, 0.5 mm; B, E, F, 0.2 mm.
</paragraph>
</caption>
<caption id="6FB26646FFCDD534D27FFBC1FEFBF852" ID-DOI="http://doi.org/10.5281/zenodo.5393644" ID-Zenodo-Dep="5393644" httpUri="https://zenodo.org/record/5393644/files/figure.png" pageId="12" pageNumber="429" startId="12.[113,124,1073,1090]" targetBox="[117,1206,215,1027]" targetPageId="12">
<paragraph id="3B7236CEFFCDD534D27FFBC1FEFBF852" blockId="12.[113,1211,1073,1135]" pageId="12" pageNumber="429">
FIG. 7. —
<taxonomicName id="FCCD4D4DFFCDD534D2B3FBC1FE3DF87F" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[189,408,1073,1090]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="429" phylum="Arthropoda" rank="species" species="mediobliteratus" status="sp. nov.">
<emphasis id="09B9EADCFFCDD534D2B3FBC1FE3DF87F" box="[189,408,1073,1090]" italics="true" pageId="12" pageNumber="429">Glyphiulus mediobliteratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFCDD534D395FBC1FE69F87F" box="[411,460,1073,1090]" pageId="12" pageNumber="429" rank="species">n. sp.</taxonomicNameLabel>
,paratype ♂:
<emphasis id="09B9EADCFFCDD534D038FBC1FDE6F87F" bold="true" box="[566,579,1073,1090]" pageId="12" pageNumber="429">A</emphasis>
, midbody segment section;
<emphasis id="09B9EADCFFCDD534D124FBC1FC92F87F" bold="true" box="[810,823,1073,1090]" pageId="12" pageNumber="429">B</emphasis>
, both gonopods
<emphasis id="09B9EADCFFCDD534D1CCFBC1FC57F87F" box="[962,1010,1073,1090]" italics="true" pageId="12" pageNumber="429">in situ</emphasis>
, frontal view;
<emphasis id="09B9EADCFFCDD534D66CFBC1FBD5F87F" bold="true" box="[1122,1136,1073,1090]" pageId="12" pageNumber="429">C</emphasis>
, anterior gonopod, frontal view;
<emphasis id="09B9EADCFFCDD534D339FBB7FEE0F865" bold="true" box="[311,325,1095,1112]" pageId="12" pageNumber="429">D</emphasis>
, anterior gonopod, caudal view;
<emphasis id="09B9EADCFFCDD534D051FBB7FDCEF865" bold="true" box="[607,619,1095,1112]" pageId="12" pageNumber="429">E</emphasis>
, posterior gonopod, subfrontal view;
<emphasis id="09B9EADCFFCDD534D1A3FBB7FC1DF865" bold="true" box="[941,952,1095,1112]" pageId="12" pageNumber="429">F</emphasis>
, claw. Scale bars: A, 0.5 mm; B-D, 0.1 mm; E, F, 0.05 mm.
</paragraph>
</caption>
<subSubSection id="73D76545FFCDD534D27FFAE3FEF0F979" pageId="12" pageNumber="429" type="etymology">
<paragraph id="3B7236CEFFCDD534D27FFAE3FEF0F979" blockId="12.[113,646,1298,1349]" pageId="12" pageNumber="429">ETYMOLOGY. — To emphasize the strongly obliterated collum and axial crests.</paragraph>
</subSubSection>
<subSubSection id="73D76545FFCDD534D27FFAADFE9DF9C5" pageId="12" pageNumber="429" type="diagnosis">
<paragraph id="3B7236CEFFCDD534D27FFAADFE9DF9C5" blockId="12.[113,647,1372,1529]" pageId="12" pageNumber="429">DIAGNOSIS. — Differs from congeners by the unpigmented ocelli and, in part, body, coupled with the nearly smooth collum, the rudimentary axial crests, the lack of a notch on each anterior gonopod half, the highly conspicuous, slightly plumose flagellum of the posterior gonopod, etc. (see also key below).</paragraph>
</subSubSection>
<subSubSection id="73D76545FFCDD535D27FF9DBFC08FA38" lastPageId="13" lastPageNumber="430" pageId="12" pageNumber="429" type="description">
<paragraph id="3B7236CEFFCDD534D27FF9DBFEA7FA7E" blockId="12.[113,646,1579,1700]" box="[113,258,1579,1605]" pageId="12" pageNumber="429">DESCRIPTION</paragraph>
<paragraph id="3B7236CEFFCDD534D27FF9BAFDF6FA99" blockId="12.[113,646,1579,1700]" pageId="12" pageNumber="429">
<materialsCitation id="8BA53C93FFCDD534D27FF9BAFDF6FA99" collectionCode="T" pageId="12" pageNumber="429" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="E476886CFFCDD534D27FF9BAFF72FA59" box="[113,215,1610,1636]" pageId="12" pageNumber="429" type="holotype">Holotype</typeStatus>
with 61p+1a+
<collectionCode id="5DDCAE0BFFCDD534D37AF9BAFE26FA59" box="[372,387,1610,1636]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="429">T</collectionCode>
, about
<quantity id="FC359B2BFFCDD534D3DAF9BAFD87FA59" box="[468,546,1610,1636]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="12" pageNumber="429" unit="mm" value="43.0">43 mm</quantity>
long and 2.0 mm wide; adult
<typeStatus id="E476886CFFCDD534D334F99AFE3CFAB9" box="[314,409,1642,1668]" pageId="12" pageNumber="429" type="paratype">paratypes</typeStatus>
(
<specimenCount id="2DCBFD47FFCDD534D3A8F99FFE19FABE" box="[422,444,1647,1667]" pageId="12" pageNumber="429" type="male"></specimenCount>
,
<specimenCount id="2DCBFD47FFCDD534D3C8F99FFE7CFABE" box="[454,473,1647,1667]" pageId="12" pageNumber="429" type="female"></specimenCount>
) with 51-62p+2- 1a+
<collectionCode id="5DDCAE0BFFCDD534D297F97AFF0DFA99" box="[153,168,1674,1700]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="429">T</collectionCode>
, length
<quantity id="FC359B2BFFCDD534D30FF979FEDDFA99" box="[257,376,1673,1700]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" metricValueMax="4.2" metricValueMin="3.2" pageId="12" pageNumber="429" unit="mm" value="37.0" valueMax="42.0" valueMin="32.0">32-42 mm</quantity>
, width 1.8-2.0 mm.
</materialsCitation>
</paragraph>
<paragraph id="3B7236CEFFCDD534D0B5FB5EFC4AF93A" blockId="12.[677,1210,1198,1700]" pageId="12" pageNumber="429">Coloration entirely pallid to pale brown, only tips of tubercles of tergal crests often brownish to dark brown. Ocelli not visible.</paragraph>
<paragraph id="3B7236CEFFCDD534D0B5FAFDFB1CFA99" blockId="12.[677,1210,1198,1700]" pageId="12" pageNumber="429">
Antennae (
<figureCitation id="A3F62A4BFFCDD534D121FAFDFC26F91A" box="[815,899,1293,1319]" captionStart="FIG" captionStartId="11.[113,124,1635,1652]" captionTargetBox="[117,1205,215,1535]" captionTargetId="figure-109@11.[654,745,613,1104]" captionTargetPageId="11" captionText="FIG. 6. — Glyphiulus mediobliteratus n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, telson, ventral view. Scale bars: A, C, D, 0.5 mm; B, E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393642" httpUri="https://zenodo.org/record/5393642/files/figure.png" pageId="12" pageNumber="429">Figs 6A</figureCitation>
;
<figureCitation id="A3F62A4BFFCDD534D182FAFDFC0BF91A" box="[908,942,1293,1319]" captionStart="FIG" captionStartId="13.[113,124,1198,1215]" captionTargetBox="[94,1236,194,1174]" captionTargetId="figure-19@13.[94,1236,194,1174]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, antenna;B, gnathochilarium; C, leg 1, frontal view;D, leg 2; E, leg 3; F, leg 9; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393646" httpUri="https://zenodo.org/record/5393646/files/figure.png" pageId="12" pageNumber="429">8A</figureCitation>
), gnathochilarium (mentum divided, n = 5) (
<figureCitation id="A3F62A4BFFCDD534D1AEFADDFC5FF97A" box="[928,1018,1325,1351]" captionStart="FIG" captionStartId="13.[113,124,1198,1215]" captionTargetBox="[94,1236,194,1174]" captionTargetId="figure-19@13.[94,1236,194,1174]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, antenna;B, gnathochilarium; C, leg 1, frontal view;D, leg 2; E, leg 3; F, leg 9; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393646" httpUri="https://zenodo.org/record/5393646/files/figure.png" pageId="12" pageNumber="429">Fig. 8B</figureCitation>
), segment shape (
<figureCitation id="A3F62A4BFFCDD534D0A2FABCFD58F95A" box="[684,765,1356,1383]" captionStart="FIG" captionStartId="12.[113,124,1073,1090]" captionTargetBox="[117,1206,215,1027]" captionTargetId="figure-51@12.[389,662,576,758]" captionTargetPageId="12" captionText="FIG. 7. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, midbody segment section;B, both gonopods in situ, frontal view;C, anterior gonopod, frontal view; D, anterior gonopod, caudal view; E, posterior gonopod, subfrontal view; F, claw. Scale bars: A, 0.5 mm; B-D, 0.1 mm; E, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393644" httpUri="https://zenodo.org/record/5393644/files/figure.png" pageId="12" pageNumber="429">Fig. 7A</figureCitation>
), epiproct (
<figureCitation id="A3F62A4BFFCDD534D17EFABCFC18F95A" box="[880,957,1356,1383]" captionStart="FIG" captionStartId="11.[113,124,1635,1652]" captionTargetBox="[117,1205,215,1535]" captionTargetId="figure-109@11.[654,745,613,1104]" captionTargetPageId="11" captionText="FIG. 6. — Glyphiulus mediobliteratus n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, telson, ventral view. Scale bars: A, C, D, 0.5 mm; B, E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393642" httpUri="https://zenodo.org/record/5393642/files/figure.png" pageId="12" pageNumber="429">Fig. 6E</figureCitation>
), paraprocts, hypoproct, male legs 2 and 3 (
<figureCitation id="A3F62A4BFFCDD534D17BFA9CFC4BF9BA" box="[885,1006,1388,1415]" captionStart="FIG" captionStartId="13.[113,124,1198,1215]" captionTargetBox="[94,1236,194,1174]" captionTargetId="figure-19@13.[94,1236,194,1174]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, antenna;B, gnathochilarium; C, leg 1, frontal view;D, leg 2; E, leg 3; F, leg 9; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393646" httpUri="https://zenodo.org/record/5393646/files/figure.png" pageId="12" pageNumber="429">Fig. 8D, E</figureCitation>
), texture (
<figureCitation id="A3F62A4BFFCDD534D651FA9CFB38F9BB" box="[1119,1181,1388,1414]" captionStart="FIG" captionStartId="11.[113,124,1635,1652]" captionTargetBox="[117,1205,215,1535]" captionTargetId="figure-109@11.[654,745,613,1104]" captionTargetPageId="11" captionText="FIG. 6. — Glyphiulus mediobliteratus n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, telson, ventral view. Scale bars: A, C, D, 0.5 mm; B, E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393642" httpUri="https://zenodo.org/record/5393642/files/figure.png" pageId="12" pageNumber="429">Fig. 6</figureCitation>
A- E) generally as in
<taxonomicName id="FCCD4D4DFFCDD534D164FA7CFC5DF99B" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[874,1016,1420,1446]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="429" phylum="Arthropoda" rank="species" species="obliteratus" status="sp. nov.">
<emphasis id="09B9EADCFFCDD534D164FA7CFC5DF99B" box="[874,1016,1420,1446]" italics="true" pageId="12" pageNumber="429">G. obliteratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFCDD534D60FFA7CFB9AF99B" box="[1025,1087,1420,1446]" pageId="12" pageNumber="429" rank="species">n. sp.</taxonomicNameLabel>
, but sparse striation present in front of stricture both laterally and dorsally; conspicuous pleurosternal crests on segments 2-4 wanting (
<figureCitation id="A3F62A4BFFCDD534D1AEFA1BFC50FA3B" box="[928,1013,1515,1542]" captionStart="FIG" captionStartId="11.[113,124,1635,1652]" captionTargetBox="[117,1205,215,1535]" captionTargetId="figure-109@11.[654,745,613,1104]" captionTargetPageId="11" captionText="FIG. 6. — Glyphiulus mediobliteratus n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, telson, ventral view. Scale bars: A, C, D, 0.5 mm; B, E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393642" httpUri="https://zenodo.org/record/5393642/files/figure.png" pageId="12" pageNumber="429">Fig. 6A</figureCitation>
); collum (
<figureCitation id="A3F62A4BFFCDD534D66CFA1BFD12FA18" captionStart="FIG" captionStartId="11.[113,124,1635,1652]" captionTargetBox="[117,1205,215,1535]" captionTargetId="figure-109@11.[654,745,613,1104]" captionTargetPageId="11" captionText="FIG. 6. — Glyphiulus mediobliteratus n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, telson, ventral view. Scale bars: A, C, D, 0.5 mm; B, E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393642" httpUri="https://zenodo.org/record/5393642/files/figure.png" pageId="12" pageNumber="429">Fig. 6A, B</figureCitation>
) with 1a-4a+ma+4a-1a very weak undulations in caudal 1/3, undulation ma being shortest; tergal crests higher, even adding another lateral crest below ozoporiferous one; ozoporiferous cones especially high, about as high as wide (
<figureCitation id="A3F62A4BFFCDD534D1EAF979FB9EFA9E" box="[996,1083,1673,1699]" captionStart="FIG" captionStartId="12.[113,124,1073,1090]" captionTargetBox="[117,1206,215,1027]" captionTargetId="figure-51@12.[389,662,576,758]" captionTargetPageId="12" captionText="FIG. 7. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, midbody segment section;B, both gonopods in situ, frontal view;C, anterior gonopod, frontal view; D, anterior gonopod, caudal view; E, posterior gonopod, subfrontal view; F, claw. Scale bars: A, 0.5 mm; B-D, 0.1 mm; E, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393644" httpUri="https://zenodo.org/record/5393644/files/figure.png" pageId="12" pageNumber="429">Fig. 7A</figureCitation>
), each crest
</paragraph>
<paragraph id="3B7236CEFFCCD535D286FFBEFEB8FC62" blockId="13.[136,285,78,95]" box="[136,285,78,95]" pageId="13" pageNumber="430">
Golovatch S. I.
<emphasis id="09B9EADCFFCCD535D2F7FFBEFEB8FC62" box="[249,285,78,95]" italics="true" pageId="13" pageNumber="430">et al.</emphasis>
</paragraph>
<caption id="6FB26646FFCCD535D27FFB5EFB8FF8EB" ID-DOI="http://doi.org/10.5281/zenodo.5393646" ID-Zenodo-Dep="5393646" httpUri="https://zenodo.org/record/5393646/files/figure.png" pageId="13" pageNumber="430" startId="13.[113,124,1198,1215]" targetBox="[94,1236,194,1174]" targetPageId="13">
<paragraph id="3B7236CEFFCCD535D27FFB5EFB8FF8EB" blockId="13.[113,1210,1198,1238]" pageId="13" pageNumber="430">
FIG. 8. —
<taxonomicName id="FCCD4D4DFFCCD535D2B3FB5EFE3DF882" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[189,408,1198,1215]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="430" phylum="Arthropoda" rank="species" species="mediobliteratus" status="sp. nov.">
<emphasis id="09B9EADCFFCCD535D2B3FB5EFE3DF882" box="[189,408,1198,1215]" italics="true" pageId="13" pageNumber="430">Glyphiulus mediobliteratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFCCD535D395FB5EFE69F882" box="[411,460,1198,1215]" pageId="13" pageNumber="430" rank="species">n. sp.</taxonomicNameLabel>
,paratype ♂:
<emphasis id="09B9EADCFFCCD535D038FB5EFDE6F882" bold="true" box="[566,579,1198,1215]" pageId="13" pageNumber="430">A</emphasis>
, antenna;
<emphasis id="09B9EADCFFCCD535D096FB5EFD00F882" bold="true" box="[664,677,1198,1215]" pageId="13" pageNumber="430">B</emphasis>
, gnathochilarium;
<emphasis id="09B9EADCFFCCD535D133FB5EFCEEF882" bold="true" box="[829,843,1198,1215]" pageId="13" pageNumber="430">C</emphasis>
, leg 1, frontal view;
<emphasis id="09B9EADCFFCCD535D1E4FB5EFC5DF882" bold="true" box="[1002,1016,1198,1215]" pageId="13" pageNumber="430">D</emphasis>
, leg 2;
<emphasis id="09B9EADCFFCCD535D63FFB5EFB98F882" bold="true" box="[1073,1085,1198,1215]" pageId="13" pageNumber="430">E</emphasis>
, leg 3;
<emphasis id="09B9EADCFFCCD535D679FB5EFB27F882" bold="true" box="[1143,1154,1198,1215]" pageId="13" pageNumber="430">F</emphasis>
, leg 9;
<emphasis id="09B9EADCFFCCD535D27FFB35FFDAF8EB" bold="true" box="[113,127,1221,1238]" pageId="13" pageNumber="430">G</emphasis>
, anterior gonopods, caudal view;
<emphasis id="09B9EADCFFCCD535D395FB35FE0CF8EB" bold="true" box="[411,425,1221,1238]" pageId="13" pageNumber="430">H</emphasis>
, right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm.
</paragraph>
</caption>
<paragraph id="3B7236CEFFCCD535D27FFAFDFEFCFA79" blockId="13.[113,647,1293,1700]" pageId="13" pageNumber="430">
distinctly subdivided into two usually subtriangular halves, except for nearly absent axial crests (m); carinotaxy pattern thus 2/2+I/i+1/1+m+1/1+I/ i+2/2 (
<figureCitation id="A3F62A4BFFCCD535D2CEFA9CFEA0F9BB" box="[192,261,1388,1414]" captionStart="FIG" captionStartId="11.[113,124,1635,1652]" captionTargetBox="[117,1205,215,1535]" captionTargetId="figure-109@11.[654,745,613,1104]" captionTargetPageId="11" captionText="FIG. 6. — Glyphiulus mediobliteratus n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, telson, ventral view. Scale bars: A, C, D, 0.5 mm; B, E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393642" httpUri="https://zenodo.org/record/5393642/files/figure.png" pageId="13" pageNumber="430">Figs 6</figureCitation>
A-E; 7A); a small accessory spine present at base of claw on legs 2 and 3 only (
<figureCitation id="A3F62A4BFFCCD535D052FA7CFF13F9FB" captionStart="FIG" captionStartId="13.[113,124,1198,1215]" captionTargetBox="[94,1236,194,1174]" captionTargetId="figure-19@13.[94,1236,194,1174]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, antenna;B, gnathochilarium; C, leg 1, frontal view;D, leg 2; E, leg 3; F, leg 9; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393646" httpUri="https://zenodo.org/record/5393646/files/figure.png" pageId="13" pageNumber="430">Fig. 8D, E</figureCitation>
), thereafter virtually missing (
<figureCitation id="A3F62A4BFFCCD535D3F0FA5BFDE8F9FB" box="[510,589,1451,1478]" captionStart="FIG" captionStartId="12.[113,124,1073,1090]" captionTargetBox="[117,1206,215,1027]" captionTargetId="figure-51@12.[389,662,576,758]" captionTargetPageId="12" captionText="FIG. 7. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, midbody segment section;B, both gonopods in situ, frontal view;C, anterior gonopod, frontal view; D, anterior gonopod, caudal view; E, posterior gonopod, subfrontal view; F, claw. Scale bars: A, 0.5 mm; B-D, 0.1 mm; E, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393644" httpUri="https://zenodo.org/record/5393644/files/figure.png" pageId="13" pageNumber="430">Fig. 7F</figureCitation>
,
<figureCitation id="A3F62A4BFFCCD535D057FA5BFDDDF9F8" box="[601,632,1451,1477]" captionStart="FIG" captionStartId="13.[113,124,1198,1215]" captionTargetBox="[94,1236,194,1174]" captionTargetId="figure-19@13.[94,1236,194,1174]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, antenna;B, gnathochilarium; C, leg 1, frontal view;D, leg 2; E, leg 3; F, leg 9; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393646" httpUri="https://zenodo.org/record/5393646/files/figure.png" pageId="13" pageNumber="430">8F</figureCitation>
); legs slightly shorter than body diameter (
<figureCitation id="A3F62A4BFFCCD535D028FA3BFDDFF9D8" box="[550,634,1483,1509]" captionStart="FIG" captionStartId="12.[113,124,1073,1090]" captionTargetBox="[117,1206,215,1027]" captionTargetId="figure-51@12.[389,662,576,758]" captionTargetPageId="12" captionText="FIG. 7. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, midbody segment section;B, both gonopods in situ, frontal view;C, anterior gonopod, frontal view; D, anterior gonopod, caudal view; E, posterior gonopod, subfrontal view; F, claw. Scale bars: A, 0.5 mm; B-D, 0.1 mm; E, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393644" httpUri="https://zenodo.org/record/5393644/files/figure.png" pageId="13" pageNumber="430">Fig. 7A</figureCitation>
); male legs 1 with strongly developed paramedian coxal outgrowths and 5-segmented, slightly reduced, telopodites (
<figureCitation id="A3F62A4BFFCCD535D2F9F9DAFEE9FA79" box="[247,332,1578,1604]" captionStart="FIG" captionStartId="13.[113,124,1198,1215]" captionTargetBox="[94,1236,194,1174]" captionTargetId="figure-19@13.[94,1236,194,1174]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, antenna;B, gnathochilarium; C, leg 1, frontal view;D, leg 2; E, leg 3; F, leg 9; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393646" httpUri="https://zenodo.org/record/5393646/files/figure.png" pageId="13" pageNumber="430">Fig. 8C</figureCitation>
).
</paragraph>
<paragraph id="3B7236CEFFCCD535D286F9BAFCA9F97A" blockId="13.[113,647,1293,1700]" lastBlockId="13.[677,1210,1293,1541]" pageId="13" pageNumber="430">
Head width = segment 2 = 6 &lt;collum = 7(8)&gt; segment 3 = 5 &lt;7 &lt;8 &lt;9 &lt;10 &lt;11 = midbody segments; body gradually but then abruptly tapering toward telson. Postcollar constriction modest (
<figureCitation id="A3F62A4BFFCCD535D0A2FADDFCA5F97A" box="[684,768,1324,1351]" captionStart="FIG" captionStartId="11.[113,124,1635,1652]" captionTargetBox="[117,1205,215,1535]" captionTargetId="figure-109@11.[654,745,613,1104]" captionTargetPageId="11" captionText="FIG. 6. — Glyphiulus mediobliteratus n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, telson, ventral view. Scale bars: A, C, D, 0.5 mm; B, E, F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5393642" httpUri="https://zenodo.org/record/5393642/files/figure.png" pageId="13" pageNumber="430">Fig. 6B</figureCitation>
).
</paragraph>
<paragraph id="3B7236CEFFCCD535D0B5FABDFB83F9BB" blockId="13.[677,1210,1293,1541]" pageId="13" pageNumber="430">
Penes broad, rounded, each with three or four strong setae distolaterally (
<figureCitation id="A3F62A4BFFCCD535D1CFFA9CFBBDF9BB" box="[961,1048,1388,1414]" captionStart="FIG" captionStartId="13.[113,124,1198,1215]" captionTargetBox="[94,1236,194,1174]" captionTargetId="figure-19@13.[94,1236,194,1174]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, antenna;B, gnathochilarium; C, leg 1, frontal view;D, leg 2; E, leg 3; F, leg 9; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393646" httpUri="https://zenodo.org/record/5393646/files/figure.png" pageId="13" pageNumber="430">Fig. 8D</figureCitation>
).
</paragraph>
<paragraph id="3B7236CEFFCCD535D0B5FA7CFC08FA38" blockId="13.[677,1210,1293,1541]" pageId="13" pageNumber="430">
Anterior gonopods (
<figureCitation id="A3F62A4BFFCCD535D1ACFA7CFC4DF99B" box="[930,1000,1420,1446]" captionStart="FIG" captionStartId="12.[113,124,1073,1090]" captionTargetBox="[117,1206,215,1027]" captionTargetId="figure-51@12.[389,662,576,758]" captionTargetPageId="12" captionText="FIG. 7. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, midbody segment section;B, both gonopods in situ, frontal view;C, anterior gonopod, frontal view; D, anterior gonopod, caudal view; E, posterior gonopod, subfrontal view; F, claw. Scale bars: A, 0.5 mm; B-D, 0.1 mm; E, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393644" httpUri="https://zenodo.org/record/5393644/files/figure.png" pageId="13" pageNumber="430">Figs 7</figureCitation>
B-D; 8G) without notches, processes (d) especially high. Posterior gonopods (
<figureCitation id="A3F62A4BFFCCD535D112FA3BFCD4F9DB" box="[796,881,1483,1510]" captionStart="FIG" captionStartId="12.[113,124,1073,1090]" captionTargetBox="[117,1206,215,1027]" captionTargetId="figure-51@12.[389,662,576,758]" captionTargetPageId="12" captionText="FIG. 7. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, midbody segment section;B, both gonopods in situ, frontal view;C, anterior gonopod, frontal view; D, anterior gonopod, caudal view; E, posterior gonopod, subfrontal view; F, claw. Scale bars: A, 0.5 mm; B-D, 0.1 mm; E, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393644" httpUri="https://zenodo.org/record/5393644/files/figure.png" pageId="13" pageNumber="430">Figs 7E</figureCitation>
;
<figureCitation id="A3F62A4BFFCCD535D172FA3BFC05F9D8" box="[892,928,1483,1509]" captionStart="FIG" captionStartId="13.[113,124,1198,1215]" captionTargetBox="[94,1236,194,1174]" captionTargetId="figure-19@13.[94,1236,194,1174]" captionTargetPageId="13" captionText="FIG. 8. — Glyphiulus mediobliteratus n. sp.,paratype ♂: A, antenna;B, gnathochilarium; C, leg 1, frontal view;D, leg 2; E, leg 3; F, leg 9; G, anterior gonopods, caudal view; H, right posterior gonopod, frontal view. Scale bar: A-F, 0.3 mm; G, H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393646" httpUri="https://zenodo.org/record/5393646/files/figure.png" pageId="13" pageNumber="430">8H</figureCitation>
) with characteristic, long, distally plumose flagella.
</paragraph>
</subSubSection>
<subSubSection id="73D76545FFCCD535D0ABF9DBFB03FA9E" pageId="13" pageNumber="430" type="discussion">
<paragraph id="3B7236CEFFCCD535D0ABF9DBFCACFA7E" blockId="13.[677,1210,1579,1700]" box="[677,777,1579,1605]" pageId="13" pageNumber="430">REMARKS</paragraph>
<paragraph id="3B7236CEFFCCD535D0ABF9BAFB03FA9E" blockId="13.[677,1210,1579,1700]" pageId="13" pageNumber="430">The presence of completely unpigmented ocelli suggests that this species may be troglobitic, but further observations are needed to confirm this.</paragraph>
</subSubSection>
</treatment>
</document>