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<document id="128DEB71E9417BA55EBA2EB63B6836F9" ID-CLB-Dataset="78128" ID-DOI="10.11646/zootaxa.5155.2.1" ID-GBIF-Dataset="be7f6620-4608-4fa7-9764-785fd401901a" ID-ISSN="1175-5326" ID-Zenodo-Dep="6673332" ID-ZooBank="F23CDF54-115F-4861-8EDB-DAFDF7810586" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1655793259292" checkinUser="plazi" docAuthor="Pinto-Da-Rocha, Ricardo &amp; Cabra-García, Jimmy" docDate="2022" docId="B3264F4BFF93705CADB9FA30FCF2FEE4" docLanguage="en" docName="zootaxa.5155.2.1.pdf" docOrigin="Zootaxa 5155 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tityus moralensis Pinto-Da-Rocha &amp; Cabra-García 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="178" masterDocId="4F1F3733FF847047AD2EFFD8FFC8FFA5" masterDocTitle="On the Tityus forcipula species group: redescription of Tityus forcipula (Scorpiones, Buthidae), description of a new Andean species, and notes on the taxonomy of the group" masterLastPageNumber="186" masterPageNumber="151" pageNumber="174" updateTime="1699349182649" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A425DB6E043BD6C7A2E2C339B87DDC29">On the Tityus forcipula species group: redescription of Tityus forcipula (Scorpiones, Buthidae), description of a new Andean species, and notes on the taxonomy of the group</mods:title>
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<mods:namePart id="7716618846EA67B8AD853EA7190C1962">Pinto-Da-Rocha, Ricardo</mods:namePart>
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<mods:affiliation id="FE61FBE0D05728A1AD5E89F88D4FC94C">Instituto de Biociências - Universidade de São Paulo, Departamento de Zoologia. Rua do Matão, travessa 14, 321, 005508 - 900, São Paulo, SP, Brazil &amp; ricrocha @ usp. br; https: // orcid. org / 0000 - 0002 - 3959 - 2205</mods:affiliation>
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<mods:namePart id="193BDAF7791FFBF727024A28F9EEB4DF">Cabra-García, Jimmy</mods:namePart>
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<mods:affiliation id="37482CEDEBDC01F63B938B56C1ED1A16">Departamento de Biología, Universidad del Valle, Ciudad Universitaria Meléndez, Calle 13 # 100 - 00, 760032, Cali, Colombia. Corresponding author: jimmy. cabra @ correounivalle. edu. co; https: // orcid. org / 0000 - 0001 - 9263 - 4390</mods:affiliation>
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<figureCitation id="A3B4E2D8FF937050ADB9F9E8FEC9F9EF" box="[151,257,1584,1610]" captionStart="FIGURE 1" captionStartId="4.[151,250,1859,1883]" captionTargetBox="[286,1301,662,1817]" captionTargetId="figure-141@4.[251,1336,630,1838]" captionTargetPageId="4" captionText="FIGURE 1. Phylogenetic relationships of Tityus species inferred from a combined dataset of 28S and COI sequences. The topology is recovered from the IQ-TREE analysis. Squares at nodes indicate the Shimodaira–Hasegawa approximate likelihood-ratio test (left), the ultrafast bootstrap (middle) and the parsimony bootstrap (right). All sequences obtained in this study are in bold." figureDoi="http://doi.org/10.5281/zenodo.6673334" httpUri="https://zenodo.org/record/6673334/files/figure.png" pageId="23" pageNumber="174">Figures 1</figureCitation>
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<figureCitation id="A3B4E2D8FF937050AC3EF9E8FE9EF9EF" box="[272,342,1584,1610]" captionStart="FIGURE 3" captionStartId="8.[151,250,1927,1951]" captionTargetBox="[162,1419,202,1891]" captionTargetId="figure-31@8.[151,1436,181,1903]" captionTargetPageId="8" captionText="FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm." figureDoi="http://doi.org/10.5281/zenodo.6673342" httpUri="https://zenodo.org/record/6673342/files/figure.png" pageId="23" pageNumber="174">3A–D</figureCitation>
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<figureCitation id="A3B4E2D8FF937050AC4DF9E8FE60F9EF" box="[355,424,1584,1610]" captionStart="FIGURE 4" captionStartId="10.[151,250,1375,1399]" captionTargetBox="[371,1216,309,1352]" captionTargetId="figure-86@10.[371,1216,309,1352]" captionTargetPageId="10" captionText="FIGURE 4. Tityus (Atreus) Gervais, 1843, carapace sculpturing, dorsal aspect. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673348" httpUri="https://zenodo.org/record/6673348/files/figure.png" pageId="23" pageNumber="174">4C–D</figureCitation>
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<figureCitation id="A3B4E2D8FF937050AC9AF9E8FE30F9EE" box="[436,504,1584,1611]" captionStart="FIGURE 5" captionStartId="13.[151,250,1447,1471]" captionTargetBox="[151,1437,309,1424]" captionTargetId="figure-91@13.[151,1437,309,1424]" captionTargetPageId="13" captionText="FIGURE 5. Tityus (Atreus) Gervais, 1843, femur sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673350" httpUri="https://zenodo.org/record/6673350/files/figure.png" pageId="23" pageNumber="174">5E–H</figureCitation>
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<figureCitation id="A3B4E2D8FF937050AF2AF9E8FD80F9EF" box="[516,584,1584,1610]" captionStart="FIGURE 6" captionStartId="14.[151,250,1863,1887]" captionTargetBox="[151,1436,561,1840]" captionTargetId="figure-228@14.[151,1436,561,1840]" captionTargetPageId="14" captionText="FIGURE 6. Tityus (Atreus) Gervais, 1843, patella sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673354" httpUri="https://zenodo.org/record/6673354/files/figure.png" pageId="23" pageNumber="174">6E–H</figureCitation>
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<figureCitation id="A3B4E2D8FF937050AF7AF9E8FD5CF9EF" box="[596,660,1584,1610]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[261,1326,181,1920]" captionTargetId="figure-19@15.[261,1326,181,1920]" captionTargetPageId="15" captionText="FIGURE 7. Tityus (Atreus) Gervais, 1843,female chela sculpturing.A–C. Tityus(Atreus) forcipula (Gervais,1843) (MUSENUV- Ar 436). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (ICN-1402). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673356" httpUri="https://zenodo.org/record/6673356/files/figure.png" pageId="23" pageNumber="174">7D–F</figureCitation>
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<figureCitation id="A3B4E2D8FF937050AF8EF9E8FD28F9EF" box="[672,736,1584,1610]" captionStart="FIGURE 8" captionStartId="16.[151,250,1944,1968]" captionTargetBox="[364,1224,181,1920]" captionTargetId="figure-31@16.[364,1224,181,1920]" captionTargetPageId="16" captionText="FIGURE 8. Tityus (Atreus) Gervais, 1843, male chela sculpturing. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673358" httpUri="https://zenodo.org/record/6673358/files/figure.png" pageId="23" pageNumber="174">8D–F</figureCitation>
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<figureCitation id="A3B4E2D8FF937050AFC3F9E8FCFBF9EF" box="[749,819,1584,1610]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="23" pageNumber="174">9C, D</figureCitation>
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<figureCitation id="A3B4E2D8FF937050AE11F9E8FC5BF9EF" box="[831,915,1584,1610]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[221,1367,181,1920]" captionTargetId="figure-19@19.[221,1367,181,1920]" captionTargetPageId="19" captionText="FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673364" httpUri="https://zenodo.org/record/6673364/files/figure.png" pageId="23" pageNumber="174">10C, D</figureCitation>
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<figureCitation id="A3B4E2D8FF937050AEB1F9E8FC25F9EF" box="[927,1005,1584,1610]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="23" pageNumber="174">11D–F</figureCitation>
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<figureCitation id="A3B4E2D8FF937050AED6F9E8FB85F9EF" box="[1016,1101,1584,1610]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="23" pageNumber="174">12C, D</figureCitation>
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<figureCitation id="A3B4E2D8FF937050A977F9E8FB65F9EF" box="[1113,1197,1584,1610]" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="23" pageNumber="174">13C, D</figureCitation>
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<figureCitation id="A3B4E2D8FF937050A997F9E8FAC5F9EF" box="[1209,1293,1584,1610]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="23" pageNumber="174">14C, D</figureCitation>
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<figureCitation id="A3B4E2D8FF937050A837F9E8FAA5F9EF" box="[1305,1389,1584,1611]" captionStart="FIGURE 15" captionStartId="24.[151,250,1090,1114]" captionTargetBox="[151,1436,345,1067]" captionTargetId="figure-116@24.[151,1436,345,1067]" captionTargetPageId="24" captionText="FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673376" httpUri="https://zenodo.org/record/6673376/files/figure.png" pageId="23" pageNumber="174">15C, D</figureCitation>
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<figureCitation id="A3B4E2D8FF937050A857F9E8FA5FF9EF" box="[1401,1431,1584,1610]" captionStart="FIGURE 16" captionStartId="27.[151,250,1428,1452]" captionTargetBox="[184,1412,353,1404]" captionTargetId="figure-95@27.[176,1412,345,1404]" captionTargetPageId="27" captionText="FIGURE 16. Distribution map of the Tityus forcipula species group. A. South America. B. Records area. C. Species records." figureDoi="http://doi.org/10.5281/zenodo.6673378" httpUri="https://zenodo.org/record/6673378/files/figure.png" pageId="23" pageNumber="174">16</figureCitation>
, 
<figureCitation id="A3B4E2D8FF937050ADB9F98CFF22F9CB" box="[151,234,1620,1646]" captionStart="FIGURE 17" captionStartId="28.[151,250,1797,1821]" captionTargetBox="[192,1395,181,1774]" captionTargetId="figure-31@28.[192,1395,181,1774]" captionTargetPageId="28" captionText="FIGURE 17. Tityus (Atreus) moralensis sp. nov. A. Male holotype, live habitus. B. Microhabitat in El Cerrito, El Moral pathway in the Cordillera Central, Colombia." figureDoi="http://doi.org/10.5281/zenodo.6673380" httpUri="https://zenodo.org/record/6673380/files/figure.png" pageId="23" pageNumber="174">17A, B</figureCitation>
; 
<tableCitation id="760DCBE6FF937050ADD8F98CFE9EF9CB" box="[246,342,1620,1646]" captionStart="TABLE 2" captionStartId="3.[151,239,151,175]" captionTargetPageId="3" captionText="TABLE 2. Taxon sampling and GenBank accession numbers. Entries in bold were generated in this study. Abbreviations: Not applicable (NA)." httpUri="http://table.plazi.org/id/6FF0AED5FF877044ADB9FF4FFEB2FF71" pageId="23" pageNumber="174" tableUuid="6FF0AED5FF877044ADB9FF4FFEB2FF71">Tables 2</tableCitation>
, 
<tableCitation id="760DCBE6FF937050AC4CF98CFEBAF9CB" box="[354,370,1620,1646]" captionStart="TABLE 4" captionStartId="5.[151,239,574,600]" captionTargetPageId="5" captionText="TABLE 4. Percent uncorrected pairwise distances between COI sequences of Tityus species. Species codes: 1: Tityus argentinus, 2: T. bahiensis, 3: T. brazilae, 4: T. cerroazul, 5: T. clathrathus, 6-7: T. curupi, 8: T. discrepans, 9: T. engelkei, 10: T. forcipula, 11: T. metuendus, 12: T. moralensis sp. nov., 13: T. nematochirus, 14: T. obscurus, 15: T. paraguayensis, 16: T. sastrei, 17: T. serrulatus, 18: T. soratensis, 19-20: T. uruguayensis. Abbreviations: GBA: GenBank accession number." httpUri="http://table.plazi.org/id/6FF0AED5FF817042ADB9FDE6F9CDFD05" pageId="23" pageNumber="174" tableUuid="6FF0AED5FF817042ADB9FDE6F9CDFD05">4</tableCitation>
, 
<tableCitation id="760DCBE6FF937050AC50F98CFE46F9CA" box="[382,398,1620,1647]" captionStart="TABLE 5" captionStartId="11.[151,239,440,464]" captionTargetBox="[167,1416,516,1987]" captionTargetPageId="11" captionText="TABLE 5. Measurements (mm) of Tityus forcipula and Tityus moralensis sp. nov. Observation: all specimens, but 1402" httpUri="http://table.plazi.org/id/6FF0AED5FF8F704CADB9FE60FAC8FE51" pageId="23" pageNumber="174" tableUuid="6FF0AED5FF8F704CADB9FE60FAC8FE51">5</tableCitation>
, 
<tableCitation id="760DCBE6FF937050ACB4F98CFE63F9CB" box="[410,427,1620,1646]" captionStart="TABLE 6" captionStartId="26.[151,238,1052,1076]" captionTargetPageId="26" captionText="TABLE 6. Variation in the number of setae of the ventrosubmedian setal rows on telotarsi I–IV of Tityus forcipula and Tityus moralensis sp. nov. Abbreviations: L, left leg; Pl, prolateral row; Rl, retrolateral row; R, right leg. Observation: all specimens, but 1402 and 1403 deposited at ICN, are deposited at MUSENUV. Bold numbers correspond to the museum codes." httpUri="http://table.plazi.org/id/6FF0AED5FF9E705DADB9FBC4FA45FBD9" pageId="23" pageNumber="174" tableUuid="6FF0AED5FF9E705DADB9FBC4FA45FBD9">6</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7395ADD6FF937050ADB9F944FE88F8CF" pageId="23" pageNumber="174" type="materials_examined">
<paragraph id="3B30FE5DFF937050ADB9F944FE88F8CF" blockId="23.[151,1437,1692,2042]" pageId="23" pageNumber="174">
<emphasis id="09FB224FFF937050ADB9F944FE8DF913" bold="true" box="[151,325,1692,1718]" pageId="23" pageNumber="174">Type material.</emphasis>
<materialsCitation id="8BE7F400FF937050AC62F944FCCBF95B" ID-GBIF-Occurrence="3816447305" collectingDate="2021-06-12" collectorName="El Cerrito &amp; El Moral Sidewalk &amp; La Floresta Farm &amp; A. Betancourt &amp; S. Forero &amp; J. Cabra" country="Colombia" county="El Cerrito" elevation="2154" latitude="3.6783056" location="El Moral Sidewalk" longLatPrecision="1" longitude="-76.07486" municipality="Tenerife Pathway" pageId="23" pageNumber="174" specimenCount="1" stateProvince="Valle del Cauca" typeStatus="holotype">
<typeStatus id="E43440FFFF937050AC62F944FE71F913" box="[332,441,1692,1718]" pageId="23" pageNumber="174" type="holotype">Holotype</typeStatus>
: 
<collectingCountry id="4398BECDFF937050ACEDF944FD95F913" box="[451,605,1692,1718]" name="Colombia" pageId="23" pageNumber="174">COLOMBIA</collectingCountry>
: 
<emphasis id="09FB224FFF937050AF49F944FC57F913" box="[615,927,1692,1718]" italics="true" pageId="23" pageNumber="174">
<collectingRegion id="F94B30BFFF937050AF49F944FC57F913" box="[615,927,1692,1718]" country="Colombia" name="Valle del Cauca" pageId="23" pageNumber="174">Valle del Cauca department</collectingRegion>
</emphasis>
: adult male, 
<collectorName id="967A9B8BFF937050A91EF944FB57F913" box="[1072,1183,1692,1718]" pageId="23" pageNumber="174">
<collectingCounty id="D25186D1FF937050A91EF944FB57F913" box="[1072,1183,1692,1718]" pageId="23" pageNumber="174">El Cerrito</collectingCounty>
</collectorName>
municipality, 
<collectingMunicipality id="DB546427FF937050A811F944FF31F97F" pageId="23" pageNumber="174">Tenerife Pathway</collectingMunicipality>
, 
<collectorName id="967A9B8BFF937050AC28F918FE29F97F" box="[262,481,1728,1754]" pageId="23" pageNumber="174">
<location id="3E50A886FF937050AC28F918FE29F97F" LSID="urn:lsid:plazi:treatment:B3264F4BFF93705CADB9FA30FCF2FEE4:3E50A886FF937050AC28F918FE29F97F" box="[262,481,1728,1754]" country="Colombia" county="El Cerrito" latitude="3.6783056" longLatPrecision="1" longitude="-76.07486" municipality="Tenerife Pathway" name="El Moral Sidewalk" pageId="23" pageNumber="174" stateProvince="Valle del Cauca">El Moral Sidewalk</location>
</collectorName>
, 
<collectorName id="967A9B8BFF937050ACC0F918FD7EF97F" box="[494,694,1728,1754]" pageId="23" pageNumber="174">
<location id="3E50A886FF937050ACC0F918FD7EF97F" LSID="urn:lsid:plazi:treatment:B3264F4BFF93705CADB9FA30FCF2FEE4:3E50A886FF937050ACC0F918FD7EF97F" box="[494,694,1728,1754]" country="Colombia" county="El Cerrito" latitude="3.6783056" longLatPrecision="1" longitude="-76.07486" municipality="Tenerife Pathway" name="La Floresta Farm" pageId="23" pageNumber="174" stateProvince="Valle del Cauca">La Floresta Farm</location>
</collectorName>
, 
<geoCoordinate id="5EBB989AFF937050AFEDF918FC94F97F" box="[707,860,1728,1754]" degrees="3" direction="north" minutes="40" orientation="latitude" pageId="23" pageNumber="174" precision="1" seconds="41.9" value="3.6783056">3°40’41.9” N</geoCoordinate>
<geoCoordinate id="5EBB989AFF937050AE48F918FBDBF97F" box="[870,1043,1728,1755]" degrees="76" direction="west" minutes="04" orientation="longitude" pageId="23" pageNumber="174" precision="1" seconds="29.5" value="-76.07486">76°04’29.5” W</geoCoordinate>
, 
<quantity id="FC7753B8FF937050A90EF918FB52F97F" box="[1056,1178,1728,1755]" metricMagnitude="3" metricUnit="m" metricValue="2.154" pageId="23" pageNumber="174" unit="m" value="2154.0">
<elevation id="B0A2196EFF937050A90EF918FB52F97F" box="[1056,1178,1728,1755]" metricMagnitude="3" metricUnit="m" metricValue="2.154" pageId="23" pageNumber="174" unit="m" value="2154.0">2154 masl</elevation>
</quantity>
, 
<date id="4F31D89DFF937050A989F918FAE9F97F" box="[1191,1313,1728,1754]" pageId="23" pageNumber="174" value="2021-06-12">
<collectingDate id="5F752175FF937050A989F918FAE9F97F" box="[1191,1313,1728,1754]" pageId="23" pageNumber="174" value="2021-06-12">12.vi.2021</collectingDate>
</date>
, 
<collectorName id="967A9B8BFF937050A803F918FF1BF95B" pageId="23" pageNumber="174">A. Betancourt</collectorName>
, 
<collectorName id="967A9B8BFF937050ADF1F93CFE80F95B" box="[223,328,1764,1790]" pageId="23" pageNumber="174">S. Forero</collectorName>
&amp; 
<collectorName id="967A9B8BFF937050AC45F93CFE0FF95B" box="[363,455,1764,1790]" pageId="23" pageNumber="174">J. Cabra</collectorName>
leg. (MUSENUV-Ar 2104)
</materialsCitation>
. 
<typeStatus id="E43440FFFF937050AE23F93CFCB6F95B" box="[781,894,1764,1790]" pageId="23" pageNumber="174" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="2D8935D4FF937050AEA7F93CFC3AF95B" box="[905,1010,1764,1790]" pageId="23" pageNumber="174" type="adult">two adult</specimenCount>
females (MUSENUV-Ar 2109, 2111) and 
<materialsCitation id="8BE7F400FF937050ADE8F8D0FCB7F887" ID-GBIF-Occurrence="3816447314" box="[198,895,1800,1826]" collectingDate="2021-06-12" collectorName="El Cerrito &amp; El Moral Sidewalk &amp; La Floresta Farm &amp; A. Betancourt &amp; S. Forero &amp; J. Cabra" country="Colombia" county="El Cerrito" elevation="2154" latitude="3.6783056" location="El Moral Sidewalk" longLatPrecision="1" longitude="-76.07486" municipality="Tenerife Pathway" pageId="23" pageNumber="174" specimenCount="1" specimenCount-juvenile="1" stateProvince="Valle del Cauca" typeStatus="paratype">
<specimenCount id="2D8935D4FF937050ADE8F8D0FE86F887" box="[198,334,1800,1826]" pageId="23" pageNumber="174" type="juvenile">one juvenile</specimenCount>
(MUSENUV-Ar 2110), same data as the holotype
</materialsCitation>
; 
<specimenCount id="2D8935D4FF937050AEA7F8D0FBDEF887" box="[905,1046,1800,1826]" pageId="23" pageNumber="174" type="subadult">one subadult</specimenCount>
female (MUSENUV-Ar 2107) and 
<materialsCitation id="8BE7F400FF937050ADB9F8F4FE88F8CF" ID-GBIF-Occurrence="3816447310" collectingDate="2019-07-05" collectionCode="ICN" collectorName="D. Garrido &amp; J. Cabra" country="Colombia" county="El Cerrito" elevation="2154" latitude="3.6783056" location="El Moral Sidewalk" longLatPrecision="1" longitude="-76.07486" municipality="Tenerife Pathway" pageId="23" pageNumber="174" specimenCode="ICN 1402, 1403" specimenCount="1" specimenCount-adult="1" stateProvince="Valle del Cauca" typeStatus="paratype">
<specimenCount id="2D8935D4FF937050ADB9F8F4FEC7F8E3" box="[151,271,1836,1862]" pageId="23" pageNumber="174" type="adult">three adult</specimenCount>
females (
<specimenCode id="6B295626FF937050AC53F8F4FE3AF8E3" box="[381,498,1836,1862]" collectionCode="ICN" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="23" pageNumber="174">ICN 1402</specimenCode>
, 
<specimenCode id="6B295626FF937050ACD0F8F4FDF1F8E3" box="[510,569,1836,1862]" collectionCode="ICN" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="23" pageNumber="174">1403</specimenCode>
, MUSENUV-Ar 2108), same locality as the holotype, 
<date id="4F31D89DFF937050A98CF8F4FADEF8E2" box="[1186,1302,1836,1863]" pageId="23" pageNumber="174" value="2019-07-05">
<collectingDate id="5F752175FF937050A98CF8F4FADEF8E2" box="[1186,1302,1836,1863]" pageId="23" pageNumber="174" value="2019-07-05">5.vii.2019</collectingDate>
</date>
, 
<collectorName id="967A9B8BFF937050A80CF8F4FA54F8E3" box="[1314,1436,1836,1862]" pageId="23" pageNumber="174">D. Garrido</collectorName>
&amp; 
<collectorName id="967A9B8BFF937050AD9AF888FED8F8CF" box="[180,272,1872,1898]" pageId="23" pageNumber="174">J. Cabra</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="7395ADD6FF937050ADE9F8ACFAD6F873" pageId="23" pageNumber="174" type="etymology">
<paragraph id="3B30FE5DFF937050ADE9F8ACFAD6F873" blockId="23.[151,1437,1692,2042]" pageId="23" pageNumber="174">
<emphasis id="09FB224FFF937050ADE9F8ACFE86F82B" bold="true" box="[199,334,1908,1934]" pageId="23" pageNumber="174">Etymology.</emphasis>
Named after the El Moral Sidewalk, Cerrito municipality in the Colombian Central Andes where the 
<typeStatus id="E43440FFFF937050ADEFF840FF39F817" box="[193,241,1944,1970]" pageId="23" pageNumber="174">type</typeStatus>
material was collected, using the Latin suffix -ensis denoting place of origin. The name was suggested by the owners of La Floresta Farm who detected the presence of the species under fallen logs decades ago.
</paragraph>
</subSubSection>
<subSubSection id="7395ADD6FF93705FADE9F838FC42FEE4" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174" type="diagnosis">
<paragraph id="3B30FE5DFF93705FADE9F838FC42FEE4" blockId="23.[151,1437,1692,2042]" lastBlockId="24.[151,1436,150,321]" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174">
<emphasis id="09FB224FFF937050ADE9F838FE8AF85F" bold="true" box="[199,322,2016,2042]" pageId="23" pageNumber="174">Diagnosis.</emphasis>
<taxonomicName id="FC8F85DEFF937050AC64F839FDD8F85F" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[330,528,2016,2042]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov.">
<emphasis id="09FB224FFF937050AC64F839FDD8F85F" box="[330,528,2016,2042]" italics="true" pageId="23" pageNumber="174">Tityus moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF937050AF37F838FDBCF85F" bold="true" box="[537,628,2016,2042]" pageId="23" pageNumber="174">
<taxonomicNameLabel id="12C89F34FF937050AF37F838FDBCF85F" box="[537,628,2016,2042]" pageId="23" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
share with 
<taxonomicName id="FC8F85DEFF937050AFD5F839FC55F85F" authorityName="Lourenco" authorityYear="1994" box="[763,925,2016,2042]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="cuellari">
<emphasis id="09FB224FFF937050AFD5F839FC55F85F" box="[763,925,2016,2042]" italics="true" pageId="23" pageNumber="174">Tityus cuellari</emphasis>
</taxonomicName>
, 
<taxonomicName id="FC8F85DEFF937050AE83F839FBB7F85F" authorityName="Lourenco" authorityYear="2013" box="[941,1151,2016,2042]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="crassicauda">
<emphasis id="09FB224FFF937050AE83F839FBB7F85F" box="[941,1151,2016,2042]" italics="true" pageId="23" pageNumber="174">Tityus crassicauda</emphasis>
</taxonomicName>
, and 
<taxonomicName id="FC8F85DEFF937050A9EEF839FABAF85F" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[1216,1394,2016,2042]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF937050A9EEF839FABAF85F" box="[1216,1394,2016,2042]" italics="true" pageId="23" pageNumber="174">Tityus forcipula</emphasis>
</taxonomicName>
the presence of 
<emphasis id="09FB224FFF9C705FAC33FF4EFEE6FF15" bold="true" box="[285,302,150,176]" pageId="24" pageNumber="175">i)</emphasis>
a metasoma widening towards segment V in both sexes (
<figureCitation id="A3B4E2D8FF9C705FAE84FF4FFB91FF14" box="[938,1113,151,177]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="24" pageNumber="175">Figures 12C, D</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9C705FA94DFF4FFB7DFF14" box="[1123,1205,151,177]" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="24" pageNumber="175">13C, D</figureCitation>
) and 
<emphasis id="09FB224FFF9C705FA9DCFF4EFAC3FF15" bold="true" box="[1266,1291,150,176]" pageId="24" pageNumber="175">ii)</emphasis>
a male chela bulkier than female chela (
<figureCitation id="A3B4E2D8FF9C705FACECFF63FD97FF70" box="[450,607,187,213]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[261,1326,181,1920]" captionTargetId="figure-19@15.[261,1326,181,1920]" captionTargetPageId="15" captionText="FIGURE 7. Tityus (Atreus) Gervais, 1843,female chela sculpturing.A–C. Tityus(Atreus) forcipula (Gervais,1843) (MUSENUV- Ar 436). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (ICN-1402). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673356" httpUri="https://zenodo.org/record/6673356/files/figure.png" pageId="24" pageNumber="175">Figures 7D–F</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9C705FAF45FF63FD66FF70" box="[619,686,187,213]" captionStart="FIGURE 8" captionStartId="16.[151,250,1944,1968]" captionTargetBox="[364,1224,181,1920]" captionTargetId="figure-31@16.[364,1224,181,1920]" captionTargetPageId="16" captionText="FIGURE 8. Tityus (Atreus) Gervais, 1843, male chela sculpturing. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673358" httpUri="https://zenodo.org/record/6673358/files/figure.png" pageId="24" pageNumber="175">8D–F</figureCitation>
). However, 
<taxonomicName id="FC8F85DEFF9C705FAE18FF63FC33FF70" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[822,1019,187,213]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov.">
<emphasis id="09FB224FFF9C705FAE18FF63FC33FF70" box="[822,1019,187,213]" italics="true" pageId="24" pageNumber="175">Tityus moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF9C705FA92CFF63FB93FF70" bold="true" box="[1026,1115,187,213]" pageId="24" pageNumber="175">
<taxonomicNameLabel id="12C89F34FF9C705FA92CFF63FB93FF70" box="[1026,1115,187,213]" pageId="24" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be readily distinguished by the presence of moderately protruding rounded granules on the DSM carinae of metasomal segments II–IV (
<figureCitation id="A3B4E2D8FF9C705FA842FF07FEDFFEB8" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="24" pageNumber="175">Figure 14C, D</figureCitation>
). Whereas, in 
<taxonomicName id="FC8F85DEFF9C705FAC99FEDBFDE2FEB8" authorityName="Lourenco" authorityYear="1994" box="[439,554,259,285]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="cuellari">
<emphasis id="09FB224FFF9C705FAC99FEDBFDE2FEB8" box="[439,554,259,285]" italics="true" pageId="24" pageNumber="175">T. cuellari</emphasis>
</taxonomicName>
, 
<taxonomicName id="FC8F85DEFF9C705FAF16FEDBFD13FEB8" authorityName="Lourenco" authorityYear="2013" box="[568,731,259,285]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="crassicauda">
<emphasis id="09FB224FFF9C705FAF16FEDBFD13FEB8" box="[568,731,259,285]" italics="true" pageId="24" pageNumber="175">T. crassicauda</emphasis>
</taxonomicName>
, and 
<taxonomicName id="FC8F85DEFF9C705FAE37FEDBFC01FEB8" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[793,969,259,285]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF9C705FAE37FEDBFC01FEB8" box="[793,969,259,285]" italics="true" pageId="24" pageNumber="175">Tityus forcipula</emphasis>
</taxonomicName>
the DSM carinae of metasomal segments II–IV exhibit strongly protruding spinoid granules (
<figureCitation id="A3B4E2D8FF9C705FAFF9FEFFFCB5FEE4" box="[727,893,295,321]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="24" pageNumber="175">Figure 14A, B</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="6FF0AED5FF9C705FADB9FB9AFCEAFB07" ID-DOI="http://doi.org/10.5281/zenodo.6673376" ID-Zenodo-Dep="6673376" httpUri="https://zenodo.org/record/6673376/files/figure.png" pageId="24" pageNumber="175" startId="24.[151,250,1090,1114]" targetBox="[151,1436,345,1067]" targetPageId="24">
<paragraph id="3B30FE5DFF9C705FADB9FB9AFCEAFB07" blockId="24.[151,1436,1090,1187]" pageId="24" pageNumber="175">
<emphasis id="09FB224FFF9C705FADB9FB9AFEEBFBFF" bold="true" box="[151,291,1090,1114]" pageId="24" pageNumber="175">FIGURE 15.</emphasis>
<taxonomicName id="FC8F85DEFF9C705FAC03FB9BFDACFBFF" authority="Gervais, 1843" authorityName="Gervais" authorityYear="1843" box="[301,612,1090,1114]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="175" phylum="Arthropoda" rank="subGenus" subGenus="Atreus">
<emphasis id="09FB224FFF9C705FAC03FB9BFEA2FBFF" box="[301,362,1091,1114]" italics="true" pageId="24" pageNumber="175">Tityus</emphasis>
(
<emphasis id="09FB224FFF9C705FAC52FB9BFE75FBFF" box="[380,445,1091,1114]" italics="true" pageId="24" pageNumber="175">Atreus</emphasis>
) 
<bibRefCitation id="5F1E83ACFF9C705FACFEFB9AFDACFBFF" author="Gervais, P. M." box="[464,612,1090,1114]" pageId="24" pageNumber="175" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
</taxonomicName>
, telson, lateral view. 
<emphasis id="09FB224FFF9C705FAE66FB9AFC4AFBFF" bold="true" box="[840,898,1090,1114]" pageId="24" pageNumber="175">A, B.</emphasis>
<taxonomicName id="FC8F85DEFF9C705FAEA2FB9BFAF2FBFF" authority="(Gervais, 1843)" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[908,1338,1090,1114]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="forcipula" subGenus="Atreus">
<emphasis id="09FB224FFF9C705FAEA2FB9BFC01FBFF" box="[908,969,1091,1114]" italics="true" pageId="24" pageNumber="175">Tityus</emphasis>
(
<emphasis id="09FB224FFF9C705FAEF5FB9BFBD4FBFF" box="[987,1052,1091,1114]" italics="true" pageId="24" pageNumber="175">Atreus</emphasis>
) 
<emphasis id="09FB224FFF9C705FA900FB9AFB42FBFF" box="[1070,1162,1090,1114]" italics="true" pageId="24" pageNumber="175">forcipula</emphasis>
(
<bibRefCitation id="5F1E83ACFF9C705FA9B5FB9AFAFAFBFF" author="Gervais, P. M." box="[1179,1330,1090,1114]" pageId="24" pageNumber="175" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
)
</taxonomicName>
. 
<emphasis id="09FB224FFF9C705FA868FB9AFA96FBFF" bold="true" box="[1350,1374,1090,1114]" pageId="24" pageNumber="175">A.</emphasis>
Male (MUSENUV-Ar 438). 
<emphasis id="09FB224FFF9C705FAC55FBBEFE5AFBDB" bold="true" box="[379,402,1126,1150]" pageId="24" pageNumber="175">B.</emphasis>
Female (MUSENUV-Ar 436). 
<emphasis id="09FB224FFF9C705FAFE6FBBEFD36FBDB" bold="true" box="[712,766,1126,1150]" pageId="24" pageNumber="175">C, D.</emphasis>
<taxonomicName id="FC8F85DEFF9C705FAE2CFBBFFBC0FBDB" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[770,1032,1126,1150]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov." subGenus="Atreus">
<emphasis id="09FB224FFF9C705FAE2CFBBFFCF7FBDB" box="[770,831,1127,1150]" italics="true" pageId="24" pageNumber="175">Tityus</emphasis>
(
<emphasis id="09FB224FFF9C705FAE65FBBFFC44FBDB" box="[843,908,1127,1150]" italics="true" pageId="24" pageNumber="175">Atreus</emphasis>
) 
<emphasis id="09FB224FFF9C705FAEB7FBBEFBC0FBDB" box="[921,1032,1126,1150]" italics="true" pageId="24" pageNumber="175">moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF9C705FA922FBBEFBBFFBDB" bold="true" box="[1036,1143,1126,1150]" pageId="24" pageNumber="175">
<taxonomicNameLabel id="12C89F34FF9C705FA922FBBEFB92FBDB" box="[1036,1114,1126,1150]" pageId="24" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
C.
</emphasis>
Male holotype (MUSENUVAr 2104). 
<emphasis id="09FB224FFF9C705FAC2EFB52FED1FB07" bold="true" box="[256,281,1162,1186]" pageId="24" pageNumber="175">D.</emphasis>
Female paratype (ICN-1402). Scale bars= 1.5 mm.
</paragraph>
</caption>
<subSubSection id="7395ADD6FF9C705CADE9FB08FCF2FEE4" lastPageId="27" lastPageNumber="178" pageId="24" pageNumber="175" type="description">
<paragraph id="3B30FE5DFF9C705FADE9FB08FA58FB4E" blockId="24.[151,1437,1232,2015]" box="[199,1424,1232,1259]" pageId="24" pageNumber="175">
<emphasis id="09FB224FFF9C705FADE9FB08FE91FB4F" bold="true" box="[199,345,1232,1258]" pageId="24" pageNumber="175">Description.</emphasis>
Based on adult male 
<typeStatus id="E43440FFFF9C705FAF64FB08FD64FB4F" box="[586,684,1232,1258]" pageId="24" pageNumber="175" type="holotype">holotype</typeStatus>
(MUSENUV-Ar 2104) and an adult female 
<typeStatus id="E43440FFFF9C705FA9B0FB09FB36FB4E" box="[1182,1278,1233,1259]" pageId="24" pageNumber="175" type="paratype">paratype</typeStatus>
(ICN-1402).
</paragraph>
<paragraph id="3B30FE5DFF9C705FADE9FB2CFA52FAAA" blockId="24.[151,1437,1232,2015]" box="[199,1434,1268,1295]" pageId="24" pageNumber="175">
<emphasis id="09FB224FFF9C705FADE9FB2CFE90FAAA" bold="true" box="[199,344,1268,1295]" italics="true" pageId="24" pageNumber="175">Total length.</emphasis>
Male 
<typeStatus id="E43440FFFF9C705FAC8EFB2CFDCEFAAB" box="[416,518,1268,1294]" pageId="24" pageNumber="175" type="holotype">holotype</typeStatus>
: 
<quantity id="FC7753B8FF9C705FAF3FFB2CFD4BFAAA" box="[529,643,1268,1295]" metricMagnitude="-2" metricUnit="m" metricValue="5.503" pageId="24" pageNumber="175" unit="mm" value="55.03">55.03 mm</quantity>
and female 
<typeStatus id="E43440FFFF9C705FAE22FB2DFCB9FAAA" box="[780,881,1269,1295]" pageId="24" pageNumber="175" type="paratype">paratype</typeStatus>
: 
<quantity id="FC7753B8FF9C705FAE55FB2CFC25FAAA" box="[891,1005,1268,1295]" metricMagnitude="-2" metricUnit="m" metricValue="5.799" pageId="24" pageNumber="175" unit="mm" value="57.99">57.99 mm</quantity>
(additional measurements in 
<tableCitation id="760DCBE6FF9C705FA819FB2CFA44FAAA" box="[1335,1420,1268,1295]" captionStart="TABLE 5" captionStartId="11.[151,239,440,464]" captionTargetBox="[167,1416,516,1987]" captionTargetPageId="11" captionText="TABLE 5. Measurements (mm) of Tityus forcipula and Tityus moralensis sp. nov. Observation: all specimens, but 1402" httpUri="http://table.plazi.org/id/6FF0AED5FF8F704CADB9FE60FAC8FE51" pageId="24" pageNumber="175" tableUuid="6FF0AED5FF8F704CADB9FE60FAC8FE51">Table 5</tableCitation>
).
</paragraph>
<paragraph id="3B30FE5DFF9C705FADE9FAC0FD27F98B" blockId="24.[151,1437,1232,2015]" pageId="24" pageNumber="175">
<emphasis id="09FB224FFF9C705FADE9FAC0FE87FA97" bold="true" box="[199,335,1304,1330]" pageId="24" pageNumber="175">Coloration.</emphasis>
General pattern (in 70% ethanol) (
<figureCitation id="A3B4E2D8FF9C705FAFF0FAC0FCB0FA96" box="[734,888,1304,1331]" captionStart="FIGURE 3" captionStartId="8.[151,250,1927,1951]" captionTargetBox="[162,1419,202,1891]" captionTargetId="figure-31@8.[151,1436,181,1903]" captionTargetPageId="8" captionText="FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm." figureDoi="http://doi.org/10.5281/zenodo.6673342" httpUri="https://zenodo.org/record/6673342/files/figure.png" pageId="24" pageNumber="175">Figure 3A–D</figureCitation>
) dark reddish-brown. 
<emphasis id="09FB224FFF9C705FA95BFAC1FB2BFA96" box="[1141,1251,1305,1331]" italics="true" pageId="24" pageNumber="175">Carapace</emphasis>
(
<figureCitation id="A3B4E2D8FF9C705FA9DDFAC0FA46FA96" box="[1267,1422,1304,1331]" captionStart="FIGURE 3" captionStartId="8.[151,250,1927,1951]" captionTargetBox="[162,1419,202,1891]" captionTargetId="figure-31@8.[151,1436,181,1903]" captionTargetPageId="8" captionText="FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm." figureDoi="http://doi.org/10.5281/zenodo.6673342" httpUri="https://zenodo.org/record/6673342/files/figure.png" pageId="24" pageNumber="175">Figure 3A, C</figureCitation>
): dark reddish-brown with black variegated pigments; black stripe on anterior margin; lateral and median eyes, surrounded by black variegated pigments. 
<emphasis id="09FB224FFF9C705FAF60FAB8FD01FADF" box="[590,713,1376,1402]" italics="true" pageId="24" pageNumber="175">Chelicerae</emphasis>
(
<figureCitation id="A3B4E2D8FF9C705FAFF9FAB8FCA6FADE" box="[727,878,1376,1403]" captionStart="FIGURE 3" captionStartId="8.[151,250,1927,1951]" captionTargetBox="[162,1419,202,1891]" captionTargetId="figure-31@8.[151,1436,181,1903]" captionTargetPageId="8" captionText="FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm." figureDoi="http://doi.org/10.5281/zenodo.6673342" httpUri="https://zenodo.org/record/6673342/files/figure.png" pageId="24" pageNumber="175">Figure 3A, C</figureCitation>
): coxa and hand dark yellow, with abundant black pigments over the hand surface; fingers, blackish-brown. 
<emphasis id="09FB224FFF9C705FAE3AFA5CFC23FA3B" box="[788,1003,1412,1438]" italics="true" pageId="24" pageNumber="175">Coxosternal region</emphasis>
, 
<emphasis id="09FB224FFF9C705FAED6FA5DFA96FA3B" box="[1016,1374,1412,1438]" italics="true" pageId="24" pageNumber="175">
Legs, 
<taxonomicName id="FC8F85DEFF9C705FA914FA5DFB7DFA3B" box="[1082,1205,1413,1438]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="24" pageNumber="175" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
, and Pedipalps
</emphasis>
(
<figureCitation id="A3B4E2D8FF9C705FA842FA5CFEC3FA66" captionStart="FIGURE 3" captionStartId="8.[151,250,1927,1951]" captionTargetBox="[162,1419,202,1891]" captionTargetId="figure-31@8.[151,1436,181,1903]" captionTargetPageId="8" captionText="FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm." figureDoi="http://doi.org/10.5281/zenodo.6673342" httpUri="https://zenodo.org/record/6673342/files/figure.png" pageId="24" pageNumber="175">Figure 3A–D</figureCitation>
): all reddish-brown, covered with black variegated pigments (except for coxosternal region, ventral areas of the pedipalp segments, and chela manus). 
<emphasis id="09FB224FFF9C705FAFA7FA14FD04FA43" box="[649,716,1484,1510]" italics="true" pageId="24" pageNumber="175">Chela</emphasis>
(
<figureCitation id="A3B4E2D8FF9C705FAFF4FA14FCB9FA42" box="[730,881,1484,1511]" captionStart="FIGURE 3" captionStartId="8.[151,250,1927,1951]" captionTargetBox="[162,1419,202,1891]" captionTargetId="figure-31@8.[151,1436,181,1903]" captionTargetPageId="8" captionText="FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm." figureDoi="http://doi.org/10.5281/zenodo.6673342" httpUri="https://zenodo.org/record/6673342/files/figure.png" pageId="24" pageNumber="175">Figure 3A–D</figureCitation>
) fingers blackish. 
<emphasis id="09FB224FFF9C705FA910FA15FB7BFA43" box="[1086,1203,1485,1510]" italics="true" pageId="24" pageNumber="175">Metasoma</emphasis>
(
<figureCitation id="A3B4E2D8FF9C705FA9EFFA14FA93FA42" box="[1217,1371,1484,1511]" captionStart="FIGURE 3" captionStartId="8.[151,250,1927,1951]" captionTargetBox="[162,1419,202,1891]" captionTargetId="figure-31@8.[151,1436,181,1903]" captionTargetPageId="8" captionText="FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm." figureDoi="http://doi.org/10.5281/zenodo.6673342" httpUri="https://zenodo.org/record/6673342/files/figure.png" pageId="24" pageNumber="175">Figure 3A–D</figureCitation>
): segments dark reddish-brown, progressively darker towards the telson; segments IV–V blackish-brown. 
<taxonomicName id="FC8F85DEFF9C705FA9D5FA28FA8BF9AF" box="[1275,1347,1520,1546]" class="Hexanauplia" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="09FB224FFF9C705FA9D5FA28FA8BF9AF" box="[1275,1347,1520,1546]" italics="true" pageId="24" pageNumber="175">Telson</emphasis>
</taxonomicName>
(
<figureCitation id="A3B4E2D8FF9C705FA87FFA28FF29F98A" captionStart="FIGURE 3" captionStartId="8.[151,250,1927,1951]" captionTargetBox="[162,1419,202,1891]" captionTargetId="figure-31@8.[151,1436,181,1903]" captionTargetPageId="8" captionText="FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm." figureDoi="http://doi.org/10.5281/zenodo.6673342" httpUri="https://zenodo.org/record/6673342/files/figure.png" pageId="24" pageNumber="175">Figure 3A–D</figureCitation>
): blackish-brown; aculeus dark reddish-brown.
</paragraph>
<paragraph id="3B30FE5DFF9C705FADE9F9E1FEB5F8EA" blockId="24.[151,1437,1232,2015]" pageId="24" pageNumber="175">
<emphasis id="09FB224FFF9C705FADE9F9E1FEA9F9F6" bold="true" box="[199,353,1593,1619]" pageId="24" pageNumber="175">Morphology.</emphasis>
Body moderately covered with microsetae (except for the pedipalp that is densely covered with microsetae), white colored under UV light. 
<emphasis id="09FB224FFF9C705FAF59F985FD2DF9D2" box="[631,741,1629,1655]" italics="true" pageId="24" pageNumber="175">Carapace</emphasis>
(
<figureCitation id="A3B4E2D8FF9C705FAFDDF984FC41F9D2" box="[755,905,1628,1655]" captionStart="FIGURE 4" captionStartId="10.[151,250,1375,1399]" captionTargetBox="[371,1216,309,1352]" captionTargetId="figure-86@10.[371,1216,309,1352]" captionTargetPageId="10" captionText="FIGURE 4. Tityus (Atreus) Gervais, 1843, carapace sculpturing, dorsal aspect. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673348" httpUri="https://zenodo.org/record/6673348/files/figure.png" pageId="24" pageNumber="175">Figure 4C, D</figureCitation>
): densely covered with fine granulation and few coarse granules; anterior margin with deep median notch; anterior median carinae, central lateral, central median, lateral ocular, posterior, posterior median and superciliary carinae, all well-marked, and furrows (anterior median, anterior marginal, central transverse, lateral ocular, supercialiary, posterior transverse, posterior lateral and posterior marginal), all well-marked; ocular tubercle well-marked, located on the anterior half of carapace; median eyes separated by 0.5 ocular diameters; with three pairs of lateral eyes and two pairs of lateral micro-ocelli (ADMi and PDMi) (pattern 4A).
</paragraph>
<paragraph id="3B30FE5DFF9C705FADE9F880FA5BF833" blockId="24.[151,1437,1232,2015]" pageId="24" pageNumber="175">
<emphasis id="09FB224FFF9C705FADE9F880FDCEF8D7" bold="true" box="[199,518,1880,1907]" pageId="24" pageNumber="175">
<emphasis id="09FB224FFF9C705FADE9F880FE8BF8D6" bold="true" box="[199,323,1880,1907]" italics="true" pageId="24" pageNumber="175">Chelicerae</emphasis>
(
<figureCitation id="A3B4E2D8FF9C705FAC7AF880FE3FF8D7" box="[340,503,1880,1906]" captionStart="FIGURE 4" captionStartId="10.[151,250,1375,1399]" captionTargetBox="[371,1216,309,1352]" captionTargetId="figure-86@10.[371,1216,309,1352]" captionTargetPageId="10" captionText="FIGURE 4. Tityus (Atreus) Gervais, 1843, carapace sculpturing, dorsal aspect. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673348" httpUri="https://zenodo.org/record/6673348/files/figure.png" pageId="24" pageNumber="175">Figure 4C, D</figureCitation>
).
</emphasis>
Dentition characteristic of the family 
<taxonomicName id="FC8F85DEFF9C705FAE90F880FB1FF8D6" authority="(Vachon 1963)" baseAuthorityName="Vachon" baseAuthorityYear="1963" box="[958,1239,1880,1907]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="24" pageNumber="175" phylum="Arthropoda" rank="family">
Buthidae (
<bibRefCitation id="5F1E83ACFF9C705FA91AF880FB06F8D6" author="Vachon, M." box="[1076,1230,1880,1907]" pageId="24" pageNumber="175" pagination="161 - 166" refId="ref21343" refString="Vachon, M. (1963) De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris, 161 - 166." type="book chapter" year="1963">Vachon 1963</bibRefCitation>
)
</taxonomicName>
, densely covered with setae over the internal and ventral surfaces; anterior half of the manus with a transverse row of fine granules.
</paragraph>
<paragraph id="3B30FE5DFF9C705EADE9F878FAAEFE50" blockId="24.[151,1437,1232,2015]" lastBlockId="25.[151,1437,151,2013]" lastPageId="25" lastPageNumber="176" pageId="24" pageNumber="175">
<emphasis id="09FB224FFF9C705FADE9F878FEF6F81E" bold="true" box="[199,318,1952,1979]" pageId="24" pageNumber="175">
<emphasis id="09FB224FFF9C705FADE9F878FEFFF81E" bold="true" box="[199,311,1952,1979]" italics="true" pageId="24" pageNumber="175">Pedipalps</emphasis>
.
</emphasis>
Chela, bulky in male (L/W= 3.18) or slender in female (L/W= 4.35). Trichobothriotaxic pattern 
<typeStatus id="E43440FFFF9C705FA84BF879FA55F81E" box="[1381,1437,1953,1979]" pageId="24" pageNumber="175">Type</typeStatus>
A, alfa configuration (hand: Eb3:Eb2:Eb1:Esb:Est:Et, fixed finger: eb:esb:est:et:db:dt:it). 
<emphasis id="09FB224FFF9C705FA98EF81DFB23F87B" box="[1184,1259,1989,2014]" italics="true" pageId="24" pageNumber="175">Femur</emphasis>
(
<figureCitation id="A3B4E2D8FF9C705FA9D2F81CFA5CF87A" box="[1276,1428,1988,2015]" captionStart="FIGURE 5" captionStartId="13.[151,250,1447,1471]" captionTargetBox="[151,1437,309,1424]" captionTargetId="figure-91@13.[151,1437,309,1424]" captionTargetPageId="13" captionText="FIGURE 5. Tityus (Atreus) Gervais, 1843, femur sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673350" httpUri="https://zenodo.org/record/6673350/files/figure.png" pageId="24" pageNumber="175">Figure 5E–H</figureCitation>
) with five well-marked carinae: VI, DI, DE and VE crenulate, EM serratocrenulate and complete, with intercarinal areas densely covered with fine granulation and few coarse granules. 
<taxonomicName id="FC8F85DEFF9D705EAE92FF63FBC5FF70" box="[956,1037,187,213]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Aplysiida" pageId="25" pageNumber="176" phylum="Mollusca" rank="genus">
<emphasis id="09FB224FFF9D705EAE92FF63FBC5FF70" box="[956,1037,187,213]" italics="true" pageId="25" pageNumber="176">Patella</emphasis>
</taxonomicName>
(
<figureCitation id="A3B4E2D8FF9D705EA930FF63FB7DFF73" box="[1054,1205,187,214]" captionStart="FIGURE 6" captionStartId="14.[151,250,1863,1887]" captionTargetBox="[151,1436,561,1840]" captionTargetId="figure-228@14.[151,1436,561,1840]" captionTargetPageId="14" captionText="FIGURE 6. Tityus (Atreus) Gervais, 1843, patella sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673354" httpUri="https://zenodo.org/record/6673354/files/figure.png" pageId="25" pageNumber="176">Figure 6E–H</figureCitation>
) with seven carinae well-marked: VI, DI, DE and EM, crenulate and complete; VE obsolete- only present on the distal half of the segment (male) or complete (female) and crenulate; DM crenulate and incomplete; IM serratocrenulate and complete, with a short spur near the segment base; with intercarinal areas densely covered with fine granulation. 
<emphasis id="09FB224FFF9D705EA823FEFFFA98FEE4" box="[1293,1360,295,321]" italics="true" pageId="25" pageNumber="176">Chela</emphasis>
(
<emphasis id="09FB224FFF9D705EA84EFEFFFA5BFEE4" box="[1376,1427,295,321]" italics="true" pageId="25" pageNumber="176">tibia</emphasis>
) (
<figureCitation id="A3B4E2D8FF9D705EADB1FE93FEF5FEC0" box="[159,317,331,357]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[261,1326,181,1920]" captionTargetId="figure-19@15.[261,1326,181,1920]" captionTargetPageId="15" captionText="FIGURE 7. Tityus (Atreus) Gervais, 1843,female chela sculpturing.A–C. Tityus(Atreus) forcipula (Gervais,1843) (MUSENUV- Ar 436). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (ICN-1402). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673356" httpUri="https://zenodo.org/record/6673356/files/figure.png" pageId="25" pageNumber="176">Figures 7D–F</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9D705EAC67FE93FE42FEC0" box="[329,394,331,357]" captionStart="FIGURE 8" captionStartId="16.[151,250,1944,1968]" captionTargetBox="[364,1224,181,1920]" captionTargetId="figure-31@16.[364,1224,181,1920]" captionTargetPageId="16" captionText="FIGURE 8. Tityus (Atreus) Gervais, 1843, male chela sculpturing. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673358" httpUri="https://zenodo.org/record/6673358/files/figure.png" pageId="25" pageNumber="176">8D–F</figureCitation>
) with eight slightly-marked (obsolete) carinae: VI, VE, D, DS, DMA, IM and ES, crenulate and complete; SA crenulate and incomplete, only present on the anterior half of the hand. Pedipalp movable and fixed fingers each with a basal concavity followed by a well-marked basal lobe (male) (
<figureCitation id="A3B4E2D8FF9D705EA944FE4BFB17FE08" box="[1130,1247,403,429]" captionStart="FIGURE 8" captionStartId="16.[151,250,1944,1968]" captionTargetBox="[364,1224,181,1920]" captionTargetId="figure-31@16.[364,1224,181,1920]" captionTargetPageId="16" captionText="FIGURE 8. Tityus (Atreus) Gervais, 1843, male chela sculpturing. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673358" httpUri="https://zenodo.org/record/6673358/files/figure.png" pageId="25" pageNumber="176">Figure 8D</figureCitation>
) or movable and fixed fingers with obsolete basal concavities and a slightly-marked basal lobe only present on the movable finger (female) (
<figureCitation id="A3B4E2D8FF9D705EAC2BFE03FEB4FE50" box="[261,380,475,501]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[261,1326,181,1920]" captionTargetId="figure-19@15.[261,1326,181,1920]" captionTargetPageId="15" captionText="FIGURE 7. Tityus (Atreus) Gervais, 1843,female chela sculpturing.A–C. Tityus(Atreus) forcipula (Gervais,1843) (MUSENUV- Ar 436). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (ICN-1402). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673356" httpUri="https://zenodo.org/record/6673356/files/figure.png" pageId="25" pageNumber="176">Figure 7D</figureCitation>
). 
<emphasis id="09FB224FFF9D705EACBEFE03FD64FE50" box="[400,684,475,501]" italics="true" pageId="25" pageNumber="176">Oblique rows of granules</emphasis>
: Movable finger with 16–17 (male) and 14–16 (female) rows.
</paragraph>
<paragraph id="3B30FE5DFF9D705EADE9FE26FECDFD20" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="176">
<emphasis id="09FB224FFF9D705EADE9FE26FDAEFDBD" bold="true" box="[199,614,510,537]" pageId="25" pageNumber="176">
<emphasis id="09FB224FFF9D705EADE9FE26FE6FFDBC" bold="true" box="[199,423,510,537]" italics="true" pageId="25" pageNumber="176">Coxosternal region</emphasis>
(
<figureCitation id="A3B4E2D8FF9D705EAC99FE26FD90FDBD" box="[439,600,510,536]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="25" pageNumber="176">Figure 9C, D</figureCitation>
).
</emphasis>
Sternum with posterior depression, outer ridge, and apical button, wellmarked; region covered with abundant coarse and fine granulation, and abundant microsetae; genital operculum longitudinally divided, composed of two sub-triangular plates. Male genital papillae are rounded and with sclerotized tips.
</paragraph>
<paragraph id="3B30FE5DFF9D705EADE9FD57FD38FCF8" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="176">
<emphasis id="09FB224FFF9D705EADE9FD57FE2BFD0D" bold="true" box="[199,483,654,681]" pageId="25" pageNumber="176">
<emphasis id="09FB224FFF9D705EADE9FD57FEEFFD0C" bold="true" box="[199,295,655,681]" italics="true" pageId="25" pageNumber="176">Pectines</emphasis>
(
<figureCitation id="A3B4E2D8FF9D705EAC18FD56FE1CFD0D" box="[310,468,654,680]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="25" pageNumber="176">Figure 9C, D</figureCitation>
).
</emphasis>
Pectines are strongly sclerotized and brownish, with white areas (less sclerotized) on the marginal plate (
<figureCitation id="A3B4E2D8FF9D705EAC5EFD6BFDDBFD68" box="[368,531,691,717]" captionStart="FIGURE 3" captionStartId="8.[151,250,1927,1951]" captionTargetBox="[162,1419,202,1891]" captionTargetId="figure-31@8.[151,1436,181,1903]" captionTargetPageId="8" captionText="FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm." figureDoi="http://doi.org/10.5281/zenodo.6673342" httpUri="https://zenodo.org/record/6673342/files/figure.png" pageId="25" pageNumber="176">Figures 3B, D</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9D705EAF30FD6BFDADFD68" box="[542,613,691,717]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="25" pageNumber="176">9C, D</figureCitation>
). Pectinal basal piece shield-like shaped, with a deep anteromedian notch and without glandular areas (
<figureCitation id="A3B4E2D8FF9D705EACF6FD0FFDA6FD54" box="[472,622,727,753]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="25" pageNumber="176">Figure 9C, D</figureCitation>
); pectinal tooth counts of 17–17 (male) and 16–17 (female). Intermediate plate, marginal plate, and fulcra densely covered with microsetae (
<figureCitation id="A3B4E2D8FF9D705EAE56FD23FBC7FCB0" box="[888,1039,763,789]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="25" pageNumber="176">Figure 9C, D</figureCitation>
). Basal middle lamellae, not dilated (male) (
<figureCitation id="A3B4E2D8FF9D705EADC0FCC7FEA9FC9C" box="[238,353,799,825]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="25" pageNumber="176">Figure 9C</figureCitation>
) or dilated and ovoid shaped (female) (
<figureCitation id="A3B4E2D8FF9D705EAE38FCC7FC45FC9C" box="[790,909,799,825]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="25" pageNumber="176">Figure 9D</figureCitation>
). Female basal middle lamellae with a vestigial distal glandular area, white colored (
<figureCitation id="A3B4E2D8FF9D705EAF1EFC9BFD79FCF8" box="[560,689,835,861]" captionStart="FIGURE 3" captionStartId="8.[151,250,1927,1951]" captionTargetBox="[162,1419,202,1891]" captionTargetId="figure-31@8.[151,1436,181,1903]" captionTargetPageId="8" captionText="FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm." figureDoi="http://doi.org/10.5281/zenodo.6673342" httpUri="https://zenodo.org/record/6673342/files/figure.png" pageId="25" pageNumber="176">Figures 3D</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9D705EAF93FC9BFD2BFCF8" box="[701,739,835,861]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="25" pageNumber="176">9D</figureCitation>
).
</paragraph>
<paragraph id="3B30FE5DFF9D705EADE9FCBFFACAFC48" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="176">
<emphasis id="09FB224FFF9D705EADE9FCBFFECDFC24" bold="true" box="[199,261,871,897]" pageId="25" pageNumber="176">
<emphasis id="09FB224FFF9D705EADE9FCBFFF35FC24" bold="true" box="[199,253,871,897]" italics="true" pageId="25" pageNumber="176">Legs</emphasis>
.
</emphasis>
Carinae present; intercarinal areas with fine granulation; ventral telotarsi setae distributed in two rows of ventrosubmedian setae (
<typeStatus id="E43440FFFF9D705EAC88FC53FE10FC00" box="[422,472,907,933]" pageId="25" pageNumber="176">type</typeStatus>
II), acute and stout; telotarsi, counts of ventral macrosetae in the left (L) and right (R) legs on prolateral (pro) and retrolateral (retro) rows from leg I to IV (L (pro/retro) R (pro/retro)): 8/8 8/8; 7/8 8/8; 8/8 8/9; 8/11 8/11 (male) and 7/8 7/7; 6/7 6/8; 7/8 -/-; 8/9 7/7 (female). Claws short and symmetrical.
</paragraph>
<paragraph id="3B30FE5DFF9D705EADE9FC2FFCE7FAA8" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="176">
<emphasis id="09FB224FFF9D705EADE9FC2FFE81FBB4" bold="true" box="[199,329,1015,1041]" pageId="25" pageNumber="176">
<taxonomicName id="FC8F85DEFF9D705EADE9FC2FFE8AFBB4" box="[199,322,1015,1041]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="25" pageNumber="176" phylum="Arthropoda" rank="genus">
<emphasis id="09FB224FFF9D705EADE9FC2FFE8AFBB4" bold="true" box="[199,322,1015,1041]" italics="true" pageId="25" pageNumber="176">Mesosoma</emphasis>
</taxonomicName>
.
</emphasis>
Tergites I–VI (
<figureCitation id="A3B4E2D8FF9D705EACDFFC2FFD4FFBB4" box="[497,647,1015,1041]" captionStart="FIGURE 3" captionStartId="8.[151,250,1927,1951]" captionTargetBox="[162,1419,202,1891]" captionTargetId="figure-31@8.[151,1436,181,1903]" captionTargetPageId="8" captionText="FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm." figureDoi="http://doi.org/10.5281/zenodo.6673342" httpUri="https://zenodo.org/record/6673342/files/figure.png" pageId="25" pageNumber="176">Figure 3A, C</figureCitation>
), moderately covered with coarse and fine granulation; pre-tergite well defined, with median carina visible on the posterior margin of the post-tergite; tergite VII with DSM and DL carinae complete and crenulate, and median carina composed of a crenulate anteromedian eminence present on the anterior half of the post-tergite. Sternites densely covered with coarse and fine granulation (
<figureCitation id="A3B4E2D8FF9D705EA919FBBBFB14FBD8" box="[1079,1244,1123,1149]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[221,1367,181,1920]" captionTargetId="figure-19@19.[221,1367,181,1920]" captionTargetPageId="19" captionText="FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673364" httpUri="https://zenodo.org/record/6673364/files/figure.png" pageId="25" pageNumber="176">Figure 10C, D</figureCitation>
); sternites III–VI with a pair of elliptic spiracles on the posterior half, which are progressively larger (
<figureCitation id="A3B4E2D8FF9D705EA917FB5FFB15FB04" box="[1081,1245,1159,1185]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[221,1367,181,1920]" captionTargetId="figure-19@19.[221,1367,181,1920]" captionTargetPageId="19" captionText="FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673364" httpUri="https://zenodo.org/record/6673364/files/figure.png" pageId="25" pageNumber="176">Figure 10C, D</figureCitation>
); sternites III–VI with a median longitudinal hyaline suture; sternite V with a large hyaline subtriangular area on the posterior margin (
<figureCitation id="A3B4E2D8FF9D705EADB1FB17FE8DFB4C" box="[159,325,1231,1257]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[221,1367,181,1920]" captionTargetId="figure-19@19.[221,1367,181,1920]" captionTargetPageId="19" captionText="FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673364" httpUri="https://zenodo.org/record/6673364/files/figure.png" pageId="25" pageNumber="176">Figure 10C, D</figureCitation>
); sternite VI with VSM carinae crenulate, present on posterior half; sternite VII with VSM and VL carinae crenulate, present on posterior 2/3 (
<figureCitation id="A3B4E2D8FF9D705EAF55FB2BFCEAFAA8" box="[635,802,1267,1293]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[221,1367,181,1920]" captionTargetId="figure-19@19.[221,1367,181,1920]" captionTargetPageId="19" captionText="FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673364" httpUri="https://zenodo.org/record/6673364/files/figure.png" pageId="25" pageNumber="176">Figure 10C, D</figureCitation>
).
</paragraph>
<paragraph id="3B30FE5DFF9D705EADE9FACEFDE0F9D0" blockId="25.[151,1437,151,2013]" pageId="25" pageNumber="176">
<emphasis id="09FB224FFF9D705EADE9FACEFD49FA94" bold="true" box="[199,641,1302,1329]" pageId="25" pageNumber="176">
<emphasis id="09FB224FFF9D705EADE9FACEFE7BFA94" bold="true" box="[199,435,1302,1329]" italics="true" pageId="25" pageNumber="176">Hemispermatophore</emphasis>
(
<figureCitation id="A3B4E2D8FF9D705EACE8FACEFDB9FA94" box="[454,625,1302,1329]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="25" pageNumber="176">Figure 11D–E</figureCitation>
).
</emphasis>
Thin and sclerotized; pars reflexa, curved but not folded over itself (
<figureCitation id="A3B4E2D8FF9D705EADB1FAE3FEF7FAF0" box="[159,319,1339,1365]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="25" pageNumber="176">Figure 11D, E</figureCitation>
); foot narrow and flat (
<figureCitation id="A3B4E2D8FF9D705EAF12FAE3FD15FAF0" box="[572,733,1339,1365]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="25" pageNumber="176">Figure 11D–F</figureCitation>
); pedal flexure inconspicuous (
<figureCitation id="A3B4E2D8FF9D705EA91BFAE3FB1FFAF0" box="[1077,1239,1339,1365]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="25" pageNumber="176">Figure 11D–F</figureCitation>
); body occupying more than 2/3 of the hemispermatophore total length (
<figureCitation id="A3B4E2D8FF9D705EAFC1FA87FC59FADC" box="[751,913,1375,1401]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="25" pageNumber="176">Figure 11D–F</figureCitation>
). Capsular region with a furrow connecting the internal lobe with the medial area of the region (
<figureCitation id="A3B4E2D8FF9D705EAF95FA5BFC88FA38" box="[699,832,1411,1437]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="25" pageNumber="176">Figure 11D</figureCitation>
); internal lobe with rounded tip forming an 80º angle (
<figureCitation id="A3B4E2D8FF9D705EADB1FA7FFEE9FA64" box="[159,289,1447,1473]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="25" pageNumber="176">Figure 11E</figureCitation>
); median lobe inconspicuous (
<figureCitation id="A3B4E2D8FF9D705EAF56FA7FFCD1FA64" box="[632,793,1447,1473]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="25" pageNumber="176">Figure 11E, F</figureCitation>
); external lobe thin and not overpassing the internal lobe level and with a translucent area between the middle of the basal lobe and the base of the internal lobe (
<figureCitation id="A3B4E2D8FF9D705EA821FA13FA47FA40" box="[1295,1423,1483,1509]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="25" pageNumber="176">Figure 11F</figureCitation>
); translucent area narrow but widened basally and distally; basal lobe bifurcate and with a moderate sized projection subovate-shaped (in posterior and ventral views) (
<figureCitation id="A3B4E2D8FF9D705EAFE4F9CBFCA6F988" box="[714,878,1555,1581]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="25" pageNumber="176">Figure 11D, F</figureCitation>
); basal lobe with a conspicuous visible surface in lateral view (
<figureCitation id="A3B4E2D8FF9D705EAC05F9EFFE66F9F4" box="[299,430,1591,1617]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="25" pageNumber="176">Figure 11E</figureCitation>
); basal lobe with a “U’’-like shaped curvature between its base and the body (in anterior or posterior views) (
<figureCitation id="A3B4E2D8FF9D705EAC57F983FDD3F9D0" box="[377,539,1627,1653]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="25" pageNumber="176">Figure 11D, F</figureCitation>
).
</paragraph>
<paragraph id="3B30FE5DFF9D705DADE9F9A7FE6BFF5C" blockId="25.[151,1437,151,2013]" lastBlockId="26.[151,1437,151,1005]" lastPageId="26" lastPageNumber="177" pageId="25" pageNumber="176">
<emphasis id="09FB224FFF9D705EADE9F9A7FD15F93D" bold="true" box="[199,733,1662,1689]" pageId="25" pageNumber="176">
<emphasis id="09FB224FFF9D705EADE9F9A7FEF7F93C" bold="true" box="[199,319,1663,1689]" italics="true" pageId="25" pageNumber="176">Metasoma</emphasis>
(
<figureCitation id="A3B4E2D8FF9D705EAC61F9A6FDCFF93C" box="[335,519,1662,1689]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="25" pageNumber="176">Figures 12C, D</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9D705EAF3BF9A6FDA2F93C" box="[533,618,1662,1689]" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="25" pageNumber="176">13C, D</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9D705EAF56F9A6FD06F93D" box="[632,718,1662,1688]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="25" pageNumber="176">14C, D</figureCitation>
).
</emphasis>
Strongly widening towards segment V (
<figureCitation id="A3B4E2D8FF9D705EA989F9A7FA92F93C" box="[1191,1370,1663,1689]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="25" pageNumber="176">Figures 12C, D</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9D705EA848F9A7FF67F918" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="25" pageNumber="176">13C, D</figureCitation>
). Segments II–V short and robust (male- L/W ratio: II= 1.28; III= 1.29; IV= 1.09; V= 1.08/ female- L/W ratio: II= 1.24; III= 1.24; IV= 1.09; V= 1.10). Segments I–II (
<figureCitation id="A3B4E2D8FF9D705EAFCEF91FFC59F944" box="[736,913,1735,1761]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="25" pageNumber="176">Figures 12C, D</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9D705EAEB3F91FFC39F944" box="[925,1009,1735,1761]" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="25" pageNumber="176">13C, D</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9D705EAED2F91FFB86F944" box="[1020,1102,1735,1761]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="25" pageNumber="176">14C, D</figureCitation>
) with eight complete carinae, parallel and crenulate (paired DL, ML, VL and VSM) and two serratocrenulate (paired DSM); ML of segment II represented by coarse granules; intercarinal areas densely covered with fine granulation and moderately covered with coarse granulation; segments III–IV (
<figureCitation id="A3B4E2D8FF9D705EAF5FF8EBFCE9F8E8" box="[625,801,1843,1869]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="25" pageNumber="176">Figures 12C, D</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9D705EAE03F8EBFC48F8E8" box="[813,896,1843,1869]" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="25" pageNumber="176">13C, D</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9D705EAEA5F8EBFC15F8E8" box="[907,989,1843,1869]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="25" pageNumber="176">14C, D</figureCitation>
) with six complete carinae, parallel and crenulate (paired DL, VL and VSM) and two serratocrenulate (paired DSM), intercarinal areas densely covered with fine granulation and moderately covered with coarse granulation; segment V (
<figureCitation id="A3B4E2D8FF9D705EAED8F8A3FB6CF830" box="[1014,1188,1915,1941]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="25" pageNumber="176">Figures 12C, D</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9D705EA980F8A3FAC9F830" box="[1198,1281,1915,1941]" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="25" pageNumber="176">13C, D</figureCitation>
, 
<figureCitation id="A3B4E2D8FF9D705EA825F8A3FA94F830" box="[1291,1372,1915,1941]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="25" pageNumber="176">14C, D</figureCitation>
) with five complete carinae and crenulate (VM, paired VL and VSM carinae) and two serratocrenulate (paired DSM), intercarinal areas densely covered with coarse granulation and moderately covered with fine granulation. Segments II–V (
<figureCitation id="A3B4E2D8FF9E705DADF5FF4FFE48FF14" box="[219,384,151,177]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="26" pageNumber="177">Figure 14C, D</figureCitation>
) with DSM carinae, composed of moderately protruding rounded granules that progressively increase distally (except on segment V), ending in an enlarged distoterminal granule (distoterminal granule not enlarged in segment V).
</paragraph>
<paragraph id="3B30FE5DFF9E705DADE9FEDAFBB2FE50" blockId="26.[151,1437,151,1005]" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DADE9FEDAFE3FFEB8" bold="true" box="[199,503,258,285]" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DADE9FEDAFE38FEB8" bold="true" box="[199,496,258,285]" italics="true" pageId="26" pageNumber="177">Metasomal macrosetation</emphasis>
.
</emphasis>
Segments I–IV each with two pairs of VSM macrosetae (2/2): pair VSM1 located in the anterior 1/3, and pair VSM3 located near the posterior margin of the segment, and two pairs of VL macrosetae (2/2): pair VL1 located near the anterior margin, and pair VL2 located in the posterior 2/3 of the segment. Segment V with two pairs of VSM macrosetae (2/2), two pairs of VL macrosetae (2/2), and a single pair of ML macrosetae; pair VSM1 and VL1 located near the anterior margin of the segment; VL2 located on the posterior 2/3, and pair ML1 located dorsolaterally near the posterior margin (behind DSM carinae); anal arch with two pairs of setae on the intercrestal area, one pair of VSM macrosetae (1/1) and one pair of VL macrosetae (1/1).
</paragraph>
<paragraph id="3B30FE5DFF9E705DADE9FE26FC7EFD68" blockId="26.[151,1437,151,1005]" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DADE9FE26FE29FDBD" bold="true" box="[199,481,510,537]" pageId="26" pageNumber="177">
<taxonomicName id="FC8F85DEFF9E705DADE9FE26FEDAFDBC" box="[199,274,510,537]" class="Hexanauplia" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="09FB224FFF9E705DADE9FE26FEDAFDBC" bold="true" box="[199,274,510,537]" italics="true" pageId="26" pageNumber="177">Telson</emphasis>
</taxonomicName>
(
<figureCitation id="A3B4E2D8FF9E705DAC0DFE26FE1AFDBD" box="[291,466,510,536]" captionStart="FIGURE 15" captionStartId="24.[151,250,1090,1114]" captionTargetBox="[151,1436,345,1067]" captionTargetId="figure-116@24.[151,1436,345,1067]" captionTargetPageId="24" captionText="FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673376" httpUri="https://zenodo.org/record/6673376/files/figure.png" pageId="26" pageNumber="177">Figure 15C, D</figureCitation>
).
</emphasis>
Covered with abundant microsetae, white colored under UV light (
<figureCitation id="A3B4E2D8FF9E705DA9CBFE27FA58FDBC" box="[1253,1424,510,537]" captionStart="FIGURE 15" captionStartId="24.[151,250,1090,1114]" captionTargetBox="[151,1436,345,1067]" captionTargetId="figure-116@24.[151,1436,345,1067]" captionTargetPageId="24" captionText="FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673376" httpUri="https://zenodo.org/record/6673376/files/figure.png" pageId="26" pageNumber="177">Figure 15C, D</figureCitation>
). Vesicle spherical, not elongated (L/H= 1.21 (male)/ L/W= 1.31 (female)), with dorsal surface smooth and a lateral longitudinal furrow on each side; with VM, paired VSM, VL, and DL carinae, vestigial and slightly marked. Subaculear tubercle vestigial pyramidal, with a rounded apex directed to the middle portion of the aculeus (
<figureCitation id="A3B4E2D8FF9E705DA83BFDB3FF66FD0C" captionStart="FIGURE 15" captionStartId="24.[151,250,1090,1114]" captionTargetBox="[151,1436,345,1067]" captionTargetId="figure-116@24.[151,1436,345,1067]" captionTargetPageId="24" captionText="FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673376" httpUri="https://zenodo.org/record/6673376/files/figure.png" pageId="26" pageNumber="177">Figure 15C, D</figureCitation>
); subaculear tubercle with a ventral pair of small rounded granules, pointing to the basal portion of the aculeus; aculeus strongly curved, shorter than vesicle and with a ventral groove.
</paragraph>
<paragraph id="3B30FE5DFF9E705DADE9FD0EFDB6FC48" blockId="26.[151,1437,151,1005]" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DADE9FD0EFE85FD55" bold="true" box="[199,333,726,752]" pageId="26" pageNumber="177">Variability.</emphasis>
<emphasis id="09FB224FFF9E705DAC78FD0FFDCBFD54" box="[342,515,727,753]" italics="true" pageId="26" pageNumber="177">Morphometrics</emphasis>
. 
<emphasis id="09FB224FFF9E705DAF3DFD0FFD53FD54" box="[531,667,727,753]" italics="true" pageId="26" pageNumber="177">Total length</emphasis>
(
<emphasis id="09FB224FFF9E705DAF83FD0FFCACFD54" box="[685,868,727,753]" italics="true" pageId="26" pageNumber="177">including telson</emphasis>
): 53.38–61.55 (female) (n=5, mean= 57.28, standard deviation (SD)= 3.41) and 55.03 (male) (n=1). 
<emphasis id="09FB224FFF9E705DAFCAFD23FC53FCB1" box="[740,923,763,789]" italics="true" pageId="26" pageNumber="177">Chela L/W ratio</emphasis>
: 4.36–4.71 (female) (n= 4; mean= 4.21; SD= 0.59) and 3.18 (male) (n=1). 
<emphasis id="09FB224FFF9E705DACC8FCC7FC87FC9D" box="[486,847,799,825]" italics="true" pageId="26" pageNumber="177">Metasomal segment I L/W ratio</emphasis>
: 0.98–1.11 (female) (n= 5; mean= 1.05; SD= 0.05) and 1.08 (male) (n= 1). 
<emphasis id="09FB224FFF9E705DAC82FC9BFCD5FCF9" box="[428,797,835,861]" italics="true" pageId="26" pageNumber="177">Metasomal segment V L/W ratio</emphasis>
: 1.09–1.17 (female) (n= 5; mean= 1.12; SD= 0.03) and 1.08 (male) (n= 1). 
<emphasis id="09FB224FFF9E705DAC59FCBFFD8EFC24" box="[375,582,871,897]" italics="true" pageId="26" pageNumber="177">
<taxonomicName id="FC8F85DEFF9E705DAC59FCBFFDDBFC24" box="[375,531,871,897]" class="Hexanauplia" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="vesicle">Telson vesicle</taxonomicName>
L/H
</emphasis>
: 1.50–1.68 (female) (n= 5; mean= 1.62; SD= 0.07) and 1.60 (male) (n= 1). 
<emphasis id="09FB224FFF9E705DADB9FC53FE2AFC01" box="[151,482,907,933]" italics="true" pageId="26" pageNumber="177">Counts. Pectinal tooth counts</emphasis>
: females 15–17 (n= 5, mode= 16) and male 17 (n=2, mode= 17). 
<emphasis id="09FB224FFF9E705DA9EEFC53FE07FC6D" italics="true" pageId="26" pageNumber="177">Number of movable finger oblique granule rows</emphasis>
: females 14–16 (n= 5, mode= 15) and male 16–17 (n= 2). The variation of the telotarsi ventrosubmedian setae counts is in 
<tableCitation id="760DCBE6FF9E705DAF0AFC0BFDB2FC48" box="[548,634,979,1005]" captionStart="TABLE 6" captionStartId="26.[151,238,1052,1076]" captionTargetPageId="26" captionText="TABLE 6. Variation in the number of setae of the ventrosubmedian setal rows on telotarsi I–IV of Tityus forcipula and Tityus moralensis sp. nov. Abbreviations: L, left leg; Pl, prolateral row; Rl, retrolateral row; R, right leg. Observation: all specimens, but 1402 and 1403 deposited at ICN, are deposited at MUSENUV. Bold numbers correspond to the museum codes." httpUri="http://table.plazi.org/id/6FF0AED5FF9E705DADB9FBC4FA45FBD9" pageId="26" pageNumber="177" tableUuid="6FF0AED5FF9E705DADB9FBC4FA45FBD9">Table 6</tableCitation>
.
</paragraph>
<caption id="6FF0AED5FF9E705DADB9FBC4FA45FBD9" ID-Table-UUID="6FF0AED5FF9E705DADB9FBC4FA45FBD9" httpUri="http://table.plazi.org/id/6FF0AED5FF9E705DADB9FBC4FA45FBD9" pageId="26" pageNumber="177" startId="26.[151,238,1052,1076]" targetBox="[167,1380,1164,2029]" targetIsTable="true" targetPageId="26">
<paragraph id="3B30FE5DFF9E705DADB9FBC4FA45FBD9" blockId="26.[151,1436,1052,1149]" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DADB9FBC4FECFFB91" bold="true" box="[151,263,1052,1076]" pageId="26" pageNumber="177">TABLE 6.</emphasis>
Variation in the number of setae of the ventrosubmedian setal rows on telotarsi I–IV of 
<taxonomicName id="FC8F85DEFF9E705DA9A5FBC5FAE5FB91" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[1163,1325,1052,1076]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF9E705DA9A5FBC5FAE5FB91" box="[1163,1325,1052,1076]" italics="true" pageId="26" pageNumber="177">Tityus forcipula</emphasis>
</taxonomicName>
and 
<taxonomicName id="FC8F85DEFF9E705DA870FBC5FEC2FBFD" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov.">
<emphasis id="09FB224FFF9E705DA870FBC5FEC2FBFD" italics="true" pageId="26" pageNumber="177">Tityus moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF9E705DAC3FFB99FEAEFBFC" bold="true" box="[273,358,1089,1113]" pageId="26" pageNumber="177">
<taxonomicNameLabel id="12C89F34FF9E705DAC3FFB99FEAEFBFC" box="[273,358,1089,1113]" pageId="26" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Abbreviations: L, left leg; Pl, prolateral row; Rl, retrolateral row; R, right leg. Observation: all specimens, but 1402 and 1403 deposited at ICN, are deposited at MUSENUV. Bold numbers correspond to the museum codes.
</paragraph>
</caption>
<paragraph id="3B30FE5DFF9E705DAD89FB54FAD5F848" pageId="26" pageNumber="177">
<table id="498F0CFDFF9E8FB8AD89FB54FAACF848" box="[167,1380,1164,2029]" gridcols="7" gridrows="12" pageId="26" pageNumber="177">
<tr id="85BFFC1FFF9E8FB8AD89FB54FAACFB01" box="[167,1380,1164,1188]" gridrow="0" pageId="26" pageNumber="177" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-6="1">
<th id="C66E9563FF9E8FB8AD89FB54FEDFFB01" box="[167,279,1164,1188]" gridcol="0" gridrow="0" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAD89FB54FEDFFB01" bold="true" box="[167,279,1164,1188]" pageId="26" pageNumber="177">Telotarsus</emphasis>
</th>
<th id="C66E9563FF9E8FB8AFD7FB54FC3FFB01" box="[761,1015,1164,1188]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="26" pageNumber="177">
<taxonomicName id="FC8F85DEFF9E705DAE14FB54FC28FB01" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[826,992,1164,1188]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF9E705DAE14FB54FC28FB01" bold="true" box="[826,992,1164,1188]" italics="true" pageId="26" pageNumber="177">Tityus forcipula</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="85BFFC1FFF9E8FB8AD89FB6FFAACFB74" box="[167,1380,1207,1233]" gridrow="1" pageId="26" pageNumber="177" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-6="1">
<td id="C66E9563FF9E8FB8ACA3FB6FFE2EFB74" box="[397,486,1207,1233]" gridcol="1" gridrow="1" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DACA3FB60FE0EFB74" bold="true" box="[397,454,1208,1233]" pageId="26" pageNumber="177">438♂</emphasis>
</td>
<td id="C66E9563FF9E8FB8AFD7FB6FFC9AFB74" box="[761,850,1207,1233]" gridcol="3" gridrow="1" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAFD7FB6FFCD7FB6A" bold="true" box="[761,799,1207,1231]" pageId="26" pageNumber="177">436</emphasis>
♀
</td>
<td id="C66E9563FF9E8FB8A94BFB6FFB76FB74" box="[1125,1214,1207,1233]" gridcol="5" gridrow="1" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DA94BFB60FB63FB74" bold="true" box="[1125,1195,1208,1233]" pageId="26" pageNumber="177">2103♀</emphasis>
</td>
</tr>
<tr id="85BFFC1FFF9E8FB8AD89FB38FAACFB5D" box="[167,1380,1248,1272]" gridrow="2" pageId="26" pageNumber="177" rowspan-0="1">
<td id="C66E9563FF9E8FB8ACA3FB38FE2EFB5D" box="[397,486,1248,1272]" gridcol="1" gridrow="2" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DACA3FB38FE2EFB5D" bold="true" box="[397,486,1248,1272]" pageId="26" pageNumber="177">L(Pl/Rl)</emphasis>
</td>
<td id="C66E9563FF9E8FB8AF0BFB38FD43FB5D" box="[549,651,1248,1272]" gridcol="2" gridrow="2" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAF0BFB38FD43FB5D" bold="true" box="[549,651,1248,1272]" pageId="26" pageNumber="177">R (PI/RI)</emphasis>
</td>
<td id="C66E9563FF9E8FB8AFD7FB38FC9AFB5D" box="[761,850,1248,1272]" gridcol="3" gridrow="2" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAFD7FB38FC9AFB5D" bold="true" box="[761,850,1248,1272]" pageId="26" pageNumber="177">L(Pl/Rl)</emphasis>
</td>
<td id="C66E9563FF9E8FB8AEBFFB38FC3FFB5D" box="[913,1015,1248,1272]" gridcol="4" gridrow="2" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAEBFFB38FC3FFB5D" bold="true" box="[913,1015,1248,1272]" pageId="26" pageNumber="177">R (PI/RI)</emphasis>
</td>
<td id="C66E9563FF9E8FB8A94BFB38FB76FB5D" box="[1125,1214,1248,1272]" gridcol="5" gridrow="2" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DA94BFB38FB76FB5D" bold="true" box="[1125,1214,1248,1272]" pageId="26" pageNumber="177">L(Pl/Rl)</emphasis>
</td>
<td id="C66E9563FF9E8FB8A9D3FB38FAACFB5D" box="[1277,1380,1248,1272]" gridcol="6" gridrow="2" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DA9D3FB38FAACFB5D" bold="true" box="[1277,1380,1248,1272]" pageId="26" pageNumber="177">R (PI/RI)</emphasis>
</td>
</tr>
<tr id="85BFFC1FFF9E8FB8AD89FAD2FAACFA04" box="[167,1380,1290,1441]" gridrow="3" pageId="26" pageNumber="177">
<th id="C66E9563FF9E8FB8AD89FAD2FEDFFA04" box="[167,279,1290,1441]" gridcol="0" gridrow="3" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAD89FAD2FF0BFA05" bold="true" box="[167,197,1290,1440]" pageId="26" pageNumber="177">I II III IV</emphasis>
</th>
<td id="C66E9563FF9E8FB8ACA3FAD2FE2EFA04" box="[397,486,1290,1441]" gridcol="1" gridrow="3" pageId="26" pageNumber="177">6/5 6/6 6/7 -/-</td>
<td id="C66E9563FF9E8FB8AF0BFAD2FD43FA04" box="[549,651,1290,1441]" gridcol="2" gridrow="3" pageId="26" pageNumber="177">6/6 5/7 6/7 7/8</td>
<td id="C66E9563FF9E8FB8AFD7FAD2FC9AFA04" box="[761,850,1290,1441]" gridcol="3" gridrow="3" pageId="26" pageNumber="177">6/6 6/6 7/7 6/7</td>
<td id="C66E9563FF9E8FB8AEBFFAD2FC3FFA04" box="[913,1015,1290,1441]" gridcol="4" gridrow="3" pageId="26" pageNumber="177">6/6 7/7 6/7 7/7</td>
<td id="C66E9563FF9E8FB8A94BFAD2FB76FA04" box="[1125,1214,1290,1441]" gridcol="5" gridrow="3" pageId="26" pageNumber="177">7/7 5/6 7/7 6/7</td>
<td id="C66E9563FF9E8FB8A9D3FAD2FAACFA04" box="[1277,1380,1290,1441]" gridcol="6" gridrow="3" pageId="26" pageNumber="177">6/6 7/7 6/7 7/7</td>
</tr>
<tr id="85BFFC1FFF9E8FB8AD89FA6AFAACFA6F" box="[167,1380,1458,1482]" gridrow="4" pageId="26" pageNumber="177" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-6="1">
<th id="C66E9563FF9E8FB8AD89FA6AFEDFFA6F" box="[167,279,1458,1482]" gridcol="0" gridrow="4" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAD89FA6AFEDFFA6F" bold="true" box="[167,279,1458,1482]" pageId="26" pageNumber="177">Telotarsus</emphasis>
</th>
<td id="C66E9563FF9E8FB8AFD7FA6AFC3FFA6F" box="[761,1015,1458,1482]" colspan="2" colspanRight="1" gridcol="3" gridrow="4" pageId="26" pageNumber="177">
<taxonomicName id="FC8F85DEFF9E705DAE1FFA6AFC21FA6F" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[817,1001,1458,1482]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="moralensis">
<emphasis id="09FB224FFF9E705DAE1FFA6AFC21FA6F" bold="true" box="[817,1001,1458,1482]" italics="true" pageId="26" pageNumber="177">Tityus moralensis</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="85BFFC1FFF9E8FB8AD89FA05FAACFA52" box="[167,1380,1501,1527]" gridrow="5" pageId="26" pageNumber="177" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-6="1">
<td id="C66E9563FF9E8FB8ACA3FA05FE2EFA52" box="[397,486,1501,1527]" gridcol="1" gridrow="5" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DACA3FA06FE1AFA52" bold="true" box="[397,466,1502,1527]" pageId="26" pageNumber="177">2104♂</emphasis>
</td>
<td id="C66E9563FF9E8FB8AFD7FA05FC9AFA52" box="[761,850,1501,1527]" gridcol="3" gridrow="5" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAFD7FA05FCE4FA50" bold="true" box="[761,812,1501,1525]" pageId="26" pageNumber="177">1402</emphasis>
♀
</td>
<td id="C66E9563FF9E8FB8A94BFA05FB76FA52" box="[1125,1214,1501,1527]" gridcol="5" gridrow="5" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DA94BFA06FB63FA52" bold="true" box="[1125,1195,1502,1527]" pageId="26" pageNumber="177">1403♀</emphasis>
</td>
</tr>
<tr id="85BFFC1FFF9E8FB8AD89F9DEFAACF9BB" box="[167,1380,1542,1566]" gridrow="6" pageId="26" pageNumber="177" rowspan-0="1">
<td id="C66E9563FF9E8FB8ACA3F9DEFE2EF9BB" box="[397,486,1542,1566]" gridcol="1" gridrow="6" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DACA3F9DEFE2EF9BB" bold="true" box="[397,486,1542,1566]" pageId="26" pageNumber="177">L(Pl/Rl)</emphasis>
</td>
<td id="C66E9563FF9E8FB8AF0BF9DEFD43F9BB" box="[549,651,1542,1566]" gridcol="2" gridrow="6" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAF0BF9DEFD43F9BB" bold="true" box="[549,651,1542,1566]" pageId="26" pageNumber="177">R (PI/RI)</emphasis>
</td>
<td id="C66E9563FF9E8FB8AFD7F9DEFC9AF9BB" box="[761,850,1542,1566]" gridcol="3" gridrow="6" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAFD7F9DEFC9AF9BB" bold="true" box="[761,850,1542,1566]" pageId="26" pageNumber="177">L(Pl/Rl)</emphasis>
</td>
<td id="C66E9563FF9E8FB8AEBFF9DEFC3FF9BB" box="[913,1015,1542,1566]" gridcol="4" gridrow="6" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAEBFF9DEFC3FF9BB" bold="true" box="[913,1015,1542,1566]" pageId="26" pageNumber="177">R (PI/RI)</emphasis>
</td>
<td id="C66E9563FF9E8FB8A94BF9DEFB76F9BB" box="[1125,1214,1542,1566]" gridcol="5" gridrow="6" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DA94BF9DEFB76F9BB" bold="true" box="[1125,1214,1542,1566]" pageId="26" pageNumber="177">L(Pl/Rl)</emphasis>
</td>
<td id="C66E9563FF9E8FB8A9D3F9DEFAACF9BB" box="[1277,1380,1542,1566]" gridcol="6" gridrow="6" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DA9D3F9DEFAACF9BB" bold="true" box="[1277,1380,1542,1566]" pageId="26" pageNumber="177">R (PI/RI)</emphasis>
</td>
</tr>
<tr id="85BFFC1FFF9E8FB8AD89F9E8FAACF962" box="[167,1380,1584,1735]" gridrow="7" pageId="26" pageNumber="177">
<th id="C66E9563FF9E8FB8AD89F9E8FEDFF962" box="[167,279,1584,1735]" gridcol="0" gridrow="7" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAD89F9E8FF0BF963" bold="true" box="[167,197,1584,1734]" pageId="26" pageNumber="177">I II III IV</emphasis>
</th>
<td id="C66E9563FF9E8FB8ACA3F9E8FE2EF962" box="[397,486,1584,1735]" gridcol="1" gridrow="7" pageId="26" pageNumber="177">8/8 7/8 8/8 8/11</td>
<td id="C66E9563FF9E8FB8AF0BF9E8FD43F962" box="[549,651,1584,1735]" gridcol="2" gridrow="7" pageId="26" pageNumber="177">8/8 8/8 8/9 8/11</td>
<td id="C66E9563FF9E8FB8AFD7F9E8FC9AF962" box="[761,850,1584,1735]" gridcol="3" gridrow="7" pageId="26" pageNumber="177">7/8 6/7 7/8 8/9</td>
<td id="C66E9563FF9E8FB8AEBFF9E8FC3FF962" box="[913,1015,1584,1735]" gridcol="4" gridrow="7" pageId="26" pageNumber="177">7/7 6/8 -/- 7/7</td>
<td id="C66E9563FF9E8FB8A94BF9E8FB76F962" box="[1125,1214,1584,1735]" gridcol="5" gridrow="7" pageId="26" pageNumber="177">9/8 8/7 9/9 10/10</td>
<td id="C66E9563FF9E8FB8A9D3F9E8FAACF962" box="[1277,1380,1584,1735]" gridcol="6" gridrow="7" pageId="26" pageNumber="177">8/8 9/8 8/7 8/10</td>
</tr>
<tr id="85BFFC1FFF9E8FB8AD89F900FAACF955" box="[167,1380,1752,1776]" gridrow="8" pageId="26" pageNumber="177" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-6="1">
<th id="C66E9563FF9E8FB8AD89F900FEDFF955" box="[167,279,1752,1776]" gridcol="0" gridrow="8" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAD89F900FEDFF955" bold="true" box="[167,279,1752,1776]" pageId="26" pageNumber="177">Telotarsus</emphasis>
</th>
<td id="C66E9563FF9E8FB8AFD7F900FC3FF955" box="[761,1015,1752,1776]" colspan="2" colspanRight="1" gridcol="3" gridrow="8" pageId="26" pageNumber="177">
<taxonomicName id="FC8F85DEFF9E705DAE1FF900FC21F955" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[817,1001,1752,1776]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="moralensis">
<emphasis id="09FB224FFF9E705DAE1FF900FC21F955" bold="true" box="[817,1001,1752,1776]" italics="true" pageId="26" pageNumber="177">Tityus moralensis</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="85BFFC1FFF9E8FB8AD89F8DBFAACF8B8" box="[167,1380,1795,1821]" gridrow="9" pageId="26" pageNumber="177" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-6="1">
<td id="C66E9563FF9E8FB8ACA3F8DBFE2EF8B8" box="[397,486,1795,1821]" gridcol="1" gridrow="9" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DACA3F8DCFE1AF8B8" bold="true" box="[397,466,1796,1821]" pageId="26" pageNumber="177">2108♀</emphasis>
</td>
<td id="C66E9563FF9E8FB8AFD7F8DBFC9AF8B8" box="[761,850,1795,1821]" gridcol="3" gridrow="9" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAFD7F8DBFCE4F8BE" bold="true" box="[761,812,1795,1819]" pageId="26" pageNumber="177">2109</emphasis>
♀
</td>
<td id="C66E9563FF9E8FB8A94BF8DBFB76F8B8" box="[1125,1214,1795,1821]" gridcol="5" gridrow="9" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DA94BF8DBFB5DF8BE" bold="true" box="[1125,1173,1795,1819]" pageId="26" pageNumber="177">2111</emphasis>
♀
</td>
</tr>
<tr id="85BFFC1FFF9E8FB8AD89F8F4FAACF8E0" box="[167,1380,1836,1861]" gridrow="10" pageId="26" pageNumber="177" rowspan-0="1">
<td id="C66E9563FF9E8FB8ACA3F8F4FE2EF8E0" box="[397,486,1836,1861]" gridcol="1" gridrow="10" pageId="26" pageNumber="177">L(Pl/Rl)</td>
<td id="C66E9563FF9E8FB8AF0BF8F4FD43F8E0" box="[549,651,1836,1861]" gridcol="2" gridrow="10" pageId="26" pageNumber="177">R (PI/RI)</td>
<td id="C66E9563FF9E8FB8AFD7F8F4FC9AF8E0" box="[761,850,1836,1861]" gridcol="3" gridrow="10" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAFD7F8F4FC9AF8E1" bold="true" box="[761,850,1836,1860]" pageId="26" pageNumber="177">L(Pl/Rl)</emphasis>
</td>
<td id="C66E9563FF9E8FB8AEBFF8F4FC3FF8E0" box="[913,1015,1836,1861]" gridcol="4" gridrow="10" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAEBFF8F4FC3FF8E1" bold="true" box="[913,1015,1836,1860]" pageId="26" pageNumber="177">R (PI/RI)</emphasis>
</td>
<td id="C66E9563FF9E8FB8A94BF8F4FB76F8E0" box="[1125,1214,1836,1861]" gridcol="5" gridrow="10" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DA94BF8F4FB76F8E1" bold="true" box="[1125,1214,1836,1860]" pageId="26" pageNumber="177">L(Pl/Rl)</emphasis>
</td>
<td id="C66E9563FF9E8FB8A9D3F8F4FAACF8E0" box="[1277,1380,1836,1861]" gridcol="6" gridrow="10" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DA9D3F8F4FAACF8E1" bold="true" box="[1277,1380,1836,1860]" pageId="26" pageNumber="177">R (PI/RI)</emphasis>
</td>
</tr>
<tr id="85BFFC1FFF9E8FB8AD89F88EFAACF848" box="[167,1380,1878,2029]" gridrow="11" pageId="26" pageNumber="177">
<th id="C66E9563FF9E8FB8AD89F88EFEDFF848" box="[167,279,1878,2029]" gridcol="0" gridrow="11" pageId="26" pageNumber="177">
<emphasis id="09FB224FFF9E705DAD89F88EFF0BF849" bold="true" box="[167,197,1878,2028]" pageId="26" pageNumber="177">I II III IV</emphasis>
</th>
<td id="C66E9563FF9E8FB8ACA3F88EFE2EF848" box="[397,486,1878,2029]" gridcol="1" gridrow="11" pageId="26" pageNumber="177">8/8 7/7 8/8 9/9</td>
<td id="C66E9563FF9E8FB8AF0BF88EFD43F848" box="[549,651,1878,2029]" gridcol="2" gridrow="11" pageId="26" pageNumber="177">7/7 6/7 7/7 9/9</td>
<td id="C66E9563FF9E8FB8AFD7F88EFC9AF848" box="[761,850,1878,2029]" gridcol="3" gridrow="11" pageId="26" pageNumber="177">12/12 10/9 8/7 7/10</td>
<td id="C66E9563FF9E8FB8AEBFF88EFC3FF848" box="[913,1015,1878,2029]" gridcol="4" gridrow="11" pageId="26" pageNumber="177">-/- 11/8 7/7 10/9</td>
<td id="C66E9563FF9E8FB8A94BF88EFB76F848" box="[1125,1214,1878,2029]" gridcol="5" gridrow="11" pageId="26" pageNumber="177">9/9 9/10 6/8 8/12</td>
<td id="C66E9563FF9E8FB8A9D3F88EFAACF848" box="[1277,1380,1878,2029]" gridcol="6" gridrow="11" pageId="26" pageNumber="177">10/7 8/8 10/9 8/9</td>
</tr>
</table>
</paragraph>
<paragraph id="3B30FE5DFF9F705CADD9FF4FFCF2FEE4" blockId="27.[151,1436,150,322]" pageId="27" pageNumber="178">
<emphasis id="09FB224FFF9F705CADD9FF4FFE7DFF15" bold="true" box="[247,437,150,177]" pageId="27" pageNumber="178">Natural history.</emphasis>
<materialsCitation id="8BE7F400FF9F705CAC92FF4FFCFEFEE4" ID-GBIF-Occurrence="3816447311" collectorName="El Cerrito &amp; El Moral Sidewalk" country="Colombia" location="Valle del Cauca department" pageId="27" pageNumber="178" specimenCount="2" stateProvince="Valle del Cauca">
The 
<typeStatus id="E43440FFFF9F705CACC1FF4FFDD7FF14" box="[495,543,151,177]" pageId="27" pageNumber="178">type</typeStatus>
specimens recorded herein were hand-captured at day in small patches of montane forests surrounded by deforested grassland. The specimens were collected under large fallen logs or under the loose bark of rotten logs. The new species was found in sympatry with the chactid 
<taxonomicName id="FC8F85DEFF9F705CA918FF07FAE5FF5C" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[1078,1325,223,249]" class="Arachnida" family="Chactidae" genus="Chactas" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="vanbenedenii">
<emphasis id="09FB224FFF9F705CA918FF07FAE5FF5C" box="[1078,1325,223,249]" italics="true" pageId="27" pageNumber="178">Chactas vanbenedenii</emphasis>
</taxonomicName>
. 
<emphasis id="09FB224FFF9F705CADE9FEDAFE24FEB9" bold="true" box="[199,492,258,284]" pageId="27" pageNumber="178">
Distribution (
<figureCitation id="A3B4E2D8FF9F705CAC46FEDAFE15FEB9" box="[360,477,258,284]" captionStart="FIGURE 16" captionStartId="27.[151,250,1428,1452]" captionTargetBox="[184,1412,353,1404]" captionTargetId="figure-95@27.[176,1412,345,1404]" captionTargetPageId="27" captionText="FIGURE 16. Distribution map of the Tityus forcipula species group. A. South America. B. Records area. C. Species records." figureDoi="http://doi.org/10.5281/zenodo.6673378" httpUri="https://zenodo.org/record/6673378/files/figure.png" pageId="27" pageNumber="178">Figure 16</figureCitation>
).
</emphasis>
Only known from its 
<typeStatus id="E43440FFFF9F705CAFCCFEDBFCDAFEB8" box="[738,786,259,285]" pageId="27" pageNumber="178">type</typeStatus>
locality in the Cordillera Central (
<collectorName id="967A9B8BFF9F705CA9BCFEDBFACAFEB8" box="[1170,1282,259,285]" pageId="27" pageNumber="178">El Cerrito</collectorName>
municipality, 
<collectorName id="967A9B8BFF9F705CADB9FEFFFEB8FEE7" box="[151,368,295,322]" pageId="27" pageNumber="178">El Moral Sidewalk</collectorName>
), 
<collectingRegion id="F94B30BFFF9F705CACACFEFFFD71FEE4" box="[386,697,295,321]" country="Colombia" name="Valle del Cauca" pageId="27" pageNumber="178">Valle del Cauca department</collectingRegion>
, 
<collectingCountry id="4398BECDFF9F705CAFEAFEFFFCFEFEE4" box="[708,822,295,321]" name="Colombia" pageId="27" pageNumber="178">Colombia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>