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<document id="82BF55C8EF31CC252F5BA66FC0C30CA9" ID-CLB-Dataset="78128" ID-DOI="10.11646/zootaxa.5155.2.1" ID-GBIF-Dataset="be7f6620-4608-4fa7-9764-785fd401901a" ID-ISSN="1175-5326" ID-Zenodo-Dep="6673332" ID-ZooBank="F23CDF54-115F-4861-8EDB-DAFDF7810586" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1655793259292" checkinUser="plazi" docAuthor="Pinto-Da-Rocha, Ricardo &amp; Cabra-García, Jimmy" docDate="2022" docId="B3264F4BFF8D7051ADB9FB73FC47F816" docLanguage="en" docName="zootaxa.5155.2.1.pdf" docOrigin="Zootaxa 5155 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tityus forcipula" docType="treatment" docVersion="9" lastPageNumber="173" masterDocId="4F1F3733FF847047AD2EFFD8FFC8FFA5" masterDocTitle="On the Tityus forcipula species group: redescription of Tityus forcipula (Scorpiones, Buthidae), description of a new Andean species, and notes on the taxonomy of the group" masterLastPageNumber="186" masterPageNumber="151" pageNumber="160" updateTime="1702001334737" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F7553B1B58C87129B8B782339DA0BE25">On the Tityus forcipula species group: redescription of Tityus forcipula (Scorpiones, Buthidae), description of a new Andean species, and notes on the taxonomy of the group</mods:title>
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<mods:namePart id="F0D0E7BC22603CE923545F8989CFF798">Pinto-Da-Rocha, Ricardo</mods:namePart>
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<mods:affiliation id="242EFE581FBAA51C81837C33DED46F2F">Instituto de Biociências - Universidade de São Paulo, Departamento de Zoologia. Rua do Matão, travessa 14, 321, 005508 - 900, São Paulo, SP, Brazil &amp; ricrocha @ usp. br; https: // orcid. org / 0000 - 0002 - 3959 - 2205</mods:affiliation>
<mods:nameIdentifier id="C70AE57355198EC921C567DD17DDE46A" type="email">ricrocha@usp.br</mods:nameIdentifier>
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<mods:namePart id="345615A95D905ADD060691B947437CF6">Cabra-García, Jimmy</mods:namePart>
<mods:nameIdentifier id="2C8B012429E1FEA3DC3297841185E887" type="ORCID">0000-0001-9263-4390</mods:nameIdentifier>
<mods:affiliation id="277B6C16E691171A7183CCE57D972320">Departamento de Biología, Universidad del Valle, Ciudad Universitaria Meléndez, Calle 13 # 100 - 00, 760032, Cali, Colombia. Corresponding author: jimmy. cabra @ correounivalle. edu. co; https: // orcid. org / 0000 - 0001 - 9263 - 4390</mods:affiliation>
<mods:nameIdentifier id="E7C2CEFC1C0A1623FFC113FC8C6DE0D6" type="email">jimmy.cabra@correounivalle.edu.co</mods:nameIdentifier>
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<treatment id="B3264F4BFF8D7051ADB9FB73FC47F816" ID-DOI="http://doi.org/10.5281/zenodo.6677778" ID-GBIF-Taxon="196095741" ID-Zenodo-Dep="6677778" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816" httpUri="http://treatment.plazi.org/id/B3264F4BFF8D7051ADB9FB73FC47F816" lastPageId="22" lastPageNumber="173" pageId="9" pageNumber="160">
<subSubSection id="7395ADD6FF8D704EADB9FB73FDD5FB61" box="[151,541,1194,1221]" pageId="9" pageNumber="160" type="nomenclature">
<paragraph id="3B30FE5DFF8D704EADB9FB73FDD5FB61" blockId="9.[151,1436,1194,1293]" box="[151,541,1194,1221]" pageId="9" pageNumber="160">
<heading id="60784931FF8D704EADB9FB73FDD5FB61" bold="true" box="[151,541,1194,1221]" fontSize="11" level="1" pageId="9" pageNumber="160" reason="1">
<taxonomicName id="FC8F85DEFF8D704EADB9FB73FDD5FB61" ID-CoL="7CDM5" authority="(Gervais, 1843)" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[151,541,1194,1221]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF8D704EADB9FB73FDD5FB61" bold="true" box="[151,541,1194,1221]" pageId="9" pageNumber="160">
<emphasis id="09FB224FFF8D704EADB9FB73FE9FFB60" bold="true" box="[151,343,1194,1221]" italics="true" pageId="9" pageNumber="160">Tityus forcipula</emphasis>
(
<bibRefCitation id="5F1E83ACFF8D704EAC49FB72FDDDFB61" author="Gervais, P. M." box="[359,533,1194,1220]" pageId="9" pageNumber="160" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
)
</emphasis>
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<subSubSection id="7395ADD6FF8D704EADB9FB17FE98FAA8" pageId="9" pageNumber="160" type="description">
<paragraph id="3B30FE5DFF8D704EADB9FB17FE98FAA8" blockId="9.[151,1436,1194,1293]" pageId="9" pageNumber="160">
<figureCitation id="A3B4E2D8FF8D704EADB9FB17FEC8FB4C" box="[151,256,1231,1257]" captionStart="FIGURE 1" captionStartId="4.[151,250,1859,1883]" captionTargetBox="[286,1301,662,1817]" captionTargetId="figure-141@4.[251,1336,630,1838]" captionTargetPageId="4" captionText="FIGURE 1. Phylogenetic relationships of Tityus species inferred from a combined dataset of 28S and COI sequences. The topology is recovered from the IQ-TREE analysis. Squares at nodes indicate the Shimodaira–Hasegawa approximate likelihood-ratio test (left), the ultrafast bootstrap (middle) and the parsimony bootstrap (right). All sequences obtained in this study are in bold." figureDoi="http://doi.org/10.5281/zenodo.6673334" httpUri="https://zenodo.org/record/6673334/files/figure.png" pageId="9" pageNumber="160">Figures 1</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EAC23FB17FE9DFB4C" box="[269,341,1231,1257]" captionStart="FIGURE 2" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[180,1404,509,1847]" captionTargetId="figure-177@7.[164,1424,489,1864]" captionTargetPageId="7" captionText="FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6673340" httpUri="https://zenodo.org/record/6673340/files/figure.png" pageId="9" pageNumber="160">2A–D</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EAC71FB17FE6CFB4C" box="[351,420,1231,1257]" captionStart="FIGURE 4" captionStartId="10.[151,250,1375,1399]" captionTargetBox="[371,1216,309,1352]" captionTargetId="figure-86@10.[371,1216,309,1352]" captionTargetPageId="10" captionText="FIGURE 4. Tityus (Atreus) Gervais, 1843, carapace sculpturing, dorsal aspect. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673348" httpUri="https://zenodo.org/record/6673348/files/figure.png" pageId="9" pageNumber="160">4A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EAC81FB16FE3EFB4C" box="[431,502,1230,1257]" captionStart="FIGURE 5" captionStartId="13.[151,250,1447,1471]" captionTargetBox="[151,1437,309,1424]" captionTargetId="figure-91@13.[151,1437,309,1424]" captionTargetPageId="13" captionText="FIGURE 5. Tityus (Atreus) Gervais, 1843, femur sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673350" httpUri="https://zenodo.org/record/6673350/files/figure.png" pageId="9" pageNumber="160">5A–D</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EAF2FFB17FD81FB4C" box="[513,585,1231,1257]" captionStart="FIGURE 6" captionStartId="14.[151,250,1863,1887]" captionTargetBox="[151,1436,561,1840]" captionTargetId="figure-228@14.[151,1436,561,1840]" captionTargetPageId="14" captionText="FIGURE 6. Tityus (Atreus) Gervais, 1843, patella sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673354" httpUri="https://zenodo.org/record/6673354/files/figure.png" pageId="9" pageNumber="160">6A–D</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EAF7AFB17FD51FB4C" box="[596,665,1231,1257]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[261,1326,181,1920]" captionTargetId="figure-19@15.[261,1326,181,1920]" captionTargetPageId="15" captionText="FIGURE 7. Tityus (Atreus) Gervais, 1843,female chela sculpturing.A–C. Tityus(Atreus) forcipula (Gervais,1843) (MUSENUV- Ar 436). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (ICN-1402). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673356" httpUri="https://zenodo.org/record/6673356/files/figure.png" pageId="9" pageNumber="160">7A–C</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EAF8AFB17FD21FB4C" box="[676,745,1231,1257]" captionStart="FIGURE 8" captionStartId="16.[151,250,1944,1968]" captionTargetBox="[364,1224,181,1920]" captionTargetId="figure-31@16.[364,1224,181,1920]" captionTargetPageId="16" captionText="FIGURE 8. Tityus (Atreus) Gervais, 1843, male chela sculpturing. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673358" httpUri="https://zenodo.org/record/6673358/files/figure.png" pageId="9" pageNumber="160">8A–C</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EAFDAFB17FCF1FB4C" box="[756,825,1231,1257]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="9" pageNumber="160">9A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EAE6BFB17FC5FFB4C" box="[837,919,1231,1257]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[221,1367,181,1920]" captionTargetId="figure-19@19.[221,1367,181,1920]" captionTargetPageId="19" captionText="FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673364" httpUri="https://zenodo.org/record/6673364/files/figure.png" pageId="9" pageNumber="160">10A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EAE8DFB17FC3DFB4C" box="[931,1013,1231,1257]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="9" pageNumber="160">11A–C</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EA92EFB17FB9BFB4C" box="[1024,1107,1231,1257]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="9" pageNumber="160">12A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EA970FB17FB79FB4C" box="[1118,1201,1231,1257]" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="9" pageNumber="160">13A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EA992FB17FAC7FB4C" box="[1212,1295,1231,1257]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="9" pageNumber="160">14A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EA834FB16FAA5FB4C" box="[1306,1389,1230,1257]" captionStart="FIGURE 15" captionStartId="24.[151,250,1090,1114]" captionTargetBox="[151,1436,345,1067]" captionTargetId="figure-116@24.[151,1436,345,1067]" captionTargetPageId="24" captionText="FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673376" httpUri="https://zenodo.org/record/6673376/files/figure.png" pageId="9" pageNumber="160">15A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF8D704EA856FB17FA5FFB4C" box="[1400,1431,1231,1257]" captionStart="FIGURE 16" captionStartId="27.[151,250,1428,1452]" captionTargetBox="[184,1412,353,1404]" captionTargetId="figure-95@27.[176,1412,345,1404]" captionTargetPageId="27" captionText="FIGURE 16. Distribution map of the Tityus forcipula species group. A. South America. B. Records area. C. Species records." figureDoi="http://doi.org/10.5281/zenodo.6673378" httpUri="https://zenodo.org/record/6673378/files/figure.png" pageId="9" pageNumber="160">16</figureCitation>
;
<tableCitation id="760DCBE6FF8D704EADB9FB2BFF3FFAA8" box="[151,247,1267,1293]" captionStart="TABLE 2" captionStartId="3.[151,239,151,175]" captionTargetPageId="3" captionText="TABLE 2. Taxon sampling and GenBank accession numbers. Entries in bold were generated in this study. Abbreviations: Not applicable (NA)." httpUri="http://table.plazi.org/id/6FF0AED5FF877044ADB9FF4FFEB2FF71" pageId="9" pageNumber="160" tableUuid="6FF0AED5FF877044ADB9FF4FFEB2FF71">Tables 2</tableCitation>
,
<tableCitation id="760DCBE6FF8D704EAC2DFB2BFEDBFAA8" box="[259,275,1267,1293]" captionStart="TABLE 4" captionStartId="5.[151,239,574,600]" captionTargetPageId="5" captionText="TABLE 4. Percent uncorrected pairwise distances between COI sequences of Tityus species. Species codes: 1: Tityus argentinus, 2: T. bahiensis, 3: T. brazilae, 4: T. cerroazul, 5: T. clathrathus, 6-7: T. curupi, 8: T. discrepans, 9: T. engelkei, 10: T. forcipula, 11: T. metuendus, 12: T. moralensis sp. nov., 13: T. nematochirus, 14: T. obscurus, 15: T. paraguayensis, 16: T. sastrei, 17: T. serrulatus, 18: T. soratensis, 19-20: T. uruguayensis. Abbreviations: GBA: GenBank accession number." httpUri="http://table.plazi.org/id/6FF0AED5FF817042ADB9FDE6F9CDFD05" pageId="9" pageNumber="160" tableUuid="6FF0AED5FF817042ADB9FDE6F9CDFD05">4</tableCitation>
,
<tableCitation id="760DCBE6FF8D704EAC31FB2AFEE7FAA8" box="[287,303,1266,1293]" captionStart="TABLE 5" captionStartId="11.[151,239,440,464]" captionTargetBox="[167,1416,516,1987]" captionTargetPageId="11" captionText="TABLE 5. Measurements (mm) of Tityus forcipula and Tityus moralensis sp. nov. Observation: all specimens, but 1402" httpUri="http://table.plazi.org/id/6FF0AED5FF8F704CADB9FE60FAC8FE51" pageId="9" pageNumber="160" tableUuid="6FF0AED5FF8F704CADB9FE60FAC8FE51">5</tableCitation>
,
<tableCitation id="760DCBE6FF8D704EAC15FB2BFE84FAA8" box="[315,332,1267,1293]" captionStart="TABLE 6" captionStartId="26.[151,238,1052,1076]" captionTargetPageId="26" captionText="TABLE 6. Variation in the number of setae of the ventrosubmedian setal rows on telotarsi I–IV of Tityus forcipula and Tityus moralensis sp. nov. Abbreviations: L, left leg; Pl, prolateral row; Rl, retrolateral row; R, right leg. Observation: all specimens, but 1402 and 1403 deposited at ICN, are deposited at MUSENUV. Bold numbers correspond to the museum codes." httpUri="http://table.plazi.org/id/6FF0AED5FF9E705DADB9FBC4FA45FBD9" pageId="9" pageNumber="160" tableUuid="6FF0AED5FF9E705DADB9FBC4FA45FBD9">6</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7395ADD6FF8D704EADB9FAE2FD4AF988" pageId="9" pageNumber="160" type="reference_group">
<paragraph id="3B30FE5DFF8D704EADB9FAE2FD4AF988" blockId="9.[151,1437,1338,2013]" pageId="9" pageNumber="160">
<emphasis id="09FB224FFF8D704EADB9FAE2FD43FAF1" bold="true" box="[151,651,1338,1365]" pageId="9" pageNumber="160">
References after
<bibRefCitation id="5F1E83ACFF8D704EAC4AFAE3FD4EFAF1" author="Fet, V. &amp; Lowe, G." box="[356,646,1338,1365]" pageId="9" pageNumber="160" pagination="54 - 286" refId="ref17140" refString="Fet, V. &amp; Lowe, G. (2000) Family Buthidae C. L. Koch, 1837. In: Catalog of the Scorpions of the World. The New York Entomological Society, New York, New York, pp. 54 - 286." type="book chapter" year="2000">Fet &amp; Lowe (2000: 245)</bibRefCitation>
.
</emphasis>
<taxonomicName id="FC8F85DEFF8D704EAFBAFAE3FC35FAF0" ID-CoL="7CDM5" authority="Gervais (1843)" authorityName="Gervais" authorityYear="1843" box="[660,1021,1339,1365]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF8D704EAFBAFAE3FC8DFAF0" box="[660,837,1339,1365]" italics="true" pageId="9" pageNumber="160">Tityus forcipula</emphasis>
<bibRefCitation id="5F1E83ACFF8D704EAE60FAE3FC35FAF0" author="Gervais, P. M." box="[846,1021,1339,1365]" pageId="9" pageNumber="160" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais (1843)</bibRefCitation>
</taxonomicName>
: 130;
<bibRefCitation id="5F1E83ACFF8D704EA96DFAE3FACEFAF0" author="Lourenco, W. R." box="[1091,1286,1339,1365]" pageId="9" pageNumber="160" pagination="1 - 6" refId="ref18737" refString="Lourenco, W. R. (1999) A new species of Tityus Koch, 1836 (Chelicerata, Scorpiones, Buthidae) from Department Cesar in Colombia. Revue Arachnologique, 13, 1 - 6." type="journal article" year="1999">Lourenço (1999)</bibRefCitation>
: 4;
<bibRefCitation id="5F1E83ACFF8D704EA81FFAE3FF2EFADC" author="Lourenco, W. R." pageId="9" pageNumber="160" pagination="449 - 461" refId="ref18776" refString="Lourenco, W. R. (2000) Synopsis of the colombian species of Tityus Koch (Chelicerata, Scorpiones, Buthidae), with descriptions of three new species. Journal of Natural History, 34, 449 - 461. https: // doi. org / 10.1080 / 002229300299561" type="journal article" year="2000">Lourenço (2000)</bibRefCitation>
: 454, 456, 458, 459, figs. 11, 13;
<bibRefCitation id="5F1E83ACFF8D704EAF58FA87FC70FADC" author="Saldarriaga, M. M. &amp; Otero, R." box="[630,952,1375,1401]" pageId="9" pageNumber="160" pagination="17 - 23" refId="ref20631" refString="Saldarriaga, M. M. &amp; Otero, R. (2000) Los escorpiones: aspectos ecologicos, biologicos y toxinologicos. MedUNAB, 3, 17 - 23." type="journal article" year="2000">Saldarriaga &amp; Otero (2000)</bibRefCitation>
: 18, table 1;
<bibRefCitation id="5F1E83ACFF8D704EA97AFA87FB3DFADC" author="Florez, D. E." box="[1108,1269,1375,1401]" pageId="9" pageNumber="160" pagination="25 - 30" refId="ref17190" refString="Florez, D. E. (2001) Escorpiones de la familia Buthidae (Chelicerata: Scorpiones) de Colombia. Biota Colombiana, 2, 25 - 30." type="journal article" year="2001">Flórez (2001)</bibRefCitation>
: 28;
<bibRefCitation id="5F1E83ACFF8D704EA81FFA87FF3CFA3B" author="Lourenco, W. R." pageId="9" pageNumber="160" pagination="127 - 141" refId="ref18854" refString="Lourenco, W. R. (2002 b) Nouvelles additions a la faune de scorpions neotropicaux (Arachnida). Revue suisse de Zoologie, 109, 127 - 141. https: // doi. org / 10.5962 / bhl. part. 79582" type="journal article" year="2002">Lourenço (2002b)</bibRefCitation>
: 133;
<bibRefCitation id="5F1E83ACFF8D704EAC16FA5BFE31FA38" author="Lourenco, W. R." box="[312,505,1410,1437]" pageId="9" pageNumber="160" pagination="221 - 227" refId="ref18901" refString="Lourenco, W. R. (2005) A new species of Tityus Koch, 1836 (Scorpiones, Buthidae) from Ecuador. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14, 221 - 227." type="journal article" year="2005">Lourenço (2005)</bibRefCitation>
: 226;
<bibRefCitation id="5F1E83ACFF8D704EAF13FA5BFD36FA3B" author="Lourenco, W. R." box="[573,766,1411,1438]" pageId="9" pageNumber="160" pagination="55 - 67" refId="ref18940" refString="Lourenco, W. R. (2006) Une nouvelle proposition de decoupage sous-generique du genre &quot; Tityus &quot; C. L. Koch, 1836 (Scorpiones, Buthidae). Boletin Sociedad Entomologica Aragonesa, 39, 55 - 67." type="journal article" year="2006">Lourenço (2006)</bibRefCitation>
: 57, 58, 61;
<bibRefCitation id="5F1E83ACFF8D704EAEA6FA5BFB38FA38" author="Botero-Trujillo, R. &amp; Fagua, G." box="[904,1264,1411,1437]" pageId="9" pageNumber="160" pagination="129 - 134" refId="ref16743" refString="Botero-Trujillo, R. &amp; Fagua, G. (2007) Additions to the knowledge of the geographical distribution of some Colombian scorpions (Buthidae: Ananteris, Rhopalurus, Tityus). Revista Iberica de Aracnologia, 14, 129 - 134." type="journal article" year="2007">Botero-Trujillo &amp; Fagua (2007)</bibRefCitation>
: 129, 133, figs. 11, 16, 17;
<bibRefCitation id="5F1E83ACFF8D704EAC3CFA7FFE1AFA64" author="Lourenco, W. R." box="[274,466,1447,1473]" pageId="9" pageNumber="160" pagination="375 - 385" refId="ref18984" refString="Lourenco, W. R. (2007) A new species of Tityus C. L. Koch, 1836 from Ecuador: the first element of the ' Tityus androcottoides ' subgroup for this country. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 176, 375 - 385." type="journal article" year="2007">Lourenço (2007)</bibRefCitation>
: 382;
<bibRefCitation id="5F1E83ACFF8D704EAF3AFA7FFC8EFA64" author="Lourenco, W. R. &amp; Leguin, E. - A." box="[532,838,1447,1473]" pageId="9" pageNumber="160" pagination="1 - 9" refId="ref19189" refString="Lourenco, W. R. &amp; Leguin, E. - A. (2008) The true identity of Scorpio (Atreus) obscurus Gervais, 1843 (Scorpiones, Buthidae). Euscorpius, 2008, 1 - 9. https: // doi. org / 10.18590 / euscorpius. 2008. vol 2008. iss 75.1" type="journal article" year="2008">Lourenço &amp; Leguin (2008)</bibRefCitation>
: 1, 3;
<bibRefCitation id="5F1E83ACFF8D704EAEA9FA7FFB4CFA64" author="Lourenco, W. R. &amp; Leguin, E. - A. &amp; Cloudsley-Thompson, J. L." box="[903,1156,1447,1473]" pageId="9" pageNumber="160" pagination="423 - 427" refId="ref19482" refString="Lourenco, W. R., Leguin, E. - A. &amp; Cloudsley-Thompson, J. L. (2009) The status of Tityus (Atreus) rufofuscus Pocock, 1897 (Scorpiones, Buthidae), an enigmatic scorpion described from Brazil. Boletin Sociedad Entomologica Aragonesa, 45, 423 - 427." type="journal article" year="2009">
Lourenço
<emphasis id="09FB224FFF8D704EAED6FA7FFBF8FA64" box="[1016,1072,1447,1473]" italics="true" pageId="9" pageNumber="160">et al.</emphasis>
(2009)
</bibRefCitation>
: 423;
<bibRefCitation id="5F1E83ACFF8D704EA9E8FA7FFAA9FA64" author="Florez, D. E." box="[1222,1377,1447,1473]" pageId="9" pageNumber="160" pagination="763 - 767" refId="ref17221" refString="Florez, D. E. (2011) Escorpiones en el Choco Biogeografico. In: Colombia Diversidad Biotica IV. El Choco biogeografico / Costa pacifica. Universidad Nacional de Colombia, Bogota D. C., Colombia, Bogota, pp. 763 - 767." type="book chapter" year="2011">Flórez (2011)</bibRefCitation>
: 764, 765, table 56;
<bibRefCitation id="5F1E83ACFF8D704EAC10FA13FD93FA40" author="Gonzalez-Sponga, M. A." box="[318,603,1483,1509]" pageId="9" pageNumber="160" pagination="65 - 114" refId="ref17561" refString="Gonzalez-Sponga, M. A. (2011) Escorpiones, caracteristicas, distribucion geografica y comentarios generales. In: Emergencia por animales ponzonosos en las Americas. Instituto Bioclon, SA de CV, Mexico, D. F., pp. 65 - 114." type="book chapter" year="2011">González-Sponga (2011)</bibRefCitation>
: 103;
<bibRefCitation id="5F1E83ACFF8D704EAF8DFA13FC1DFA40" author="Lourenco, W. R. &amp; Ythier, E." box="[675,981,1483,1509]" pageId="9" pageNumber="160" pagination="1 - 13" refId="ref19366" refString="Lourenco, W. R. &amp; Ythier, E. (2013) The remarkable scorpion diversity in the Ecuadorian Andes and description of a new species of Tityus C. L. Koch, 1836 (Scorpiones, Buthidae). ZooKeys, 307, 1 - 13. https: // doi. org / 10.3897 / zookeys. 307.5334" type="journal article" year="2013">Lourenço &amp; Ythier (2013)</bibRefCitation>
: 3, figs. 13, 18, table I;
<bibRefCitation id="5F1E83ACFF8D704EA9C2FA13FF2EF9AC" author="Brito, G. &amp; Borges, A." pageId="9" pageNumber="160" pagination="1 - 17" refId="ref16788" refString="Brito, G. &amp; Borges, A. (2015) A checklist of the scorpions of Ecuador (Arachnida: Scorpiones), with notes on the distribution and medical significance of some species. Journal of Venomous Animals and Toxins Including Tropical Diseases, 21, 1 - 17. https: // doi. org / 10.1186 / s 40409 - 015 - 0023 - x" type="journal article" year="2015">Brito &amp; Borges (2015)</bibRefCitation>
: 2, 7, 9, 14, figs. 1, 4G;
<bibRefCitation id="5F1E83ACFF8D704EACC1FA37FCD0F9AC" author="Lourenco, W. R. &amp; Ythier, E." box="[495,792,1519,1545]" pageId="9" pageNumber="160" pagination="18 - 34" refId="ref19429" refString="Lourenco, W. R. &amp; Ythier, E. (2017) Description of Tityus (Atreus) cisandinus sp. n. from Ecuadorian Amazonia, with comments on some related species (Scorpiones: Buthidae). Arachnida - Rivista Aracnologica Italiana, 15, 18 - 34." type="journal article" year="2017">Lourenço &amp; Ythier (2017)</bibRefCitation>
: 29;
<bibRefCitation id="5F1E83ACFF8D704EAE65FA37FC20F9AC" author="Ythier, E." box="[843,1000,1519,1545]" pageId="9" pageNumber="160" pagination="18 - 23" refId="ref21562" refString="Ythier, E. (2018) On the contribution of Wilson R. Lourenco to the knowledge of the scorpion fauna of Ecuador. Arachnida - Rivista Aracnologica Italiana, 17, 18 - 23." type="journal article" year="2018">Ythier (2018)</bibRefCitation>
: 20;
<bibRefCitation id="5F1E83ACFF8D704EA932FA37FB03F9AC" author="Francke, O. F." box="[1052,1227,1519,1545]" pageId="9" pageNumber="160" pagination="1 - 56" refId="ref17366" refString="Francke, O. F. (2019) Conspectus Genericus Scorpionorum 1758 - 1985 (Arachnida: Scorpiones) updated through 2018. Zootaxa, 4657 (1), 1 - 56. https: // doi. org / 10.11646 / zootaxa. 4657.1.1" type="journal article" year="2019">Francke (2019)</bibRefCitation>
: 24;
<bibRefCitation id="5F1E83ACFF8D704EA82EFA37FF2EF988" author="Miranda, R. J. &amp; Bermudez, S. &amp; Florez, D. E. &amp; Armas, L. F. de" pageId="9" pageNumber="160" pagination="197 - 204" refId="ref19680" refString="Miranda, R. J., Bermudez, S., Florez, D. E. &amp; Armas, L. F. de (2020) A new species of Tityus from Panama and Costa Rica previously identified as Tityus pachyurus Pocock, 1897 (Scorpiones: Buthidae). Revista Iberica de Aracnologia, 37, 197 - 204." type="journal article" year="2020">
<collectingRegion id="F94B30BFFF8D704EA82EFA37FA97F9AC" box="[1280,1375,1519,1545]" country="Venezuela" name="Miranda" pageId="9" pageNumber="160">Miranda</collectingRegion>
<emphasis id="09FB224FFF8D704EA84BFA37FA55F9AC" box="[1381,1437,1519,1545]" italics="true" pageId="9" pageNumber="160">et al.</emphasis>
(2020)
</bibRefCitation>
: 197, 203, fig. 27;
<bibRefCitation id="5F1E83ACFF8D704EAC97F9CBFD9CF988" author="Dupre, G." box="[441,596,1555,1581]" pageId="9" pageNumber="160" pagination="2 - 111" refId="ref16858" refString="Dupre, G. (2021) Checklist of scorpions' taxa 1758 - 2020. Arachnides, 100, 2 - 111." type="journal article" year="2021">Dupré (2021)</bibRefCitation>
: 52.
</paragraph>
</subSubSection>
<subSubSection id="7395ADD6FF8D704DADE9F9EFFD9DFEB8" lastPageId="10" lastPageNumber="161" pageId="9" pageNumber="160" type="materials_examined">
<paragraph id="3B30FE5DFF8D704EADE9F9EFFE51F9D0" blockId="9.[151,1437,1338,2013]" pageId="9" pageNumber="160">
<emphasis id="09FB224FFF8D704EADE9F9EFFEBEF9F5" bold="true" box="[199,374,1590,1617]" pageId="9" pageNumber="160">Type material.</emphasis>
<materialsCitation id="8BE7F400FF8D704EAC50F9EFFB9CF9F7" ID-GBIF-Occurrence="3816447302" box="[382,1108,1591,1618]" collectionCode="NHM" country="Colombia" location="Colombia" pageId="9" pageNumber="160" specimenCode="NHM 1846.20" specimenCount="1" typeStatus="holotype">
<typeStatus id="E43440FFFF8D704EAC50F9EFFE23F9F4" box="[382,491,1591,1617]" pageId="9" pageNumber="160" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="4398BECDFF8D704EACD9F9EFFD59F9F4" box="[503,657,1591,1617]" name="Colombia" pageId="9" pageNumber="160">COLOMBIA</collectingCountry>
: adult male,
<location id="3E50A886FF8D704EAE0CF9EFFC58F9F4" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF8D704EAE0CF9EFFC58F9F4" box="[802,912,1591,1617]" country="Colombia" name="Colombia" pageId="9" pageNumber="160">Colombia</location>
(
<specimenCode id="6B295626FF8D704EAE8EF9EFFB84F9F7" box="[928,1100,1591,1618]" collectionCode="NHM" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="9" pageNumber="160" type="Herbarium">NHM 1846.20</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="8BE7F400FF8D704EA94EF9EFFE5DF9D0" ID-GBIF-Occurrence="3816447312" collectionCode="MNHN" country="Colombia" pageId="9" pageNumber="160" specimenCount="1" typeStatus="paratype">
<typeStatus id="E43440FFFF8D704EA94EF9EFFB0EF9F4" box="[1120,1222,1591,1617]" pageId="9" pageNumber="160" type="paratype">Paratype</typeStatus>
: unknow sex, unkwon locality (
<collectionCode id="5D9E6698FF8D704EAC1FF983FE45F9D0" box="[305,397,1627,1653]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="160">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="3B30FE5DFF8D704DADE9F9A6FD9DFEB8" blockId="9.[151,1437,1338,2013]" lastBlockId="10.[151,1436,151,285]" lastPageId="10" lastPageNumber="161" pageId="9" pageNumber="160">
<emphasis id="09FB224FFF8D704EADE9F9A6FE7CF93D" bold="true" box="[199,436,1662,1688]" pageId="9" pageNumber="160">Examined material.</emphasis>
<materialsCitation id="8BE7F400FF8D704EAC92F9A7FE27F918" ID-GBIF-Occurrence="3816447321" collectingDate="1979-06" collectingDateMax="2021-02" collectingDateMin="1979-06" country="Colombia" county="Cali" elevation="1500" latitude="3.87975" location="Cali" longLatPrecision="1" longitude="-76.43875" municipality="Km" pageId="9" pageNumber="160" specimenCode="IBALCC-RPDR 00256" specimenCount="3" specimenCount-adult="3" stateProvince="Valle del Cauca">
<collectingCountry id="4398BECDFF8D704EAC92F9A7FD9EF93C" box="[444,598,1663,1689]" name="Colombia" pageId="9" pageNumber="160">COLOMBIA</collectingCountry>
:
<emphasis id="09FB224FFF8D704EAF4FF9A7FC53F93C" box="[609,923,1663,1689]" italics="true" pageId="9" pageNumber="160">
<collectingRegion id="F94B30BFFF8D704EAF4FF9A7FC53F93C" box="[609,923,1663,1689]" country="Colombia" name="Valle del Cauca" pageId="9" pageNumber="160">Valle del Cauca department</collectingRegion>
</emphasis>
: adult male,
<collectingCounty id="D25186D1FF8D704EA901F9A7FBA8F93C" box="[1071,1120,1663,1689]" pageId="9" pageNumber="160">Cali</collectingCounty>
, 18
<collectingMunicipality id="DB546427FF8D704EA9BEF9A7FB72F93C" box="[1168,1210,1663,1689]" pageId="9" pageNumber="160">Km</collectingMunicipality>
highway to the sea,
<date id="4F31D89DFF8D704EADB9F97AFF22F918" box="[151,234,1698,1725]" pageId="9" pageNumber="160" value="1985-04">
<collectingDate id="5F752175FF8D704EADB9F97AFF22F918" box="[151,234,1698,1725]" pageId="9" pageNumber="160" value="1985-04">iv.1985</collectingDate>
</date>
(MUSENUV-Ar 438);
</materialsCitation>
<materialsCitation id="8BE7F400FF8D704EACD9F97BFF1AF947" ID-GBIF-Occurrence="3816447322" collectingDate="1979-06" collectingDateMax="2021-02" collectingDateMin="1979-06" country="Colombia" county="Cali" elevation="1500" latitude="3.87975" location="Cali" longLatPrecision="1" longitude="-76.43875" municipality="Km" pageId="9" pageNumber="160" specimenCode="IBALCC-RPDR 00256" specimenCount="3" specimenCount-adult="3" stateProvince="Valle del Cauca">
adult female,
<location id="3E50A886FF8D704EAFBDF97BFD0CF918" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF8D704EAFBDF97BFD0CF918" box="[659,708,1699,1725]" country="Colombia" county="Cali" latitude="3.87975" longLatPrecision="1" longitude="-76.43875" municipality="Km" name="Cali" pageId="9" pageNumber="160" stateProvince="Valle del Cauca">Cali</location>
,
<location id="3E50A886FF8D704EAFFFF97BFC13F918" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF8D704EAFFFF97BFC13F918" box="[721,987,1699,1725]" country="Colombia" county="Cali" latitude="3.87975" longLatPrecision="1" longitude="-76.43875" municipality="Km" name="Villa Carmelo Pathway" pageId="9" pageNumber="160" stateProvince="Valle del Cauca">Villa Carmelo Pathway</location>
,
<quantity id="FC7753B8FF8D704EAEC9F97AFB97F918" box="[999,1119,1698,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="160" unit="m" value="1500.0">
<elevation id="B0A2196EFF8D704EAEC9F97AFB97F918" box="[999,1119,1698,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="160" unit="m" value="1500.0">1500 masl</elevation>
</quantity>
,
<date id="4F31D89DFF8D704EA942F97BFB14F918" box="[1132,1244,1699,1725]" pageId="9" pageNumber="160" value="1982-10-13">
<collectingDate id="5F752175FF8D704EA942F97BFB14F918" box="[1132,1244,1699,1725]" pageId="9" pageNumber="160" value="1982-10-13">13.x.1982</collectingDate>
</date>
(MUSENUV-Ar 436);
</materialsCitation>
<materialsCitation id="8BE7F400FF8D704EADF7F91FFC0CF944" ID-GBIF-Occurrence="3816447318" box="[217,964,1735,1761]" collectingDate="1979-06" collectingDateMax="2021-02" collectingDateMin="1979-06" country="Colombia" county="Cali" elevation="1500" latitude="3.87975" location="Cali" longLatPrecision="1" longitude="-76.43875" municipality="Km" pageId="9" pageNumber="160" specimenCode="IBALCC-RPDR 00256" specimenCount="3" specimenCount-adult="3" stateProvince="Valle del Cauca">
adult female,
<location id="3E50A886FF8D704EAC5CF91FFE6BF944" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF8D704EAC5CF91FFE6BF944" box="[370,419,1735,1761]" country="Colombia" county="Cali" latitude="3.87975" longLatPrecision="1" longitude="-76.43875" municipality="Km" name="Cali" pageId="9" pageNumber="160" stateProvince="Valle del Cauca">Cali</location>
,
<location id="3E50A886FF8D704EAC80F91FFDACF944" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF8D704EAC80F91FFDACF944" box="[430,612,1735,1761]" country="Colombia" county="Cali" latitude="3.87975" longLatPrecision="1" longitude="-76.43875" municipality="Km" name="Felidia Pathway" pageId="9" pageNumber="160" stateProvince="Valle del Cauca">Felidia Pathway</location>
,
<date id="4F31D89DFF8D704EAF41F91FFD0CF944" box="[623,708,1735,1761]" pageId="9" pageNumber="160" value="1979-06">
<collectingDate id="5F752175FF8D704EAF41F91FFD0CF944" box="[623,708,1735,1761]" pageId="9" pageNumber="160" value="1979-06">vi.1979</collectingDate>
</date>
(MUSENUV-Ar 434);
</materialsCitation>
<materialsCitation id="8BE7F400FF8D704EAEE5F91FFD7FF8A0" ID-GBIF-Occurrence="3816447306" collectingDate="1979-06" collectingDateMax="2021-02" collectingDateMin="1979-06" collectorName="G. Salcedo" country="Colombia" county="Cali" elevation="1500" latitude="3.87975" location="Cali" longLatPrecision="1" longitude="-76.43875" municipality="Km" pageId="9" pageNumber="160" specimenCode="IBALCC-RPDR 00256" specimenCount="3" specimenCount-adult="3" stateProvince="Valle del Cauca">
adult female,
<location id="3E50A886FF8D704EA94DF91FFB5CF944" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF8D704EA94DF91FFB5CF944" box="[1123,1172,1735,1761]" country="Colombia" county="Cali" latitude="3.87975" longLatPrecision="1" longitude="-76.43875" municipality="Km" name="Cali" pageId="9" pageNumber="160" stateProvince="Valle del Cauca">Cali</location>
,
<collectorName id="967A9B8BFF8D704EA98EF91FFA92F944" box="[1184,1370,1735,1761]" pageId="9" pageNumber="160">
<location id="3E50A886FF8D704EA98EF91FFA92F944" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF8D704EA98EF91FFA92F944" box="[1184,1370,1735,1761]" country="Colombia" county="Cali" latitude="3.87975" longLatPrecision="1" longitude="-76.43875" municipality="Km" name="El Pato Pathway" pageId="9" pageNumber="160" stateProvince="Valle del Cauca">El Pato Pathway</location>
</collectorName>
,
<quantity id="FC7753B8FF8D704EA84AF91FFF07F8A0" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="160" unit="m" value="2000.0">
<elevation id="B0A2196EFF8D704EA84AF91FFF07F8A0" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="160" unit="m" value="2000.0">2000 masl</elevation>
</quantity>
,
<date id="4F31D89DFF8D704EADF7F933FEFAF8A0" box="[217,306,1771,1797]" pageId="9" pageNumber="160" value="1988-03">
<collectingDate id="5F752175FF8D704EADF7F933FEFAF8A0" box="[217,306,1771,1797]" pageId="9" pageNumber="160" value="1988-03">iii.1988</collectingDate>
</date>
,
<collectorName id="967A9B8BFF8D704EAC13F933FE7FF8A0" box="[317,439,1771,1797]" pageId="9" pageNumber="160">G. Salcedo</collectorName>
(MUSENUV-Ar 418);
</materialsCitation>
<materialsCitation id="8BE7F400FF8D704EAF93F933FA95F88F" ID-GBIF-Occurrence="3816447323" collectingDate="1979-06" collectingDateMax="2021-02" collectingDateMin="1979-06" collectorName="J. Cabra-Garcia" country="Colombia" county="Cali" elevation="1500" latitude="3.87975" location="Cali" longLatPrecision="1" longitude="-76.43875" municipality="Km" pageId="9" pageNumber="160" specimenCode="IBALCC-RPDR 00256" specimenCount="3" specimenCount-adult="3" stateProvince="Valle del Cauca">
adult female,
<location id="3E50A886FF8D704EAE7AF933FC4DF8A0" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF8D704EAE7AF933FC4DF8A0" box="[852,901,1771,1797]" country="Colombia" county="Cali" latitude="3.87975" longLatPrecision="1" longitude="-76.43875" municipality="Km" name="Cali" pageId="9" pageNumber="160" stateProvince="Valle del Cauca">Cali</location>
,
<collectorName id="967A9B8BFF8D704EAEBEF933FBBBF8A0" box="[912,1139,1771,1797]" pageId="9" pageNumber="160">
<location id="3E50A886FF8D704EAEBEF933FBBBF8A0" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF8D704EAEBEF933FBBBF8A0" box="[912,1139,1771,1797]" country="Colombia" county="Cali" latitude="3.87975" longLatPrecision="1" longitude="-76.43875" municipality="Km" name="El Saladito Pathway" pageId="9" pageNumber="160" stateProvince="Valle del Cauca">El Saladito Pathway</location>
</collectorName>
,
<location id="3E50A886FF8D704EA953F933FA9FF8A0" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF8D704EA953F933FA9FF8A0" box="[1149,1367,1771,1797]" country="Colombia" county="Cali" latitude="3.87975" longLatPrecision="1" longitude="-76.43875" municipality="Km" name="San Antonio Forest" pageId="9" pageNumber="160" stateProvince="Valle del Cauca">San Antonio Forest</location>
, highway to the sea,
<geoCoordinate id="5EBB989AFF8D704EAC6DF8D7FE2AF88C" box="[323,482,1807,1833]" degrees="3" direction="north" minutes="29" orientation="latitude" pageId="9" pageNumber="160" precision="1" seconds="34.22" value="3.4928389">3º29’34.22’’N</geoCoordinate>
<geoCoordinate id="5EBB989AFF8D704EACC7F8D7FD55F88C" box="[489,669,1807,1833]" degrees="76" direction="west" minutes="36" orientation="longitude" pageId="9" pageNumber="160" precision="1" seconds="49.71" value="-76.61381">76º36’49.71’’W</geoCoordinate>
,
<quantity id="FC7753B8FF8D704EAF86F8D7FCD7F88C" box="[680,799,1807,1833]" metricMagnitude="3" metricUnit="m" metricValue="1.813" pageId="9" pageNumber="160" unit="m" value="1813.0">
<elevation id="B0A2196EFF8D704EAF86F8D7FCD7F88C" box="[680,799,1807,1833]" metricMagnitude="3" metricUnit="m" metricValue="1.813" pageId="9" pageNumber="160" unit="m" value="1813.0">1813 masl</elevation>
</quantity>
,
<date id="4F31D89DFF8D704EAE04F8D7FC5EF88C" box="[810,918,1807,1833]" pageId="9" pageNumber="160" value="2019-01-21">
<collectingDate id="5F752175FF8D704EAE04F8D7FC5EF88C" box="[810,918,1807,1833]" pageId="9" pageNumber="160" value="2019-01-21">21.i.2019</collectingDate>
</date>
,
<collectorName id="967A9B8BFF8D704EAE8FF8D7FB98F88C" box="[929,1104,1807,1833]" pageId="9" pageNumber="160">J. Cabra-García</collectorName>
(MUSENUV-Ar 2116);
</materialsCitation>
<materialsCitation id="8BE7F400FF8D704EA84AF8D7FBC6F8E8" ID-GBIF-Occurrence="3816447315" collectingDate="1979-06" collectingDateMax="2021-02" collectingDateMin="1979-06" collectorName="L. Flórez" country="Colombia" county="Cali" elevation="1500" latitude="3.87975" location="Cali" longLatPrecision="1" longitude="-76.43875" municipality="Km" pageId="9" pageNumber="160" specimenCode="IBALCC-RPDR 00256" specimenCount="3" specimenCount-adult="3" stateProvince="Valle del Cauca">
adult female,
<collectorName id="967A9B8BFF8D704EADDFF8EBFEBBF8E8" box="[241,371,1843,1869]" pageId="9" pageNumber="160">La Cumbre</collectorName>
,
<collectorName id="967A9B8BFF8D704EAC50F8EBFDF9F8E8" box="[382,561,1843,1869]" pageId="9" pageNumber="160">Bitaco Pathway</collectorName>
,
<date id="4F31D89DFF8D704EAF15F8EBFD45F8E8" box="[571,653,1843,1869]" pageId="9" pageNumber="160" value="2021-02">
<collectingDate id="5F752175FF8D704EAF15F8EBFD45F8E8" box="[571,653,1843,1869]" pageId="9" pageNumber="160" value="2021-02">ii.2021</collectingDate>
</date>
,
<collectorName id="967A9B8BFF8D704EAFB6F8EBFD37F8E8" box="[664,767,1843,1869]" pageId="9" pageNumber="160">L. Flórez</collectorName>
(MUSENUV-Ar 2103);
</materialsCitation>
<materialsCitation id="8BE7F400FF8D704EA93BF8EBFB6AF8D4" ID-GBIF-Occurrence="3816447316" collectingDate="1979-06" collectingDateMax="2021-02" collectingDateMin="1979-06" collectorName="C. Sepulveda" country="Colombia" county="Cali" elevation="1500" latitude="3.87975" location="Cali" longLatPrecision="1" longitude="-76.43875" municipality="Km" pageId="9" pageNumber="160" specimenCode="IBALCC-RPDR 00256" specimenCount="3" specimenCount-adult="3" stateProvince="Valle del Cauca">
adult female,
<collectorName id="967A9B8BFF8D704EA981F8EBFAF8F8E8" box="[1199,1328,1843,1869]" pageId="9" pageNumber="160">La Cumbre</collectorName>
,
<collectorName id="967A9B8BFF8D704EA812F8EBFF31F8D4" pageId="9" pageNumber="160">Chicoral Pathway</collectorName>
, 28
<collectorName id="967A9B8BFF8D704EAC0AF88FFE86F8D4" box="[292,334,1879,1905]" pageId="9" pageNumber="160">Km</collectorName>
highway to the sea,
<quantity id="FC7753B8FF8D704EAF03F88EFD6BF8D4" box="[557,675,1878,1905]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="160" unit="m" value="1500.0">
<elevation id="B0A2196EFF8D704EAF03F88EFD6BF8D4" box="[557,675,1878,1905]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="160" unit="m" value="1500.0">1500 masl</elevation>
</quantity>
,
<date id="4F31D89DFF8D704EAF83F88FFCCDF8D4" box="[685,773,1879,1905]" pageId="9" pageNumber="160" value="1988-03">
<collectingDate id="5F752175FF8D704EAF83F88FFCCDF8D4" box="[685,773,1879,1905]" pageId="9" pageNumber="160" value="1988-03">iii.1988</collectingDate>
</date>
,
<collectorName id="967A9B8BFF8D704EAE3EF88FFC6CF8D4" box="[784,932,1879,1905]" pageId="9" pageNumber="160">C. Sepulveda</collectorName>
(MUSENUV-Ar 409);
</materialsCitation>
<materialsCitation id="8BE7F400FF8D704EA986F88FFCF5F830" ID-GBIF-Occurrence="3816447317" collectingDate="1979-06" collectingDateMax="2021-02" collectingDateMin="1979-06" collectorName="Palmira &amp; Panorama Farm" country="Colombia" county="Cali" elevation="1500" latitude="3.87975" location="Cali" longLatPrecision="1" longitude="-76.43875" municipality="Km" pageId="9" pageNumber="160" specimenCode="IBALCC-RPDR 00256" specimenCount="3" specimenCount-adult="3" stateProvince="Valle del Cauca">
adult female,
<collectorName id="967A9B8BFF8D704EA810F88FFA50F8D4" box="[1342,1432,1879,1905]" pageId="9" pageNumber="160">Palmira</collectorName>
,
<collectorName id="967A9B8BFF8D704EADB9F8A3FE83F830" box="[151,331,1915,1941]" pageId="9" pageNumber="160">Panorama Farm</collectorName>
,
<quantity id="FC7753B8FF8D704EAC7BF8A3FE02F830" box="[341,458,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="160" unit="m" value="2100.0">
<elevation id="B0A2196EFF8D704EAC7BF8A3FE02F830" box="[341,458,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="160" unit="m" value="2100.0">2100 masl</elevation>
</quantity>
,
<date id="4F31D89DFF8D704EACFAF8A2FDF7F830" box="[468,575,1914,1941]" pageId="9" pageNumber="160" value="2005-03-09">
<collectingDate id="5F752175FF8D704EACFAF8A2FDF7F830" box="[468,575,1914,1941]" pageId="9" pageNumber="160" value="2005-03-09">9.iii.2005</collectingDate>
</date>
(MUSENUV-Ar 432);
</materialsCitation>
<materialsCitation id="8BE7F400FF8D704EAE6AF8A3FCD2F81C" ID-GBIF-Occurrence="3816447303" collectingDate="1979-06" collectingDateMax="2021-02" collectingDateMin="1979-06" collectionCode="IBALCC-RPDR" collectorName="J. A. Moreno" country="Colombia" county="Cali" elevation="1500" latitude="3.87975" location="Cali" longLatPrecision="1" longitude="-76.43875" municipality="Km" pageId="9" pageNumber="160" specimenCode="IBALCC-RPDR 00256" specimenCount="3" specimenCount-adult="3" stateProvince="Valle del Cauca">
adult female,
<collectorName id="967A9B8BFF8D704EAEF7F8A3FBE5F830" box="[985,1069,1915,1941]" pageId="9" pageNumber="160">Yumbo</collectorName>
,
<collectorName id="967A9B8BFF8D704EA919F8A3FBBDF830" box="[1079,1141,1915,1941]" pageId="9" pageNumber="160">Dapa</collectorName>
,
<location id="3E50A886FF8D704EA950F8A3FA50F830" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF8D704EA950F8A3FA50F830" box="[1150,1432,1915,1941]" country="Colombia" county="Cali" latitude="3.87975" longLatPrecision="1" longitude="-76.43875" municipality="Km" name="Bocatoma del Acueducto" pageId="9" pageNumber="160" stateProvince="Valle del Cauca">Bocatoma del Acueducto</location>
,
<date id="4F31D89DFF8D704EADB9F847FE83F81C" box="[151,331,1951,1977]" pageId="9" pageNumber="160" value="2016-08-17" valueMax="2016-08-18" valueMin="2016-08-17">
<collectingDate id="5F752175FF8D704EADB9F847FE83F81C" box="[151,331,1951,1977]" pageId="9" pageNumber="160" value="2016-08-17" valueMax="2016-08-18" valueMin="2016-08-17">17–18.viii.2016</collectingDate>
</date>
,
<collectorName id="967A9B8BFF8D704EAC76F847FE27F81C" box="[344,495,1951,1977]" pageId="9" pageNumber="160">J. A. Moreno</collectorName>
(
<specimenCode id="6B295626FF8D704EACD1F847FCC6F81C" box="[511,782,1950,1977]" collectionCode="IBALCC-RPDR" pageId="9" pageNumber="160">IBALCC-RPDR 00256</specimenCode>
);
</materialsCitation>
<materialsCitation id="8BE7F400FF8D704EAE0DF847FAE1F878" ID-GBIF-Occurrence="3816447304" collectingDate="1979-06" collectingDateMax="2021-02" collectingDateMin="1979-06" collectorName="D. Garrido &amp; B. Ospina" country="Colombia" county="Cali" elevation="1500" latitude="3.87975" location="Cali" longLatPrecision="1" longitude="-76.43875" municipality="Km" pageId="9" pageNumber="160" specimenCode="IBALCC-RPDR 00256" specimenCount="3" specimenCount-adult="3" stateProvince="Valle del Cauca">
<specimenCount id="2D8935D4FF8D704EAE0DF847FC54F81C" box="[803,924,1951,1977]" pageId="9" pageNumber="160" type="adult">three adult</specimenCount>
females and
<specimenCount id="2D8935D4FF8D704EA918F847FB68F81C" box="[1078,1184,1951,1977]" pageId="9" pageNumber="160" type="adult">one adult</specimenCount>
male, Yotoco, Yotoco RNF,
<geoCoordinate id="5EBB989AFF8D704EADF5F81AFE95F878" box="[219,349,1986,2013]" degrees="3" direction="north" minutes="52.785" orientation="latitude" pageId="9" pageNumber="160" precision="1" value="3.87975">3º52.785’N</geoCoordinate>
<geoCoordinate id="5EBB989AFF8D704EAC4BF81AFE32F878" box="[357,506,1986,2013]" degrees="76" direction="west" minutes="26.325" orientation="longitude" pageId="9" pageNumber="160" precision="1" value="-76.43875">76º26.325’W</geoCoordinate>
,
<quantity id="FC7753B8FF8D704EAF29F81BFDB7F878" box="[519,639,1987,2013]" metricMagnitude="3" metricUnit="m" metricValue="1.644" pageId="9" pageNumber="160" unit="m" value="1644.0">
<elevation id="B0A2196EFF8D704EAF29F81BFDB7F878" box="[519,639,1987,2013]" metricMagnitude="3" metricUnit="m" metricValue="1.644" pageId="9" pageNumber="160" unit="m" value="1644.0">1644 masl</elevation>
</quantity>
,
<date id="4F31D89DFF8D704EAFA2F81BFD30F878" box="[652,760,1987,2013]" pageId="9" pageNumber="160" value="2019-11-02">
<collectingDate id="5F752175FF8D704EAFA2F81BFD30F878" box="[652,760,1987,2013]" pageId="9" pageNumber="160" value="2019-11-02">2.xi.2019</collectingDate>
</date>
,
<collectorName id="967A9B8BFF8D704EAE2BF81BFC48F878" box="[773,896,1987,2013]" pageId="9" pageNumber="160">D. Garrido</collectorName>
&amp;
<collectorName id="967A9B8BFF8D704EAE89F81BFBD0F878" box="[935,1048,1987,2013]" pageId="9" pageNumber="160">B. Ospina</collectorName>
(MUSENUV-Ar 2117);
</materialsCitation>
<materialsCitation id="8BE7F400FF8D704DA81CF81BFBE0FF14" ID-GBIF-Occurrence="3816447319" collectingDate="1979-06" collectingDateMax="2021-02" collectingDateMin="1979-06" country="Colombia" county="Cali" elevation="1500" lastPageId="10" lastPageNumber="161" latitude="3.87975" location="Cali" longLatPrecision="1" longitude="-76.43875" municipality="Km" pageId="9" pageNumber="160" specimenCode="IBALCC-RPDR 00256" specimenCount="3" specimenCount-adult="3" stateProvince="Valle del Cauca">
<specimenCount id="2D8935D4FF8D704EA81CF81BFA54F878" box="[1330,1436,1987,2013]" pageId="9" pageNumber="160" type="adult">two adult</specimenCount>
females, same locality as the previous record,
<date id="4F31D89DFF8E704DAF82FF4FFCDEFF14" box="[684,790,151,177]" pageId="10" pageNumber="161" value="2019-11-01">
<collectingDate id="5F752175FF8E704DAF82FF4FFCDEFF14" box="[684,790,151,177]" pageId="10" pageNumber="161" value="2019-11-01">1.xi.2019</collectingDate>
</date>
(MUSENUV-Ar 2118);
</materialsCitation>
<materialsCitation id="8BE7F400FF8E704DA91FFF4FFE27FF70" ID-GBIF-Occurrence="3816447307" collectingDate="1979-06" collectingDateMax="2021-02" collectingDateMin="1979-06" country="Colombia" county="Cali" elevation="1500" latitude="3.87975" location="Cali" longLatPrecision="1" longitude="-76.43875" municipality="Km" pageId="10" pageNumber="161" specimenCode="IBALCC-RPDR 00256" specimenCount="3" specimenCount-adult="3" stateProvince="Valle del Cauca">
adult female, Tuluá,
<quantity id="FC7753B8FF8E704DA831FF4FFA50FF14" box="[1311,1432,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="161" unit="m" value="1000.0">
<elevation id="B0A2196EFF8E704DA831FF4FFA50FF14" box="[1311,1432,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="161" unit="m" value="1000.0">1000 masl</elevation>
</quantity>
,
<date id="4F31D89DFF8E704DADB9FF63FF3CFF70" box="[151,244,187,213]" pageId="10" pageNumber="161" value="1984-12">
<collectingDate id="5F752175FF8E704DADB9FF63FF3CFF70" box="[151,244,187,213]" pageId="10" pageNumber="161" value="1984-12">xii.1984</collectingDate>
</date>
(MUSENUV-Ar 437)
</materialsCitation>
.
<materialsCitation id="8BE7F400FF8E704DACD5FF63FD99FEB8" ID-GBIF-Occurrence="3816447313" collectingDate="2012-10-17" collectionCode="MZSP" collectorName="J. A. Moreno" country="Colombia" county="Santuario" elevation="2158" latitude="5.126111" location="San Rafael Plains" longLatPrecision="15" longitude="-76.00733" municipality="San Rafael Plains Natural Regional" pageId="10" pageNumber="161" specimenCount="2" specimenCount-adult="2" stateProvince="Risaralda">
<emphasis id="09FB224FFF8E704DACD5FF63FD39FF70" box="[507,753,187,213]" italics="true" pageId="10" pageNumber="161">
<collectingRegion id="F94B30BFFF8E704DACD5FF63FD39FF70" box="[507,753,187,213]" country="Colombia" name="Risaralda" pageId="10" pageNumber="161">Risaralda department</collectingRegion>
</emphasis>
:
<specimenCount id="2D8935D4FF8E704DAFD1FF63FCBEFF70" box="[767,886,187,213]" pageId="10" pageNumber="161" type="adult">three adult</specimenCount>
males and
<specimenCount id="2D8935D4FF8E704DAEDAFF63FB95FF70" box="[1012,1117,187,213]" pageId="10" pageNumber="161" type="adult">two adult</specimenCount>
females,
<collectingCounty id="D25186D1FF8E704DA9E4FF63FAF0FF70" box="[1226,1336,187,213]" pageId="10" pageNumber="161">Santuario</collectingCounty>
,
<collectingMunicipality id="DB546427FF8E704DA86DFF63FE1EFF5C" pageId="10" pageNumber="161">
<location id="3E50A886FF8E704DA86DFF63FEC5FF5C" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF8E704DA86DFF63FEC5FF5C" country="Colombia" county="Santuario" latitude="5.126111" longLatPrecision="15" longitude="-76.00733" municipality="San Rafael Plains Natural Regional" name="San Rafael Plains" pageId="10" pageNumber="161" stateProvince="Risaralda">
San
<location id="3E50A886FF8E704DA85AFF63FEC5FF5C" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF8E704DA85AFF63FEC5FF5C" country="Colombia" county="Santuario" latitude="5.126111" longLatPrecision="15" longitude="-76.00733" municipality="San Rafael Plains Natural Regional" name="Rafael Plains Natural Regional Park" pageId="10" pageNumber="161" stateProvince="Risaralda">Rafael Plains</location>
</location>
Natural Regional
</collectingMunicipality>
Park [Parque Natural Regional Planes de San Rafael],
<geoCoordinate id="5EBB989AFF8E704DA966FF06FB7EFF5C" box="[1096,1206,222,249]" degrees="5" direction="north" minutes="7" orientation="latitude" pageId="10" pageNumber="161" precision="15" seconds="34" value="5.126111">5°7’34”N</geoCoordinate>
<geoCoordinate id="5EBB989AFF8E704DA993FF07FA90FF5C" box="[1213,1368,223,249]" degrees="76" direction="west" minutes="0" orientation="longitude" pageId="10" pageNumber="161" precision="1" seconds="26.4" value="-76.00733">76°0’26.4’’W</geoCoordinate>
,
<quantity id="FC7753B8FF8E704DA84AFF06FF07FEB8" metricMagnitude="3" metricUnit="m" metricValue="2.158" pageId="10" pageNumber="161" unit="m" value="2158.0">
<elevation id="B0A2196EFF8E704DA84AFF06FF07FEB8" metricMagnitude="3" metricUnit="m" metricValue="2.158" pageId="10" pageNumber="161" unit="m" value="2158.0">2158 masl</elevation>
</quantity>
,
<date id="4F31D89DFF8E704DADF4FEDBFE84FEB8" box="[218,332,259,285]" pageId="10" pageNumber="161" value="2012-10-17">
<collectingDate id="5F752175FF8E704DADF4FEDBFE84FEB8" box="[218,332,259,285]" pageId="10" pageNumber="161" value="2012-10-17">17.x.2012</collectingDate>
</date>
,
<collectorName id="967A9B8BFF8E704DAC76FEDBFE23FEB8" box="[344,491,259,285]" pageId="10" pageNumber="161">J. A. Moreno</collectorName>
(
<collectionCode id="5D9E6698FF8E704DACD7FEDBFD80FEB8" box="[505,584,259,285]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="10" pageNumber="161" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="6FF0AED5FF8E704DADB9FA87FC66FA1A" ID-DOI="http://doi.org/10.5281/zenodo.6673348" ID-Zenodo-Dep="6673348" httpUri="https://zenodo.org/record/6673348/files/figure.png" pageId="10" pageNumber="161" startId="10.[151,250,1375,1399]" targetBox="[371,1216,309,1352]" targetPageId="10">
<paragraph id="3B30FE5DFF8E704DADB9FA87FC66FA1A" blockId="10.[151,1437,1375,1472]" pageId="10" pageNumber="161">
<emphasis id="09FB224FFF8E704DADB9FA87FEDBFAD2" bold="true" box="[151,275,1375,1399]" pageId="10" pageNumber="161">FIGURE 4.</emphasis>
<taxonomicName id="FC8F85DEFF8E704DAC37FAB8FD8DFAD2" authority="Gervais, 1843" authorityName="Gervais" authorityYear="1843" box="[281,581,1375,1399]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="subGenus" subGenus="Atreus">
<emphasis id="09FB224FFF8E704DAC37FAB8FE9EFAD2" box="[281,342,1376,1399]" italics="true" pageId="10" pageNumber="161">Tityus</emphasis>
(
<emphasis id="09FB224FFF8E704DAC4BFA87FE6EFAD3" box="[357,422,1375,1398]" italics="true" pageId="10" pageNumber="161">Atreus</emphasis>
)
<bibRefCitation id="5F1E83ACFF8E704DAC9BFA87FD8DFAD2" author="Gervais, P. M." box="[437,581,1375,1399]" pageId="10" pageNumber="161" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
</taxonomicName>
, carapace sculpturing, dorsal aspect.
<emphasis id="09FB224FFF8E704DAE93FA87FC3BFAD2" bold="true" box="[957,1011,1375,1399]" pageId="10" pageNumber="161">A, B.</emphasis>
<taxonomicName id="FC8F85DEFF8E704DAED7FAB8FA51FAD2" authority="(Gervais, 1843)" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[1017,1433,1375,1399]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="forcipula" subGenus="Atreus">
<emphasis id="09FB224FFF8E704DAED7FAB8FBFEFAD2" box="[1017,1078,1376,1399]" italics="true" pageId="10" pageNumber="161">Tityus</emphasis>
(
<emphasis id="09FB224FFF8E704DA96BFA87FB4EFAD3" box="[1093,1158,1375,1398]" italics="true" pageId="10" pageNumber="161">Atreus</emphasis>
)
<emphasis id="09FB224FFF8E704DA9BBFA87FB39FAD2" box="[1173,1265,1375,1399]" italics="true" pageId="10" pageNumber="161">forcipula</emphasis>
(
<bibRefCitation id="5F1E83ACFF8E704DA9D0FA87FA59FAD2" author="Gervais, P. M." box="[1278,1425,1375,1399]" pageId="10" pageNumber="161" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="09FB224FFF8E704DADB9FA5BFF78FA3E" bold="true" box="[151,176,1411,1435]" pageId="10" pageNumber="161">A.</emphasis>
Male (MUSENUV-Ar 438).
<emphasis id="09FB224FFF8E704DACF0FA5BFE3DFA3E" bold="true" box="[478,501,1411,1435]" pageId="10" pageNumber="161">B.</emphasis>
Female (MUSENUV-Ar 436).
<emphasis id="09FB224FFF8E704DAE17FA5BFCBAFA3E" bold="true" box="[825,882,1411,1435]" pageId="10" pageNumber="161">C, D.</emphasis>
<taxonomicName id="FC8F85DEFF8E704DAE57FA5CFB4EFA3E" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[889,1158,1411,1435]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov." subGenus="Atreus">
<emphasis id="09FB224FFF8E704DAE57FA5CFC7EFA3E" box="[889,950,1412,1435]" italics="true" pageId="10" pageNumber="161">Tityus</emphasis>
(
<emphasis id="09FB224FFF8E704DAEE8FA5BFBCFFA3F" box="[966,1031,1411,1434]" italics="true" pageId="10" pageNumber="161">Atreus</emphasis>
)
<emphasis id="09FB224FFF8E704DA939FA5BFB4EFA3E" box="[1047,1158,1411,1435]" italics="true" pageId="10" pageNumber="161">moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF8E704DA9A0FA5BFAC8FA3E" bold="true" box="[1166,1280,1411,1435]" pageId="10" pageNumber="161">
<taxonomicNameLabel id="12C89F34FF8E704DA9A0FA5BFB28FA3E" box="[1166,1248,1411,1435]" pageId="10" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
C.
</emphasis>
Male holotype (MUSENUV-Ar 2104).
<emphasis id="09FB224FFF8E704DACA2FA7FFE6DFA1A" bold="true" box="[396,421,1447,1471]" pageId="10" pageNumber="161">D.</emphasis>
Female paratype (ICN-1402). Scale bars= 2.5 mm.
</paragraph>
</caption>
<subSubSection id="7395ADD6FF8E704CADE9FA35FDC2FEE4" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161" type="diagnosis">
<paragraph id="3B30FE5DFF8E704DADE9FA35FE5FF919" blockId="10.[151,1437,1517,2012]" pageId="10" pageNumber="161">
<emphasis id="09FB224FFF8E704DADE9FA35FE8AF9A2" bold="true" box="[199,322,1517,1543]" pageId="10" pageNumber="161">Diagnosis.</emphasis>
<taxonomicName id="FC8F85DEFF8E704DAC69FA36FE3EF9A2" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[327,502,1517,1543]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF8E704DAC69FA36FE3EF9A2" box="[327,502,1517,1543]" italics="true" pageId="10" pageNumber="161">Tityus forcipula</emphasis>
</taxonomicName>
share a metasoma widening towards segment V in both sexes (
<figureCitation id="A3B4E2D8FF8E704DA983FA35FA92F9AD" box="[1197,1370,1517,1544]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="10" pageNumber="161">Figures 12A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF8E704DA84BFA35FF62F989" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="10" pageNumber="161">13A, B</figureCitation>
) and a male chela bulkier than female chela (
<figureCitation id="A3B4E2D8FF8E704DAF87F9C9FC84F989" box="[681,844,1553,1580]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[261,1326,181,1920]" captionTargetId="figure-19@15.[261,1326,181,1920]" captionTargetPageId="15" captionText="FIGURE 7. Tityus (Atreus) Gervais, 1843,female chela sculpturing.A–C. Tityus(Atreus) forcipula (Gervais,1843) (MUSENUV- Ar 436). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (ICN-1402). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673356" httpUri="https://zenodo.org/record/6673356/files/figure.png" pageId="10" pageNumber="161">Figures 7A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF8E704DAE76F9C9FC53F989" box="[856,923,1553,1580]" captionStart="FIGURE 8" captionStartId="16.[151,250,1944,1968]" captionTargetBox="[364,1224,181,1920]" captionTargetId="figure-31@16.[364,1224,181,1920]" captionTargetPageId="16" captionText="FIGURE 8. Tityus (Atreus) Gervais, 1843, male chela sculpturing. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673358" httpUri="https://zenodo.org/record/6673358/files/figure.png" pageId="10" pageNumber="161">8A, B</figureCitation>
) with
<taxonomicName id="FC8F85DEFF8E704DAECAF9CAFB4EF98E" authorityName="Lourenco" authorityYear="1994" box="[996,1158,1553,1579]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="cuellari">
<emphasis id="09FB224FFF8E704DAECAF9CAFB4EF98E" box="[996,1158,1553,1579]" italics="true" pageId="10" pageNumber="161">Tityus cuellari</emphasis>
</taxonomicName>
,
<taxonomicName id="FC8F85DEFF8E704DA9BAF9CAFAADF98E" authorityName="Lourenco" authorityYear="2013" box="[1172,1381,1553,1579]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="crassicauda">
<emphasis id="09FB224FFF8E704DA9BAF9CAFAADF98E" box="[1172,1381,1553,1579]" italics="true" pageId="10" pageNumber="161">Tityus crassicauda</emphasis>
</taxonomicName>
, and
<taxonomicName id="FC8F85DEFF8E704DADB9F9EEFE94F9EA" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[151,348,1589,1615]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov.">
<emphasis id="09FB224FFF8E704DADB9F9EEFE94F9EA" box="[151,348,1589,1615]" italics="true" pageId="10" pageNumber="161">Tityus moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF8E704DAC4DF9EDFE74F9EA" bold="true" box="[355,444,1589,1615]" pageId="10" pageNumber="161">
<taxonomicNameLabel id="12C89F34FF8E704DAC4DF9EDFE74F9EA" box="[355,444,1589,1615]" pageId="10" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. However,
<emphasis id="09FB224FFF8E704DAF13F9EEFC0CF9EA" box="[573,964,1589,1615]" italics="true" pageId="10" pageNumber="161">
<taxonomicName id="FC8F85DEFF8E704DAF13F9EEFCD9F9EA" authorityName="Lourenco" authorityYear="2013" box="[573,785,1589,1615]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="crassicauda">Tityus crassicauda</taxonomicName>
,
<taxonomicName id="FC8F85DEFF8E704DAE33F9EEFC08F9EA" authorityName="Lourenco" authorityYear="1994" box="[797,960,1589,1615]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="cuellari">Tityus cuellari</taxonomicName>
,
</emphasis>
and
<taxonomicName id="FC8F85DEFF8E704DAED5F9EEFB62F9EA" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[1019,1194,1589,1615]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF8E704DAED5F9EEFB62F9EA" box="[1019,1194,1589,1615]" italics="true" pageId="10" pageNumber="161">Tityus forcipula</emphasis>
</taxonomicName>
share the presence of strongly protruding spinoid granules on the DSM carinae of metasomal segments II–IV (
<figureCitation id="A3B4E2D8FF8E704DA950F981FAECF9D1" box="[1150,1316,1625,1652]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="10" pageNumber="161">Figure 14A, B</figureCitation>
), whereas,
<taxonomicName id="FC8F85DEFF8E704DADB9F9A6FE97F932" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[151,351,1661,1687]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="moralensis">
<emphasis id="09FB224FFF8E704DADB9F9A6FE97F932" box="[151,351,1661,1687]" italics="true" pageId="10" pageNumber="161">Tityus moralensis</emphasis>
</taxonomicName>
, present moderately protruding rounded granules on the DSM carinae of metasomal segments II–IV (
<figureCitation id="A3B4E2D8FF8E704DADCDF979FE42F919" box="[227,394,1697,1724]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="10" pageNumber="161">Figure 14C, D</figureCitation>
).
</paragraph>
<paragraph id="3B30FE5DFF8E704DADE9F91DFAF2F836" blockId="10.[151,1437,1517,2012]" pageId="10" pageNumber="161">
On the other hand,
<taxonomicName id="FC8F85DEFF8E704DAC8FF91EFD99F97A" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[417,593,1733,1759]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF8E704DAC8FF91EFD99F97A" box="[417,593,1733,1759]" italics="true" pageId="10" pageNumber="161">Tityus forcipula</emphasis>
</taxonomicName>
and
<taxonomicName id="FC8F85DEFF8E704DAFA6F91EFCE2F97A" authorityName="Lourenco" authorityYear="1994" box="[648,810,1733,1759]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="cuellari">
<emphasis id="09FB224FFF8E704DAFA6F91EFCE2F97A" box="[648,810,1733,1759]" italics="true" pageId="10" pageNumber="161">Tityus cuellari</emphasis>
</taxonomicName>
share the presence of metasomal segment V with DSM carinae composed of moderately to strongly protruding spinoid granules (e.g.,
<figureCitation id="A3B4E2D8FF8E704DAEDAF931FB5CF8A1" box="[1012,1172,1769,1796]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="10" pageNumber="161">Figure 14A, B</figureCitation>
), whereas
<taxonomicName id="FC8F85DEFF8E704DA826F932FF13F882" authorityName="Lourenco" authorityYear="2013" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="crassicauda">
<emphasis id="09FB224FFF8E704DA826F932FF13F882" italics="true" pageId="10" pageNumber="161">Tityus crassicauda</emphasis>
</taxonomicName>
presents a metasomal segment V with DSM carinae composed of slightly protruding granules (
<bibRefCitation id="5F1E83ACFF8E704DA83CF8D5FEE0F8EE" author="Lourenco, W. R. &amp; Ythier, E." pageId="10" pageNumber="161" pagination="1 - 13" refId="ref19366" refString="Lourenco, W. R. &amp; Ythier, E. (2013) The remarkable scorpion diversity in the Ecuadorian Andes and description of a new species of Tityus C. L. Koch, 1836 (Scorpiones, Buthidae). ZooKeys, 307, 1 - 13. https: // doi. org / 10.3897 / zookeys. 307.5334" type="journal article" year="2013">Lourenço &amp; Ythier, 2013</bibRefCitation>
: figs 1–4). Finally,
<taxonomicName id="FC8F85DEFF8E704DAF2EF8EAFD78F8EE" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[512,688,1841,1867]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF8E704DAF2EF8EAFD78F8EE" box="[512,688,1841,1867]" italics="true" pageId="10" pageNumber="161">Tityus forcipula</emphasis>
</taxonomicName>
presents a medium-sized subaculear tubercle with a medium-sized dorsal area between the subaculear tubercle and the aculeus base (
<figureCitation id="A3B4E2D8FF8E704DAE55F88DFBE8F8D5" box="[891,1056,1877,1904]" captionStart="FIGURE 15" captionStartId="24.[151,250,1090,1114]" captionTargetBox="[151,1436,345,1067]" captionTargetId="figure-116@24.[151,1436,345,1067]" captionTargetPageId="24" captionText="FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673376" httpUri="https://zenodo.org/record/6673376/files/figure.png" pageId="10" pageNumber="161">Figure 15A, B</figureCitation>
), whereas
<taxonomicName id="FC8F85DEFF8E704DA9B6F88EFAF1F8CA" authorityName="Lourenco" authorityYear="1994" box="[1176,1337,1877,1903]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="cuellari">
<emphasis id="09FB224FFF8E704DA9B6F88EFAF1F8CA" box="[1176,1337,1877,1903]" italics="true" pageId="10" pageNumber="161">Tityus cuellari</emphasis>
</taxonomicName>
presents a small subaculear tubercle with a small dorsal area between the subaculear tubercle and the aculeus base.
</paragraph>
<paragraph id="3B30FE5DFF8E704CADE9F845FDC2FEE4" blockId="10.[151,1437,1517,2012]" lastBlockId="11.[151,1437,151,393]" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161">
<emphasis id="09FB224FFF8E704DADE9F845FE76F812" bold="true" box="[199,446,1949,1975]" pageId="10" pageNumber="161">Taxonomic remarks.</emphasis>
As mentioned by
<bibRefCitation id="5F1E83ACFF8E704DAFA0F845FC59F81D" author="Brito, G. &amp; Borges, A." box="[654,913,1949,1976]" pageId="10" pageNumber="161" pagination="1 - 17" refId="ref16788" refString="Brito, G. &amp; Borges, A. (2015) A checklist of the scorpions of Ecuador (Arachnida: Scorpiones), with notes on the distribution and medical significance of some species. Journal of Venomous Animals and Toxins Including Tropical Diseases, 21, 1 - 17. https: // doi. org / 10.1186 / s 40409 - 015 - 0023 - x" type="journal article" year="2015">Brito &amp; Borges (2015)</bibRefCitation>
,
<taxonomicName id="FC8F85DEFF8E704DAEB0F846FB8EF812" authorityName="Pocock" authorityYear="1898" box="[926,1094,1950,1975]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="spinatus">
<emphasis id="09FB224FFF8E704DAEB0F846FB8EF812" box="[926,1094,1950,1975]" italics="true" pageId="10" pageNumber="161">Tityus spinatus</emphasis>
</taxonomicName>
was synonymized with
<taxonomicName id="FC8F85DEFF8E704DA877F846FF35F87E" baseAuthorityName="Gervais" baseAuthorityYear="1843" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF8E704DA877F846FF35F87E" italics="true" pageId="10" pageNumber="161">Tityus forcipula</emphasis>
</taxonomicName>
by
<bibRefCitation id="5F1E83ACFF8E704DAC04F81AFE24F87E" author="Lourenco, W. R." box="[298,492,1985,2012]" pageId="10" pageNumber="161" pagination="729 - 739" refId="ref18482" refString="Lourenco, W. R. (1984) Etude systematique de quelques especes appartenant au complexe Tityus forcipula (Gervais, 1844) (Scorpiones, Buthidae). Bulletin du Museum national d'histoire naturelle. Section A, Zoologie, biologie et ecologie animales, 6, 729 - 739." type="journal article" year="1984">Lourenço (1984)</bibRefCitation>
. However, in recent papers such as
<bibRefCitation id="5F1E83ACFF8E704DAEA7F81AFB67F87E" author="Lourenco, W. R. &amp; Ythier, E." box="[905,1199,1985,2012]" pageId="10" pageNumber="161" pagination="1 - 13" refId="ref19366" refString="Lourenco, W. R. &amp; Ythier, E. (2013) The remarkable scorpion diversity in the Ecuadorian Andes and description of a new species of Tityus C. L. Koch, 1836 (Scorpiones, Buthidae). ZooKeys, 307, 1 - 13. https: // doi. org / 10.3897 / zookeys. 307.5334" type="journal article" year="2013">Lourenço &amp; Ythier (2013</bibRefCitation>
,
<bibRefCitation id="5F1E83ACFF8E704DA992F819FB3EF87E" author="Lourenco, W. R. &amp; Ythier, E." box="[1212,1270,1985,2011]" pageId="10" pageNumber="161" pagination="18 - 34" refId="ref19429" refString="Lourenco, W. R. &amp; Ythier, E. (2017) Description of Tityus (Atreus) cisandinus sp. n. from Ecuadorian Amazonia, with comments on some related species (Scorpiones: Buthidae). Arachnida - Rivista Aracnologica Italiana, 15, 18 - 34." type="journal article" year="2017">2017</bibRefCitation>
)
<taxonomicName id="FC8F85DEFF8E704DA828F81AFA48F87E" authorityName="Pocock" authorityYear="1898" box="[1286,1408,1986,2011]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="spinatus">
<emphasis id="09FB224FFF8E704DA828F81AFA48F87E" box="[1286,1408,1986,2011]" italics="true" pageId="10" pageNumber="161">T. spinatus</emphasis>
</taxonomicName>
is mentioned without any comments about its validity. In fact,
<bibRefCitation id="5F1E83ACFF8F704CAE6EFF4FFBA2FF14" author="Lourenco, W. R. &amp; Ythier, E." box="[832,1130,151,177]" pageId="11" pageNumber="162" pagination="1 - 13" refId="ref19366" refString="Lourenco, W. R. &amp; Ythier, E. (2013) The remarkable scorpion diversity in the Ecuadorian Andes and description of a new species of Tityus C. L. Koch, 1836 (Scorpiones, Buthidae). ZooKeys, 307, 1 - 13. https: // doi. org / 10.3897 / zookeys. 307.5334" type="journal article" year="2013">Lourenço &amp; Ythier (2013)</bibRefCitation>
did not present any formal species revalidation and only provided a single drawing of the metasomal segments II–V + telson of the female
<typeStatus id="E43440FFFF8F704CADB9FF07FF35FF5C" box="[151,253,223,249]" pageId="11" pageNumber="162" type="holotype">holotype</typeStatus>
. For this reason, we followed
<bibRefCitation id="5F1E83ACFF8F704CAF63FF07FCC2FF5C" author="Lourenco, W. R." box="[589,778,223,249]" pageId="11" pageNumber="162" pagination="681 - 697" refId="ref18535" refString="Lourenco, W. R. (1988) La faune des Scorpions de l'Equateur. I. Les Buthidae. Systematique et biogeographie. Revue Suisse de Zoologie, 95, 681 - 697. https: // doi. org / 10.5962 / bhl. part. 81928" type="journal article" year="1988">Lourenço (1988)</bibRefCitation>
and
<bibRefCitation id="5F1E83ACFF8F704CAE6FFF07FBF6FF5C" author="Brito, G. &amp; Borges, A." box="[833,1086,222,249]" pageId="11" pageNumber="162" pagination="1 - 17" refId="ref16788" refString="Brito, G. &amp; Borges, A. (2015) A checklist of the scorpions of Ecuador (Arachnida: Scorpiones), with notes on the distribution and medical significance of some species. Journal of Venomous Animals and Toxins Including Tropical Diseases, 21, 1 - 17. https: // doi. org / 10.1186 / s 40409 - 015 - 0023 - x" type="journal article" year="2015">Brito &amp; Borges (2015)</bibRefCitation>
and considered
<taxonomicName id="FC8F85DEFF8F704CA9DBFF07FA54FF5D" authorityName="Pocock" authorityYear="1898" box="[1269,1436,223,248]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="spinatus">
<emphasis id="09FB224FFF8F704CA9DBFF07FA54FF5D" box="[1269,1436,223,248]" italics="true" pageId="11" pageNumber="162">Tityus spinatus</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="FC8F85DEFF8F704CAC76FEDBFE13FEB8" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[344,475,259,285]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF8F704CAC76FEDBFE13FEB8" box="[344,475,259,285]" italics="true" pageId="11" pageNumber="162">T. forcipula</emphasis>
</taxonomicName>
as proposed by
<bibRefCitation id="5F1E83ACFF8F704CAFBAFEDBFC9CFEB8" author="Lourenco, W. R." box="[660,852,259,285]" pageId="11" pageNumber="162" pagination="729 - 739" refId="ref18482" refString="Lourenco, W. R. (1984) Etude systematique de quelques especes appartenant au complexe Tityus forcipula (Gervais, 1844) (Scorpiones, Buthidae). Bulletin du Museum national d'histoire naturelle. Section A, Zoologie, biologie et ecologie animales, 6, 729 - 739." type="journal article" year="1984">Lourenço (1984)</bibRefCitation>
, until additional evidence is presented and a formal species revalidation is published.
</paragraph>
</subSubSection>
<subSubSection id="7395ADD6FF8F7055ADE9FE92FB7EF9F7" lastPageId="18" lastPageNumber="169" pageId="11" pageNumber="162" type="description">
<paragraph id="3B30FE5DFF8F704CADE9FE92FA8AFEC3" blockId="11.[151,1437,151,393]" box="[199,1346,330,358]" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CADE9FE92FE91FEC1" bold="true" box="[199,345,330,356]" pageId="11" pageNumber="162">Description.</emphasis>
Based on an adult male (MUSENUV-Ar 438) and an adult female (MUSENUV-Ar 436).
</paragraph>
<paragraph id="3B30FE5DFF8F704CADE9FEB6FB0AFE2C" blockId="11.[151,1437,151,393]" box="[199,1218,366,393]" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CADE9FEB6FE90FE2C" bold="true" box="[199,344,366,393]" italics="true" pageId="11" pageNumber="162">Total length.</emphasis>
Male:
<quantity id="FC7753B8FF8F704CAC86FEB6FDD2FE2C" box="[424,538,366,393]" metricMagnitude="-2" metricUnit="m" metricValue="5.381" pageId="11" pageNumber="162" unit="mm" value="53.81">53.81 mm</quantity>
and female
<quantity id="FC7753B8FF8F704CAF8DFEB7FCDDFE2C" box="[675,789,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="6.173" pageId="11" pageNumber="162" unit="mm" value="61.73">61.73 mm</quantity>
(additional measurements in
<tableCitation id="760DCBE6FF8F704CA971FEB7FB7CFE2C" box="[1119,1204,366,393]" captionStart="TABLE 5" captionStartId="11.[151,239,440,464]" captionTargetBox="[167,1416,516,1987]" captionTargetPageId="11" captionText="TABLE 5. Measurements (mm) of Tityus forcipula and Tityus moralensis sp. nov. Observation: all specimens, but 1402" httpUri="http://table.plazi.org/id/6FF0AED5FF8F704CADB9FE60FAC8FE51" pageId="11" pageNumber="162" tableUuid="6FF0AED5FF8F704CADB9FE60FAC8FE51">Table 5</tableCitation>
).
</paragraph>
<caption id="6FF0AED5FF8F704CADB9FE60FAC8FE51" ID-Table-UUID="6FF0AED5FF8F704CADB9FE60FAC8FE51" httpUri="http://table.plazi.org/id/6FF0AED5FF8F704CADB9FE60FAC8FE51" pageId="11" pageNumber="162" startId="11.[151,239,440,464]" targetBox="[167,1416,516,1987]" targetIsTable="true" targetPageId="11">
<paragraph id="3B30FE5DFF8F704CADB9FE60FAC8FE51" blockId="11.[151,1436,440,501]" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CADB9FE60FEC2FE74" bold="true" box="[151,266,440,465]" pageId="11" pageNumber="162">TABLE 5.</emphasis>
Measurements (mm) of
<taxonomicName id="FC8F85DEFF8F704CAF3DFE61FD73FE75" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[531,699,440,464]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF8F704CAF3DFE61FD73FE75" box="[531,699,440,464]" italics="true" pageId="11" pageNumber="162">Tityus forcipula</emphasis>
</taxonomicName>
and
<taxonomicName id="FC8F85DEFF8F704CAFC1FE61FC63FE75" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[751,939,440,464]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov.">
<emphasis id="09FB224FFF8F704CAFC1FE61FC63FE75" box="[751,939,440,464]" italics="true" pageId="11" pageNumber="162">Tityus moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF8F704CAE9CFE61FBC0FE74" bold="true" box="[946,1032,441,465]" pageId="11" pageNumber="162">
<taxonomicNameLabel id="12C89F34FF8F704CAE9CFE61FBC0FE74" box="[946,1032,441,465]" pageId="11" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Observation: all specimens, but 1402 and 1403 deposited at ICN, are deposited at MUSENUV. Bold numbers correspond to the museum codes.
</paragraph>
</caption>
<paragraph id="3B30FE5DFF8F704CAD89FDDCFA40F866" pageId="11" pageNumber="162">
<table id="498F0CFDFF8F8FB8AD89FDDCFA40F866" box="[167,1416,516,1987]" colsContinueIn="12.[167,1416,203,1063]" gridcols="11" gridrows="33" pageId="11" pageNumber="162">
<tr id="85BFFC1FFF8F8FB8AD89FDDCFA40FDB9" box="[167,1416,516,540]" gridrow="0" pageId="11" pageNumber="162" rowspan-2="1" rowspan-6="1" rowspan-7="1">
<th id="C66E9563FF8F8FB8AD89FDDCFEF9FDB9" box="[167,305,516,540]" gridcol="0" gridrow="0" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CAD89FDDCFEC7FDB9" bold="true" box="[167,271,516,540]" pageId="11" pageNumber="162">Structure</emphasis>
</th>
<th id="C66E9563FF8F8FB8AC50FDDCFDD1FDB9" box="[382,537,516,540]" gridcol="1" gridrow="0" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CAC50FDDCFE14FDB9" bold="true" box="[382,476,516,540]" pageId="11" pageNumber="162">Measure</emphasis>
</th>
<th id="C66E9563FF8F8FB8AF8BFDDCFC55FDB9" box="[677,925,516,540]" colspan="3" colspanRight="2" gridcol="3" gridrow="0" pageId="11" pageNumber="162">
<taxonomicName id="FC8F85DEFF8F704CAFEAFDDCFCB4FDB9" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[708,892,516,540]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="moralensis">
<emphasis id="09FB224FFF8F704CAFEAFDDCFCB4FDB9" bold="true" box="[708,892,516,540]" italics="true" pageId="11" pageNumber="162">Tityus moralensis</emphasis>
</taxonomicName>
</th>
<th id="C66E9563FF8F8FB8A9AAFDDCFA40FDB9" box="[1156,1416,516,540]" colspan="3" colspanRight="2" gridcol="8" gridrow="0" pageId="11" pageNumber="162">
<taxonomicName id="FC8F85DEFF8F704CA99BFDDCFA93FDB9" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[1205,1371,516,540]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF8F704CA99BFDDCFA93FDB9" bold="true" box="[1205,1371,516,540]" italics="true" pageId="11" pageNumber="162">Tityus forcipula</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FDF6FA40FDC8" box="[167,1416,558,621]" gridrow="1" pageId="11" pageNumber="162" rowspan-0="1" rowspan-1="1">
<td id="C66E9563FF8F8FB8AF15FDF6FDBCFDC8" box="[571,628,558,621]" gridcol="2" gridrow="1" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CAF15FDF6FDA6FDE3" bold="true" box="[571,622,558,582]" pageId="11" pageNumber="162">2104</emphasis>
♂
</td>
<td id="C66E9563FF8F8FB8AF8BFDF6FD16FDC8" box="[677,734,558,621]" gridcol="3" gridrow="1" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CAF8BFDF6FD70FDC8" bold="true" box="[677,728,558,621]" pageId="11" pageNumber="162">1402 ♀</emphasis>
</td>
<td id="C66E9563FF8F8FB8AE2AFDF6FCF5FDC8" box="[772,829,558,621]" gridcol="4" gridrow="1" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CAE2AFDF6FCDFFDC8" bold="true" box="[772,823,558,621]" pageId="11" pageNumber="162">1403 ♀</emphasis>
</td>
<td id="C66E9563FF8F8FB8AE4AFDF6FC55FDC8" box="[868,925,558,621]" gridcol="5" gridrow="1" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CAE4AFDF6FCBFFDC8" bold="true" box="[868,919,558,621]" pageId="11" pageNumber="162">2108 ♀</emphasis>
</td>
<td id="C66E9563FF8F8FB8AEEAFDF6FC35FDC8" box="[964,1021,558,621]" gridcol="6" gridrow="1" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CAEEAFDF6FC1FFDC8" bold="true" box="[964,1015,558,621]" pageId="11" pageNumber="162">2109 ♀</emphasis>
</td>
<td id="C66E9563FF8F8FB8A90AFDF6FB95FDC8" box="[1060,1117,558,621]" gridcol="7" gridrow="1" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CA90AFDF6FBFFFDC8" bold="true" box="[1060,1108,558,621]" pageId="11" pageNumber="162">2111 ♀</emphasis>
</td>
<td id="C66E9563FF8F8FB8A9AAFDF6FB75FDC8" box="[1156,1213,558,621]" gridcol="8" gridrow="1" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CA9AAFDF6FB5FFDC8" bold="true" box="[1156,1194,558,621]" pageId="11" pageNumber="162">436 ♀</emphasis>
</td>
<td id="C66E9563FF8F8FB8A9C1FDF6FAE0FDC8" box="[1263,1320,558,621]" gridcol="9" gridrow="1" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CA9C1FDF6FACAFDC8" bold="true" box="[1263,1314,558,621]" pageId="11" pageNumber="162">2103 ♀</emphasis>
</td>
<td id="C66E9563FF8F8FB8A861FDF6FA40FDC8" box="[1359,1416,558,621]" gridcol="10" gridrow="1" pageId="11" pageNumber="162">
<emphasis id="09FB224FFF8F704CA861FDF6FAAAFDC8" bold="true" box="[1359,1397,558,621]" pageId="11" pageNumber="162">438 ♂</emphasis>
</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FDA2FA40FD37" box="[167,1416,634,658]" gridrow="2" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89FDA2FEF9FD37" box="[167,305,634,658]" gridcol="0" gridrow="2" pageId="11" pageNumber="162">Body</th>
<td id="C66E9563FF8F8FB8AC50FDA2FDD1FD37" box="[382,537,634,658]" gridcol="1" gridrow="2" pageId="11" pageNumber="162">Total length</td>
<td id="C66E9563FF8F8FB8AF15FDA2FDBCFD37" box="[571,628,634,658]" gridcol="2" gridrow="2" pageId="11" pageNumber="162">55.03</td>
<td id="C66E9563FF8F8FB8AF8BFDA2FD16FD37" box="[677,734,634,658]" gridcol="3" gridrow="2" pageId="11" pageNumber="162">57.99</td>
<td id="C66E9563FF8F8FB8AE2AFDA2FCF5FD37" box="[772,829,634,658]" gridcol="4" gridrow="2" pageId="11" pageNumber="162">53.38</td>
<td id="C66E9563FF8F8FB8AE4AFDA2FC55FD37" box="[868,925,634,658]" gridcol="5" gridrow="2" pageId="11" pageNumber="162">57.93</td>
<td id="C66E9563FF8F8FB8AEEAFDA2FC35FD37" box="[964,1021,634,658]" gridcol="6" gridrow="2" pageId="11" pageNumber="162">61.55</td>
<td id="C66E9563FF8F8FB8A90AFDA2FB95FD37" box="[1060,1117,634,658]" gridcol="7" gridrow="2" pageId="11" pageNumber="162">56.25</td>
<td id="C66E9563FF8F8FB8A9AAFDA2FB75FD37" box="[1156,1213,634,658]" gridcol="8" gridrow="2" pageId="11" pageNumber="162">61.73</td>
<td id="C66E9563FF8F8FB8A9C1FDA2FAE0FD37" box="[1263,1320,634,658]" gridcol="9" gridrow="2" pageId="11" pageNumber="162">62.30</td>
<td id="C66E9563FF8F8FB8A861FDA2FA40FD37" box="[1359,1416,634,658]" gridcol="10" gridrow="2" pageId="11" pageNumber="162">53.81</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FD7CFA40FD18" box="[167,1416,676,701]" gridrow="3" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89FD7CFEF9FD18" box="[167,305,676,701]" gridcol="0" gridrow="3" pageId="11" pageNumber="162">Carapace</th>
<td id="C66E9563FF8F8FB8AC50FD7CFDD1FD18" box="[382,537,676,701]" gridcol="1" gridrow="3" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15FD7CFDBCFD18" box="[571,628,676,701]" gridcol="2" gridrow="3" pageId="11" pageNumber="162">6.00</td>
<td id="C66E9563FF8F8FB8AF8BFD7CFD16FD18" box="[677,734,676,701]" gridcol="3" gridrow="3" pageId="11" pageNumber="162">6.29</td>
<td id="C66E9563FF8F8FB8AE2AFD7CFCF5FD18" box="[772,829,676,701]" gridcol="4" gridrow="3" pageId="11" pageNumber="162">5.71</td>
<td id="C66E9563FF8F8FB8AE4AFD7CFC55FD18" box="[868,925,676,701]" gridcol="5" gridrow="3" pageId="11" pageNumber="162">6.43</td>
<td id="C66E9563FF8F8FB8AEEAFD7CFC35FD18" box="[964,1021,676,701]" gridcol="6" gridrow="3" pageId="11" pageNumber="162">6.50</td>
<td id="C66E9563FF8F8FB8A90AFD7CFB95FD18" box="[1060,1117,676,701]" gridcol="7" gridrow="3" pageId="11" pageNumber="162">6.35</td>
<td id="C66E9563FF8F8FB8A9AAFD7CFB75FD18" box="[1156,1213,676,701]" gridcol="8" gridrow="3" pageId="11" pageNumber="162">7.00</td>
<td id="C66E9563FF8F8FB8A9C1FD7CFAE0FD18" box="[1263,1320,676,701]" gridcol="9" gridrow="3" pageId="11" pageNumber="162">7.00</td>
<td id="C66E9563FF8F8FB8A861FD7CFA40FD18" box="[1359,1416,676,701]" gridcol="10" gridrow="3" pageId="11" pageNumber="162">5.71</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FD16FA40FD42" box="[167,1416,718,743]" gridrow="4" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50FD16FDD1FD42" box="[382,537,718,743]" gridcol="1" gridrow="4" pageId="11" pageNumber="162">Anterior width</td>
<td id="C66E9563FF8F8FB8AF15FD16FDBCFD42" box="[571,628,718,743]" gridcol="2" gridrow="4" pageId="11" pageNumber="162">4.14</td>
<td id="C66E9563FF8F8FB8AF8BFD16FD16FD42" box="[677,734,718,743]" gridcol="3" gridrow="4" pageId="11" pageNumber="162">4.57</td>
<td id="C66E9563FF8F8FB8AE2AFD16FCF5FD42" box="[772,829,718,743]" gridcol="4" gridrow="4" pageId="11" pageNumber="162">4.00</td>
<td id="C66E9563FF8F8FB8AE4AFD16FC55FD42" box="[868,925,718,743]" gridcol="5" gridrow="4" pageId="11" pageNumber="162">4.71</td>
<td id="C66E9563FF8F8FB8AEEAFD16FC35FD42" box="[964,1021,718,743]" gridcol="6" gridrow="4" pageId="11" pageNumber="162">4.65</td>
<td id="C66E9563FF8F8FB8A90AFD16FB95FD42" box="[1060,1117,718,743]" gridcol="7" gridrow="4" pageId="11" pageNumber="162">4.75</td>
<td id="C66E9563FF8F8FB8A9AAFD16FB75FD42" box="[1156,1213,718,743]" gridcol="8" gridrow="4" pageId="11" pageNumber="162">5.00</td>
<td id="C66E9563FF8F8FB8A9C1FD16FAE0FD42" box="[1263,1320,718,743]" gridcol="9" gridrow="4" pageId="11" pageNumber="162">4.86</td>
<td id="C66E9563FF8F8FB8A861FD16FA40FD42" box="[1359,1416,718,743]" gridcol="10" gridrow="4" pageId="11" pageNumber="162">4.14</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FD20FA40FCB4" box="[167,1416,760,785]" gridrow="5" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50FD20FDD1FCB4" box="[382,537,760,785]" gridcol="1" gridrow="5" pageId="11" pageNumber="162">Posterior width</td>
<td id="C66E9563FF8F8FB8AF15FD20FDBCFCB4" box="[571,628,760,785]" gridcol="2" gridrow="5" pageId="11" pageNumber="162">6.29</td>
<td id="C66E9563FF8F8FB8AF8BFD20FD16FCB4" box="[677,734,760,785]" gridcol="3" gridrow="5" pageId="11" pageNumber="162">7.00</td>
<td id="C66E9563FF8F8FB8AE2AFD20FCF5FCB4" box="[772,829,760,785]" gridcol="4" gridrow="5" pageId="11" pageNumber="162">6.57</td>
<td id="C66E9563FF8F8FB8AE4AFD20FC55FCB4" box="[868,925,760,785]" gridcol="5" gridrow="5" pageId="11" pageNumber="162">7.29</td>
<td id="C66E9563FF8F8FB8AEEAFD20FC35FCB4" box="[964,1021,760,785]" gridcol="6" gridrow="5" pageId="11" pageNumber="162">7.90</td>
<td id="C66E9563FF8F8FB8A90AFD20FB95FCB4" box="[1060,1117,760,785]" gridcol="7" gridrow="5" pageId="11" pageNumber="162">8.00</td>
<td id="C66E9563FF8F8FB8A9AAFD20FB75FCB4" box="[1156,1213,760,785]" gridcol="8" gridrow="5" pageId="11" pageNumber="162">7.71</td>
<td id="C66E9563FF8F8FB8A9C1FD20FAE0FCB4" box="[1263,1320,760,785]" gridcol="9" gridrow="5" pageId="11" pageNumber="162">7.86</td>
<td id="C66E9563FF8F8FB8A861FD20FA40FCB4" box="[1359,1416,760,785]" gridcol="10" gridrow="5" pageId="11" pageNumber="162">6.29</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FCFAFA40FC9E" box="[167,1416,802,827]" gridrow="6" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50FCFAFDD1FC9E" box="[382,537,802,827]" gridcol="1" gridrow="6" pageId="11" pageNumber="162">Eye diameter</td>
<td id="C66E9563FF8F8FB8AF15FCFAFDBCFC9E" box="[571,628,802,827]" gridcol="2" gridrow="6" pageId="11" pageNumber="162">0.50</td>
<td id="C66E9563FF8F8FB8AF8BFCFAFD16FC9E" box="[677,734,802,827]" gridcol="3" gridrow="6" pageId="11" pageNumber="162">0.50</td>
<td id="C66E9563FF8F8FB8AE2AFCFAFCF5FC9E" box="[772,829,802,827]" gridcol="4" gridrow="6" pageId="11" pageNumber="162">0.50</td>
<td id="C66E9563FF8F8FB8AE4AFCFAFC55FC9E" box="[868,925,802,827]" gridcol="5" gridrow="6" pageId="11" pageNumber="162">0.53</td>
<td id="C66E9563FF8F8FB8AEEAFCFAFC35FC9E" box="[964,1021,802,827]" gridcol="6" gridrow="6" pageId="11" pageNumber="162">0.60</td>
<td id="C66E9563FF8F8FB8A90AFCFAFB95FC9E" box="[1060,1117,802,827]" gridcol="7" gridrow="6" pageId="11" pageNumber="162">0.55</td>
<td id="C66E9563FF8F8FB8A9AAFCFAFB75FC9E" box="[1156,1213,802,827]" gridcol="8" gridrow="6" pageId="11" pageNumber="162">0.63</td>
<td id="C66E9563FF8F8FB8A9C1FCFAFAE0FC9E" box="[1263,1320,802,827]" gridcol="9" gridrow="6" pageId="11" pageNumber="162">0.63</td>
<td id="C66E9563FF8F8FB8A861FCFAFA40FC9E" box="[1359,1416,802,827]" gridcol="10" gridrow="6" pageId="11" pageNumber="162">0.60</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FC94FA40FC2D" box="[167,1416,844,904]" gridrow="7" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50FC94FDD1FC2D" box="[382,537,844,904]" gridcol="1" gridrow="7" pageId="11" pageNumber="162">Interocular distance</td>
<td id="C66E9563FF8F8FB8AF15FC94FDBCFC2D" box="[571,628,844,904]" gridcol="2" gridrow="7" pageId="11" pageNumber="162">0.57</td>
<td id="C66E9563FF8F8FB8AF8BFC94FD16FC2D" box="[677,734,844,904]" gridcol="3" gridrow="7" pageId="11" pageNumber="162">0.70</td>
<td id="C66E9563FF8F8FB8AE2AFC94FCF5FC2D" box="[772,829,844,904]" gridcol="4" gridrow="7" pageId="11" pageNumber="162">0.60</td>
<td id="C66E9563FF8F8FB8AE4AFC94FC55FC2D" box="[868,925,844,904]" gridcol="5" gridrow="7" pageId="11" pageNumber="162">0.67</td>
<td id="C66E9563FF8F8FB8AEEAFC94FC35FC2D" box="[964,1021,844,904]" gridcol="6" gridrow="7" pageId="11" pageNumber="162">0.70</td>
<td id="C66E9563FF8F8FB8A90AFC94FB95FC2D" box="[1060,1117,844,904]" gridcol="7" gridrow="7" pageId="11" pageNumber="162">0.70</td>
<td id="C66E9563FF8F8FB8A9AAFC94FB75FC2D" box="[1156,1213,844,904]" gridcol="8" gridrow="7" pageId="11" pageNumber="162">0.57</td>
<td id="C66E9563FF8F8FB8A9C1FC94FAE0FC2D" box="[1263,1320,844,904]" gridcol="9" gridrow="7" pageId="11" pageNumber="162">0.57</td>
<td id="C66E9563FF8F8FB8A861FC94FA40FC2D" box="[1359,1416,844,904]" gridcol="10" gridrow="7" pageId="11" pageNumber="162">0.60</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FC41FA40FC70" box="[167,1416,921,981]" gridrow="8" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50FC41FDD1FC70" box="[382,537,921,981]" gridcol="1" gridrow="8" pageId="11" pageNumber="162">ocular diad width</td>
<td id="C66E9563FF8F8FB8AF15FC41FDBCFC70" box="[571,628,921,981]" gridcol="2" gridrow="8" pageId="11" pageNumber="162">1.23</td>
<td id="C66E9563FF8F8FB8AF8BFC41FD16FC70" box="[677,734,921,981]" gridcol="3" gridrow="8" pageId="11" pageNumber="162">1.43</td>
<td id="C66E9563FF8F8FB8AE2AFC41FCF5FC70" box="[772,829,921,981]" gridcol="4" gridrow="8" pageId="11" pageNumber="162">1.30</td>
<td id="C66E9563FF8F8FB8AE4AFC41FC55FC70" box="[868,925,921,981]" gridcol="5" gridrow="8" pageId="11" pageNumber="162">1.43</td>
<td id="C66E9563FF8F8FB8AEEAFC41FC35FC70" box="[964,1021,921,981]" gridcol="6" gridrow="8" pageId="11" pageNumber="162">1.40</td>
<td id="C66E9563FF8F8FB8A90AFC41FB95FC70" box="[1060,1117,921,981]" gridcol="7" gridrow="8" pageId="11" pageNumber="162">1.40</td>
<td id="C66E9563FF8F8FB8A9AAFC41FB75FC70" box="[1156,1213,921,981]" gridcol="8" gridrow="8" pageId="11" pageNumber="162">1.57</td>
<td id="C66E9563FF8F8FB8A9C1FC41FAE0FC70" box="[1263,1320,921,981]" gridcol="9" gridrow="8" pageId="11" pageNumber="162">1.53</td>
<td id="C66E9563FF8F8FB8A861FC41FA40FC70" box="[1359,1416,921,981]" gridcol="10" gridrow="8" pageId="11" pageNumber="162">1.37</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FC3DFA40FC58" box="[167,1416,997,1021]" gridrow="9" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89FC3DFEF9FC58" box="[167,305,997,1021]" gridcol="0" gridrow="9" pageId="11" pageNumber="162">Tergite I</th>
<td id="C66E9563FF8F8FB8AC50FC3DFDD1FC58" box="[382,537,997,1021]" gridcol="1" gridrow="9" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15FC3DFDBCFC58" box="[571,628,997,1021]" gridcol="2" gridrow="9" pageId="11" pageNumber="162">1.00</td>
<td id="C66E9563FF8F8FB8AF8BFC3DFD16FC58" box="[677,734,997,1021]" gridcol="3" gridrow="9" pageId="11" pageNumber="162">1.07</td>
<td id="C66E9563FF8F8FB8AE2AFC3DFCF5FC58" box="[772,829,997,1021]" gridcol="4" gridrow="9" pageId="11" pageNumber="162">1.07</td>
<td id="C66E9563FF8F8FB8AE4AFC3DFC55FC58" box="[868,925,997,1021]" gridcol="5" gridrow="9" pageId="11" pageNumber="162">1.13</td>
<td id="C66E9563FF8F8FB8AEEAFC3DFC35FC58" box="[964,1021,997,1021]" gridcol="6" gridrow="9" pageId="11" pageNumber="162">1.50</td>
<td id="C66E9563FF8F8FB8A90AFC3DFB95FC58" box="[1060,1117,997,1021]" gridcol="7" gridrow="9" pageId="11" pageNumber="162">1.05</td>
<td id="C66E9563FF8F8FB8A9AAFC3DFB75FC58" box="[1156,1213,997,1021]" gridcol="8" gridrow="9" pageId="11" pageNumber="162">1.13</td>
<td id="C66E9563FF8F8FB8A9C1FC3DFAE0FC58" box="[1263,1320,997,1021]" gridcol="9" gridrow="9" pageId="11" pageNumber="162">1.27</td>
<td id="C66E9563FF8F8FB8A861FC3DFA40FC58" box="[1359,1416,997,1021]" gridcol="10" gridrow="9" pageId="11" pageNumber="162">1.00</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FBD7FA40FB82" box="[167,1416,1039,1063]" gridrow="10" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89FBD7FEF9FB82" box="[167,305,1039,1063]" gridcol="0" gridrow="10" pageId="11" pageNumber="162">Tergite II</th>
<td id="C66E9563FF8F8FB8AC50FBD7FDD1FB82" box="[382,537,1039,1063]" gridcol="1" gridrow="10" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15FBD7FDBCFB82" box="[571,628,1039,1063]" gridcol="2" gridrow="10" pageId="11" pageNumber="162">1.33</td>
<td id="C66E9563FF8F8FB8AF8BFBD7FD16FB82" box="[677,734,1039,1063]" gridcol="3" gridrow="10" pageId="11" pageNumber="162">1.53</td>
<td id="C66E9563FF8F8FB8AE2AFBD7FCF5FB82" box="[772,829,1039,1063]" gridcol="4" gridrow="10" pageId="11" pageNumber="162">1.47</td>
<td id="C66E9563FF8F8FB8AE4AFBD7FC55FB82" box="[868,925,1039,1063]" gridcol="5" gridrow="10" pageId="11" pageNumber="162">1.53</td>
<td id="C66E9563FF8F8FB8AEEAFBD7FC35FB82" box="[964,1021,1039,1063]" gridcol="6" gridrow="10" pageId="11" pageNumber="162">1.55</td>
<td id="C66E9563FF8F8FB8A90AFBD7FB95FB82" box="[1060,1117,1039,1063]" gridcol="7" gridrow="10" pageId="11" pageNumber="162">1.00</td>
<td id="C66E9563FF8F8FB8A9AAFBD7FB75FB82" box="[1156,1213,1039,1063]" gridcol="8" gridrow="10" pageId="11" pageNumber="162">1.47</td>
<td id="C66E9563FF8F8FB8A9C1FBD7FAE0FB82" box="[1263,1320,1039,1063]" gridcol="9" gridrow="10" pageId="11" pageNumber="162">1.60</td>
<td id="C66E9563FF8F8FB8A861FBD7FA40FB82" box="[1359,1416,1039,1063]" gridcol="10" gridrow="10" pageId="11" pageNumber="162">1.20</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FBE1FA40FBF4" box="[167,1416,1081,1105]" gridrow="11" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89FBE1FEF9FBF4" box="[167,305,1081,1105]" gridcol="0" gridrow="11" pageId="11" pageNumber="162">Tergite III</th>
<td id="C66E9563FF8F8FB8AC50FBE1FDD1FBF4" box="[382,537,1081,1105]" gridcol="1" gridrow="11" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15FBE1FDBCFBF4" box="[571,628,1081,1105]" gridcol="2" gridrow="11" pageId="11" pageNumber="162">1.73</td>
<td id="C66E9563FF8F8FB8AF8BFBE1FD16FBF4" box="[677,734,1081,1105]" gridcol="3" gridrow="11" pageId="11" pageNumber="162">2.00</td>
<td id="C66E9563FF8F8FB8AE2AFBE1FCF5FBF4" box="[772,829,1081,1105]" gridcol="4" gridrow="11" pageId="11" pageNumber="162">1.80</td>
<td id="C66E9563FF8F8FB8AE4AFBE1FC55FBF4" box="[868,925,1081,1105]" gridcol="5" gridrow="11" pageId="11" pageNumber="162">2.07</td>
<td id="C66E9563FF8F8FB8AEEAFBE1FC35FBF4" box="[964,1021,1081,1105]" gridcol="6" gridrow="11" pageId="11" pageNumber="162">2.15</td>
<td id="C66E9563FF8F8FB8A90AFBE1FB95FBF4" box="[1060,1117,1081,1105]" gridcol="7" gridrow="11" pageId="11" pageNumber="162">2.00</td>
<td id="C66E9563FF8F8FB8A9AAFBE1FB75FBF4" box="[1156,1213,1081,1105]" gridcol="8" gridrow="11" pageId="11" pageNumber="162">1.80</td>
<td id="C66E9563FF8F8FB8A9C1FBE1FAE0FBF4" box="[1263,1320,1081,1105]" gridcol="9" gridrow="11" pageId="11" pageNumber="162">2.07</td>
<td id="C66E9563FF8F8FB8A861FBE1FA40FBF4" box="[1359,1416,1081,1105]" gridcol="10" gridrow="11" pageId="11" pageNumber="162">1.53</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FBBBFA40FBDE" box="[167,1416,1123,1147]" gridrow="12" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89FBBBFEF9FBDE" box="[167,305,1123,1147]" gridcol="0" gridrow="12" pageId="11" pageNumber="162">Tergite IV</th>
<td id="C66E9563FF8F8FB8AC50FBBBFDD1FBDE" box="[382,537,1123,1147]" gridcol="1" gridrow="12" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15FBBBFDBCFBDE" box="[571,628,1123,1147]" gridcol="2" gridrow="12" pageId="11" pageNumber="162">2.20</td>
<td id="C66E9563FF8F8FB8AF8BFBBBFD16FBDE" box="[677,734,1123,1147]" gridcol="3" gridrow="12" pageId="11" pageNumber="162">2.33</td>
<td id="C66E9563FF8F8FB8AE2AFBBBFCF5FBDE" box="[772,829,1123,1147]" gridcol="4" gridrow="12" pageId="11" pageNumber="162">2.27</td>
<td id="C66E9563FF8F8FB8AE4AFBBBFC55FBDE" box="[868,925,1123,1147]" gridcol="5" gridrow="12" pageId="11" pageNumber="162">2.53</td>
<td id="C66E9563FF8F8FB8AEEAFBBBFC35FBDE" box="[964,1021,1123,1147]" gridcol="6" gridrow="12" pageId="11" pageNumber="162">2.50</td>
<td id="C66E9563FF8F8FB8A90AFBBBFB95FBDE" box="[1060,1117,1123,1147]" gridcol="7" gridrow="12" pageId="11" pageNumber="162">2.00</td>
<td id="C66E9563FF8F8FB8A9AAFBBBFB75FBDE" box="[1156,1213,1123,1147]" gridcol="8" gridrow="12" pageId="11" pageNumber="162">2.40</td>
<td id="C66E9563FF8F8FB8A9C1FBBBFAE0FBDE" box="[1263,1320,1123,1147]" gridcol="9" gridrow="12" pageId="11" pageNumber="162">2.67</td>
<td id="C66E9563FF8F8FB8A861FBBBFA40FBDE" box="[1359,1416,1123,1147]" gridcol="10" gridrow="12" pageId="11" pageNumber="162">1.93</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FB55FA40FB00" box="[167,1416,1165,1189]" gridrow="13" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89FB55FEF9FB00" box="[167,305,1165,1189]" gridcol="0" gridrow="13" pageId="11" pageNumber="162">Tergite V</th>
<td id="C66E9563FF8F8FB8AC50FB55FDD1FB00" box="[382,537,1165,1189]" gridcol="1" gridrow="13" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15FB55FDBCFB00" box="[571,628,1165,1189]" gridcol="2" gridrow="13" pageId="11" pageNumber="162">2.33</td>
<td id="C66E9563FF8F8FB8AF8BFB55FD16FB00" box="[677,734,1165,1189]" gridcol="3" gridrow="13" pageId="11" pageNumber="162">2.73</td>
<td id="C66E9563FF8F8FB8AE2AFB55FCF5FB00" box="[772,829,1165,1189]" gridcol="4" gridrow="13" pageId="11" pageNumber="162">2.53</td>
<td id="C66E9563FF8F8FB8AE4AFB55FC55FB00" box="[868,925,1165,1189]" gridcol="5" gridrow="13" pageId="11" pageNumber="162">2.80</td>
<td id="C66E9563FF8F8FB8AEEAFB55FC35FB00" box="[964,1021,1165,1189]" gridcol="6" gridrow="13" pageId="11" pageNumber="162">3.00</td>
<td id="C66E9563FF8F8FB8A90AFB55FB95FB00" box="[1060,1117,1165,1189]" gridcol="7" gridrow="13" pageId="11" pageNumber="162">2.50</td>
<td id="C66E9563FF8F8FB8A9AAFB55FB75FB00" box="[1156,1213,1165,1189]" gridcol="8" gridrow="13" pageId="11" pageNumber="162">2.80</td>
<td id="C66E9563FF8F8FB8A9C1FB55FAE0FB00" box="[1263,1320,1165,1189]" gridcol="9" gridrow="13" pageId="11" pageNumber="162">2.80</td>
<td id="C66E9563FF8F8FB8A861FB55FA40FB00" box="[1359,1416,1165,1189]" gridcol="10" gridrow="13" pageId="11" pageNumber="162">2.13</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FB6FFA40FB6A" box="[167,1416,1207,1231]" gridrow="14" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89FB6FFEF9FB6A" box="[167,305,1207,1231]" gridcol="0" gridrow="14" pageId="11" pageNumber="162">Tergite VI</th>
<td id="C66E9563FF8F8FB8AC50FB6FFDD1FB6A" box="[382,537,1207,1231]" gridcol="1" gridrow="14" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15FB6FFDBCFB6A" box="[571,628,1207,1231]" gridcol="2" gridrow="14" pageId="11" pageNumber="162">2.47</td>
<td id="C66E9563FF8F8FB8AF8BFB6FFD16FB6A" box="[677,734,1207,1231]" gridcol="3" gridrow="14" pageId="11" pageNumber="162">3.00</td>
<td id="C66E9563FF8F8FB8AE2AFB6FFCF5FB6A" box="[772,829,1207,1231]" gridcol="4" gridrow="14" pageId="11" pageNumber="162">2.80</td>
<td id="C66E9563FF8F8FB8AE4AFB6FFC55FB6A" box="[868,925,1207,1231]" gridcol="5" gridrow="14" pageId="11" pageNumber="162">3.00</td>
<td id="C66E9563FF8F8FB8AEEAFB6FFC35FB6A" box="[964,1021,1207,1231]" gridcol="6" gridrow="14" pageId="11" pageNumber="162">3.60</td>
<td id="C66E9563FF8F8FB8A90AFB6FFB95FB6A" box="[1060,1117,1207,1231]" gridcol="7" gridrow="14" pageId="11" pageNumber="162">2.50</td>
<td id="C66E9563FF8F8FB8A9AAFB6FFB75FB6A" box="[1156,1213,1207,1231]" gridcol="8" gridrow="14" pageId="11" pageNumber="162">3.07</td>
<td id="C66E9563FF8F8FB8A9C1FB6FFAE0FB6A" box="[1263,1320,1207,1231]" gridcol="9" gridrow="14" pageId="11" pageNumber="162">3.07</td>
<td id="C66E9563FF8F8FB8A861FB6FFA40FB6A" box="[1359,1416,1207,1231]" gridcol="10" gridrow="14" pageId="11" pageNumber="162">2.27</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FB39FA40FB5C" box="[167,1416,1249,1273]" gridrow="15" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89FB39FEF9FB5C" box="[167,305,1249,1273]" gridcol="0" gridrow="15" pageId="11" pageNumber="162">Tergite VII</th>
<td id="C66E9563FF8F8FB8AC50FB39FDD1FB5C" box="[382,537,1249,1273]" gridcol="1" gridrow="15" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15FB39FDBCFB5C" box="[571,628,1249,1273]" gridcol="2" gridrow="15" pageId="11" pageNumber="162">4.27</td>
<td id="C66E9563FF8F8FB8AF8BFB39FD16FB5C" box="[677,734,1249,1273]" gridcol="3" gridrow="15" pageId="11" pageNumber="162">4.73</td>
<td id="C66E9563FF8F8FB8AE2AFB39FCF5FB5C" box="[772,829,1249,1273]" gridcol="4" gridrow="15" pageId="11" pageNumber="162">4.33</td>
<td id="C66E9563FF8F8FB8AE4AFB39FC55FB5C" box="[868,925,1249,1273]" gridcol="5" gridrow="15" pageId="11" pageNumber="162">4.73</td>
<td id="C66E9563FF8F8FB8AEEAFB39FC35FB5C" box="[964,1021,1249,1273]" gridcol="6" gridrow="15" pageId="11" pageNumber="162">4.75</td>
<td id="C66E9563FF8F8FB8A90AFB39FB95FB5C" box="[1060,1117,1249,1273]" gridcol="7" gridrow="15" pageId="11" pageNumber="162">4.25</td>
<td id="C66E9563FF8F8FB8A9AAFB39FB75FB5C" box="[1156,1213,1249,1273]" gridcol="8" gridrow="15" pageId="11" pageNumber="162">4.87</td>
<td id="C66E9563FF8F8FB8A9C1FB39FAE0FB5C" box="[1263,1320,1249,1273]" gridcol="9" gridrow="15" pageId="11" pageNumber="162">4.93</td>
<td id="C66E9563FF8F8FB8A861FB39FA40FB5C" box="[1359,1416,1249,1273]" gridcol="10" gridrow="15" pageId="11" pageNumber="162">4.13</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FAD3FA40FA86" box="[167,1416,1291,1315]" gridrow="16" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89FAD3FEF9FA86" box="[167,305,1291,1315]" gridcol="0" gridrow="16" pageId="11" pageNumber="162">Mesosoma</th>
<td id="C66E9563FF8F8FB8AC50FAD3FDD1FA86" box="[382,537,1291,1315]" gridcol="1" gridrow="16" pageId="11" pageNumber="162">Total Length</td>
<td id="C66E9563FF8F8FB8AF15FAD3FDBCFA86" box="[571,628,1291,1315]" gridcol="2" gridrow="16" pageId="11" pageNumber="162">15.33</td>
<td id="C66E9563FF8F8FB8AF8BFAD3FD16FA86" box="[677,734,1291,1315]" gridcol="3" gridrow="16" pageId="11" pageNumber="162">17.40</td>
<td id="C66E9563FF8F8FB8AE2AFAD3FCF5FA86" box="[772,829,1291,1315]" gridcol="4" gridrow="16" pageId="11" pageNumber="162">16.27</td>
<td id="C66E9563FF8F8FB8AE4AFAD3FC55FA86" box="[868,925,1291,1315]" gridcol="5" gridrow="16" pageId="11" pageNumber="162">17.80</td>
<td id="C66E9563FF8F8FB8AEEAFAD3FC35FA86" box="[964,1021,1291,1315]" gridcol="6" gridrow="16" pageId="11" pageNumber="162">19.05</td>
<td id="C66E9563FF8F8FB8A90AFAD3FB95FA86" box="[1060,1117,1291,1315]" gridcol="7" gridrow="16" pageId="11" pageNumber="162">15.30</td>
<td id="C66E9563FF8F8FB8A9AAFAD3FB75FA86" box="[1156,1213,1291,1315]" gridcol="8" gridrow="16" pageId="11" pageNumber="162">17.53</td>
<td id="C66E9563FF8F8FB8A9C1FAD3FAE0FA86" box="[1263,1320,1291,1315]" gridcol="9" gridrow="16" pageId="11" pageNumber="162">18.40</td>
<td id="C66E9563FF8F8FB8A861FAD3FA40FA86" box="[1359,1416,1291,1315]" gridcol="10" gridrow="16" pageId="11" pageNumber="162">14.20</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FAEDFA40FAE8" box="[167,1416,1333,1357]" gridrow="17" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89FAEDFEF9FAE8" box="[167,305,1333,1357]" gridcol="0" gridrow="17" pageId="11" pageNumber="162">Metasoma I</th>
<td id="C66E9563FF8F8FB8AC50FAEDFDD1FAE8" box="[382,537,1333,1357]" gridcol="1" gridrow="17" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15FAEDFDBCFAE8" box="[571,628,1333,1357]" gridcol="2" gridrow="17" pageId="11" pageNumber="162">4.20</td>
<td id="C66E9563FF8F8FB8AF8BFAEDFD16FAE8" box="[677,734,1333,1357]" gridcol="3" gridrow="17" pageId="11" pageNumber="162">4.20</td>
<td id="C66E9563FF8F8FB8AE2AFAEDFCF5FAE8" box="[772,829,1333,1357]" gridcol="4" gridrow="17" pageId="11" pageNumber="162">4.00</td>
<td id="C66E9563FF8F8FB8AE4AFAEDFC55FAE8" box="[868,925,1333,1357]" gridcol="5" gridrow="17" pageId="11" pageNumber="162">4.10</td>
<td id="C66E9563FF8F8FB8AEEAFAEDFC35FAE8" box="[964,1021,1333,1357]" gridcol="6" gridrow="17" pageId="11" pageNumber="162">4.00</td>
<td id="C66E9563FF8F8FB8A90AFAEDFB95FAE8" box="[1060,1117,1333,1357]" gridcol="7" gridrow="17" pageId="11" pageNumber="162">4.50</td>
<td id="C66E9563FF8F8FB8A9AAFAEDFB75FAE8" box="[1156,1213,1333,1357]" gridcol="8" gridrow="17" pageId="11" pageNumber="162">4.30</td>
<td id="C66E9563FF8F8FB8A9C1FAEDFAE0FAE8" box="[1263,1320,1333,1357]" gridcol="9" gridrow="17" pageId="11" pageNumber="162">4.50</td>
<td id="C66E9563FF8F8FB8A861FAEDFA40FAE8" box="[1359,1416,1333,1357]" gridcol="10" gridrow="17" pageId="11" pageNumber="162">4.10</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FA87FA40FAD2" box="[167,1416,1375,1399]" gridrow="18" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50FA87FDD1FAD2" box="[382,537,1375,1399]" gridcol="1" gridrow="18" pageId="11" pageNumber="162">Width</td>
<td id="C66E9563FF8F8FB8AF15FA87FDBCFAD2" box="[571,628,1375,1399]" gridcol="2" gridrow="18" pageId="11" pageNumber="162">3.90</td>
<td id="C66E9563FF8F8FB8AF8BFA87FD16FAD2" box="[677,734,1375,1399]" gridcol="3" gridrow="18" pageId="11" pageNumber="162">3.90</td>
<td id="C66E9563FF8F8FB8AE2AFA87FCF5FAD2" box="[772,829,1375,1399]" gridcol="4" gridrow="18" pageId="11" pageNumber="162">3.60</td>
<td id="C66E9563FF8F8FB8AE4AFA87FC55FAD2" box="[868,925,1375,1399]" gridcol="5" gridrow="18" pageId="11" pageNumber="162">4.00</td>
<td id="C66E9563FF8F8FB8AEEAFA87FC35FAD2" box="[964,1021,1375,1399]" gridcol="6" gridrow="18" pageId="11" pageNumber="162">4.10</td>
<td id="C66E9563FF8F8FB8A90AFA87FB95FAD2" box="[1060,1117,1375,1399]" gridcol="7" gridrow="18" pageId="11" pageNumber="162">4.30</td>
<td id="C66E9563FF8F8FB8A9AAFA87FB75FAD2" box="[1156,1213,1375,1399]" gridcol="8" gridrow="18" pageId="11" pageNumber="162">4.00</td>
<td id="C66E9563FF8F8FB8A9C1FA87FAE0FAD2" box="[1263,1320,1375,1399]" gridcol="9" gridrow="18" pageId="11" pageNumber="162">4.10</td>
<td id="C66E9563FF8F8FB8A861FA87FA40FAD2" box="[1359,1416,1375,1399]" gridcol="10" gridrow="18" pageId="11" pageNumber="162">3.70</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FA51FA40FA04" box="[167,1416,1417,1441]" gridrow="19" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50FA51FDD1FA04" box="[382,537,1417,1441]" gridcol="1" gridrow="19" pageId="11" pageNumber="162">Height</td>
<td id="C66E9563FF8F8FB8AF15FA51FDBCFA04" box="[571,628,1417,1441]" gridcol="2" gridrow="19" pageId="11" pageNumber="162">3.20</td>
<td id="C66E9563FF8F8FB8AF8BFA51FD16FA04" box="[677,734,1417,1441]" gridcol="3" gridrow="19" pageId="11" pageNumber="162">3.50</td>
<td id="C66E9563FF8F8FB8AE2AFA51FCF5FA04" box="[772,829,1417,1441]" gridcol="4" gridrow="19" pageId="11" pageNumber="162">3.20</td>
<td id="C66E9563FF8F8FB8AE4AFA51FC55FA04" box="[868,925,1417,1441]" gridcol="5" gridrow="19" pageId="11" pageNumber="162">3.40</td>
<td id="C66E9563FF8F8FB8AEEAFA51FC35FA04" box="[964,1021,1417,1441]" gridcol="6" gridrow="19" pageId="11" pageNumber="162">3.50</td>
<td id="C66E9563FF8F8FB8A90AFA51FB95FA04" box="[1060,1117,1417,1441]" gridcol="7" gridrow="19" pageId="11" pageNumber="162">3.50</td>
<td id="C66E9563FF8F8FB8A9AAFA51FB75FA04" box="[1156,1213,1417,1441]" gridcol="8" gridrow="19" pageId="11" pageNumber="162">3.60</td>
<td id="C66E9563FF8F8FB8A9C1FA51FAE0FA04" box="[1263,1320,1417,1441]" gridcol="9" gridrow="19" pageId="11" pageNumber="162">3.80</td>
<td id="C66E9563FF8F8FB8A861FA51FA40FA04" box="[1359,1416,1417,1441]" gridcol="10" gridrow="19" pageId="11" pageNumber="162">3.60</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FA6BFA40FA6E" box="[167,1416,1459,1483]" gridrow="20" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89FA6BFEF9FA6E" box="[167,305,1459,1483]" gridcol="0" gridrow="20" pageId="11" pageNumber="162">Metasoma II</th>
<td id="C66E9563FF8F8FB8AC50FA6BFDD1FA6E" box="[382,537,1459,1483]" gridcol="1" gridrow="20" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15FA6BFDBCFA6E" box="[571,628,1459,1483]" gridcol="2" gridrow="20" pageId="11" pageNumber="162">5.10</td>
<td id="C66E9563FF8F8FB8AF8BFA6BFD16FA6E" box="[677,734,1459,1483]" gridcol="3" gridrow="20" pageId="11" pageNumber="162">5.10</td>
<td id="C66E9563FF8F8FB8AE2AFA6BFCF5FA6E" box="[772,829,1459,1483]" gridcol="4" gridrow="20" pageId="11" pageNumber="162">4.80</td>
<td id="C66E9563FF8F8FB8AE4AFA6BFC55FA6E" box="[868,925,1459,1483]" gridcol="5" gridrow="20" pageId="11" pageNumber="162">5.00</td>
<td id="C66E9563FF8F8FB8AEEAFA6BFC35FA6E" box="[964,1021,1459,1483]" gridcol="6" gridrow="20" pageId="11" pageNumber="162">5.15</td>
<td id="C66E9563FF8F8FB8A90AFA6BFB95FA6E" box="[1060,1117,1459,1483]" gridcol="7" gridrow="20" pageId="11" pageNumber="162">4.80</td>
<td id="C66E9563FF8F8FB8A9AAFA6BFB75FA6E" box="[1156,1213,1459,1483]" gridcol="8" gridrow="20" pageId="11" pageNumber="162">5.40</td>
<td id="C66E9563FF8F8FB8A9C1FA6BFAE0FA6E" box="[1263,1320,1459,1483]" gridcol="9" gridrow="20" pageId="11" pageNumber="162">5.50</td>
<td id="C66E9563FF8F8FB8A861FA6BFA40FA6E" box="[1359,1416,1459,1483]" gridcol="10" gridrow="20" pageId="11" pageNumber="162">4.90</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89FA05FA40FA50" box="[167,1416,1501,1525]" gridrow="21" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50FA05FDD1FA50" box="[382,537,1501,1525]" gridcol="1" gridrow="21" pageId="11" pageNumber="162">Width</td>
<td id="C66E9563FF8F8FB8AF15FA05FDBCFA50" box="[571,628,1501,1525]" gridcol="2" gridrow="21" pageId="11" pageNumber="162">4.00</td>
<td id="C66E9563FF8F8FB8AF8BFA05FD16FA50" box="[677,734,1501,1525]" gridcol="3" gridrow="21" pageId="11" pageNumber="162">4.10</td>
<td id="C66E9563FF8F8FB8AE2AFA05FCF5FA50" box="[772,829,1501,1525]" gridcol="4" gridrow="21" pageId="11" pageNumber="162">3.70</td>
<td id="C66E9563FF8F8FB8AE4AFA05FC55FA50" box="[868,925,1501,1525]" gridcol="5" gridrow="21" pageId="11" pageNumber="162">4.10</td>
<td id="C66E9563FF8F8FB8AEEAFA05FC35FA50" box="[964,1021,1501,1525]" gridcol="6" gridrow="21" pageId="11" pageNumber="162">4.40</td>
<td id="C66E9563FF8F8FB8A90AFA05FB95FA50" box="[1060,1117,1501,1525]" gridcol="7" gridrow="21" pageId="11" pageNumber="162">4.45</td>
<td id="C66E9563FF8F8FB8A9AAFA05FB75FA50" box="[1156,1213,1501,1525]" gridcol="8" gridrow="21" pageId="11" pageNumber="162">4.10</td>
<td id="C66E9563FF8F8FB8A9C1FA05FAE0FA50" box="[1263,1320,1501,1525]" gridcol="9" gridrow="21" pageId="11" pageNumber="162">4.30</td>
<td id="C66E9563FF8F8FB8A861FA05FA40FA50" box="[1359,1416,1501,1525]" gridcol="10" gridrow="21" pageId="11" pageNumber="162">3.80</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89F9DFFA40F9BA" box="[167,1416,1543,1567]" gridrow="22" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50F9DFFDD1F9BA" box="[382,537,1543,1567]" gridcol="1" gridrow="22" pageId="11" pageNumber="162">Height</td>
<td id="C66E9563FF8F8FB8AF15F9DFFDBCF9BA" box="[571,628,1543,1567]" gridcol="2" gridrow="22" pageId="11" pageNumber="162">3.40</td>
<td id="C66E9563FF8F8FB8AF8BF9DFFD16F9BA" box="[677,734,1543,1567]" gridcol="3" gridrow="22" pageId="11" pageNumber="162">3.60</td>
<td id="C66E9563FF8F8FB8AE2AF9DFFCF5F9BA" box="[772,829,1543,1567]" gridcol="4" gridrow="22" pageId="11" pageNumber="162">3.30</td>
<td id="C66E9563FF8F8FB8AE4AF9DFFC55F9BA" box="[868,925,1543,1567]" gridcol="5" gridrow="22" pageId="11" pageNumber="162">3.40</td>
<td id="C66E9563FF8F8FB8AEEAF9DFFC35F9BA" box="[964,1021,1543,1567]" gridcol="6" gridrow="22" pageId="11" pageNumber="162">3.60</td>
<td id="C66E9563FF8F8FB8A90AF9DFFB95F9BA" box="[1060,1117,1543,1567]" gridcol="7" gridrow="22" pageId="11" pageNumber="162">3.50</td>
<td id="C66E9563FF8F8FB8A9AAF9DFFB75F9BA" box="[1156,1213,1543,1567]" gridcol="8" gridrow="22" pageId="11" pageNumber="162">3.70</td>
<td id="C66E9563FF8F8FB8A9C1F9DFFAE0F9BA" box="[1263,1320,1543,1567]" gridcol="9" gridrow="22" pageId="11" pageNumber="162">3.90</td>
<td id="C66E9563FF8F8FB8A861F9DFFA40F9BA" box="[1359,1416,1543,1567]" gridcol="10" gridrow="22" pageId="11" pageNumber="162">3.50</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89F9E9FA40F9EC" box="[167,1416,1585,1609]" gridrow="23" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89F9E9FEF9F9EC" box="[167,305,1585,1609]" gridcol="0" gridrow="23" pageId="11" pageNumber="162">Metasoma III</th>
<td id="C66E9563FF8F8FB8AC50F9E9FDD1F9EC" box="[382,537,1585,1609]" gridcol="1" gridrow="23" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15F9E9FDBCF9EC" box="[571,628,1585,1609]" gridcol="2" gridrow="23" pageId="11" pageNumber="162">5.80</td>
<td id="C66E9563FF8F8FB8AF8BF9E9FD16F9EC" box="[677,734,1585,1609]" gridcol="3" gridrow="23" pageId="11" pageNumber="162">5.70</td>
<td id="C66E9563FF8F8FB8AE2AF9E9FCF5F9EC" box="[772,829,1585,1609]" gridcol="4" gridrow="23" pageId="11" pageNumber="162">5.20</td>
<td id="C66E9563FF8F8FB8AE4AF9E9FC55F9EC" box="[868,925,1585,1609]" gridcol="5" gridrow="23" pageId="11" pageNumber="162">5.60</td>
<td id="C66E9563FF8F8FB8AEEAF9E9FC35F9EC" box="[964,1021,1585,1609]" gridcol="6" gridrow="23" pageId="11" pageNumber="162">6.00</td>
<td id="C66E9563FF8F8FB8A90AF9E9FB95F9EC" box="[1060,1117,1585,1609]" gridcol="7" gridrow="23" pageId="11" pageNumber="162">6.00</td>
<td id="C66E9563FF8F8FB8A9AAF9E9FB75F9EC" box="[1156,1213,1585,1609]" gridcol="8" gridrow="23" pageId="11" pageNumber="162">6.10</td>
<td id="C66E9563FF8F8FB8A9C1F9E9FAE0F9EC" box="[1263,1320,1585,1609]" gridcol="9" gridrow="23" pageId="11" pageNumber="162">6.00</td>
<td id="C66E9563FF8F8FB8A861F9E9FA40F9EC" box="[1359,1416,1585,1609]" gridcol="10" gridrow="23" pageId="11" pageNumber="162">5.50</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89F983FA40F9D6" box="[167,1416,1627,1651]" gridrow="24" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50F983FDD1F9D6" box="[382,537,1627,1651]" gridcol="1" gridrow="24" pageId="11" pageNumber="162">Width</td>
<td id="C66E9563FF8F8FB8AF15F983FDBCF9D6" box="[571,628,1627,1651]" gridcol="2" gridrow="24" pageId="11" pageNumber="162">4.50</td>
<td id="C66E9563FF8F8FB8AF8BF983FD16F9D6" box="[677,734,1627,1651]" gridcol="3" gridrow="24" pageId="11" pageNumber="162">4.60</td>
<td id="C66E9563FF8F8FB8AE2AF983FCF5F9D6" box="[772,829,1627,1651]" gridcol="4" gridrow="24" pageId="11" pageNumber="162">4.10</td>
<td id="C66E9563FF8F8FB8AE4AF983FC55F9D6" box="[868,925,1627,1651]" gridcol="5" gridrow="24" pageId="11" pageNumber="162">4.70</td>
<td id="C66E9563FF8F8FB8AEEAF983FC35F9D6" box="[964,1021,1627,1651]" gridcol="6" gridrow="24" pageId="11" pageNumber="162">4.90</td>
<td id="C66E9563FF8F8FB8A90AF983FB95F9D6" box="[1060,1117,1627,1651]" gridcol="7" gridrow="24" pageId="11" pageNumber="162">4.75</td>
<td id="C66E9563FF8F8FB8A9AAF983FB75F9D6" box="[1156,1213,1627,1651]" gridcol="8" gridrow="24" pageId="11" pageNumber="162">4.40</td>
<td id="C66E9563FF8F8FB8A9C1F983FAE0F9D6" box="[1263,1320,1627,1651]" gridcol="9" gridrow="24" pageId="11" pageNumber="162">4.50</td>
<td id="C66E9563FF8F8FB8A861F983FA40F9D6" box="[1359,1416,1627,1651]" gridcol="10" gridrow="24" pageId="11" pageNumber="162">4.00</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89F95DFA40F938" box="[167,1416,1669,1693]" gridrow="25" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50F95DFDD1F938" box="[382,537,1669,1693]" gridcol="1" gridrow="25" pageId="11" pageNumber="162">Height</td>
<td id="C66E9563FF8F8FB8AF15F95DFDBCF938" box="[571,628,1669,1693]" gridcol="2" gridrow="25" pageId="11" pageNumber="162">3.60</td>
<td id="C66E9563FF8F8FB8AF8BF95DFD16F938" box="[677,734,1669,1693]" gridcol="3" gridrow="25" pageId="11" pageNumber="162">4.00</td>
<td id="C66E9563FF8F8FB8AE2AF95DFCF5F938" box="[772,829,1669,1693]" gridcol="4" gridrow="25" pageId="11" pageNumber="162">3.60</td>
<td id="C66E9563FF8F8FB8AE4AF95DFC55F938" box="[868,925,1669,1693]" gridcol="5" gridrow="25" pageId="11" pageNumber="162">4.10</td>
<td id="C66E9563FF8F8FB8AEEAF95DFC35F938" box="[964,1021,1669,1693]" gridcol="6" gridrow="25" pageId="11" pageNumber="162">3.75</td>
<td id="C66E9563FF8F8FB8A90AF95DFB95F938" box="[1060,1117,1669,1693]" gridcol="7" gridrow="25" pageId="11" pageNumber="162">4.00</td>
<td id="C66E9563FF8F8FB8A9AAF95DFB75F938" box="[1156,1213,1669,1693]" gridcol="8" gridrow="25" pageId="11" pageNumber="162">4.10</td>
<td id="C66E9563FF8F8FB8A9C1F95DFAE0F938" box="[1263,1320,1669,1693]" gridcol="9" gridrow="25" pageId="11" pageNumber="162">4.40</td>
<td id="C66E9563FF8F8FB8A861F95DFA40F938" box="[1359,1416,1669,1693]" gridcol="10" gridrow="25" pageId="11" pageNumber="162">3.80</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89F977FA40F962" box="[167,1416,1711,1735]" gridrow="26" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89F977FEF9F962" box="[167,305,1711,1735]" gridcol="0" gridrow="26" pageId="11" pageNumber="162">Metasoma IV</th>
<td id="C66E9563FF8F8FB8AC50F977FDD1F962" box="[382,537,1711,1735]" gridcol="1" gridrow="26" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15F977FDBCF962" box="[571,628,1711,1735]" gridcol="2" gridrow="26" pageId="11" pageNumber="162">6.30</td>
<td id="C66E9563FF8F8FB8AF8BF977FD16F962" box="[677,734,1711,1735]" gridcol="3" gridrow="26" pageId="11" pageNumber="162">6.30</td>
<td id="C66E9563FF8F8FB8AE2AF977FCF5F962" box="[772,829,1711,1735]" gridcol="4" gridrow="26" pageId="11" pageNumber="162">5.50</td>
<td id="C66E9563FF8F8FB8AE4AF977FC55F962" box="[868,925,1711,1735]" gridcol="5" gridrow="26" pageId="11" pageNumber="162">6.40</td>
<td id="C66E9563FF8F8FB8AEEAF977FC35F962" box="[964,1021,1711,1735]" gridcol="6" gridrow="26" pageId="11" pageNumber="162">6.20</td>
<td id="C66E9563FF8F8FB8A90AF977FB95F962" box="[1060,1117,1711,1735]" gridcol="7" gridrow="26" pageId="11" pageNumber="162">6.75</td>
<td id="C66E9563FF8F8FB8A9AAF977FB75F962" box="[1156,1213,1711,1735]" gridcol="8" gridrow="26" pageId="11" pageNumber="162">7.00</td>
<td id="C66E9563FF8F8FB8A9C1F977FAE0F962" box="[1263,1320,1711,1735]" gridcol="9" gridrow="26" pageId="11" pageNumber="162">6.90</td>
<td id="C66E9563FF8F8FB8A861F977FA40F962" box="[1359,1416,1711,1735]" gridcol="10" gridrow="26" pageId="11" pageNumber="162">6.10</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89F901FA40F954" box="[167,1416,1753,1777]" gridrow="27" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50F901FDD1F954" box="[382,537,1753,1777]" gridcol="1" gridrow="27" pageId="11" pageNumber="162">Width</td>
<td id="C66E9563FF8F8FB8AF15F901FDBCF954" box="[571,628,1753,1777]" gridcol="2" gridrow="27" pageId="11" pageNumber="162">5.80</td>
<td id="C66E9563FF8F8FB8AF8BF901FD16F954" box="[677,734,1753,1777]" gridcol="3" gridrow="27" pageId="11" pageNumber="162">5.80</td>
<td id="C66E9563FF8F8FB8AE2AF901FCF5F954" box="[772,829,1753,1777]" gridcol="4" gridrow="27" pageId="11" pageNumber="162">5.20</td>
<td id="C66E9563FF8F8FB8AE4AF901FC55F954" box="[868,925,1753,1777]" gridcol="5" gridrow="27" pageId="11" pageNumber="162">5.80</td>
<td id="C66E9563FF8F8FB8AEEAF901FC35F954" box="[964,1021,1753,1777]" gridcol="6" gridrow="27" pageId="11" pageNumber="162">6.10</td>
<td id="C66E9563FF8F8FB8A90AF901FB95F954" box="[1060,1117,1753,1777]" gridcol="7" gridrow="27" pageId="11" pageNumber="162">5.75</td>
<td id="C66E9563FF8F8FB8A9AAF901FB75F954" box="[1156,1213,1753,1777]" gridcol="8" gridrow="27" pageId="11" pageNumber="162">4.80</td>
<td id="C66E9563FF8F8FB8A9C1F901FAE0F954" box="[1263,1320,1753,1777]" gridcol="9" gridrow="27" pageId="11" pageNumber="162">5.00</td>
<td id="C66E9563FF8F8FB8A861F901FA40F954" box="[1359,1416,1753,1777]" gridcol="10" gridrow="27" pageId="11" pageNumber="162">4.40</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89F8DBFA40F8BE" box="[167,1416,1795,1819]" gridrow="28" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50F8DBFDD1F8BE" box="[382,537,1795,1819]" gridcol="1" gridrow="28" pageId="11" pageNumber="162">Height</td>
<td id="C66E9563FF8F8FB8AF15F8DBFDBCF8BE" box="[571,628,1795,1819]" gridcol="2" gridrow="28" pageId="11" pageNumber="162">4.20</td>
<td id="C66E9563FF8F8FB8AF8BF8DBFD16F8BE" box="[677,734,1795,1819]" gridcol="3" gridrow="28" pageId="11" pageNumber="162">4.30</td>
<td id="C66E9563FF8F8FB8AE2AF8DBFCF5F8BE" box="[772,829,1795,1819]" gridcol="4" gridrow="28" pageId="11" pageNumber="162">3.80</td>
<td id="C66E9563FF8F8FB8AE4AF8DBFC55F8BE" box="[868,925,1795,1819]" gridcol="5" gridrow="28" pageId="11" pageNumber="162">4.40</td>
<td id="C66E9563FF8F8FB8AEEAF8DBFC35F8BE" box="[964,1021,1795,1819]" gridcol="6" gridrow="28" pageId="11" pageNumber="162">4.75</td>
<td id="C66E9563FF8F8FB8A90AF8DBFB95F8BE" box="[1060,1117,1795,1819]" gridcol="7" gridrow="28" pageId="11" pageNumber="162">4.90</td>
<td id="C66E9563FF8F8FB8A9AAF8DBFB75F8BE" box="[1156,1213,1795,1819]" gridcol="8" gridrow="28" pageId="11" pageNumber="162">4.10</td>
<td id="C66E9563FF8F8FB8A9C1F8DBFAE0F8BE" box="[1263,1320,1795,1819]" gridcol="9" gridrow="28" pageId="11" pageNumber="162">4.50</td>
<td id="C66E9563FF8F8FB8A861F8DBFA40F8BE" box="[1359,1416,1795,1819]" gridcol="10" gridrow="28" pageId="11" pageNumber="162">4.00</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89F8F5FA40F8E0" box="[167,1416,1837,1861]" gridrow="29" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89F8F5FEF9F8E0" box="[167,305,1837,1861]" gridcol="0" gridrow="29" pageId="11" pageNumber="162">Metasoma V</th>
<td id="C66E9563FF8F8FB8AC50F8F5FDD1F8E0" box="[382,537,1837,1861]" gridcol="1" gridrow="29" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15F8F5FDBCF8E0" box="[571,628,1837,1861]" gridcol="2" gridrow="29" pageId="11" pageNumber="162">6.50</td>
<td id="C66E9563FF8F8FB8AF8BF8F5FD16F8E0" box="[677,734,1837,1861]" gridcol="3" gridrow="29" pageId="11" pageNumber="162">6.60</td>
<td id="C66E9563FF8F8FB8AE2AF8F5FCF5F8E0" box="[772,829,1837,1861]" gridcol="4" gridrow="29" pageId="11" pageNumber="162">6.20</td>
<td id="C66E9563FF8F8FB8AE4AF8F5FC55F8E0" box="[868,925,1837,1861]" gridcol="5" gridrow="29" pageId="11" pageNumber="162">6.60</td>
<td id="C66E9563FF8F8FB8AEEAF8F5FC35F8E0" box="[964,1021,1837,1861]" gridcol="6" gridrow="29" pageId="11" pageNumber="162">6.90</td>
<td id="C66E9563FF8F8FB8A90AF8F5FB95F8E0" box="[1060,1117,1837,1861]" gridcol="7" gridrow="29" pageId="11" pageNumber="162">7.00</td>
<td id="C66E9563FF8F8FB8A9AAF8F5FB75F8E0" box="[1156,1213,1837,1861]" gridcol="8" gridrow="29" pageId="11" pageNumber="162">7.50</td>
<td id="C66E9563FF8F8FB8A9C1F8F5FAE0F8E0" box="[1263,1320,1837,1861]" gridcol="9" gridrow="29" pageId="11" pageNumber="162">7.40</td>
<td id="C66E9563FF8F8FB8A861F8F5FA40F8E0" box="[1359,1416,1837,1861]" gridcol="10" gridrow="29" pageId="11" pageNumber="162">6.90</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89F88FFA40F8CA" box="[167,1416,1879,1903]" gridrow="30" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50F88FFDD1F8CA" box="[382,537,1879,1903]" gridcol="1" gridrow="30" pageId="11" pageNumber="162">Width</td>
<td id="C66E9563FF8F8FB8AF15F88FFDBCF8CA" box="[571,628,1879,1903]" gridcol="2" gridrow="30" pageId="11" pageNumber="162">6.00</td>
<td id="C66E9563FF8F8FB8AF8BF88FFD16F8CA" box="[677,734,1879,1903]" gridcol="3" gridrow="30" pageId="11" pageNumber="162">6.00</td>
<td id="C66E9563FF8F8FB8AE2AF88FFCF5F8CA" box="[772,829,1879,1903]" gridcol="4" gridrow="30" pageId="11" pageNumber="162">5.70</td>
<td id="C66E9563FF8F8FB8AE4AF88FFC55F8CA" box="[868,925,1879,1903]" gridcol="5" gridrow="30" pageId="11" pageNumber="162">5.90</td>
<td id="C66E9563FF8F8FB8AEEAF88FFC35F8CA" box="[964,1021,1879,1903]" gridcol="6" gridrow="30" pageId="11" pageNumber="162">6.20</td>
<td id="C66E9563FF8F8FB8A90AF88FFB95F8CA" box="[1060,1117,1879,1903]" gridcol="7" gridrow="30" pageId="11" pageNumber="162">6.00</td>
<td id="C66E9563FF8F8FB8A9AAF88FFB75F8CA" box="[1156,1213,1879,1903]" gridcol="8" gridrow="30" pageId="11" pageNumber="162">4.90</td>
<td id="C66E9563FF8F8FB8A9C1F88FFAE0F8CA" box="[1263,1320,1879,1903]" gridcol="9" gridrow="30" pageId="11" pageNumber="162">4.90</td>
<td id="C66E9563FF8F8FB8A861F88FFA40F8CA" box="[1359,1416,1879,1903]" gridcol="10" gridrow="30" pageId="11" pageNumber="162">4.50</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89F859FA40F83C" box="[167,1416,1921,1945]" gridrow="31" pageId="11" pageNumber="162" rowspan-0="1">
<td id="C66E9563FF8F8FB8AC50F859FDD1F83C" box="[382,537,1921,1945]" gridcol="1" gridrow="31" pageId="11" pageNumber="162">Height</td>
<td id="C66E9563FF8F8FB8AF15F859FDBCF83C" box="[571,628,1921,1945]" gridcol="2" gridrow="31" pageId="11" pageNumber="162">4.00</td>
<td id="C66E9563FF8F8FB8AF8BF859FD16F83C" box="[677,734,1921,1945]" gridcol="3" gridrow="31" pageId="11" pageNumber="162">3.90</td>
<td id="C66E9563FF8F8FB8AE2AF859FCF5F83C" box="[772,829,1921,1945]" gridcol="4" gridrow="31" pageId="11" pageNumber="162">3.70</td>
<td id="C66E9563FF8F8FB8AE4AF859FC55F83C" box="[868,925,1921,1945]" gridcol="5" gridrow="31" pageId="11" pageNumber="162">4.40</td>
<td id="C66E9563FF8F8FB8AEEAF859FC35F83C" box="[964,1021,1921,1945]" gridcol="6" gridrow="31" pageId="11" pageNumber="162">4.15</td>
<td id="C66E9563FF8F8FB8A90AF859FB95F83C" box="[1060,1117,1921,1945]" gridcol="7" gridrow="31" pageId="11" pageNumber="162">4.00</td>
<td id="C66E9563FF8F8FB8A9AAF859FB75F83C" box="[1156,1213,1921,1945]" gridcol="8" gridrow="31" pageId="11" pageNumber="162">4.10</td>
<td id="C66E9563FF8F8FB8A9C1F859FAE0F83C" box="[1263,1320,1921,1945]" gridcol="9" gridrow="31" pageId="11" pageNumber="162">3.90</td>
<td id="C66E9563FF8F8FB8A861F859FA40F83C" box="[1359,1416,1921,1945]" gridcol="10" gridrow="31" pageId="11" pageNumber="162">3.70</td>
</tr>
<tr id="85BFFC1FFF8F8FB8AD89F873FA40F866" box="[167,1416,1963,1987]" gridrow="32" pageId="11" pageNumber="162">
<th id="C66E9563FF8F8FB8AD89F873FEF9F866" box="[167,305,1963,1987]" gridcol="0" gridrow="32" pageId="11" pageNumber="162">Metasoma</th>
<td id="C66E9563FF8F8FB8AC50F873FDD1F866" box="[382,537,1963,1987]" gridcol="1" gridrow="32" pageId="11" pageNumber="162">Length</td>
<td id="C66E9563FF8F8FB8AF15F873FDBCF866" box="[571,628,1963,1987]" gridcol="2" gridrow="32" pageId="11" pageNumber="162">27.90</td>
<td id="C66E9563FF8F8FB8AF8BF873FD16F866" box="[677,734,1963,1987]" gridcol="3" gridrow="32" pageId="11" pageNumber="162">27.90</td>
<td id="C66E9563FF8F8FB8AE2AF873FCF5F866" box="[772,829,1963,1987]" gridcol="4" gridrow="32" pageId="11" pageNumber="162">25.70</td>
<td id="C66E9563FF8F8FB8AE4AF873FC55F866" box="[868,925,1963,1987]" gridcol="5" gridrow="32" pageId="11" pageNumber="162">27.70</td>
<td id="C66E9563FF8F8FB8AEEAF873FC35F866" box="[964,1021,1963,1987]" gridcol="6" gridrow="32" pageId="11" pageNumber="162">28.25</td>
<td id="C66E9563FF8F8FB8A90AF873FB95F866" box="[1060,1117,1963,1987]" gridcol="7" gridrow="32" pageId="11" pageNumber="162">29.05</td>
<td id="C66E9563FF8F8FB8A9AAF873FB75F866" box="[1156,1213,1963,1987]" gridcol="8" gridrow="32" pageId="11" pageNumber="162">30.30</td>
<td id="C66E9563FF8F8FB8A9C1F873FAE0F866" box="[1263,1320,1963,1987]" gridcol="9" gridrow="32" pageId="11" pageNumber="162">30.30</td>
<td id="C66E9563FF8F8FB8A861F873FA40F866" box="[1359,1416,1963,1987]" gridcol="10" gridrow="32" pageId="11" pageNumber="162">27.50</td>
</tr>
</table>
</paragraph>
<paragraph id="3B30FE5DFF8F704CA978F80DFA45F848" blockId="11.[1110,1421,2005,2029]" box="[1110,1421,2005,2029]" pageId="11" pageNumber="162">
<tableNote id="C669FFD3FF8F704CA978F80DFA45F848" box="[1110,1421,2005,2029]" pageId="11" pageNumber="162" targetBox="[167,1416,516,1987]" targetPageId="11">......continued on the next page</tableNote>
</paragraph>
<paragraph id="3B30FE5DFF88704BAD89FF79FE55FF1C" blockId="12.[167,413,161,186]" box="[167,413,161,186]" pageId="12" pageNumber="163">TABLE 5. (Continued)</paragraph>
<paragraph id="3B30FE5DFF88704BAD89FF13FAB3FB82" pageId="12" pageNumber="163">
<table id="498F0CFDFF888FB8AD89FF13FA40FB82" box="[167,1416,203,1063]" colsContinueFrom="11.[167,1416,516,1987]" gridcols="11" gridrows="18" pageId="12" pageNumber="163">
<tr id="85BFFC1FFF888FB8AD89FF13FA40FF41" box="[167,1416,203,228]" gridrow="0" pageId="12" pageNumber="163" rowspan-2="1" rowspan-6="1" rowspan-7="1">
<th id="C66E9563FF888FB8AD89FF13FE9FFF41" box="[167,343,203,228]" gridcol="0" gridrow="0" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BAD89FF13FEC7FF46" bold="true" box="[167,271,203,227]" pageId="12" pageNumber="163">Structure</emphasis>
</th>
<th id="C66E9563FF888FB8AC50FF13FDC7FF41" box="[382,527,203,228]" gridcol="1" gridrow="0" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BAC50FF14FE14FF41" bold="true" box="[382,476,204,228]" pageId="12" pageNumber="163">Measure</emphasis>
</th>
<th id="C66E9563FF888FB8AF8BFF13FC55FF41" box="[677,925,203,228]" colspan="3" colspanRight="2" gridcol="3" gridrow="0" pageId="12" pageNumber="163">
<taxonomicName id="FC8F85DEFF88704BAFEAFF13FCB4FF46" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[708,892,203,227]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="moralensis">
<emphasis id="09FB224FFF88704BAFEAFF13FCB4FF46" bold="true" box="[708,892,203,227]" italics="true" pageId="12" pageNumber="163">Tityus moralensis</emphasis>
</taxonomicName>
</th>
<th id="C66E9563FF888FB8A9AAFF13FA40FF41" box="[1156,1416,203,228]" colspan="3" colspanRight="2" gridcol="8" gridrow="0" pageId="12" pageNumber="163">
<taxonomicName id="FC8F85DEFF88704BA99BFF13FA93FF46" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[1205,1371,203,227]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF88704BA99BFF13FA93FF46" bold="true" box="[1205,1371,203,227]" italics="true" pageId="12" pageNumber="163">Tityus forcipula</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="85BFFC1FFF888FB8AD89FF2DFA40FE91" box="[167,1416,245,308]" gridrow="1" pageId="12" pageNumber="163" rowspan-0="1" rowspan-1="1">
<td id="C66E9563FF888FB8AF15FF2DFDBCFE91" box="[571,628,245,308]" gridcol="2" gridrow="1" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BAF15FF2DFDA6FEA8" bold="true" box="[571,622,245,269]" pageId="12" pageNumber="163">2104</emphasis>
♂
</td>
<td id="C66E9563FF888FB8AF8BFF2DFD16FE91" box="[677,734,245,308]" gridcol="3" gridrow="1" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BAF8BFF2DFD70FE91" bold="true" box="[677,728,245,308]" pageId="12" pageNumber="163">1402 ♀</emphasis>
</td>
<td id="C66E9563FF888FB8AE2AFF2DFCF5FE91" box="[772,829,245,308]" gridcol="4" gridrow="1" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BAE2AFF2DFCDFFE91" bold="true" box="[772,823,245,308]" pageId="12" pageNumber="163">1403 ♀</emphasis>
</td>
<td id="C66E9563FF888FB8AE4AFF2DFC55FE91" box="[868,925,245,308]" gridcol="5" gridrow="1" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BAE4AFF2DFCBFFE91" bold="true" box="[868,919,245,308]" pageId="12" pageNumber="163">2108 ♀</emphasis>
</td>
<td id="C66E9563FF888FB8AEEAFF2DFC35FE91" box="[964,1021,245,308]" gridcol="6" gridrow="1" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BAEEAFF2DFC1FFE91" bold="true" box="[964,1015,245,308]" pageId="12" pageNumber="163">2109 ♀</emphasis>
</td>
<td id="C66E9563FF888FB8A90AFF2DFB95FE91" box="[1060,1117,245,308]" gridcol="7" gridrow="1" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BA90AFF2DFBFFFE91" bold="true" box="[1060,1108,245,308]" pageId="12" pageNumber="163">2111 ♀</emphasis>
</td>
<td id="C66E9563FF888FB8A9AAFF2DFB75FE91" box="[1156,1213,245,308]" gridcol="8" gridrow="1" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BA9AAFF2DFB5FFE91" bold="true" box="[1156,1194,245,308]" pageId="12" pageNumber="163">436 ♀</emphasis>
</td>
<td id="C66E9563FF888FB8A9C1FF2DFAE0FE91" box="[1263,1320,245,308]" gridcol="9" gridrow="1" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BA9C1FF2DFACAFE91" bold="true" box="[1263,1314,245,308]" pageId="12" pageNumber="163">2103 ♀</emphasis>
</td>
<td id="C66E9563FF888FB8A861FF2DFA40FE91" box="[1359,1416,245,308]" gridcol="10" gridrow="1" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BA861FF2DFAAAFE91" bold="true" box="[1359,1397,245,308]" pageId="12" pageNumber="163">438 ♂</emphasis>
</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FE9AFA40FEFF" box="[167,1416,322,346]" gridrow="2" pageId="12" pageNumber="163">
<th id="C66E9563FF888FB8AD89FE9AFE9FFEFF" box="[167,343,322,346]" gridcol="0" gridrow="2" pageId="12" pageNumber="163">Telson</th>
<td id="C66E9563FF888FB8AC50FE9AFDC7FEFF" box="[382,527,322,346]" gridcol="1" gridrow="2" pageId="12" pageNumber="163">vesicle length</td>
<td id="C66E9563FF888FB8AF15FE9AFDBCFEFF" box="[571,628,322,346]" gridcol="2" gridrow="2" pageId="12" pageNumber="163">4.00</td>
<td id="C66E9563FF888FB8AF8BFE9AFD16FEFF" box="[677,734,322,346]" gridcol="3" gridrow="2" pageId="12" pageNumber="163">4.20</td>
<td id="C66E9563FF888FB8AE2AFE9AFCF5FEFF" box="[772,829,322,346]" gridcol="4" gridrow="2" pageId="12" pageNumber="163">3.70</td>
<td id="C66E9563FF888FB8AE4AFE9AFC55FEFF" box="[868,925,322,346]" gridcol="5" gridrow="2" pageId="12" pageNumber="163">3.90</td>
<td id="C66E9563FF888FB8AEEAFE9AFC35FEFF" box="[964,1021,322,346]" gridcol="6" gridrow="2" pageId="12" pageNumber="163">4.50</td>
<td id="C66E9563FF888FB8A90AFE9AFB95FEFF" box="[1060,1117,322,346]" gridcol="7" gridrow="2" pageId="12" pageNumber="163">4.55</td>
<td id="C66E9563FF888FB8A9AAFE9AFB75FEFF" box="[1156,1213,322,346]" gridcol="8" gridrow="2" pageId="12" pageNumber="163">4.80</td>
<td id="C66E9563FF888FB8A9C1FE9AFAE0FEFF" box="[1263,1320,322,346]" gridcol="9" gridrow="2" pageId="12" pageNumber="163">4.50</td>
<td id="C66E9563FF888FB8A861FE9AFA40FEFF" box="[1359,1416,322,346]" gridcol="10" gridrow="2" pageId="12" pageNumber="163">4.20</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FEB4FA40FE21" box="[167,1416,364,388]" gridrow="3" pageId="12" pageNumber="163" rowspan-0="1">
<td id="C66E9563FF888FB8AC50FEB4FDC7FE21" box="[382,527,364,388]" gridcol="1" gridrow="3" pageId="12" pageNumber="163">vesicle width</td>
<td id="C66E9563FF888FB8AF15FEB4FDBCFE21" box="[571,628,364,388]" gridcol="2" gridrow="3" pageId="12" pageNumber="163">3.30</td>
<td id="C66E9563FF888FB8AF8BFEB4FD16FE21" box="[677,734,364,388]" gridcol="3" gridrow="3" pageId="12" pageNumber="163">3.20</td>
<td id="C66E9563FF888FB8AE2AFEB4FCF5FE21" box="[772,829,364,388]" gridcol="4" gridrow="3" pageId="12" pageNumber="163">3.20</td>
<td id="C66E9563FF888FB8AE4AFEB4FC55FE21" box="[868,925,364,388]" gridcol="5" gridrow="3" pageId="12" pageNumber="163">3.30</td>
<td id="C66E9563FF888FB8AEEAFEB4FC35FE21" box="[964,1021,364,388]" gridcol="6" gridrow="3" pageId="12" pageNumber="163">3.50</td>
<td id="C66E9563FF888FB8A90AFEB4FB95FE21" box="[1060,1117,364,388]" gridcol="7" gridrow="3" pageId="12" pageNumber="163">3.40</td>
<td id="C66E9563FF888FB8A9AAFEB4FB75FE21" box="[1156,1213,364,388]" gridcol="8" gridrow="3" pageId="12" pageNumber="163">3.20</td>
<td id="C66E9563FF888FB8A9C1FEB4FAE0FE21" box="[1263,1320,364,388]" gridcol="9" gridrow="3" pageId="12" pageNumber="163">3.20</td>
<td id="C66E9563FF888FB8A861FEB4FA40FE21" box="[1359,1416,364,388]" gridcol="10" gridrow="3" pageId="12" pageNumber="163">3.00</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FE4EFA40FE0B" box="[167,1416,406,430]" gridrow="4" pageId="12" pageNumber="163" rowspan-0="1">
<td id="C66E9563FF888FB8AC50FE4EFDC7FE0B" box="[382,527,406,430]" gridcol="1" gridrow="4" pageId="12" pageNumber="163">vesicle height</td>
<td id="C66E9563FF888FB8AF15FE4EFDBCFE0B" box="[571,628,406,430]" gridcol="2" gridrow="4" pageId="12" pageNumber="163">2.50</td>
<td id="C66E9563FF888FB8AF8BFE4EFD16FE0B" box="[677,734,406,430]" gridcol="3" gridrow="4" pageId="12" pageNumber="163">2.50</td>
<td id="C66E9563FF888FB8AE2AFE4EFCF5FE0B" box="[772,829,406,430]" gridcol="4" gridrow="4" pageId="12" pageNumber="163">2.30</td>
<td id="C66E9563FF888FB8AE4AFE4EFC55FE0B" box="[868,925,406,430]" gridcol="5" gridrow="4" pageId="12" pageNumber="163">2.60</td>
<td id="C66E9563FF888FB8AEEAFE4EFC35FE0B" box="[964,1021,406,430]" gridcol="6" gridrow="4" pageId="12" pageNumber="163">2.70</td>
<td id="C66E9563FF888FB8A90AFE4EFB95FE0B" box="[1060,1117,406,430]" gridcol="7" gridrow="4" pageId="12" pageNumber="163">2.80</td>
<td id="C66E9563FF888FB8A9AAFE4EFB75FE0B" box="[1156,1213,406,430]" gridcol="8" gridrow="4" pageId="12" pageNumber="163">2.60</td>
<td id="C66E9563FF888FB8A9C1FE4EFAE0FE0B" box="[1263,1320,406,430]" gridcol="9" gridrow="4" pageId="12" pageNumber="163">2.50</td>
<td id="C66E9563FF888FB8A861FE4EFA40FE0B" box="[1359,1416,406,430]" gridcol="10" gridrow="4" pageId="12" pageNumber="163">2.40</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FE18FA40FE7D" box="[167,1416,448,472]" gridrow="5" pageId="12" pageNumber="163" rowspan-0="1">
<td id="C66E9563FF888FB8AC50FE18FDC7FE7D" box="[382,527,448,472]" gridcol="1" gridrow="5" pageId="12" pageNumber="163">aculeus lenght</td>
<td id="C66E9563FF888FB8AF15FE18FDBCFE7D" box="[571,628,448,472]" gridcol="2" gridrow="5" pageId="12" pageNumber="163">2.50</td>
<td id="C66E9563FF888FB8AF8BFE18FD16FE7D" box="[677,734,448,472]" gridcol="3" gridrow="5" pageId="12" pageNumber="163">2.80</td>
<td id="C66E9563FF888FB8AE2AFE18FCF5FE7D" box="[772,829,448,472]" gridcol="4" gridrow="5" pageId="12" pageNumber="163">2.60</td>
<td id="C66E9563FF888FB8AE4AFE18FC55FE7D" box="[868,925,448,472]" gridcol="5" gridrow="5" pageId="12" pageNumber="163">2.70</td>
<td id="C66E9563FF888FB8AEEAFE18FC35FE7D" box="[964,1021,448,472]" gridcol="6" gridrow="5" pageId="12" pageNumber="163">3.25</td>
<td id="C66E9563FF888FB8A90AFE18FB95FE7D" box="[1060,1117,448,472]" gridcol="7" gridrow="5" pageId="12" pageNumber="163">1.00</td>
<td id="C66E9563FF888FB8A9AAFE18FB75FE7D" box="[1156,1213,448,472]" gridcol="8" gridrow="5" pageId="12" pageNumber="163">3.00</td>
<td id="C66E9563FF888FB8A9C1FE18FAE0FE7D" box="[1263,1320,448,472]" gridcol="9" gridrow="5" pageId="12" pageNumber="163">3.00</td>
<td id="C66E9563FF888FB8A861FE18FA40FE7D" box="[1359,1416,448,472]" gridcol="10" gridrow="5" pageId="12" pageNumber="163">2.90</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FE32FA40FDA7" box="[167,1416,490,514]" gridrow="6" pageId="12" pageNumber="163" rowspan-0="1">
<td id="C66E9563FF888FB8AC50FE32FDC7FDA7" box="[382,527,490,514]" gridcol="1" gridrow="6" pageId="12" pageNumber="163">total length</td>
<td id="C66E9563FF888FB8AF15FE32FDBCFDA7" box="[571,628,490,514]" gridcol="2" gridrow="6" pageId="12" pageNumber="163">5.80</td>
<td id="C66E9563FF888FB8AF8BFE32FD16FDA7" box="[677,734,490,514]" gridcol="3" gridrow="6" pageId="12" pageNumber="163">6.40</td>
<td id="C66E9563FF888FB8AE2AFE32FCF5FDA7" box="[772,829,490,514]" gridcol="4" gridrow="6" pageId="12" pageNumber="163">5.70</td>
<td id="C66E9563FF888FB8AE4AFE32FC55FDA7" box="[868,925,490,514]" gridcol="5" gridrow="6" pageId="12" pageNumber="163">6.00</td>
<td id="C66E9563FF888FB8AEEAFE32FC35FDA7" box="[964,1021,490,514]" gridcol="6" gridrow="6" pageId="12" pageNumber="163">7.75</td>
<td id="C66E9563FF888FB8A90AFE32FB95FDA7" box="[1060,1117,490,514]" gridcol="7" gridrow="6" pageId="12" pageNumber="163">5.55</td>
<td id="C66E9563FF888FB8A9AAFE32FB75FDA7" box="[1156,1213,490,514]" gridcol="8" gridrow="6" pageId="12" pageNumber="163">6.90</td>
<td id="C66E9563FF888FB8A9C1FE32FAE0FDA7" box="[1263,1320,490,514]" gridcol="9" gridrow="6" pageId="12" pageNumber="163">6.60</td>
<td id="C66E9563FF888FB8A861FE32FA40FDA7" box="[1359,1416,490,514]" gridcol="10" gridrow="6" pageId="12" pageNumber="163">6.40</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FDCCFA40FD89" box="[167,1416,532,556]" gridrow="7" pageId="12" pageNumber="163">
<th id="C66E9563FF888FB8AD89FDCCFE9FFD89" box="[167,343,532,556]" gridcol="0" gridrow="7" pageId="12" pageNumber="163">Metasoma+Telson</th>
<td id="C66E9563FF888FB8AC50FDCCFDC7FD89" box="[382,527,532,556]" gridcol="1" gridrow="7" pageId="12" pageNumber="163">Total length</td>
<td id="C66E9563FF888FB8AF15FDCCFDBCFD89" box="[571,628,532,556]" gridcol="2" gridrow="7" pageId="12" pageNumber="163">33.70</td>
<td id="C66E9563FF888FB8AF8BFDCCFD16FD89" box="[677,734,532,556]" gridcol="3" gridrow="7" pageId="12" pageNumber="163">34.30</td>
<td id="C66E9563FF888FB8AE2AFDCCFCF5FD89" box="[772,829,532,556]" gridcol="4" gridrow="7" pageId="12" pageNumber="163">31.40</td>
<td id="C66E9563FF888FB8AE4AFDCCFC55FD89" box="[868,925,532,556]" gridcol="5" gridrow="7" pageId="12" pageNumber="163">33.70</td>
<td id="C66E9563FF888FB8AEEAFDCCFC35FD89" box="[964,1021,532,556]" gridcol="6" gridrow="7" pageId="12" pageNumber="163">36.00</td>
<td id="C66E9563FF888FB8A90AFDCCFB95FD89" box="[1060,1117,532,556]" gridcol="7" gridrow="7" pageId="12" pageNumber="163">34.60</td>
<td id="C66E9563FF888FB8A9AAFDCCFB75FD89" box="[1156,1213,532,556]" gridcol="8" gridrow="7" pageId="12" pageNumber="163">37.20</td>
<td id="C66E9563FF888FB8A9C1FDCCFAE0FD89" box="[1263,1320,532,556]" gridcol="9" gridrow="7" pageId="12" pageNumber="163">36.90</td>
<td id="C66E9563FF888FB8A861FDCCFA40FD89" box="[1359,1416,532,556]" gridcol="10" gridrow="7" pageId="12" pageNumber="163">33.90</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FD88FA40FDCC" box="[167,1416,592,617]" gridrow="8" pageId="12" pageNumber="163">
<th id="C66E9563FF888FB8AD89FD88FE9FFDCC" box="[167,343,592,617]" gridcol="0" gridrow="8" pageId="12" pageNumber="163">Femur</th>
<td id="C66E9563FF888FB8AC50FD88FDC7FDCC" box="[382,527,592,617]" gridcol="1" gridrow="8" pageId="12" pageNumber="163">Length</td>
<td id="C66E9563FF888FB8AF15FD88FDBCFDCC" box="[571,628,592,617]" gridcol="2" gridrow="8" pageId="12" pageNumber="163">6.00</td>
<td id="C66E9563FF888FB8AF8BFD88FD16FDCC" box="[677,734,592,617]" gridcol="3" gridrow="8" pageId="12" pageNumber="163">5.90</td>
<td id="C66E9563FF888FB8AE2AFD88FCF5FDCC" box="[772,829,592,617]" gridcol="4" gridrow="8" pageId="12" pageNumber="163">5.50</td>
<td id="C66E9563FF888FB8AE4AFD88FC55FDCC" box="[868,925,592,617]" gridcol="5" gridrow="8" pageId="12" pageNumber="163">6.10</td>
<td id="C66E9563FF888FB8AEEAFD88FC35FDCC" box="[964,1021,592,617]" gridcol="6" gridrow="8" pageId="12" pageNumber="163">6.50</td>
<td id="C66E9563FF888FB8A90AFD88FB95FDCC" box="[1060,1117,592,617]" gridcol="7" gridrow="8" pageId="12" pageNumber="163">6.30</td>
<td id="C66E9563FF888FB8A9AAFD88FB75FDCC" box="[1156,1213,592,617]" gridcol="8" gridrow="8" pageId="12" pageNumber="163">6.90</td>
<td id="C66E9563FF888FB8A9C1FD88FAE0FDCC" box="[1263,1320,592,617]" gridcol="9" gridrow="8" pageId="12" pageNumber="163">7.00</td>
<td id="C66E9563FF888FB8A861FD88FA40FDCC" box="[1359,1416,592,617]" gridcol="10" gridrow="8" pageId="12" pageNumber="163">5.90</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FDA2FA40FD36" box="[167,1416,634,659]" gridrow="9" pageId="12" pageNumber="163" rowspan-0="1">
<td id="C66E9563FF888FB8AC50FDA2FDC7FD36" box="[382,527,634,659]" gridcol="1" gridrow="9" pageId="12" pageNumber="163">Width</td>
<td id="C66E9563FF888FB8AF15FDA2FDBCFD36" box="[571,628,634,659]" gridcol="2" gridrow="9" pageId="12" pageNumber="163">1.80</td>
<td id="C66E9563FF888FB8AF8BFDA2FD16FD36" box="[677,734,634,659]" gridcol="3" gridrow="9" pageId="12" pageNumber="163">1.90</td>
<td id="C66E9563FF888FB8AE2AFDA2FCF5FD36" box="[772,829,634,659]" gridcol="4" gridrow="9" pageId="12" pageNumber="163">1.80</td>
<td id="C66E9563FF888FB8AE4AFDA2FC55FD36" box="[868,925,634,659]" gridcol="5" gridrow="9" pageId="12" pageNumber="163">1.90</td>
<td id="C66E9563FF888FB8AEEAFDA2FC35FD36" box="[964,1021,634,659]" gridcol="6" gridrow="9" pageId="12" pageNumber="163">1.75</td>
<td id="C66E9563FF888FB8A90AFDA2FB95FD36" box="[1060,1117,634,659]" gridcol="7" gridrow="9" pageId="12" pageNumber="163">2.00</td>
<td id="C66E9563FF888FB8A9AAFDA2FB75FD36" box="[1156,1213,634,659]" gridcol="8" gridrow="9" pageId="12" pageNumber="163">1.90</td>
<td id="C66E9563FF888FB8A9C1FDA2FAE0FD36" box="[1263,1320,634,659]" gridcol="9" gridrow="9" pageId="12" pageNumber="163">2.00</td>
<td id="C66E9563FF888FB8A861FDA2FA40FD36" box="[1359,1416,634,659]" gridcol="10" gridrow="9" pageId="12" pageNumber="163">1.80</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FD7DFA40FD18" box="[167,1416,677,701]" gridrow="10" pageId="12" pageNumber="163">
<th id="C66E9563FF888FB8AD89FD7DFE9FFD18" box="[167,343,677,701]" gridcol="0" gridrow="10" pageId="12" pageNumber="163">Patella</th>
<td id="C66E9563FF888FB8AC50FD7DFDC7FD18" box="[382,527,677,701]" gridcol="1" gridrow="10" pageId="12" pageNumber="163">Lenght</td>
<td id="C66E9563FF888FB8AF15FD7DFDBCFD18" box="[571,628,677,701]" gridcol="2" gridrow="10" pageId="12" pageNumber="163">6.60</td>
<td id="C66E9563FF888FB8AF8BFD7DFD16FD18" box="[677,734,677,701]" gridcol="3" gridrow="10" pageId="12" pageNumber="163">6.60</td>
<td id="C66E9563FF888FB8AE2AFD7DFCF5FD18" box="[772,829,677,701]" gridcol="4" gridrow="10" pageId="12" pageNumber="163">6.00</td>
<td id="C66E9563FF888FB8AE4AFD7DFC55FD18" box="[868,925,677,701]" gridcol="5" gridrow="10" pageId="12" pageNumber="163">6.80</td>
<td id="C66E9563FF888FB8AEEAFD7DFC35FD18" box="[964,1021,677,701]" gridcol="6" gridrow="10" pageId="12" pageNumber="163">7.10</td>
<td id="C66E9563FF888FB8A90AFD7DFB95FD18" box="[1060,1117,677,701]" gridcol="7" gridrow="10" pageId="12" pageNumber="163">7.10</td>
<td id="C66E9563FF888FB8A9AAFD7DFB75FD18" box="[1156,1213,677,701]" gridcol="8" gridrow="10" pageId="12" pageNumber="163">7.60</td>
<td id="C66E9563FF888FB8A9C1FD7DFAE0FD18" box="[1263,1320,677,701]" gridcol="9" gridrow="10" pageId="12" pageNumber="163">7.60</td>
<td id="C66E9563FF888FB8A861FD7DFA40FD18" box="[1359,1416,677,701]" gridcol="10" gridrow="10" pageId="12" pageNumber="163">6.60</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FD16FA40FD42" box="[167,1416,718,743]" gridrow="11" pageId="12" pageNumber="163" rowspan-0="1">
<td id="C66E9563FF888FB8AC50FD16FDC7FD42" box="[382,527,718,743]" gridcol="1" gridrow="11" pageId="12" pageNumber="163">Width</td>
<td id="C66E9563FF888FB8AF15FD16FDBCFD42" box="[571,628,718,743]" gridcol="2" gridrow="11" pageId="12" pageNumber="163">2.50</td>
<td id="C66E9563FF888FB8AF8BFD16FD16FD42" box="[677,734,718,743]" gridcol="3" gridrow="11" pageId="12" pageNumber="163">2.50</td>
<td id="C66E9563FF888FB8AE2AFD16FCF5FD42" box="[772,829,718,743]" gridcol="4" gridrow="11" pageId="12" pageNumber="163">2.40</td>
<td id="C66E9563FF888FB8AE4AFD16FC55FD42" box="[868,925,718,743]" gridcol="5" gridrow="11" pageId="12" pageNumber="163">2.60</td>
<td id="C66E9563FF888FB8AEEAFD16FC35FD42" box="[964,1021,718,743]" gridcol="6" gridrow="11" pageId="12" pageNumber="163">3.10</td>
<td id="C66E9563FF888FB8A90AFD16FB95FD42" box="[1060,1117,718,743]" gridcol="7" gridrow="11" pageId="12" pageNumber="163">2.75</td>
<td id="C66E9563FF888FB8A9AAFD16FB75FD42" box="[1156,1213,718,743]" gridcol="8" gridrow="11" pageId="12" pageNumber="163">2.60</td>
<td id="C66E9563FF888FB8A9C1FD16FAE0FD42" box="[1263,1320,718,743]" gridcol="9" gridrow="11" pageId="12" pageNumber="163">2.80</td>
<td id="C66E9563FF888FB8A861FD16FA40FD42" box="[1359,1416,718,743]" gridcol="10" gridrow="11" pageId="12" pageNumber="163">2.40</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FD20FA40FCB4" box="[167,1416,760,785]" gridrow="12" pageId="12" pageNumber="163">
<th id="C66E9563FF888FB8AD89FD20FE9FFCB4" box="[167,343,760,785]" gridcol="0" gridrow="12" pageId="12" pageNumber="163">Chela</th>
<td id="C66E9563FF888FB8AC50FD20FDC7FCB4" box="[382,527,760,785]" gridcol="1" gridrow="12" pageId="12" pageNumber="163">Length</td>
<td id="C66E9563FF888FB8AF15FD20FDBCFCB4" box="[571,628,760,785]" gridcol="2" gridrow="12" pageId="12" pageNumber="163">12.40</td>
<td id="C66E9563FF888FB8AF8BFD20FD16FCB4" box="[677,734,760,785]" gridcol="3" gridrow="12" pageId="12" pageNumber="163">12.20</td>
<td id="C66E9563FF888FB8AE2AFD20FCF5FCB4" box="[772,829,760,785]" gridcol="4" gridrow="12" pageId="12" pageNumber="163">11.30</td>
<td id="C66E9563FF888FB8AE4AFD20FC55FCB4" box="[868,925,760,785]" gridcol="5" gridrow="12" pageId="12" pageNumber="163">12.20</td>
<td id="C66E9563FF888FB8AEEAFD20FC35FCB4" box="[964,1021,760,785]" gridcol="6" gridrow="12" pageId="12" pageNumber="163">13.20</td>
<td id="C66E9563FF888FB8A90AFD20FB95FCB4" box="[1060,1117,760,785]" gridcol="7" gridrow="12" pageId="12" pageNumber="163">13.50</td>
<td id="C66E9563FF888FB8A9AAFD20FB75FCB4" box="[1156,1213,760,785]" gridcol="8" gridrow="12" pageId="12" pageNumber="163">13.80</td>
<td id="C66E9563FF888FB8A9C1FD20FAE0FCB4" box="[1263,1320,760,785]" gridcol="9" gridrow="12" pageId="12" pageNumber="163">13.40</td>
<td id="C66E9563FF888FB8A861FD20FA40FCB4" box="[1359,1416,760,785]" gridcol="10" gridrow="12" pageId="12" pageNumber="163">11.50</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FCFAFA40FC9E" box="[167,1416,802,827]" gridrow="13" pageId="12" pageNumber="163" rowspan-0="1">
<td id="C66E9563FF888FB8AC50FCFAFDC7FC9E" box="[382,527,802,827]" gridcol="1" gridrow="13" pageId="12" pageNumber="163">Width</td>
<td id="C66E9563FF888FB8AF15FCFAFDBCFC9E" box="[571,628,802,827]" gridcol="2" gridrow="13" pageId="12" pageNumber="163">3.90</td>
<td id="C66E9563FF888FB8AF8BFCFAFD16FC9E" box="[677,734,802,827]" gridcol="3" gridrow="13" pageId="12" pageNumber="163">2.80</td>
<td id="C66E9563FF888FB8AE2AFCFAFCF5FC9E" box="[772,829,802,827]" gridcol="4" gridrow="13" pageId="12" pageNumber="163">2.40</td>
<td id="C66E9563FF888FB8AE4AFCFAFC55FC9E" box="[868,925,802,827]" gridcol="5" gridrow="13" pageId="12" pageNumber="163">2.80</td>
<td id="C66E9563FF888FB8AEEAFCFAFC35FC9E" box="[964,1021,802,827]" gridcol="6" gridrow="13" pageId="12" pageNumber="163">3.00</td>
<td id="C66E9563FF888FB8A90AFCFAFB95FC9E" box="[1060,1117,802,827]" gridcol="7" gridrow="13" pageId="12" pageNumber="163">3.05</td>
<td id="C66E9563FF888FB8A9AAFCFAFB75FC9E" box="[1156,1213,802,827]" gridcol="8" gridrow="13" pageId="12" pageNumber="163">2.60</td>
<td id="C66E9563FF888FB8A9C1FCFAFAE0FC9E" box="[1263,1320,802,827]" gridcol="9" gridrow="13" pageId="12" pageNumber="163">2.60</td>
<td id="C66E9563FF888FB8A861FCFAFA40FC9E" box="[1359,1416,802,827]" gridcol="10" gridrow="13" pageId="12" pageNumber="163">3.10</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FC94FA40FCC0" box="[167,1416,844,869]" gridrow="14" pageId="12" pageNumber="163" rowspan-0="1">
<td id="C66E9563FF888FB8AC50FC94FDC7FCC0" box="[382,527,844,869]" gridcol="1" gridrow="14" pageId="12" pageNumber="163">Height</td>
<td id="C66E9563FF888FB8AF15FC94FDBCFCC0" box="[571,628,844,869]" gridcol="2" gridrow="14" pageId="12" pageNumber="163">4.00</td>
<td id="C66E9563FF888FB8AF8BFC94FD16FCC0" box="[677,734,844,869]" gridcol="3" gridrow="14" pageId="12" pageNumber="163">2.90</td>
<td id="C66E9563FF888FB8AE2AFC94FCF5FCC0" box="[772,829,844,869]" gridcol="4" gridrow="14" pageId="12" pageNumber="163">2.50</td>
<td id="C66E9563FF888FB8AE4AFC94FC55FCC0" box="[868,925,844,869]" gridcol="5" gridrow="14" pageId="12" pageNumber="163">2.90</td>
<td id="C66E9563FF888FB8AEEAFC94FC35FCC0" box="[964,1021,844,869]" gridcol="6" gridrow="14" pageId="12" pageNumber="163">2.75</td>
<td id="C66E9563FF888FB8A90AFC94FB95FCC0" box="[1060,1117,844,869]" gridcol="7" gridrow="14" pageId="12" pageNumber="163">2.65</td>
<td id="C66E9563FF888FB8A9AAFC94FB75FCC0" box="[1156,1213,844,869]" gridcol="8" gridrow="14" pageId="12" pageNumber="163">2.60</td>
<td id="C66E9563FF888FB8A9C1FC94FAE0FCC0" box="[1263,1320,844,869]" gridcol="9" gridrow="14" pageId="12" pageNumber="163">2.80</td>
<td id="C66E9563FF888FB8A861FC94FA40FCC0" box="[1359,1416,844,869]" gridcol="10" gridrow="14" pageId="12" pageNumber="163">3.10</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FCAEFA40FC16" box="[167,1416,886,947]" gridrow="15" pageId="12" pageNumber="163" rowspan-0="1">
<td id="C66E9563FF888FB8AC50FCAEFDC7FC16" box="[382,527,886,947]" gridcol="1" gridrow="15" pageId="12" pageNumber="163">Length mov. fing.</td>
<td id="C66E9563FF888FB8AF15FCAEFDBCFC16" box="[571,628,886,947]" gridcol="2" gridrow="15" pageId="12" pageNumber="163">7.90</td>
<td id="C66E9563FF888FB8AF8BFCAEFD16FC16" box="[677,734,886,947]" gridcol="3" gridrow="15" pageId="12" pageNumber="163">7.90</td>
<td id="C66E9563FF888FB8AE2AFCAEFCF5FC16" box="[772,829,886,947]" gridcol="4" gridrow="15" pageId="12" pageNumber="163">7.40</td>
<td id="C66E9563FF888FB8AE4AFCAEFC55FC16" box="[868,925,886,947]" gridcol="5" gridrow="15" pageId="12" pageNumber="163">8.20</td>
<td id="C66E9563FF888FB8AEEAFCAEFC35FC16" box="[964,1021,886,947]" gridcol="6" gridrow="15" pageId="12" pageNumber="163">8.25</td>
<td id="C66E9563FF888FB8A90AFCAEFB95FC16" box="[1060,1117,886,947]" gridcol="7" gridrow="15" pageId="12" pageNumber="163">8.70</td>
<td id="C66E9563FF888FB8A9AAFCAEFB75FC16" box="[1156,1213,886,947]" gridcol="8" gridrow="15" pageId="12" pageNumber="163">9.50</td>
<td id="C66E9563FF888FB8A9C1FCAEFAE0FC16" box="[1263,1320,886,947]" gridcol="9" gridrow="15" pageId="12" pageNumber="163">8.90</td>
<td id="C66E9563FF888FB8A861FCAEFA40FC16" box="[1359,1416,886,947]" gridcol="10" gridrow="15" pageId="12" pageNumber="163">7.20</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FC1BFA40FC5A" box="[167,1416,963,1023]" gridrow="16" pageId="12" pageNumber="163" rowspan-0="1">
<td id="C66E9563FF888FB8AC50FC1BFDC7FC5A" box="[382,527,963,1023]" gridcol="1" gridrow="16" pageId="12" pageNumber="163">Length fix. fing.</td>
<td id="C66E9563FF888FB8AF15FC1BFDBCFC5A" box="[571,628,963,1023]" gridcol="2" gridrow="16" pageId="12" pageNumber="163">6.40</td>
<td id="C66E9563FF888FB8AF8BFC1BFD16FC5A" box="[677,734,963,1023]" gridcol="3" gridrow="16" pageId="12" pageNumber="163">6.70</td>
<td id="C66E9563FF888FB8AE2AFC1BFCF5FC5A" box="[772,829,963,1023]" gridcol="4" gridrow="16" pageId="12" pageNumber="163">6.40</td>
<td id="C66E9563FF888FB8AE4AFC1BFC55FC5A" box="[868,925,963,1023]" gridcol="5" gridrow="16" pageId="12" pageNumber="163">6.90</td>
<td id="C66E9563FF888FB8AEEAFC1BFC35FC5A" box="[964,1021,963,1023]" gridcol="6" gridrow="16" pageId="12" pageNumber="163">8.00</td>
<td id="C66E9563FF888FB8A90AFC1BFB95FC5A" box="[1060,1117,963,1023]" gridcol="7" gridrow="16" pageId="12" pageNumber="163">6.75</td>
<td id="C66E9563FF888FB8A9AAFC1BFB75FC5A" box="[1156,1213,963,1023]" gridcol="8" gridrow="16" pageId="12" pageNumber="163">8.20</td>
<td id="C66E9563FF888FB8A9C1FC1BFAE0FC5A" box="[1263,1320,963,1023]" gridcol="9" gridrow="16" pageId="12" pageNumber="163">7.70</td>
<td id="C66E9563FF888FB8A861FC1BFA40FC5A" box="[1359,1416,963,1023]" gridcol="10" gridrow="16" pageId="12" pageNumber="163">5.90</td>
</tr>
<tr id="85BFFC1FFF888FB8AD89FBD7FA40FB82" box="[167,1416,1039,1063]" gridrow="17" pageId="12" pageNumber="163" rowspan-0="1">
<td id="C66E9563FF888FB8AC50FBD7FDC7FB82" box="[382,527,1039,1063]" gridcol="1" gridrow="17" pageId="12" pageNumber="163">Length palm</td>
<td id="C66E9563FF888FB8AF15FBD7FDBCFB82" box="[571,628,1039,1063]" gridcol="2" gridrow="17" pageId="12" pageNumber="163">5.10</td>
<td id="C66E9563FF888FB8AF8BFBD7FD16FB82" box="[677,734,1039,1063]" gridcol="3" gridrow="17" pageId="12" pageNumber="163">4.20</td>
<td id="C66E9563FF888FB8AE2AFBD7FCF5FB82" box="[772,829,1039,1063]" gridcol="4" gridrow="17" pageId="12" pageNumber="163">4.00</td>
<td id="C66E9563FF888FB8AE4AFBD7FC55FB82" box="[868,925,1039,1063]" gridcol="5" gridrow="17" pageId="12" pageNumber="163">4.20</td>
<td id="C66E9563FF888FB8AEEAFBD7FC35FB82" box="[964,1021,1039,1063]" gridcol="6" gridrow="17" pageId="12" pageNumber="163">4.50</td>
<td id="C66E9563FF888FB8A90AFBD7FB95FB82" box="[1060,1117,1039,1063]" gridcol="7" gridrow="17" pageId="12" pageNumber="163">5.25</td>
<td id="C66E9563FF888FB8A9AAFBD7FB75FB82" box="[1156,1213,1039,1063]" gridcol="8" gridrow="17" pageId="12" pageNumber="163">4.60</td>
<td id="C66E9563FF888FB8A9C1FBD7FAE0FB82" box="[1263,1320,1039,1063]" gridcol="9" gridrow="17" pageId="12" pageNumber="163">4.50</td>
<td id="C66E9563FF888FB8A861FBD7FA40FB82" box="[1359,1416,1039,1063]" gridcol="10" gridrow="17" pageId="12" pageNumber="163">4.60</td>
</tr>
</table>
</paragraph>
<paragraph id="3B30FE5DFF88704BADE9FB80FD27FACB" blockId="12.[151,1437,1112,2039]" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BADE9FB80FE87FBD7" bold="true" box="[199,335,1112,1138]" pageId="12" pageNumber="163">Coloration.</emphasis>
General pattern (in 70% ethanol) (
<figureCitation id="A3B4E2D8FF88704BAFF0FB80FCBFFBD6" box="[734,887,1112,1139]" captionStart="FIGURE 2" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[180,1404,509,1847]" captionTargetId="figure-177@7.[164,1424,489,1864]" captionTargetPageId="7" captionText="FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6673340" httpUri="https://zenodo.org/record/6673340/files/figure.png" pageId="12" pageNumber="163">Figure 2A–D</figureCitation>
) dark reddish-brown.
<emphasis id="09FB224FFF88704BA95BFB81FB2BFBD6" box="[1141,1251,1113,1139]" italics="true" pageId="12" pageNumber="163">Carapace</emphasis>
(
<figureCitation id="A3B4E2D8FF88704BA9DDFB80FA46FBD6" box="[1267,1422,1112,1139]" captionStart="FIGURE 2" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[180,1404,509,1847]" captionTargetId="figure-177@7.[164,1424,489,1864]" captionTargetPageId="7" captionText="FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6673340" httpUri="https://zenodo.org/record/6673340/files/figure.png" pageId="12" pageNumber="163">Figure 2A, C</figureCitation>
): dark reddish-brown with black variegated pigments; black stripe on anterior margin; lateral and median eyes, surrounded by black variegated pigments.
<emphasis id="09FB224FFF88704BAF60FB78FD01FB1F" box="[590,713,1184,1210]" italics="true" pageId="12" pageNumber="163">Chelicerae</emphasis>
(
<figureCitation id="A3B4E2D8FF88704BAFF9FB78FCA6FB1E" box="[727,878,1184,1211]" captionStart="FIGURE 2" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[180,1404,509,1847]" captionTargetId="figure-177@7.[164,1424,489,1864]" captionTargetPageId="7" captionText="FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6673340" httpUri="https://zenodo.org/record/6673340/files/figure.png" pageId="12" pageNumber="163">Figure 2A, C</figureCitation>
): coxa and hand dark yellow, with abundant black pigments over the hand surface; fingers, blackish-brown.
<emphasis id="09FB224FFF88704BAE16FB1CFBDCFB7B" box="[824,1044,1220,1246]" italics="true" pageId="12" pageNumber="163">Coxosternal region</emphasis>
,
<emphasis id="09FB224FFF88704BA908FB1DFB93FB7B" box="[1062,1115,1221,1246]" italics="true" pageId="12" pageNumber="163">Legs</emphasis>
,
<taxonomicName id="FC8F85DEFF88704BA943FB1DFB2DFB7B" box="[1133,1253,1221,1246]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="09FB224FFF88704BA943FB1DFB2DFB7B" box="[1133,1253,1221,1246]" italics="true" pageId="12" pageNumber="163">Mesosoma</emphasis>
</taxonomicName>
,
<emphasis id="09FB224FFF88704BA9D9FB1CFA54FB7B" box="[1271,1436,1220,1246]" italics="true" pageId="12" pageNumber="163">and Pedipalps</emphasis>
(
<figureCitation id="A3B4E2D8FF88704BADB1FB30FEF3FAA6" box="[159,315,1256,1283]" captionStart="FIGURE 2" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[180,1404,509,1847]" captionTargetId="figure-177@7.[164,1424,489,1864]" captionTargetPageId="7" captionText="FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6673340" httpUri="https://zenodo.org/record/6673340/files/figure.png" pageId="12" pageNumber="163">Figure 2A–D</figureCitation>
): all reddish-brown, covered with black variegated pigments (except for the coxosternal region and ventral areas of the pedipalp segments).
<emphasis id="09FB224FFF88704BAF42FAD4FD67FA83" box="[620,687,1292,1318]" italics="true" pageId="12" pageNumber="163">Chela</emphasis>
(
<figureCitation id="A3B4E2D8FF88704BAFEFFAD4FC93FA82" box="[705,859,1292,1319]" captionStart="FIGURE 2" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[180,1404,509,1847]" captionTargetId="figure-177@7.[164,1424,489,1864]" captionTargetPageId="7" captionText="FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6673340" httpUri="https://zenodo.org/record/6673340/files/figure.png" pageId="12" pageNumber="163">Figure 2A–D</figureCitation>
) fingers blackish.
<emphasis id="09FB224FFF88704BA91DFAD5FB60FA83" box="[1075,1192,1293,1318]" italics="true" pageId="12" pageNumber="163">Metasoma</emphasis>
(
<figureCitation id="A3B4E2D8FF88704BA997FAD4FA9FFA82" box="[1209,1367,1292,1319]" captionStart="FIGURE 2" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[180,1404,509,1847]" captionTargetId="figure-177@7.[164,1424,489,1864]" captionTargetPageId="7" captionText="FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6673340" httpUri="https://zenodo.org/record/6673340/files/figure.png" pageId="12" pageNumber="163">Figure 2A–D</figureCitation>
): segments dark reddish-brown, progressively darker towards the telson; segments IV–V blackish-brown.
<taxonomicName id="FC8F85DEFF88704BA9D5FAE8FA8BFAEF" box="[1275,1347,1328,1354]" class="Hexanauplia" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="09FB224FFF88704BA9D5FAE8FA8BFAEF" box="[1275,1347,1328,1354]" italics="true" pageId="12" pageNumber="163">Telson</emphasis>
</taxonomicName>
(
<figureCitation id="A3B4E2D8FF88704BA87FFAE8FF29FACA" captionStart="FIGURE 2" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[180,1404,509,1847]" captionTargetId="figure-177@7.[164,1424,489,1864]" captionTargetPageId="7" captionText="FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6673340" httpUri="https://zenodo.org/record/6673340/files/figure.png" pageId="12" pageNumber="163">Figure 2A–D</figureCitation>
): blackish-brown; aculeus dark reddish-brown.
</paragraph>
<paragraph id="3B30FE5DFF88704BADE9FAA1FB48F9CE" blockId="12.[151,1437,1112,2039]" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BADE9FAA1FEA9FA36" bold="true" box="[199,353,1401,1427]" pageId="12" pageNumber="163">Morphology.</emphasis>
Body hirsute, covered with abundant microsetae, white colored under UV light.
<emphasis id="09FB224FFF88704BA9C0FAA1FA94FA36" box="[1262,1372,1401,1427]" italics="true" pageId="12" pageNumber="163">Carapace</emphasis>
(
<figureCitation id="A3B4E2D8FF88704BA842FAA0FEC2FA12" captionStart="FIGURE 4" captionStartId="10.[151,250,1375,1399]" captionTargetBox="[371,1216,309,1352]" captionTargetId="figure-86@10.[371,1216,309,1352]" captionTargetPageId="10" captionText="FIGURE 4. Tityus (Atreus) Gervais, 1843, carapace sculpturing, dorsal aspect. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673348" httpUri="https://zenodo.org/record/6673348/files/figure.png" pageId="12" pageNumber="163">Figure 4A, B</figureCitation>
): moderately covered with fine granulation and few coarse granules; anterior margin with deep median notch; anterior median carinae, central lateral, central median, lateral ocular, posterior, posterior median and superciliary carinae, all well-marked, and furrows (anterior median, anterior marginal, central transverse, lateral ocular, supercialiary, posterior transverse, posterior lateral and posterior marginal), all well-marked; ocular tubercle wellmarked, located on the anterior half of carapace; median eyes separated by about 0.6 ocular diameters; with three pairs of lateral eyes and two pairs of lateral micro-ocelli (ADMi and PDMi) (pattern 4A).
</paragraph>
<paragraph id="3B30FE5DFF88704BADE9F9ACFF04F973" blockId="12.[151,1437,1112,2039]" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BADE9F9ACFDCCF92B" bold="true" box="[199,516,1652,1679]" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BADE9F9ACFE8BF92A" bold="true" box="[199,323,1652,1679]" italics="true" pageId="12" pageNumber="163">Chelicerae</emphasis>
(
<figureCitation id="A3B4E2D8FF88704BAC7AF9ACFE3DF92B" box="[340,501,1652,1678]" captionStart="FIGURE 4" captionStartId="10.[151,250,1375,1399]" captionTargetBox="[371,1216,309,1352]" captionTargetId="figure-86@10.[371,1216,309,1352]" captionTargetPageId="10" captionText="FIGURE 4. Tityus (Atreus) Gervais, 1843, carapace sculpturing, dorsal aspect. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673348" httpUri="https://zenodo.org/record/6673348/files/figure.png" pageId="12" pageNumber="163">Figure 4A, B</figureCitation>
).
</emphasis>
Dentition characteristic of the family
<taxonomicName id="FC8F85DEFF88704BAE90F9ACFB1EF92A" authority="(Vachon 1963)" baseAuthorityName="Vachon" baseAuthorityYear="1963" box="[958,1238,1652,1679]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="163" phylum="Arthropoda" rank="family">
Buthidae (
<bibRefCitation id="5F1E83ACFF88704BA91AF9ACFB06F92A" author="Vachon, M." box="[1076,1230,1652,1679]" pageId="12" pageNumber="163" pagination="161 - 166" refId="ref21343" refString="Vachon, M. (1963) De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris, 161 - 166." type="book chapter" year="1963">Vachon 1963</bibRefCitation>
)
</taxonomicName>
, densely covered with setae over the internal and ventral surfaces; anterior half of the manus with a transverse row of coarse granules.
</paragraph>
<paragraph id="3B30FE5DFF88704AADE9F938FAC8FEB8" blockId="12.[151,1437,1112,2039]" lastBlockId="13.[151,1437,151,286]" lastPageId="13" lastPageNumber="164" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BADE9F938FEF6F95E" bold="true" box="[199,318,1760,1787]" pageId="12" pageNumber="163">
<emphasis id="09FB224FFF88704BADE9F938FEFFF95E" bold="true" box="[199,311,1760,1787]" italics="true" pageId="12" pageNumber="163">Pedipalps</emphasis>
.
</emphasis>
Chela, bulky in male (L/W= 3.71) or slender in female (L/W= 5.31). Trichobothriotaxic pattern
<typeStatus id="E43440FFFF88704BA84BF939FA55F95E" box="[1381,1437,1761,1787]" pageId="12" pageNumber="163">Type</typeStatus>
A, alfa configuration (hand: Eb3:Eb1:Eb2:Esb:Est:Et, fixed finger: eb:esb:est:et:db:dt:it).
<emphasis id="09FB224FFF88704BA9B0F8DDFB21F8BB" box="[1182,1257,1797,1822]" italics="true" pageId="12" pageNumber="163">Femur</emphasis>
(
<figureCitation id="A3B4E2D8FF88704BA9D4F8DCFA5DF8BA" box="[1274,1429,1796,1823]" captionStart="FIGURE 5" captionStartId="13.[151,250,1447,1471]" captionTargetBox="[151,1437,309,1424]" captionTargetId="figure-91@13.[151,1437,309,1424]" captionTargetPageId="13" captionText="FIGURE 5. Tityus (Atreus) Gervais, 1843, femur sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673350" httpUri="https://zenodo.org/record/6673350/files/figure.png" pageId="12" pageNumber="163">Figure 5A–D</figureCitation>
) with five well-marked carinae: VI, DI, DE and VE crenulate, EM serratocrenulate and complete, with intercarinal areas densely covered with fine granulation and few coarse granules.
<taxonomicName id="FC8F85DEFF88704BAEE7F894FBD2F8C3" box="[969,1050,1868,1894]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Aplysiida" pageId="12" pageNumber="163" phylum="Mollusca" rank="genus">
<emphasis id="09FB224FFF88704BAEE7F894FBD2F8C3" box="[969,1050,1868,1894]" italics="true" pageId="12" pageNumber="163">Patella</emphasis>
</taxonomicName>
(
<figureCitation id="A3B4E2D8FF88704BA903F894FB00F8C2" box="[1069,1224,1868,1895]" captionStart="FIGURE 6" captionStartId="14.[151,250,1863,1887]" captionTargetBox="[151,1436,561,1840]" captionTargetId="figure-228@14.[151,1436,561,1840]" captionTargetPageId="14" captionText="FIGURE 6. Tityus (Atreus) Gervais, 1843, patella sculpturing. A–D. Tityus (Atreus) forcipula (Gervais, 1843). A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. E–H. Tityus (Atreus) moralensis sp. nov. E, F. Male holotype (MUSENUV-Ar 2104). E. Dorsal view. F. Ventral view. G, H. Female paratype (ICN-1402). G. Dorsal view. H. Ventral view. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673354" httpUri="https://zenodo.org/record/6673354/files/figure.png" pageId="12" pageNumber="163">Figure 6A–D</figureCitation>
) with seven wellmarked carinae: VI, DI, DE and EM, crenulate and complete; VE incomplete- only present on the distal half of the segment (female) or complete (male) and crenulate; DM crenulate and incomplete; IM serratocrenulate and complete, with a short spur near the segment base; with intercarinal areas densely covered with fine granulation.
<emphasis id="09FB224FFF88704BADB9F804FF12F853" box="[151,218,2012,2038]" italics="true" pageId="12" pageNumber="163">Chela</emphasis>
(
<emphasis id="09FB224FFF88704BADC5F804FED6F853" box="[235,286,2012,2038]" italics="true" pageId="12" pageNumber="163">tibia</emphasis>
) (
<figureCitation id="A3B4E2D8FF88704BAC19F804FE12F852" box="[311,474,2012,2039]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[261,1326,181,1920]" captionTargetId="figure-19@15.[261,1326,181,1920]" captionTargetPageId="15" captionText="FIGURE 7. Tityus (Atreus) Gervais, 1843,female chela sculpturing.A–C. Tityus(Atreus) forcipula (Gervais,1843) (MUSENUV- Ar 436). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (ICN-1402). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673356" httpUri="https://zenodo.org/record/6673356/files/figure.png" pageId="12" pageNumber="163">Figures 7A–C</figureCitation>
,
<figureCitation id="A3B4E2D8FF88704BACC8F805FDE2F852" box="[486,554,2013,2039]" captionStart="FIGURE 8" captionStartId="16.[151,250,1944,1968]" captionTargetBox="[364,1224,181,1920]" captionTargetId="figure-31@16.[364,1224,181,1920]" captionTargetPageId="16" captionText="FIGURE 8. Tityus (Atreus) Gervais, 1843, male chela sculpturing. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673358" httpUri="https://zenodo.org/record/6673358/files/figure.png" pageId="12" pageNumber="163">8A–C</figureCitation>
) with eight well-marked carinae: VI, VE, D, DS, DMA, IM and ES, crenulate and complete; SA crenulate and incomplete, only present on the anterior half of the hand. Pedipalp movable and fixed fingers each with a basal concavity followed by a well-marked basal lobe (male) (
<figureCitation id="A3B4E2D8FF89704AA944FF63FB28FF70" box="[1130,1248,187,213]" captionStart="FIGURE 8" captionStartId="16.[151,250,1944,1968]" captionTargetBox="[364,1224,181,1920]" captionTargetId="figure-31@16.[364,1224,181,1920]" captionTargetPageId="16" captionText="FIGURE 8. Tityus (Atreus) Gervais, 1843, male chela sculpturing. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673358" httpUri="https://zenodo.org/record/6673358/files/figure.png" pageId="13" pageNumber="164">Figure 8A</figureCitation>
) or movable and fixed fingers with obsolete basal concavities and an obsolete basal lobe only present on the movable finger (female) (
<figureCitation id="A3B4E2D8FF89704AADB1FEDBFEDEFEB8" box="[159,278,259,285]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[261,1326,181,1920]" captionTargetId="figure-19@15.[261,1326,181,1920]" captionTargetPageId="15" captionText="FIGURE 7. Tityus (Atreus) Gervais, 1843,female chela sculpturing.A–C. Tityus(Atreus) forcipula (Gervais,1843) (MUSENUV- Ar 436). A. External view. B. Dorsal view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (ICN-1402). D. External view. E. Dorsal view. F. Ventral view. Scale bars= 3 mm." figureDoi="http://doi.org/10.5281/zenodo.6673356" httpUri="https://zenodo.org/record/6673356/files/figure.png" pageId="13" pageNumber="164">Figure 7A</figureCitation>
).
<emphasis id="09FB224FFF89704AAC04FEDBFD8FFEB8" box="[298,583,259,285]" italics="true" pageId="13" pageNumber="164">Oblique rows of granules</emphasis>
: Movable finger with 16–16 (male) and 17–17 (female) rows.
</paragraph>
<caption id="6FF0AED5FF89704AADB9FA7FFC12F98E" ID-DOI="http://doi.org/10.5281/zenodo.6673350" ID-Zenodo-Dep="6673350" httpUri="https://zenodo.org/record/6673350/files/figure.png" pageId="13" pageNumber="164" startId="13.[151,250,1447,1471]" targetBox="[151,1437,309,1424]" targetPageId="13">
<paragraph id="3B30FE5DFF89704AADB9FA7FFC12F98E" blockId="13.[151,1437,1447,1580]" pageId="13" pageNumber="164">
<emphasis id="09FB224FFF89704AADB9FA7FFEDEFA1A" bold="true" box="[151,278,1447,1471]" pageId="13" pageNumber="164">FIGURE 5.</emphasis>
<taxonomicName id="FC8F85DEFF89704AAC0EFA70FD9EFA1A" authority="Gervais, 1843" authorityName="Gervais" authorityYear="1843" box="[288,598,1447,1471]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="164" phylum="Arthropoda" rank="subGenus" subGenus="Atreus">
<emphasis id="09FB224FFF89704AAC0EFA70FE95FA1A" box="[288,349,1448,1471]" italics="true" pageId="13" pageNumber="164">Tityus</emphasis>
(
<emphasis id="09FB224FFF89704AAC41FA7FFE78FA1B" box="[367,432,1447,1470]" italics="true" pageId="13" pageNumber="164">Atreus</emphasis>
)
<bibRefCitation id="5F1E83ACFF89704AACECFA7FFD9EFA1A" author="Gervais, P. M." box="[450,598,1447,1471]" pageId="13" pageNumber="164" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
</taxonomicName>
, femur sculpturing.
<emphasis id="09FB224FFF89704AAE05FA7FFCABFA1A" bold="true" box="[811,867,1447,1471]" pageId="13" pageNumber="164">A–D.</emphasis>
<taxonomicName id="FC8F85DEFF89704AAE43FA70FAD2FA1A" authority="(Gervais, 1843)" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[877,1306,1447,1471]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="forcipula" subGenus="Atreus">
<emphasis id="09FB224FFF89704AAE43FA70FC62FA1A" box="[877,938,1448,1471]" italics="true" pageId="13" pageNumber="164">Tityus</emphasis>
(
<emphasis id="09FB224FFF89704AAE95FA7FFC34FA1B" box="[955,1020,1447,1470]" italics="true" pageId="13" pageNumber="164">Atreus</emphasis>
)
<emphasis id="09FB224FFF89704AA921FA7FFBA3FA1A" box="[1039,1131,1447,1471]" italics="true" pageId="13" pageNumber="164">forcipula</emphasis>
(
<bibRefCitation id="5F1E83ACFF89704AA955FA7FFAD9FA1A" author="Gervais, P. M." box="[1147,1297,1447,1471]" pageId="13" pageNumber="164" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="09FB224FFF89704AA80BFA7FFA96FA1A" bold="true" box="[1317,1374,1447,1471]" pageId="13" pageNumber="164">A, B.</emphasis>
Male (MUSENUV-Ar 438).
<emphasis id="09FB224FFF89704AACAFFA13FE52FA46" bold="true" box="[385,410,1483,1507]" pageId="13" pageNumber="164">A.</emphasis>
Dorsal view.
<emphasis id="09FB224FFF89704AAF07FA13FD88FA46" bold="true" box="[553,576,1483,1507]" pageId="13" pageNumber="164">B.</emphasis>
Ventral view.
<emphasis id="09FB224FFF89704AAFFAFA13FCC4FA46" bold="true" box="[724,780,1483,1507]" pageId="13" pageNumber="164">C, D.</emphasis>
Female (MUSENUV-Ar 436).
<emphasis id="09FB224FFF89704AA961FA13FBAFFA46" bold="true" box="[1103,1127,1483,1507]" pageId="13" pageNumber="164">C.</emphasis>
Dorsal view.
<emphasis id="09FB224FFF89704AA9D9FA13FAC7FA46" bold="true" box="[1271,1295,1483,1507]" pageId="13" pageNumber="164">D.</emphasis>
Ventral view.
<emphasis id="09FB224FFF89704AADB9FA37FF07F9A2" bold="true" box="[151,207,1519,1543]" pageId="13" pageNumber="164">E–H.</emphasis>
<taxonomicName id="FC8F85DEFF89704AADF8FA28FE2BF9A2" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[214,483,1519,1543]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov." subGenus="Atreus">
<emphasis id="09FB224FFF89704AADF8FA28FEDBF9A2" box="[214,275,1520,1543]" italics="true" pageId="13" pageNumber="164">Tityus</emphasis>
(
<emphasis id="09FB224FFF89704AAC0DFA37FEACF9A3" box="[291,356,1519,1542]" italics="true" pageId="13" pageNumber="164">Atreus</emphasis>
)
<emphasis id="09FB224FFF89704AAC5AFA37FE2BF9A2" box="[372,483,1519,1543]" italics="true" pageId="13" pageNumber="164">moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF89704AACC5FA37FDBEF9A2" bold="true" box="[491,630,1519,1543]" pageId="13" pageNumber="164">
<taxonomicNameLabel id="12C89F34FF89704AACC5FA37FDF4F9A2" box="[491,572,1519,1543]" pageId="13" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
E, F.
</emphasis>
Male holotype (MUSENUV-Ar 2104).
<emphasis id="09FB224FFF89704AA93FFA37FBE0F9A2" bold="true" box="[1041,1064,1519,1543]" pageId="13" pageNumber="164">E.</emphasis>
Dorsal view.
<emphasis id="09FB224FFF89704AA996FA37FB03F9A2" bold="true" box="[1208,1227,1519,1543]" pageId="13" pageNumber="164">F.</emphasis>
Ventral view.
<emphasis id="09FB224FFF89704AA84FFA37FA54F9A2" bold="true" box="[1377,1436,1519,1543]" pageId="13" pageNumber="164">G, H.</emphasis>
Female paratype (ICN-1402).
<emphasis id="09FB224FFF89704AACE4F9CBFE2CF98E" bold="true" box="[458,484,1555,1579]" pageId="13" pageNumber="164">G.</emphasis>
Dorsal view.
<emphasis id="09FB224FFF89704AAF5EF9CBFD43F98E" bold="true" box="[624,651,1555,1579]" pageId="13" pageNumber="164">H.</emphasis>
Ventral view. Scale bars= 2 mm.
</paragraph>
</caption>
<paragraph id="3B30FE5DFF89704AADE9F981FECDF97A" blockId="13.[151,1437,1625,2012]" pageId="13" pageNumber="164">
<emphasis id="09FB224FFF89704AADE9F981FDADF9D6" bold="true" box="[199,613,1625,1652]" pageId="13" pageNumber="164">
<emphasis id="09FB224FFF89704AADE9F981FE6FF9D6" bold="true" box="[199,423,1625,1652]" italics="true" pageId="13" pageNumber="164">Coxosternal region</emphasis>
(
<figureCitation id="A3B4E2D8FF89704AAC99F981FD9EF9D6" box="[439,598,1625,1651]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="13" pageNumber="164">Figure 9A, B</figureCitation>
).
</emphasis>
Sternum with posterior depression, outer ridge, and apical button, wellmarked; region covered with abundant coarse and fine granulation, and abundant microsetae; genital operculum longitudinally divided, composed of two sub-triangular plates. Male genital papillae are rounded and with sclerotized tips.
</paragraph>
<paragraph id="3B30FE5DFF89704AADE9F931FD62F812" blockId="13.[151,1437,1625,2012]" pageId="13" pageNumber="164">
<emphasis id="09FB224FFF89704AADE9F931FE29F8A6" bold="true" box="[199,481,1769,1795]" pageId="13" pageNumber="164">
<emphasis id="09FB224FFF89704AADE9F931FEEFF8A6" bold="true" box="[199,295,1769,1795]" italics="true" pageId="13" pageNumber="164">Pectines</emphasis>
(
<figureCitation id="A3B4E2D8FF89704AAC18F931FE1AF8A6" box="[310,466,1769,1795]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="13" pageNumber="164">Figure 9A, B</figureCitation>
).
</emphasis>
Pectines are strongly sclerotized and brownish, with white areas (less sclerotized) on the marginal plates (
<figureCitation id="A3B4E2D8FF89704AAC57F8D5FDD2F88D" box="[377,538,1805,1832]" captionStart="FIGURE 2" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[180,1404,509,1847]" captionTargetId="figure-177@7.[164,1424,489,1864]" captionTargetPageId="7" captionText="FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6673340" httpUri="https://zenodo.org/record/6673340/files/figure.png" pageId="13" pageNumber="164">Figures 2B, D</figureCitation>
,
<figureCitation id="A3B4E2D8FF89704AAF0BF8D5FDA2F88D" box="[549,618,1805,1832]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="13" pageNumber="164">9A, B</figureCitation>
). Pectinal basal piece shield-like shaped, with a deep anteromedian notch and without glandular areas (
<figureCitation id="A3B4E2D8FF89704AACF6F8E9FDA5F8E9" box="[472,621,1841,1868]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="13" pageNumber="164">Figure 9A, B</figureCitation>
); pectinal tooth counts of 18–17 (male) and 16–18 (female). Intermediate plate, marginal plate, and fulcra densely covered with microsetae (
<figureCitation id="A3B4E2D8FF89704AAE56F88DFBC5F8D5" box="[888,1037,1877,1904]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="13" pageNumber="164">Figure 9A, B</figureCitation>
). Basal middle lamellae, not dilated (male) (
<figureCitation id="A3B4E2D8FF89704AADDFF8A1FEA1F836" box="[241,361,1913,1939]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="13" pageNumber="164">Figure 9A</figureCitation>
) or dilated and oval shaped (female) (
<figureCitation id="A3B4E2D8FF89704AAE0CF8A1FC53F836" box="[802,923,1913,1939]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="13" pageNumber="164">Figure 9B</figureCitation>
). Female basal middle lamellae with a distal glandular area, white colored (
<figureCitation id="A3B4E2D8FF89704AACC2F845FDA6F812" box="[492,622,1949,1975]" captionStart="FIGURE 2" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[180,1404,509,1847]" captionTargetId="figure-177@7.[164,1424,489,1864]" captionTargetPageId="7" captionText="FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6673340" httpUri="https://zenodo.org/record/6673340/files/figure.png" pageId="13" pageNumber="164">Figures 2D</figureCitation>
,
<figureCitation id="A3B4E2D8FF89704AAF57F845FD56F812" box="[633,670,1949,1975]" captionStart="FIGURE 9" captionStartId="17.[151,250,1245,1269]" captionTargetBox="[163,1424,181,1221]" captionTargetId="figure-19@17.[163,1424,181,1221]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="13" pageNumber="164">9B</figureCitation>
).
</paragraph>
<paragraph id="3B30FE5DFF897049ADE9F81AFB20FF5C" blockId="13.[151,1437,1625,2012]" lastBlockId="14.[151,1437,151,537]" lastPageId="14" lastPageNumber="165" pageId="13" pageNumber="164">
<emphasis id="09FB224FFF89704AADE9F81AFECDF87E" bold="true" box="[199,261,1985,2012]" pageId="13" pageNumber="164">
<emphasis id="09FB224FFF89704AADE9F81AFF35F879" bold="true" box="[199,253,1986,2012]" italics="true" pageId="13" pageNumber="164">Legs</emphasis>
.
</emphasis>
Carinae present; intercarinal areas with fine granulation; ventral telotarsi setae distributed in two rows of ventrosubmedian setae (
<typeStatus id="E43440FFFF8A7049AC88FF4FFE10FF14" box="[422,472,151,177]" pageId="14" pageNumber="165">type</typeStatus>
II), acute and stout; telotarsi, counts of ventral macrosetae in the left (L) and right (R) legs on prolateral (pro) and retrolateral (retro) rows from leg I to IV (L (pro/retro) R (pro/retro)): 6/5 6/6; 6/6 5/7; 6/7 6/7; -/- 7/8 (male) and 6/6 6/6; 6/6 7/7; 7/7 6/7; 6/7 7/7 (female). Claws short and symmetrical.
</paragraph>
<paragraph id="3B30FE5DFF8A7049ADE9FEDBFAF8FDBC" blockId="14.[151,1437,151,537]" pageId="14" pageNumber="165">
<emphasis id="09FB224FFF8A7049ADE9FEDBFE81FEB8" bold="true" box="[199,329,259,285]" pageId="14" pageNumber="165">
<taxonomicName id="FC8F85DEFF8A7049ADE9FEDBFE8AFEB8" box="[199,322,259,285]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="09FB224FFF8A7049ADE9FEDBFE8AFEB8" bold="true" box="[199,322,259,285]" italics="true" pageId="14" pageNumber="165">Mesosoma</emphasis>
</taxonomicName>
.
</emphasis>
Tergites I–VI (
<figureCitation id="A3B4E2D8FF8A7049ACDFFEDBFD4FFEB8" box="[497,647,259,285]" captionStart="FIGURE 2" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[180,1404,509,1847]" captionTargetId="figure-177@7.[164,1424,489,1864]" captionTargetPageId="7" captionText="FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6673340" httpUri="https://zenodo.org/record/6673340/files/figure.png" pageId="14" pageNumber="165">Figure 2A, C</figureCitation>
), moderately covered with coarse and fine granulation; pre-tergite well defined, with median carina visible on the posterior margin of the post-tergite; tergite VII with DSM and DL carinae complete and crenulate, and median carina composed of a crenulate anteromedian eminence present on the anterior half of the post-tergite. Sternites densely covered with coarse and fine granulation (
<figureCitation id="A3B4E2D8FF8A7049A919FEB7FB13FE2C" box="[1079,1243,367,393]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[221,1367,181,1920]" captionTargetId="figure-19@19.[221,1367,181,1920]" captionTargetPageId="19" captionText="FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673364" httpUri="https://zenodo.org/record/6673364/files/figure.png" pageId="14" pageNumber="165">Figure 10A, B</figureCitation>
); sternites III–VI with a pair of elliptic spiracles on the posterior half, which are progressively larger (
<figureCitation id="A3B4E2D8FF8A7049A917FE4BFB14FE08" box="[1081,1244,403,429]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[221,1367,181,1920]" captionTargetId="figure-19@19.[221,1367,181,1920]" captionTargetPageId="19" captionText="FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673364" httpUri="https://zenodo.org/record/6673364/files/figure.png" pageId="14" pageNumber="165">Figure 10A, B</figureCitation>
); sternites III–VI with a median longitudinal hyaline suture; sternite V with a large (male) (
<figureCitation id="A3B4E2D8FF8A7049AEC2FE6FFBBAFE74" box="[1004,1138,439,465]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[221,1367,181,1920]" captionTargetId="figure-19@19.[221,1367,181,1920]" captionTargetPageId="19" captionText="FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673364" httpUri="https://zenodo.org/record/6673364/files/figure.png" pageId="14" pageNumber="165">Figure 10A</figureCitation>
) to medium-size (female) (
<figureCitation id="A3B4E2D8FF8A7049ADB1FE03FED7FE50" box="[159,287,475,501]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[221,1367,181,1920]" captionTargetId="figure-19@19.[221,1367,181,1920]" captionTargetPageId="19" captionText="FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673364" httpUri="https://zenodo.org/record/6673364/files/figure.png" pageId="14" pageNumber="165">Figure 10B</figureCitation>
) hyaline subtriangular area on the posterior margin; sternite VI with VSM carinae crenulate, present on posterior half; sternite VII with VSM and VL carinae crenulate, present on posterior 2/3 (
<figureCitation id="A3B4E2D8FF8A7049A953FE27FAEAFDBC" box="[1149,1314,511,537]" captionStart="FIGURE 10" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[221,1367,181,1920]" captionTargetId="figure-19@19.[221,1367,181,1920]" captionTargetPageId="19" captionText="FIGURE 10. Tityus (Atreus) Gervais, 1843, sternites IV–VII. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402)." figureDoi="http://doi.org/10.5281/zenodo.6673364" httpUri="https://zenodo.org/record/6673364/files/figure.png" pageId="14" pageNumber="165">Figure 10A, B</figureCitation>
).
</paragraph>
<caption id="6FF0AED5FF8A7049ADB9F89FFC12F86E" ID-DOI="http://doi.org/10.5281/zenodo.6673354" ID-Zenodo-Dep="6673354" httpUri="https://zenodo.org/record/6673354/files/figure.png" pageId="14" pageNumber="165" startId="14.[151,250,1863,1887]" targetBox="[151,1436,561,1840]" targetPageId="14">
<paragraph id="3B30FE5DFF8A7049ADB9F89FFC12F86E" blockId="14.[151,1437,1863,1996]" pageId="14" pageNumber="165">
<emphasis id="09FB224FFF8A7049ADB9F89FFEDEF8FA" bold="true" box="[151,278,1863,1887]" pageId="14" pageNumber="165">FIGURE 6.</emphasis>
<taxonomicName id="FC8F85DEFF8A7049AC31F890FD9BF8C5" authority="Gervais, 1843" authorityName="Gervais" authorityYear="1843" box="[287,595,1864,1888]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="165" phylum="Arthropoda" rank="subGenus" subGenus="Atreus">
<emphasis id="09FB224FFF8A7049AC31F890FE94F8FA" box="[287,348,1864,1887]" italics="true" pageId="14" pageNumber="165">Tityus</emphasis>
(
<emphasis id="09FB224FFF8A7049AC43F890FE66F8FA" box="[365,430,1864,1887]" italics="true" pageId="14" pageNumber="165">Atreus</emphasis>
)
<bibRefCitation id="5F1E83ACFF8A7049ACEEF890FD9BF8C5" author="Gervais, P. M." box="[448,595,1864,1888]" pageId="14" pageNumber="165" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
</taxonomicName>
, patella sculpturing.
<emphasis id="09FB224FFF8A7049AE01F89FFCAFF8FA" bold="true" box="[815,871,1863,1887]" pageId="14" pageNumber="165">A–D.</emphasis>
<taxonomicName id="FC8F85DEFF8A7049AE5EF890FAD3F8C5" authority="(Gervais, 1843)" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[880,1307,1864,1888]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="forcipula" subGenus="Atreus">
<emphasis id="09FB224FFF8A7049AE5EF890FC65F8FA" box="[880,941,1864,1887]" italics="true" pageId="14" pageNumber="165">Tityus</emphasis>
(
<emphasis id="09FB224FFF8A7049AE90F890FC37F8FA" box="[958,1023,1864,1887]" italics="true" pageId="14" pageNumber="165">Atreus</emphasis>
)
<emphasis id="09FB224FFF8A7049A93FF890FBA5F8C5" box="[1041,1133,1864,1888]" italics="true" pageId="14" pageNumber="165">forcipula</emphasis>
(
<bibRefCitation id="5F1E83ACFF8A7049A953F890FADBF8C5" author="Gervais, P. M." box="[1149,1299,1864,1888]" pageId="14" pageNumber="165" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="09FB224FFF8A7049A808F89FFA97F8C5" bold="true" box="[1318,1375,1863,1888]" pageId="14" pageNumber="165">A, B.</emphasis>
Male (MUSENUV-Ar 438).
<emphasis id="09FB224FFF8A7049ACAFF8B3FE52F826" bold="true" box="[385,410,1899,1923]" pageId="14" pageNumber="165">A.</emphasis>
Dorsal view.
<emphasis id="09FB224FFF8A7049AF07F8B4FD88F821" bold="true" box="[553,576,1900,1924]" pageId="14" pageNumber="165">B.</emphasis>
Ventral view.
<emphasis id="09FB224FFF8A7049AFFAF8B3FCC4F821" bold="true" box="[724,780,1899,1924]" pageId="14" pageNumber="165">C, D.</emphasis>
Female (MUSENUV-Ar 436).
<emphasis id="09FB224FFF8A7049A961F8B3FBAFF826" bold="true" box="[1103,1127,1899,1923]" pageId="14" pageNumber="165">C.</emphasis>
Dorsal view.
<emphasis id="09FB224FFF8A7049A9D9F8B4FAC7F821" bold="true" box="[1271,1295,1900,1924]" pageId="14" pageNumber="165">D.</emphasis>
Ventral view.
<emphasis id="09FB224FFF8A7049ADB9F848FF07F80D" bold="true" box="[151,207,1936,1960]" pageId="14" pageNumber="165">E–H.</emphasis>
<taxonomicName id="FC8F85DEFF8A7049ADF8F848FE2BF80D" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[214,483,1936,1960]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov." subGenus="Atreus">
<emphasis id="09FB224FFF8A7049ADF8F848FEDBF802" box="[214,275,1936,1959]" italics="true" pageId="14" pageNumber="165">Tityus</emphasis>
(
<emphasis id="09FB224FFF8A7049AC0DF848FEACF802" box="[291,356,1936,1959]" italics="true" pageId="14" pageNumber="165">Atreus</emphasis>
)
<emphasis id="09FB224FFF8A7049AC5AF848FE2BF80D" box="[372,483,1936,1960]" italics="true" pageId="14" pageNumber="165">moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF8A7049ACC5F848FDBEF80D" bold="true" box="[491,630,1936,1960]" pageId="14" pageNumber="165">
<taxonomicNameLabel id="12C89F34FF8A7049ACC5F848FDF4F80D" box="[491,572,1936,1960]" pageId="14" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
E, F.
</emphasis>
Male holotype (MUSENUV-Ar 2104).
<emphasis id="09FB224FFF8A7049A93FF848FBE0F80D" bold="true" box="[1041,1064,1936,1960]" pageId="14" pageNumber="165">E.</emphasis>
Dorsal view.
<emphasis id="09FB224FFF8A7049A996F848FB03F80D" bold="true" box="[1208,1227,1936,1960]" pageId="14" pageNumber="165">F.</emphasis>
Ventral view.
<emphasis id="09FB224FFF8A7049A84FF857FA54F80D" bold="true" box="[1377,1436,1935,1960]" pageId="14" pageNumber="165">G, H.</emphasis>
Female paratype (ICN-1402).
<emphasis id="09FB224FFF8A7049ACE4F86BFE2CF86E" bold="true" box="[458,484,1971,1995]" pageId="14" pageNumber="165">G.</emphasis>
Dorsal view.
<emphasis id="09FB224FFF8A7049AF5EF86CFD43F869" bold="true" box="[624,651,1972,1996]" pageId="14" pageNumber="165">H.</emphasis>
Ventral view. Scale bars= 2 mm.
</paragraph>
</caption>
<caption id="6FF0AED5FF8B7048ADB9F84FFD12F852" ID-DOI="http://doi.org/10.5281/zenodo.6673356" ID-Zenodo-Dep="6673356" httpUri="https://zenodo.org/record/6673356/files/figure.png" pageId="15" pageNumber="166" startId="15.[151,250,1943,1967]" targetBox="[261,1326,181,1920]" targetPageId="15">
<paragraph id="3B30FE5DFF8B7048ADB9F84FFD12F852" blockId="15.[151,1436,1943,2040]" pageId="15" pageNumber="166">
<emphasis id="09FB224FFF8B7048ADB9F84FFED8F815" bold="true" box="[151,272,1943,1968]" pageId="15" pageNumber="166">FIGURE 7.</emphasis>
<taxonomicName id="FC8F85DEFF8B7048AC3AF840FDF1F815" authority="Gervais, 1843" authorityName="Gervais" authorityYear="1843" box="[276,569,1944,1968]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="166" phylum="Arthropoda" rank="subGenus" subGenus="Atreus">
<emphasis id="09FB224FFF8B7048AC3AF840FE99F80A" box="[276,337,1944,1967]" italics="true" pageId="15" pageNumber="166">Tityus</emphasis>
(
<emphasis id="09FB224FFF8B7048AC73F840FE56F80A" box="[349,414,1944,1967]" italics="true" pageId="15" pageNumber="166">Atreus</emphasis>
)
<bibRefCitation id="5F1E83ACFF8B7048AC85F840FDF1F815" author="Gervais, P. M." box="[427,569,1944,1968]" pageId="15" pageNumber="166" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
</taxonomicName>
, female chela sculpturing.
<emphasis id="09FB224FFF8B7048AE11F84FFCBEF80A" bold="true" box="[831,886,1943,1967]" pageId="15" pageNumber="166">A–C.</emphasis>
<taxonomicName id="FC8F85DEFF8B7048AE54F840FAC4F815" authority="(Gervais, 1843)" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[890,1292,1944,1968]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="forcipula" subGenus="Atreus">
<emphasis id="09FB224FFF8B7048AE54F840FC7FF80A" box="[890,951,1944,1967]" italics="true" pageId="15" pageNumber="166">Tityus</emphasis>
(
<emphasis id="09FB224FFF8B7048AEEDF840FBCCF80A" box="[963,1028,1944,1967]" italics="true" pageId="15" pageNumber="166">Atreus</emphasis>
)
<emphasis id="09FB224FFF8B7048A93EF840FBA4F815" box="[1040,1132,1944,1968]" italics="true" pageId="15" pageNumber="166">forcipula</emphasis>
(
<bibRefCitation id="5F1E83ACFF8B7048A959F840FACDF815" author="Gervais, P. M." box="[1143,1285,1944,1968]" pageId="15" pageNumber="166" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
)
</taxonomicName>
(MUSENUVAr 436).
<emphasis id="09FB224FFF8B7048ADDDF863FEC4F876" bold="true" box="[243,268,1979,2003]" pageId="15" pageNumber="166">A.</emphasis>
External view.
<emphasis id="09FB224FFF8B7048AC82F864FE0BF871" bold="true" box="[428,451,1980,2004]" pageId="15" pageNumber="166">B.</emphasis>
Dorsal view.
<emphasis id="09FB224FFF8B7048AF7CF863FDA2F876" bold="true" box="[594,618,1979,2003]" pageId="15" pageNumber="166">C.</emphasis>
Ventral view.
<emphasis id="09FB224FFF8B7048AFD0F864FCF8F871" bold="true" box="[766,816,1980,2004]" pageId="15" pageNumber="166">D–F.</emphasis>
<taxonomicName id="FC8F85DEFF8B7048AE19F864FB8AF871" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[823,1090,1980,2004]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov." subGenus="Atreus">
<emphasis id="09FB224FFF8B7048AE19F864FCBCF876" box="[823,884,1980,2003]" italics="true" pageId="15" pageNumber="166">Tityus</emphasis>
(
<emphasis id="09FB224FFF8B7048AEADF864FC0CF876" box="[899,964,1980,2003]" italics="true" pageId="15" pageNumber="166">Atreus</emphasis>
)
<emphasis id="09FB224FFF8B7048AEFDF864FB8AF871" box="[979,1090,1980,2004]" italics="true" pageId="15" pageNumber="166">moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF8B7048A964F864FB53F871" bold="true" box="[1098,1179,1980,2004]" pageId="15" pageNumber="166">
<taxonomicNameLabel id="12C89F34FF8B7048A964F864FB53F871" box="[1098,1179,1980,2004]" pageId="15" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(ICN-1402).
<emphasis id="09FB224FFF8B7048A809F864FA88F871" bold="true" box="[1319,1344,1980,2004]" pageId="15" pageNumber="166">D.</emphasis>
External view.
<emphasis id="09FB224FFF8B7048ADFDF838FF22F85D" bold="true" box="[211,234,2016,2040]" pageId="15" pageNumber="166">E.</emphasis>
Dorsal view.
<emphasis id="09FB224FFF8B7048AC59F838FE42F85D" bold="true" box="[375,394,2016,2040]" pageId="15" pageNumber="166">F.</emphasis>
Ventral view. Scale bars= 3 mm.
</paragraph>
</caption>
<caption id="6FF0AED5FF947057ADB9F840FCFFF85D" ID-DOI="http://doi.org/10.5281/zenodo.6673358" ID-Zenodo-Dep="6673358" httpUri="https://zenodo.org/record/6673358/files/figure.png" pageId="16" pageNumber="167" startId="16.[151,250,1944,1968]" targetBox="[364,1224,181,1920]" targetPageId="16">
<paragraph id="3B30FE5DFF947057ADB9F840FCFFF85D" blockId="16.[151,1436,1944,2040]" pageId="16" pageNumber="167">
<emphasis id="09FB224FFF947057ADB9F840FEDAF815" bold="true" box="[151,274,1944,1968]" pageId="16" pageNumber="167">FIGURE 8.</emphasis>
<taxonomicName id="FC8F85DEFF947057AC39F841FDF7F815" authority="Gervais, 1843" authorityName="Gervais" authorityYear="1843" box="[279,575,1944,1968]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="167" phylum="Arthropoda" rank="subGenus" subGenus="Atreus">
<emphasis id="09FB224FFF947057AC39F841FE9CF815" box="[279,340,1945,1968]" italics="true" pageId="16" pageNumber="167">Tityus</emphasis>
(
<emphasis id="09FB224FFF947057AC4FF840FE6AF80A" box="[353,418,1944,1967]" italics="true" pageId="16" pageNumber="167">Atreus</emphasis>
)
<bibRefCitation id="5F1E83ACFF947057AC9EF840FDF7F815" author="Gervais, P. M." box="[432,575,1944,1968]" pageId="16" pageNumber="167" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
</taxonomicName>
, male chela sculpturing.
<emphasis id="09FB224FFF947057AE19F840FCA7F815" bold="true" box="[823,879,1944,1968]" pageId="16" pageNumber="167">A–C.</emphasis>
<taxonomicName id="FC8F85DEFF947057AE5DF841FAC3F815" authority="(Gervais, 1843)" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[883,1291,1944,1968]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="forcipula" subGenus="Atreus">
<emphasis id="09FB224FFF947057AE5DF841FC78F815" box="[883,944,1945,1968]" italics="true" pageId="16" pageNumber="167">Tityus</emphasis>
(
<emphasis id="09FB224FFF947057AE90F840FC37F80A" box="[958,1023,1944,1967]" italics="true" pageId="16" pageNumber="167">Atreus</emphasis>
)
<emphasis id="09FB224FFF947057A922F840FBA0F815" box="[1036,1128,1944,1968]" italics="true" pageId="16" pageNumber="167">forcipula</emphasis>
(
<bibRefCitation id="5F1E83ACFF947057A95BF840FACCF815" author="Gervais, P. M." box="[1141,1284,1944,1968]" pageId="16" pageNumber="167" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
)
</taxonomicName>
(MUSENUVAr 438).
<emphasis id="09FB224FFF947057ADDFF864FEC2F871" bold="true" box="[241,266,1980,2004]" pageId="16" pageNumber="167">A.</emphasis>
External view.
<emphasis id="09FB224FFF947057AC89F864FE76F871" bold="true" box="[423,446,1980,2004]" pageId="16" pageNumber="167">B.</emphasis>
Dorsal view.
<emphasis id="09FB224FFF947057AF67F864FDAAF871" bold="true" box="[585,610,1980,2004]" pageId="16" pageNumber="167">C.</emphasis>
Ventral view.
<emphasis id="09FB224FFF947057AFDCF864FCECF871" bold="true" box="[754,804,1980,2004]" pageId="16" pageNumber="167">D–F.</emphasis>
<taxonomicName id="FC8F85DEFF947057AE04F865FBFCF871" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[810,1076,1980,2004]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov." subGenus="Atreus">
<emphasis id="09FB224FFF947057AE04F865FCAFF871" box="[810,871,1981,2004]" italics="true" pageId="16" pageNumber="167">Tityus</emphasis>
(
<emphasis id="09FB224FFF947057AE5BF864FC7EF876" box="[885,950,1980,2003]" italics="true" pageId="16" pageNumber="167">Atreus</emphasis>
)
<emphasis id="09FB224FFF947057AEEBF864FBFCF871" box="[965,1076,1980,2004]" italics="true" pageId="16" pageNumber="167">moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF947057A914F864FB42F871" bold="true" box="[1082,1162,1980,2004]" pageId="16" pageNumber="167">
<taxonomicNameLabel id="12C89F34FF947057A914F864FB42F871" box="[1082,1162,1980,2004]" pageId="16" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MUSENUV-Ar 2104).
<emphasis id="09FB224FFF947057A8AAF864FA54F871" bold="true" box="[1412,1436,1980,2004]" pageId="16" pageNumber="167">D.</emphasis>
External view.
<emphasis id="09FB224FFF947057AC1EF838FE8EF85D" bold="true" box="[304,326,2016,2040]" pageId="16" pageNumber="167">E.</emphasis>
Dorsal view.
<emphasis id="09FB224FFF947057ACFDF838FE2EF85D" bold="true" box="[467,486,2016,2040]" pageId="16" pageNumber="167">F.</emphasis>
Ventral view. Scale bars= 3 mm.
</paragraph>
</caption>
<caption id="6FF0AED5FF957056ADB9FB05FE2AFA98" ID-DOI="http://doi.org/10.5281/zenodo.6673360" ID-Zenodo-Dep="6673360" httpUri="https://zenodo.org/record/6673360/files/figure.png" pageId="17" pageNumber="168" startId="17.[151,250,1245,1269]" targetBox="[163,1424,181,1221]" targetPageId="17">
<paragraph id="3B30FE5DFF957056ADB9FB05FE2AFA98" blockId="17.[151,1437,1245,1341]" pageId="17" pageNumber="168">
<emphasis id="09FB224FFF957056ADB9FB05FEDAFB50" bold="true" box="[151,274,1245,1269]" pageId="17" pageNumber="168">FIGURE 9.</emphasis>
<taxonomicName id="FC8F85DEFF957056AC36FB06FD8AFB50" authority="Gervais, 1843" authorityName="Gervais" authorityYear="1843" box="[280,578,1245,1269]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="168" phylum="Arthropoda" rank="subGenus" subGenus="Atreus">
<emphasis id="09FB224FFF957056AC36FB06FE9DFB50" box="[280,341,1246,1269]" italics="true" pageId="17" pageNumber="168">Tityus</emphasis>
(
<emphasis id="09FB224FFF957056AC4DFB05FE6CFB51" box="[355,420,1245,1268]" italics="true" pageId="17" pageNumber="168">Atreus</emphasis>
)
<bibRefCitation id="5F1E83ACFF957056AC9CFB05FD8AFB50" author="Gervais, P. M." box="[434,578,1245,1269]" pageId="17" pageNumber="168" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
</taxonomicName>
, pectinal area.
<emphasis id="09FB224FFF957056AFF6FB05FCC6FB50" bold="true" box="[728,782,1245,1269]" pageId="17" pageNumber="168">A, B.</emphasis>
<taxonomicName id="FC8F85DEFF957056AE3DFB06FB79FB50" authority="(Gervais, 1843)" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[787,1201,1245,1269]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="forcipula" subGenus="Atreus">
<emphasis id="09FB224FFF957056AE3DFB06FC98FB50" box="[787,848,1246,1269]" italics="true" pageId="17" pageNumber="168">Tityus</emphasis>
(
<emphasis id="09FB224FFF957056AE70FB05FC57FB51" box="[862,927,1245,1268]" italics="true" pageId="17" pageNumber="168">Atreus</emphasis>
)
<emphasis id="09FB224FFF957056AE83FB05FBC1FB50" box="[941,1033,1245,1269]" italics="true" pageId="17" pageNumber="168">forcipula</emphasis>
(
<bibRefCitation id="5F1E83ACFF957056A938FB05FB60FB50" author="Gervais, P. M." box="[1046,1192,1245,1269]" pageId="17" pageNumber="168" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="09FB224FFF957056A996FB05FB19FB50" bold="true" box="[1208,1233,1245,1269]" pageId="17" pageNumber="168">A.</emphasis>
Male (MUSENUVAr 438).
<emphasis id="09FB224FFF957056ADDAFAD9FEC3FABC" bold="true" box="[244,267,1281,1305]" pageId="17" pageNumber="168">B.</emphasis>
Female (MUSENUV-Ar 436).
<emphasis id="09FB224FFF957056AF62FAD9FD4CFABC" bold="true" box="[588,644,1281,1305]" pageId="17" pageNumber="168">C, D.</emphasis>
<taxonomicName id="FC8F85DEFF957056AFA4FADAFC5DFABC" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[650,917,1281,1305]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov." subGenus="Atreus">
<emphasis id="09FB224FFF957056AFA4FADAFD0FFABC" box="[650,711,1282,1305]" italics="true" pageId="17" pageNumber="168">Tityus</emphasis>
(
<emphasis id="09FB224FFF957056AFF8FAD9FCDFFABD" box="[726,791,1281,1304]" italics="true" pageId="17" pageNumber="168">Atreus</emphasis>
)
<emphasis id="09FB224FFF957056AE08FAD9FC5DFABC" box="[806,917,1281,1305]" italics="true" pageId="17" pageNumber="168">moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF957056AEB2FAD9FBC5FABC" bold="true" box="[924,1037,1281,1305]" pageId="17" pageNumber="168">
<taxonomicNameLabel id="12C89F34FF957056AEB2FAD9FC25FABC" box="[924,1005,1281,1305]" pageId="17" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
C.
</emphasis>
Male holotype (MUSENUV-Ar 2104).
<emphasis id="09FB224FFF957056ADB9FAFDFF78FA98" bold="true" box="[151,176,1317,1341]" pageId="17" pageNumber="168">D.</emphasis>
Female paratype (ICN-1402).
</paragraph>
</caption>
<paragraph id="3B30FE5DFF957056ADE9FAB3FDEFF96C" blockId="17.[151,1437,1387,2026]" pageId="17" pageNumber="168">
<emphasis id="09FB224FFF957056ADE9FAB3FD4AFA20" bold="true" box="[199,642,1387,1414]" pageId="17" pageNumber="168">
<emphasis id="09FB224FFF957056ADE9FAB3FE7BFA23" bold="true" box="[199,435,1387,1414]" italics="true" pageId="17" pageNumber="168">Hemispermatophore</emphasis>
(
<figureCitation id="A3B4E2D8FF957056ACE8FAB3FDBDFA20" box="[454,629,1387,1413]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="17" pageNumber="168">Figure 11A–C</figureCitation>
).
</emphasis>
Thin and sclerotized; pars reflexa, curved but not folded over itself (
<figureCitation id="A3B4E2D8FF957056ADB1FA57FE8BFA0F" box="[159,323,1423,1450]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="17" pageNumber="168">Figure 11A, B</figureCitation>
); foot narrow and flat (
<figureCitation id="A3B4E2D8FF957056AF68FA57FD25FA0C" box="[582,749,1423,1449]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="17" pageNumber="168">Figure 11A–C</figureCitation>
); pedal flexure inconspicuous (
<figureCitation id="A3B4E2D8FF957056A967FA57FB27FA0C" box="[1097,1263,1423,1449]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="17" pageNumber="168">Figure 11A–C</figureCitation>
); body occupying more than 2/3 of the hemispermatophore total length (
<figureCitation id="A3B4E2D8FF957056AE38FA6BFC74FA68" box="[790,956,1459,1485]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="17" pageNumber="168">Figure 11A–C</figureCitation>
). Capsular region with a furrow connecting the internal lobe with the medial area of the region (
<figureCitation id="A3B4E2D8FF957056AFDDFA0FFCB2FA54" box="[755,890,1495,1521]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="17" pageNumber="168">Figure 11A</figureCitation>
); internal lobe with rounded tip forming an 80º angle (
<figureCitation id="A3B4E2D8FF957056ADCCFA23FEACF9B0" box="[226,356,1531,1557]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="17" pageNumber="168">Figure 11B</figureCitation>
); median lobe inconspicuous (
<figureCitation id="A3B4E2D8FF957056AF99FA23FC92F9B0" box="[695,858,1531,1557]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="17" pageNumber="168">Figure 11B, C</figureCitation>
); external lobe thin and not overpassing the internal lobe level and with a translucent area between the middle of the basal lobe and the base of the internal lobe (
<figureCitation id="A3B4E2D8FF957056A87FF9C7FF00F9F8" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="17" pageNumber="168">Figure 11C</figureCitation>
); translucent area narrow but widened basally and distally; basal lobe bifurcate and with a vestigial projection subovate-shaped (in posterior and ventral views) (
<figureCitation id="A3B4E2D8FF957056AFEAF9BFFCA0F924" box="[708,872,1639,1665]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="17" pageNumber="168">Figure 11A, C</figureCitation>
); basal lobe with no conspicuous visible surface in lateral view (
<figureCitation id="A3B4E2D8FF957056AC08F953FE60F900" box="[294,424,1675,1701]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="17" pageNumber="168">Figure 11B</figureCitation>
); basal lobe without a “U”-like shaped curvature between its base and the body (in posterior and ventral views) (
<figureCitation id="A3B4E2D8FF957056AC5BF977FDD2F96C" box="[373,538,1711,1737]" captionStart="FIGURE 11" captionStartId="20.[151,250,1886,1910]" captionTargetBox="[224,1363,181,1862]" captionTargetId="figure-31@20.[224,1363,181,1862]" captionTargetPageId="20" captionText="FIGURE 11. Tityus (Atreus) Gervais, 1843, hemispermatophore. A–C. Tityus (Atreus) forcipula (Gervais, 1843) (MUSENUV- Ar 438). A. Posterior view. B. Lateral view. C. Ventral view. D–F. Tityus (Atreus) moralensis sp. nov. (MUSENUV-Ar 2104). D. Posterior view. E. Lateral view. F. Ventral view. Abbreviations= B, body; Bl, basal lobe; El, external lobe; F, flagellum; Fo, foot; iL, internal lobe; Pref, pars reflexa; Prec, pars recta. Scale bars= 2 mm. Scale bars= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="17" pageNumber="168">Figure 11A, C</figureCitation>
).
</paragraph>
<paragraph id="3B30FE5DFF957055ADE9F90BFD05FEE4" blockId="17.[151,1437,1387,2026]" lastBlockId="18.[151,1437,151,2014]" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168">
<emphasis id="09FB224FFF957056ADE9F90BFD04F948" bold="true" box="[199,716,1747,1773]" pageId="17" pageNumber="168">
<emphasis id="09FB224FFF957056ADE9F90BFEF7F948" bold="true" box="[199,319,1747,1773]" italics="true" pageId="17" pageNumber="168">Metasoma</emphasis>
(
<figureCitation id="A3B4E2D8FF957056AC63F90BFDC9F948" box="[333,513,1747,1773]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="17" pageNumber="168">Figures 12A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF957056AF23F90BFD97F948" box="[525,607,1747,1773]" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="17" pageNumber="168">13A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF957056AF45F90BFD75F948" box="[619,701,1747,1773]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="17" pageNumber="168">14A, B</figureCitation>
).
</emphasis>
moderately widening towards segment V (
<figureCitation id="A3B4E2D8FF957056A985F90BFA92F94B" box="[1195,1370,1747,1774]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="17" pageNumber="168">Figures 12A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF957056A84BF90BFF64F8B7" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="17" pageNumber="168">13A, B</figureCitation>
). Segments II–V short and robust (male- L/W ratio: II= 1.29; III= 1.38; IV= 1.39; V= 1.53/ female- L/W ratio: II= 1.32; III= 1.39; IV= 1.46; V= 1.53). Segments I–II (
<figureCitation id="A3B4E2D8FF957056AFCEF8C3FC59F893" box="[736,913,1819,1846]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="17" pageNumber="168">Figures 12A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF957056AEB3F8C3FC38F893" box="[925,1008,1819,1846]" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="17" pageNumber="168">13A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF957056AED2F8C3FB86F893" box="[1020,1102,1819,1846]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="17" pageNumber="168">14A, B</figureCitation>
) with eight complete carinae, parallel and crenulate (paired DL, ML, VL and VSM) and two serratocrenulate (paired DSM); ML of segment II represented by coarse granules on posterior 2/3 and fine granule over the anterior third; intercarinal areas densely covered with fine granulation and moderately covered with coarse granulation; segments III–IV (
<figureCitation id="A3B4E2D8FF957056A9CBF85FFA50F807" box="[1253,1432,1927,1954]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="17" pageNumber="168">Figures 12A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF957056ADB9F873FF22F863" box="[151,234,1963,1990]" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="17" pageNumber="168">13A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF957056ADD8F873FE8FF863" box="[246,327,1963,1990]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="17" pageNumber="168">14A, B</figureCitation>
) with six complete carinae, parallel and crenulate (paired DL, VL and VSM) and two serratocrenulate (paired DSM), intercarinal areas densely covered with fine granulation and moderately covered with coarse granulation; segment V (
<figureCitation id="A3B4E2D8FF967055AC87FF4FFD90FF14" box="[425,600,151,177]" captionStart="FIGURE 12" captionStartId="21.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1724]" captionTargetId="figure-19@21.[151,1435,181,1724]" captionTargetPageId="21" captionText="FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="18" pageNumber="169">Figures 12A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF967055AF4DFF4FFD7DFF14" box="[611,693,151,177]" captionStart="FIGURE 13" captionStartId="22.[151,250,1695,1719]" captionTargetBox="[151,1436,181,1671]" captionTargetId="figure-31@22.[151,1436,181,1671]" captionTargetPageId="22" captionText="FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="18" pageNumber="169">13A, B</figureCitation>
,
<figureCitation id="A3B4E2D8FF967055AFEEFF4FFCD8FF14" box="[704,784,151,177]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="18" pageNumber="169">14A, B</figureCitation>
) with five complete carinae and crenulate (VM, paired VL and VSM carinae) and two serratocrenulate (paired DSM), intercarinal areas densely covered with coarse granulation and moderately covered with fine granulation. Segments II–V (
<figureCitation id="A3B4E2D8FF967055AEABFF07FBEEFF5C" box="[901,1062,223,249]" captionStart="FIGURE 14" captionStartId="23.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@23.[151,1436,181,1309]" captionTargetPageId="23" captionText="FIGURE 14. Tityus (Atreus) Gervais, 1843, metasoma, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673374" httpUri="https://zenodo.org/record/6673374/files/figure.png" pageId="18" pageNumber="169">Figure 14A, B</figureCitation>
) with DSM carinae, composed of strongly protruding spinoid granules that progressively increase distally, ending in an enlarged distoterminal granule (distoterminal granule not enlarged in segment V).
</paragraph>
<paragraph id="3B30FE5DFF967055ADE9FE92FBB2FD98" blockId="18.[151,1437,151,2014]" pageId="18" pageNumber="169">
<emphasis id="09FB224FFF967055ADE9FE92FE3FFEC0" bold="true" box="[199,503,330,357]" pageId="18" pageNumber="169">
<emphasis id="09FB224FFF967055ADE9FE92FE38FEC0" bold="true" box="[199,496,330,357]" italics="true" pageId="18" pageNumber="169">Metasomal macrosetation</emphasis>
.
</emphasis>
Segments I–IV each with two pairs of VSM macrosetae (2/2): pair VSM1 located in the anterior 1/3, and pair VSM3 located near the posterior margin of the segment, and two pairs of VL macrosetae (2/2): pair VL1 located near the anterior margin, and pair VL2 located in the posterior 2/3 of the segment. Segment V with two pairs of VSM macrosetae (2/2), two pairs of VL macrosetae (2/2), and a single pair of ML macrosetae; pair VSM1 and VL1 located near the anterior margin of the segment; VL2 located on the posterior 2/3, and pair ML1 located dorsolaterally near the posterior margin (behind DSM carinae); anal arch with two pairs of setae on the intercrestal area, one pair of VSM macrosetae (1/1) and one pair of VL macrosetae (1/1).
</paragraph>
<paragraph id="3B30FE5DFF967055ADE9FD9EFC7EFCB0" blockId="18.[151,1437,151,2014]" pageId="18" pageNumber="169">
<emphasis id="09FB224FFF967055ADE9FD9EFE28FDC5" bold="true" box="[199,480,582,609]" pageId="18" pageNumber="169">
<taxonomicName id="FC8F85DEFF967055ADE9FD9EFEDAFDC4" box="[199,274,582,609]" class="Hexanauplia" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="09FB224FFF967055ADE9FD9EFEDAFDC4" bold="true" box="[199,274,582,609]" italics="true" pageId="18" pageNumber="169">Telson</emphasis>
</taxonomicName>
(
<figureCitation id="A3B4E2D8FF967055AC0DFD9EFE19FDC5" box="[291,465,582,608]" captionStart="FIGURE 15" captionStartId="24.[151,250,1090,1114]" captionTargetBox="[151,1436,345,1067]" captionTargetId="figure-116@24.[151,1436,345,1067]" captionTargetPageId="24" captionText="FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673376" httpUri="https://zenodo.org/record/6673376/files/figure.png" pageId="18" pageNumber="169">Figure 15A, B</figureCitation>
).
</emphasis>
Covered with abundant microsetae, white colored under UV light (
<figureCitation id="A3B4E2D8FF967055A9CBFD9FFA46FDC4" box="[1253,1422,582,609]" captionStart="FIGURE 15" captionStartId="24.[151,250,1090,1114]" captionTargetBox="[151,1436,345,1067]" captionTargetId="figure-116@24.[151,1436,345,1067]" captionTargetPageId="24" captionText="FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673376" httpUri="https://zenodo.org/record/6673376/files/figure.png" pageId="18" pageNumber="169">Figure 15A, B</figureCitation>
). Vesicle suboval, not elongated (L/H= 1.75 (male)/ L/W= 1.85 (female)), with dorsal surface smooth and a lateral longitudinal furrow on each side; with VM, paired VSM, VL, and DL carinae, vestigial and moderately marked. Subaculear tubercle small pyramidal, with a rounded apex directed to the middle portion of the aculeus (
<figureCitation id="A3B4E2D8FF967055A83AFD6BFF64FD54" captionStart="FIGURE 15" captionStartId="24.[151,250,1090,1114]" captionTargetBox="[151,1436,345,1067]" captionTargetId="figure-116@24.[151,1436,345,1067]" captionTargetPageId="24" captionText="FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6673376" httpUri="https://zenodo.org/record/6673376/files/figure.png" pageId="18" pageNumber="169">Figure 15A, B</figureCitation>
); subaculear tubercle with a ventral pair of small rounded granules, pointing to the basal portion of the aculeus; aculeus strongly curved, shorter than vesicle and with a ventral groove.
</paragraph>
<paragraph id="3B30FE5DFF967055ADE9FCC6FB8BFBD8" blockId="18.[151,1437,151,2014]" pageId="18" pageNumber="169">
<emphasis id="09FB224FFF967055ADE9FCC6FDC5FC9C" bold="true" box="[199,525,798,825]" pageId="18" pageNumber="169">
Variability.
<emphasis id="09FB224FFF967055AC7BFCC6FDCEFC9C" bold="true" box="[341,518,798,825]" italics="true" pageId="18" pageNumber="169">Morphometrics</emphasis>
.
</emphasis>
<emphasis id="09FB224FFF967055AF3BFCC7FD54FC9C" box="[533,668,799,825]" italics="true" pageId="18" pageNumber="169">Total length</emphasis>
(
<emphasis id="09FB224FFF967055AF80FCC7FCAAFC9C" box="[686,866,799,825]" italics="true" pageId="18" pageNumber="169">including telson</emphasis>
): 52.98–62.30 (female) (n= 4, mean= 57.53, standard deviation (SD)= 5.19) and 49.24–53.81 (male) (n= 4, mean= 51.99, SD= 1.95).
<emphasis id="09FB224FFF967055A94DFC9BFAD3FCF9" box="[1123,1307,835,861]" italics="true" pageId="18" pageNumber="169">Chela L/W ratio</emphasis>
: 5.15–5.31 (female) (n= 4; mean= 5.02; SD= 0.30) and 3.61–3.71 (male) (n= 4; mean= 3.84; SD= 0.33).
<emphasis id="09FB224FFF967055A983FCBFFECBFC01" italics="true" pageId="18" pageNumber="169">Metasomal segment I L/W ratio</emphasis>
: 0.92–1.10 (female) (n= 4; mean= 1.02; SD= 0.08) and 1.05–1.15 (male) (n= 4; mean= 1.09; SD= 0.05).
<emphasis id="09FB224FFF967055ADB9FC77FDCFFC6D" box="[151,519,943,969]" italics="true" pageId="18" pageNumber="169">Metasomal segment V L/W ratio</emphasis>
: 1.42–1.53 (female) (n= 4; mean= 1.48; SD= 0.05) and 1.36–1.53 (male) (n= 4; mean= 1.46; SD= 0.08).
<emphasis id="09FB224FFF967055AC87FC0BFDBCFC48" box="[425,628,979,1005]" italics="true" pageId="18" pageNumber="169">
<taxonomicName id="FC8F85DEFF967055AC87FC0BFD8BFC48" box="[425,579,979,1005]" class="Hexanauplia" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="vesicle">Telson vesicle</taxonomicName>
L/H
</emphasis>
: 1.70–1.86 (female) (n= 4; mean= 1.80; SD= 0.08) and 1.68–1.85 (male) (n= 4; mean= 1.76; SD= 0.07).
<emphasis id="09FB224FFF967055ACDEFC2FFCF0FBB5" box="[496,824,1015,1041]" italics="true" pageId="18" pageNumber="169">Counts. Pectinal tooth counts</emphasis>
: females 14–18 (n= 8, mode= 16) and males 14–18 (n= 8, mode= 17).
<emphasis id="09FB224FFF967055AC12FBC3FC90FB91" box="[316,856,1051,1077]" italics="true" pageId="18" pageNumber="169">Number of movable finger oblique granule rows</emphasis>
: females 15–17 (n= 8, mode= 16) and males 16–17 (n= 8, mode= 16).
<emphasis id="09FB224FFF967055AC41FBE7FDBCFBFD" box="[367,628,1087,1113]" italics="true" pageId="18" pageNumber="169">Metasomal macrosetae</emphasis>
: The specimens from
<collectingRegion id="F94B30BFFF967055AE5DFBE7FBABFBFC" box="[883,1123,1087,1113]" country="Colombia" name="Risaralda" pageId="18" pageNumber="169">Risaralda department</collectingRegion>
exhibit three pairs of VSM macrosetae in segments I–IV, instead of two pairs as the other examined specimens.
</paragraph>
<paragraph id="3B30FE5DFF967055ADE9FB5FFACDFADC" blockId="18.[151,1437,151,2014]" pageId="18" pageNumber="169">
<emphasis id="09FB224FFF967055ADE9FB5FFE4FFB05" bold="true" box="[199,391,1158,1185]" pageId="18" pageNumber="169">Natural history.</emphasis>
<bibRefCitation id="5F1E83ACFF967055ACBEFB5FFCD0FB04" author="Moreno-Gonzalez, J. A. &amp; Hazzi, N. A." box="[400,792,1159,1185]" pageId="18" pageNumber="169" pagination="117 - 120" refId="ref19950" refString="Moreno-Gonzalez, J. A. &amp; Hazzi, N. A. (2012) Intraguild predation case: Tityus forcipula Gervais, 1843 (Scorpiones, Buthidae) feeding on Chactas vanbenedeni Gervais, 1843 (Scorpiones, Chactidae) in Colombia. Revista Iberica de Aracnologia, 20, 117 - 120." type="journal article" year="2012">Moreno-González &amp; Hazzi (2012)</bibRefCitation>
observed a female of this species feeding upon an adult female of
<taxonomicName id="FC8F85DEFF967055AC22FB73FD08FB60" authority="(Gervais, 1843)" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[268,704,1195,1221]" class="Arachnida" family="Chactidae" genus="Chactas" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="vanbenedenii">
<emphasis id="09FB224FFF967055AC22FB73FDCEFB60" box="[268,518,1195,1221]" italics="true" pageId="18" pageNumber="169">Chactas vanbenedenii</emphasis>
(
<bibRefCitation id="5F1E83ACFF967055AF38FB73FD70FB60" author="Gervais, P. M." box="[534,696,1195,1221]" pageId="18" pageNumber="169" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
)
</taxonomicName>
in Yotoco Forest Reserve, Yotoco,
<collectingRegion id="F94B30BFFF967055A97BFB73FA50FB60" box="[1109,1432,1195,1221]" country="Colombia" name="Valle del Cauca" pageId="18" pageNumber="169">Valle del Cauca Department</collectingRegion>
,
<collectingCountry id="4398BECDFF967055ADB9FB17FEC1FB4C" box="[151,265,1231,1257]" name="Colombia" pageId="18" pageNumber="169">Colombia</collectingCountry>
.
<bibRefCitation id="5F1E83ACFF967055AC3DFB17FE2AFB4C" author="Seiter, M. &amp; Friedl, N. &amp; Cozijn, M. A. C." box="[275,482,1231,1257]" pageId="18" pageNumber="169" pagination="161 - 168" refId="ref20906" refString="Seiter, M., Friedl, N. &amp; Cozijn, M. A. C. (2020) Life history aspects of the buthid scorpion Tityus forcipula (Gervais, 1843) with remarks on its thermal limits. Journal of Arachnology, 48, 161 - 168. https: // doi. org / 10.1636 / 0161 - 8202 - 48.2.161" type="journal article" year="2020">
Seiter
<emphasis id="09FB224FFF967055AC74FB17FE59FB4C" box="[346,401,1231,1257]" italics="true" pageId="18" pageNumber="169">et al.</emphasis>
(2020)
</bibRefCitation>
studied life story aspects of
<taxonomicName id="FC8F85DEFF967055AE32FB17FC55FB4C" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[796,925,1231,1257]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF967055AE32FB17FC55FB4C" box="[796,925,1231,1257]" italics="true" pageId="18" pageNumber="169">T. forcipula</emphasis>
</taxonomicName>
and observed that specimens reared in captivity are unable to thrive under high temperatures. Only 10% of the specimens kept at 25–27°C reached adulthood, none reproduced. Among
<specimenCount id="2D8935D4FF967055AC90FACFFD90FA94" box="[446,600,1303,1329]" pageId="18" pageNumber="169" type="generic">43 specimens</specimenCount>
kept between 23–24°C,
<specimenCount id="2D8935D4FF967055AE5CFACFFC25FA94" box="[882,1005,1303,1329]" pageId="18" pageNumber="169" type="female">21 females</specimenCount>
and
<specimenCount id="2D8935D4FF967055A909FACFFB44FA94" box="[1063,1164,1303,1329]" pageId="18" pageNumber="169" type="male">19 males</specimenCount>
reached maturity in the fifth instar, but
<specimenCount id="2D8935D4FF967055AC6EFAE3FE72FAF0" box="[320,442,1339,1365]" pageId="18" pageNumber="169" type="female">one female</specimenCount>
and
<specimenCount id="2D8935D4FF967055ACC0FAE3FD97FAF0" box="[494,607,1339,1365]" pageId="18" pageNumber="169" type="male">two males</specimenCount>
required an extra molt to reach maturity. The embryonic development took 208 days (with an average of 12 young per litter
<emphasis id="09FB224FFF967055AF9CFA87FD73FADC" box="[690,699,1375,1401]" italics="true" pageId="18" pageNumber="169">)</emphasis>
and the postembryonic development took 463 days.
</paragraph>
<paragraph id="3B30FE5DFF967055ADD9FA5BFB7EF9F7" blockId="18.[151,1437,151,2014]" pageId="18" pageNumber="169">
This species has been observed living in sympatry with
<taxonomicName id="FC8F85DEFF967055AE5BFA5BFBA6FA38" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[885,1134,1411,1437]" class="Arachnida" family="Chactidae" genus="Chactas" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="vanbenedenii">
<emphasis id="09FB224FFF967055AE5BFA5BFBA6FA38" box="[885,1134,1411,1437]" italics="true" pageId="18" pageNumber="169">Chactas vanbenedenii</emphasis>
</taxonomicName>
and
<taxonomicName id="FC8F85DEFF967055A989FA5BFEDEFA64" authority="Kraepelin, 1914" authorityName="Kraepelin" authorityYear="1914" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="09FB224FFF967055A989FA5BFA9CFA38" box="[1191,1364,1411,1437]" italics="true" pageId="18" pageNumber="169">Tityus parvulus</emphasis>
<bibRefCitation id="5F1E83ACFF967055A872FA5BFEDEFA64" author="Kraepelin, K." pageId="18" pageNumber="169" pagination="15 - 28" refId="ref18356" refString="Kraepelin, K. (1914) Beitrag zur Kenntnis der Skorpione und Pedipalpen Columbiens. Memoires de la Societe des Sciences naturelles, Neuchatel, 5, 15 - 28." type="journal article" year="1914">Kraepelin, 1914</bibRefCitation>
</taxonomicName>
across several high-altitude (over
<quantity id="FC7753B8FF967055AFB5FA7FFCDAFA67" box="[667,786,1447,1474]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="169" unit="m" value="1600.0">1600 masl</quantity>
) localities in the Cordillera Occidental in
<collectingRegion id="F94B30BFFF967055A9C5FA7FFED0FA40" country="Colombia" name="Valle del Cauca" pageId="18" pageNumber="169">Valle del Cauca department</collectingRegion>
,
<collectingCountry id="4398BECDFF967055AC0DFA13FE5DFA40" box="[291,405,1483,1509]" name="Colombia" pageId="18" pageNumber="169">Colombia</collectingCountry>
. The microhabitat that this species seems to prefer is underside rotten logs, inside leaf litter, or under rocks, and rarely it has been observed under tree bark over
<quantity id="FC7753B8FF967055AEB2FA37FC00F9AC" box="[924,968,1519,1545]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="169" unit="m" value="1.0">1 m</quantity>
above the soil level. A strong association between adult specimens and low vegetation has never been observed, maybe this relationship is due to the particular telotarsi macrosetae structure that this species exhibit (Moreno-González, J. A., pers. obs.).
</paragraph>
</subSubSection>
<subSubSection id="7395ADD6FF967051ADD9F982FC47F816" lastPageId="22" lastPageNumber="173" pageId="18" pageNumber="169" type="distribution">
<paragraph id="3B30FE5DFF967055ADD9F982FCCCF88C" blockId="18.[151,1437,151,2014]" pageId="18" pageNumber="169">
<emphasis id="09FB224FFF967055ADD9F982FDEBF9D1" bold="true" box="[247,547,1626,1652]" pageId="18" pageNumber="169">
Distribution (
<figureCitation id="A3B4E2D8FF967055ACB5F982FDDCF9D1" box="[411,532,1626,1652]" captionStart="FIGURE 16" captionStartId="27.[151,250,1428,1452]" captionTargetBox="[184,1412,353,1404]" captionTargetId="figure-95@27.[176,1412,345,1404]" captionTargetPageId="27" captionText="FIGURE 16. Distribution map of the Tityus forcipula species group. A. South America. B. Records area. C. Species records." figureDoi="http://doi.org/10.5281/zenodo.6673378" httpUri="https://zenodo.org/record/6673378/files/figure.png" pageId="18" pageNumber="169">Figure 16</figureCitation>
).
</emphasis>
We opted to follow a rather conservative approach. As such, we considered as doubtful records those specimens that we were not able to examine (i.e., Ecuadorian Andes and Central and Oriental Cordilleras from
<collectingCountry id="4398BECDFF967055AC93F97BFDE5F91B" box="[445,557,1699,1726]" name="Colombia" pageId="18" pageNumber="169">Colombia</collectingCountry>
) (
<figureCitation id="A3B4E2D8FF967055AF6DF97BFD7DF91B" box="[579,693,1699,1726]" captionStart="FIGURE 16" captionStartId="27.[151,250,1428,1452]" captionTargetBox="[184,1412,353,1404]" captionTargetId="figure-95@27.[176,1412,345,1404]" captionTargetPageId="27" captionText="FIGURE 16. Distribution map of the Tityus forcipula species group. A. South America. B. Records area. C. Species records." figureDoi="http://doi.org/10.5281/zenodo.6673378" httpUri="https://zenodo.org/record/6673378/files/figure.png" pageId="18" pageNumber="169">Figure 16</figureCitation>
). The exact type locality of
<taxonomicName id="FC8F85DEFF967055AEDCF97BFBBCF918" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[1010,1140,1699,1725]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="forcipula">
<emphasis id="09FB224FFF967055AEDCF97BFBBCF918" box="[1010,1140,1699,1725]" italics="true" pageId="18" pageNumber="169">T. forcipula</emphasis>
</taxonomicName>
is not precise, but there is consensus that the male
<typeStatus id="E43440FFFF967055AC86F91FFDC2F944" box="[424,522,1735,1761]" pageId="18" pageNumber="169" type="holotype">holotype</typeStatus>
of this species came from
<collectingCountry id="4398BECDFF967055AE18F91FFC6CF944" box="[822,932,1735,1761]" name="Colombia" pageId="18" pageNumber="169">Colombia</collectingCountry>
(
<bibRefCitation id="5F1E83ACFF967055AE9DF91FFBABF944" author="Lourenco, W. R." box="[947,1123,1735,1761]" pageId="18" pageNumber="169" pagination="729 - 739" refId="ref18482" refString="Lourenco, W. R. (1984) Etude systematique de quelques especes appartenant au complexe Tityus forcipula (Gervais, 1844) (Scorpiones, Buthidae). Bulletin du Museum national d'histoire naturelle. Section A, Zoologie, biologie et ecologie animales, 6, 729 - 739." type="journal article" year="1984">Lourenço 1984</bibRefCitation>
). Our redescription is based on specimens from the Cordillera Occidental of Cali,
<collectingRegion id="F94B30BFFF967055AFC1F933FC6AF8A0" box="[751,930,1771,1797]" country="Colombia" name="Valle del Cauca" pageId="18" pageNumber="169">Valle del Cauca</collectingRegion>
,
<collectingCountry id="4398BECDFF967055AE83F933FBD6F8A0" box="[941,1054,1771,1797]" name="Colombia" pageId="18" pageNumber="169">Colombia</collectingCountry>
, where
<bibRefCitation id="5F1E83ACFF967055A95AF933FA54F8A0" author="Lourenco, W. R. &amp; Florez, D. E." box="[1140,1436,1771,1797]" pageId="18" pageNumber="169" pagination="119 - 133" refId="ref19143" refString="Lourenco, W. R. &amp; Florez, D. E. (1990) Scorpions (Chelicerata) from Colombia III. The scorpio-fauna of pacific region (Choco), with some biogeographic considerations. Amazoniana, 11, 119 - 133." type="journal article" year="1990">Lourenço &amp; Flórez (1990)</bibRefCitation>
and
<bibRefCitation id="5F1E83ACFF967055ADE9F8D7FE4BF88C" author="Lourenco, W. R." box="[199,387,1807,1833]" pageId="18" pageNumber="169" pagination="61 - 94" refId="ref18632" refString="Lourenco, W. R. (1997) Synopsis de la faune de scorpions de Colombie, avec des considerations sur le systematique et la biogeographie des especes. Revue Suisse de Zoologie, 104, 61 - 94. https: // doi. org / 10.5962 / bhl. part. 79991" type="journal article" year="1997">Lourenço (1997)</bibRefCitation>
identified samples of this species.
</paragraph>
<paragraph id="3B30FE5DFF967055ADE9F8EAFAE3F878" blockId="18.[151,1437,151,2014]" pageId="18" pageNumber="169">
<emphasis id="09FB224FFF967055ADE9F8EAFD95F8E9" bold="true" box="[199,605,1842,1869]" pageId="18" pageNumber="169">
Confirmed records.
<collectingCountry id="4398BECDFF967055AC90F8EAFD95F8E9" box="[446,605,1842,1868]" name="Colombia" pageId="18" pageNumber="169">COLOMBIA</collectingCountry>
</emphasis>
:
<emphasis id="09FB224FFF967055AF40F8EBFC7CF8E9" box="[622,948,1843,1869]" italics="true" pageId="18" pageNumber="169">
<collectingRegion id="F94B30BFFF967055AF40F8EBFC7CF8E9" box="[622,948,1843,1869]" country="Colombia" name="Valle del Cauca" pageId="18" pageNumber="169">Valle del Cauca Department</collectingRegion>
</emphasis>
:
<collectingCounty id="D25186D1FF967055AEEBF8EBFC3CF8E8" box="[965,1012,1843,1869]" pageId="18" pageNumber="169">Cali</collectingCounty>
(Pichinde-
<collectingMunicipality id="DB546427FF967055A955F8EBFAEBF8EB" box="[1147,1315,1843,1870]" pageId="18" pageNumber="169">Peñas Blancas</collectingMunicipality>
),
<location id="3E50A886FF967055A817F8EBFA55F8E8" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF967055A817F8EBFA55F8E8" box="[1337,1437,1843,1869]" country="Colombia" county="Cali" municipality="Penas Blancas" name="Restrepo" pageId="18" pageNumber="169" stateProvince="Valle del Cauca">Restrepo</location>
(road
<location id="3E50A886FF967055ADF7F88FFE76F8D4" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF967055ADF7F88FFE76F8D4" box="[217,446,1879,1905]" country="Colombia" county="Cali" municipality="Penas Blancas" name="Buga-Buenaventura" pageId="18" pageNumber="169" stateProvince="Valle del Cauca">Buga-Buenaventura</location>
),
<location id="3E50A886FF967055ACE0F88FFDD6F8D4" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF967055ACE0F88FFDD6F8D4" box="[462,542,1879,1905]" country="Colombia" county="Cali" municipality="Penas Blancas" name="Yotoco" pageId="18" pageNumber="169" stateProvince="Valle del Cauca">Yotoco</location>
(
<location id="3E50A886FF967055AF02F88FFC41F8D4" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF967055AF02F88FFC41F8D4" box="[556,905,1879,1905]" country="Colombia" county="Cali" municipality="Penas Blancas" name="Yotoco Forest Natural Reserve" pageId="18" pageNumber="169" stateProvince="Valle del Cauca">Yotoco Forest Natural Reserve</location>
), and
<location id="3E50A886FF967055AEE7F88FFBD2F8D4" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF967055AEE7F88FFBD2F8D4" box="[969,1050,1879,1905]" country="Colombia" county="Cali" municipality="Penas Blancas" name="Yumbo" pageId="18" pageNumber="169" stateProvince="Valle del Cauca">Yumbo</location>
(
<location id="3E50A886FF967055A906F88FFBA1F8D4" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF967055A906F88FFBA1F8D4" box="[1064,1129,1879,1905]" country="Colombia" county="Cali" municipality="Penas Blancas" name="Dapa" pageId="18" pageNumber="169" stateProvince="Valle del Cauca">Dapa</location>
).
<emphasis id="09FB224FFF967055A955F88FFF01F831" bold="true" pageId="18" pageNumber="169">New Records. COLOMBIA</emphasis>
:
<emphasis id="09FB224FFF967055ADF9F8A3FE03F830" box="[215,459,1915,1941]" italics="true" pageId="18" pageNumber="169">
<collectingRegion id="F94B30BFFF967055ADF9F8A3FE03F830" box="[215,459,1915,1941]" country="Colombia" name="Risaralda" pageId="18" pageNumber="169">Risaralda department</collectingRegion>
</emphasis>
: Santuario (
<location id="3E50A886FF967055AF7CF8A3FCDEF830" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF967055AF7CF8A3FCDEF830" box="[594,790,1915,1941]" country="Colombia" name="San Rafael Plains" pageId="18" pageNumber="169" stateProvince="Risaralda">San Rafael Plains</location>
Natural Regional Park).
<emphasis id="09FB224FFF967055A905F8A3FAA8F830" box="[1067,1376,1915,1941]" italics="true" pageId="18" pageNumber="169">
<collectingRegion id="F94B30BFFF967055A905F8A3FAA8F830" box="[1067,1376,1915,1941]" country="Colombia" name="Valle del Cauca" pageId="18" pageNumber="169">Valle del Cauca department</collectingRegion>
</emphasis>
: Cali (
<collectorName id="967A9B8BFF967055ADB1F847FE96F81C" box="[159,350,1951,1977]" pageId="18" pageNumber="169">El Pato Pathway</collectorName>
,
<collectorName id="967A9B8BFF967055AC44F847FD98F81C" box="[362,592,1951,1977]" pageId="18" pageNumber="169">El Saladito Pathway</collectorName>
(
<location id="3E50A886FF967055AF4FF847FC8AF81C" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF967055AF4FF847FC8AF81C" box="[609,834,1951,1977]" country="Colombia" name="San Antonio Forest" pageId="18" pageNumber="169" stateProvince="Valle del Cauca">San Antonio Forest</location>
), Felidia Pathway, Villacarmelo Pathway and 18 Km Highway to the Sea),
<collectorName id="967A9B8BFF967055ACA5F81BFDC1F878" box="[395,521,1987,2013]" pageId="18" pageNumber="169">La Cumbre</collectorName>
(Bitaco and Chicoral Pathways), Palmira (Panorama Farm), and Tuluá.
</paragraph>
<caption id="6FF0AED5FF977054ADB9F84FFD83F85D" ID-DOI="http://doi.org/10.5281/zenodo.6673364" ID-Zenodo-Dep="6673364" httpUri="https://zenodo.org/record/6673364/files/figure.png" pageId="19" pageNumber="170" startId="19.[151,250,1943,1967]" targetBox="[221,1367,181,1920]" targetPageId="19">
<paragraph id="3B30FE5DFF977054ADB9F84FFD83F85D" blockId="19.[151,1436,1943,2040]" pageId="19" pageNumber="170">
<emphasis id="09FB224FFF977054ADB9F84FFEEDF80A" bold="true" box="[151,293,1943,1967]" pageId="19" pageNumber="170">FIGURE 10.</emphasis>
<taxonomicName id="FC8F85DEFF977054AC1FF840FDA5F815" authority="Gervais, 1843" authorityName="Gervais" authorityYear="1843" box="[305,621,1944,1968]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="170" phylum="Arthropoda" rank="subGenus" subGenus="Atreus">
<emphasis id="09FB224FFF977054AC1FF840FEA6F80A" box="[305,366,1944,1967]" italics="true" pageId="19" pageNumber="170">Tityus</emphasis>
(
<emphasis id="09FB224FFF977054ACACF840FE0BF80A" box="[386,451,1944,1967]" italics="true" pageId="19" pageNumber="170">Atreus</emphasis>
)
<bibRefCitation id="5F1E83ACFF977054ACF9F840FDA5F815" author="Gervais, P. M." box="[471,621,1944,1968]" pageId="19" pageNumber="170" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
</taxonomicName>
, sternites IV
<emphasis id="09FB224FFF977054AFD6F840FCCDF815" bold="true" box="[760,773,1944,1968]" pageId="19" pageNumber="170">–</emphasis>
VII.
<emphasis id="09FB224FFF977054AE17F84FFCBDF815" bold="true" box="[825,885,1943,1968]" pageId="19" pageNumber="170">A, B.</emphasis>
<taxonomicName id="FC8F85DEFF977054AEAFF840FAFEF815" authority="(Gervais, 1843)" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[897,1334,1944,1968]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="forcipula" subGenus="Atreus">
<emphasis id="09FB224FFF977054AEAFF840FC76F80A" box="[897,958,1944,1967]" italics="true" pageId="19" pageNumber="170">Tityus</emphasis>
(
<emphasis id="09FB224FFF977054AEFCF840FBDBF80A" box="[978,1043,1944,1967]" italics="true" pageId="19" pageNumber="170">Atreus</emphasis>
)
<emphasis id="09FB224FFF977054A909F840FB4BF815" box="[1063,1155,1944,1968]" italics="true" pageId="19" pageNumber="170">forcipula</emphasis>
(
<bibRefCitation id="5F1E83ACFF977054A9B8F840FAE6F815" author="Gervais, P. M." box="[1174,1326,1944,1968]" pageId="19" pageNumber="170" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="09FB224FFF977054A86AF84FFA95F80A" bold="true" box="[1348,1373,1943,1967]" pageId="19" pageNumber="170">A.</emphasis>
Male (MUSENUV-Ar 438).
<emphasis id="09FB224FFF977054AC55F864FE5AF871" bold="true" box="[379,402,1980,2004]" pageId="19" pageNumber="170">B.</emphasis>
Female (MUSENUV-Ar 436).
<emphasis id="09FB224FFF977054AFE6F863FD36F871" bold="true" box="[712,766,1979,2004]" pageId="19" pageNumber="170">C, D.</emphasis>
<taxonomicName id="FC8F85DEFF977054AE2CF864FBC0F871" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[770,1032,1980,2004]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov." subGenus="Atreus">
<emphasis id="09FB224FFF977054AE2CF864FCF7F876" box="[770,831,1980,2003]" italics="true" pageId="19" pageNumber="170">Tityus</emphasis>
(
<emphasis id="09FB224FFF977054AE65F864FC44F876" box="[843,908,1980,2003]" italics="true" pageId="19" pageNumber="170">Atreus</emphasis>
)
<emphasis id="09FB224FFF977054AEB7F864FBC0F871" box="[921,1032,1980,2004]" italics="true" pageId="19" pageNumber="170">moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF977054A922F864FBBFF876" bold="true" box="[1036,1143,1979,2004]" pageId="19" pageNumber="170">
<taxonomicNameLabel id="12C89F34FF977054A922F864FB92F871" box="[1036,1114,1980,2004]" pageId="19" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
C.
</emphasis>
Male holotype (MUSENUVAr 2104).
<emphasis id="09FB224FFF977054AC2EF838FED1F85D" bold="true" box="[256,281,2016,2040]" pageId="19" pageNumber="170">D.</emphasis>
Female paratype (ICN-1402).
</paragraph>
</caption>
<caption id="6FF0AED5FF907053ADB9F886FBAAF847" ID-DOI="http://doi.org/10.5281/zenodo.6673366" ID-Zenodo-Dep="6673366" httpUri="https://zenodo.org/record/6673366/files/figure.png" pageId="20" pageNumber="171" startId="20.[151,250,1886,1910]" targetBox="[224,1363,181,1862]" targetPageId="20">
<paragraph id="3B30FE5DFF907053ADB9F886FBAAF847" blockId="20.[151,1437,1886,2018]" pageId="20" pageNumber="171">
<emphasis id="09FB224FFF907053ADB9F886FED6F8D3" bold="true" box="[151,286,1886,1910]" pageId="20" pageNumber="171">FIGURE 11.</emphasis>
<taxonomicName id="FC8F85DEFF907053AC0AF887FD98F8D3" authority="Gervais, 1843" authorityName="Gervais" authorityYear="1843" box="[292,592,1886,1910]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="171" phylum="Arthropoda" rank="subGenus" subGenus="Atreus">
<emphasis id="09FB224FFF907053AC0AF887FEA9F8D3" box="[292,353,1887,1910]" italics="true" pageId="20" pageNumber="171">Tityus</emphasis>
(
<emphasis id="09FB224FFF907053AC5EF887FE79F8D3" box="[368,433,1887,1910]" italics="true" pageId="20" pageNumber="171">Atreus</emphasis>
)
<bibRefCitation id="5F1E83ACFF907053ACEEF886FD98F8D3" author="Gervais, P. M." box="[448,592,1886,1910]" pageId="20" pageNumber="171" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
</taxonomicName>
, hemispermatophore.
<emphasis id="09FB224FFF907053AE1EF886FCA0F8D3" bold="true" box="[816,872,1886,1910]" pageId="20" pageNumber="171">A–C.</emphasis>
<taxonomicName id="FC8F85DEFF907053AE40F887FAC2F8D3" authority="(Gervais, 1843)" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[878,1290,1886,1910]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="forcipula" subGenus="Atreus">
<emphasis id="09FB224FFF907053AE40F887FC63F8D3" box="[878,939,1887,1910]" italics="true" pageId="20" pageNumber="171">Tityus</emphasis>
(
<emphasis id="09FB224FFF907053AE97F887FC32F8D3" box="[953,1018,1887,1910]" italics="true" pageId="20" pageNumber="171">Atreus</emphasis>
)
<emphasis id="09FB224FFF907053A927F886FBADF8D3" box="[1033,1125,1886,1910]" italics="true" pageId="20" pageNumber="171">forcipula</emphasis>
(
<bibRefCitation id="5F1E83ACFF907053A95CF886FACBF8D3" author="Gervais, P. M." box="[1138,1283,1886,1910]" pageId="20" pageNumber="171" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
)
</taxonomicName>
(MUSENUVAr 438).
<emphasis id="09FB224FFF907053ADDDF85AFEC4F83F" bold="true" box="[243,268,1922,1946]" pageId="20" pageNumber="171">A.</emphasis>
Posterior view.
<emphasis id="09FB224FFF907053AC9CF85AFE01F83F" bold="true" box="[434,457,1922,1946]" pageId="20" pageNumber="171">B.</emphasis>
Lateral view.
<emphasis id="09FB224FFF907053AF72F85AFDBCF83F" bold="true" box="[604,628,1922,1946]" pageId="20" pageNumber="171">C.</emphasis>
Ventral view.
<emphasis id="09FB224FFF907053AE27F85AFCF3F83F" bold="true" box="[777,827,1922,1946]" pageId="20" pageNumber="171">D–F.</emphasis>
<taxonomicName id="FC8F85DEFF907053AE6CF85BFB86F83F" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[834,1102,1922,1946]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="171" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov." subGenus="Atreus">
<emphasis id="09FB224FFF907053AE6CF85BFCB7F83F" box="[834,895,1923,1946]" italics="true" pageId="20" pageNumber="171">Tityus</emphasis>
(
<emphasis id="09FB224FFF907053AEA0F85BFC07F83F" box="[910,975,1923,1946]" italics="true" pageId="20" pageNumber="171">Atreus</emphasis>
)
<emphasis id="09FB224FFF907053AEF1F85AFB86F83F" box="[991,1102,1922,1946]" italics="true" pageId="20" pageNumber="171">moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF907053A97BF85AFB6EF83F" bold="true" box="[1109,1190,1922,1946]" pageId="20" pageNumber="171">
<taxonomicNameLabel id="12C89F34FF907053A97BF85AFB6EF83F" box="[1109,1190,1922,1946]" pageId="20" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MUSENUV-Ar 2104).
<emphasis id="09FB224FFF907053ADB9F87EFF78F81B" bold="true" box="[151,176,1958,1982]" pageId="20" pageNumber="171">D.</emphasis>
Posterior view.
<emphasis id="09FB224FFF907053AC7DF87EFEA2F81B" bold="true" box="[339,362,1958,1982]" pageId="20" pageNumber="171">E.</emphasis>
Lateral view.
<emphasis id="09FB224FFF907053ACD4F87EFDC5F81B" bold="true" box="[506,525,1958,1982]" pageId="20" pageNumber="171">F.</emphasis>
Ventral view.
<emphasis id="09FB224FFF907053AFB0F87EFC8DF81B" bold="true" box="[670,837,1958,1982]" pageId="20" pageNumber="171">Abbreviations=</emphasis>
<emphasis id="09FB224FFF907053AE65F87FFC92F81B" box="[843,858,1959,1982]" italics="true" pageId="20" pageNumber="171">B</emphasis>
, body;
<emphasis id="09FB224FFF907053AE89F87EFC76F81B" box="[935,958,1958,1982]" italics="true" pageId="20" pageNumber="171">Bl</emphasis>
, basal lobe;
<emphasis id="09FB224FFF907053A912F87EFB9BF81B" box="[1084,1107,1958,1982]" italics="true" pageId="20" pageNumber="171">El</emphasis>
, external lobe;
<emphasis id="09FB224FFF907053A9DEF87FFB37F81B" box="[1264,1279,1959,1982]" italics="true" pageId="20" pageNumber="171">F</emphasis>
, flagellum;
<emphasis id="09FB224FFF907053A854F87FFA5EF81B" box="[1402,1430,1959,1982]" italics="true" pageId="20" pageNumber="171">Fo</emphasis>
, foot;
<emphasis id="09FB224FFF907053ADE3F813FF2AF847" box="[205,226,1995,2018]" italics="true" pageId="20" pageNumber="171">iL</emphasis>
, internal lobe;
<emphasis id="09FB224FFF907053AC52F812FE6FF847" box="[380,423,1994,2018]" italics="true" pageId="20" pageNumber="171">Pref</emphasis>
, pars reflexa;
<emphasis id="09FB224FFF907053AF19F813FDAEF847" box="[567,614,1995,2018]" italics="true" pageId="20" pageNumber="171">Prec</emphasis>
, pars recta. Scale bars= 2 mm. Scale bars= 2 mm.
</paragraph>
</caption>
<caption id="6FF0AED5FF917052ADB9F90CFCC7F891" ID-DOI="http://doi.org/10.5281/zenodo.6673368" ID-Zenodo-Dep="6673368" httpUri="https://zenodo.org/record/6673368/files/figure.png" pageId="21" pageNumber="172" startId="21.[151,250,1748,1772]" targetBox="[151,1435,181,1724]" targetPageId="21">
<paragraph id="3B30FE5DFF917052ADB9F90CFCC7F891" blockId="21.[151,1436,1748,1845]" pageId="21" pageNumber="172">
<emphasis id="09FB224FFF917052ADB9F90CFEE9F949" bold="true" box="[151,289,1748,1772]" pageId="21" pageNumber="172">FIGURE 12.</emphasis>
<taxonomicName id="FC8F85DEFF917052AC06F90DFD90F949" authority="Gervais, 1843" authorityName="Gervais" authorityYear="1843" box="[296,600,1748,1772]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="172" phylum="Arthropoda" rank="subGenus" subGenus="Atreus">
<emphasis id="09FB224FFF917052AC06F90DFEADF949" box="[296,357,1749,1772]" italics="true" pageId="21" pageNumber="172">Tityus</emphasis>
(
<emphasis id="09FB224FFF917052AC5BF90DFE7EF949" box="[373,438,1749,1772]" italics="true" pageId="21" pageNumber="172">Atreus</emphasis>
)
<bibRefCitation id="5F1E83ACFF917052ACE8F90CFD90F949" author="Gervais, P. M." box="[454,600,1748,1772]" pageId="21" pageNumber="172" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
</taxonomicName>
, metasoma, dorsal view.
<emphasis id="09FB224FFF917052AE75F90CFC5AF949" bold="true" box="[859,914,1748,1772]" pageId="21" pageNumber="172">A, B.</emphasis>
<taxonomicName id="FC8F85DEFF917052AEB4F90DFAF7F949" authority="(Gervais, 1843)" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[922,1343,1748,1772]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="172" phylum="Arthropoda" rank="species" species="forcipula" subGenus="Atreus">
<emphasis id="09FB224FFF917052AEB4F90DFC1FF949" box="[922,983,1749,1772]" italics="true" pageId="21" pageNumber="172">Tityus</emphasis>
(
<emphasis id="09FB224FFF917052AEC9F90DFBE0F949" box="[999,1064,1749,1772]" italics="true" pageId="21" pageNumber="172">Atreus</emphasis>
)
<emphasis id="09FB224FFF917052A916F90CFB5CF949" box="[1080,1172,1748,1772]" italics="true" pageId="21" pageNumber="172">forcipula</emphasis>
(
<bibRefCitation id="5F1E83ACFF917052A98CF90CFAFEF949" author="Gervais, P. M." box="[1186,1334,1748,1772]" pageId="21" pageNumber="172" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="09FB224FFF917052A866F90CFAA9F949" bold="true" box="[1352,1377,1748,1772]" pageId="21" pageNumber="172">A.</emphasis>
Male (MUSENUV-Ar 438).
<emphasis id="09FB224FFF917052AC55F920FE5AF8B5" bold="true" box="[379,402,1784,1808]" pageId="21" pageNumber="172">B.</emphasis>
Female (MUSENUV-Ar 436).
<emphasis id="09FB224FFF917052AFE6F920FD36F8B5" bold="true" box="[712,766,1784,1808]" pageId="21" pageNumber="172">C, D.</emphasis>
<taxonomicName id="FC8F85DEFF917052AE2CF921FBC0F8B5" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[770,1032,1784,1808]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="172" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov." subGenus="Atreus">
<emphasis id="09FB224FFF917052AE2CF921FCF7F8B5" box="[770,831,1785,1808]" italics="true" pageId="21" pageNumber="172">Tityus</emphasis>
(
<emphasis id="09FB224FFF917052AE65F921FC44F8B5" box="[843,908,1785,1808]" italics="true" pageId="21" pageNumber="172">Atreus</emphasis>
)
<emphasis id="09FB224FFF917052AEB7F920FBC0F8B5" box="[921,1032,1784,1808]" italics="true" pageId="21" pageNumber="172">moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF917052A922F920FBBFF8B5" bold="true" box="[1036,1143,1784,1808]" pageId="21" pageNumber="172">
<taxonomicNameLabel id="12C89F34FF917052A922F920FB92F8B5" box="[1036,1114,1784,1808]" pageId="21" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
C.
</emphasis>
Male holotype (MUSENUVAr 2104).
<emphasis id="09FB224FFF917052AC2EF8C4FED1F891" bold="true" box="[256,281,1820,1844]" pageId="21" pageNumber="172">D.</emphasis>
Female paratype (ICN-1402). Scale bars= 5 mm.
</paragraph>
</caption>
<caption id="6FF0AED5FF927051ADB9F947FC53F95B" ID-DOI="http://doi.org/10.5281/zenodo.6673372" ID-Zenodo-Dep="6673372" httpUri="https://zenodo.org/record/6673372/files/figure.png" pageId="22" pageNumber="173" startId="22.[151,250,1695,1719]" targetBox="[151,1436,181,1671]" targetPageId="22">
<paragraph id="3B30FE5DFF927051ADB9F947FC53F95B" blockId="22.[151,1437,1695,1791]" pageId="22" pageNumber="173">
<emphasis id="09FB224FFF927051ADB9F947FEECF912" bold="true" box="[151,292,1695,1719]" pageId="22" pageNumber="173">FIGURE 13.</emphasis>
<taxonomicName id="FC8F85DEFF927051AC01F978FDA0F912" authority="Gervais, 1843" authorityName="Gervais" authorityYear="1843" box="[303,616,1695,1719]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="173" phylum="Arthropoda" rank="subGenus" subGenus="Atreus">
<emphasis id="09FB224FFF927051AC01F978FEA4F912" box="[303,364,1696,1719]" italics="true" pageId="22" pageNumber="173">Tityus</emphasis>
(
<emphasis id="09FB224FFF927051AC51F947FE08F913" box="[383,448,1695,1718]" italics="true" pageId="22" pageNumber="173">Atreus</emphasis>
)
<bibRefCitation id="5F1E83ACFF927051ACFDF947FDA0F912" author="Gervais, P. M." box="[467,616,1695,1719]" pageId="22" pageNumber="173" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
</taxonomicName>
, metasoma, ventral view.
<emphasis id="09FB224FFF927051AEAEF947FC73F912" bold="true" box="[896,955,1695,1719]" pageId="22" pageNumber="173">A, B.</emphasis>
<taxonomicName id="FC8F85DEFF927051AEE8F978FABFF912" authority="(Gervais, 1843)" baseAuthorityName="Gervais" baseAuthorityYear="1843" box="[966,1399,1695,1719]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="forcipula" subGenus="Atreus">
<emphasis id="09FB224FFF927051AEE8F978FBCBF912" box="[966,1027,1696,1719]" italics="true" pageId="22" pageNumber="173">Tityus</emphasis>
(
<emphasis id="09FB224FFF927051A93BF947FB9EF913" box="[1045,1110,1695,1718]" italics="true" pageId="22" pageNumber="173">Atreus</emphasis>
)
<emphasis id="09FB224FFF927051A944F947FB0EF912" box="[1130,1222,1695,1719]" italics="true" pageId="22" pageNumber="173">forcipula</emphasis>
(
<bibRefCitation id="5F1E83ACFF927051A9F6F947FAA7F912" author="Gervais, P. M." box="[1240,1391,1695,1719]" pageId="22" pageNumber="173" pagination="129 - 131" refId="ref17413" refString="Gervais, P. M. (1843) Les principaux resultats d'un travail sur la famille des Scorpions. Societe Philomatique de Paris 5, 129 - 131." type="journal article" year="1843">Gervais, 1843</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="09FB224FFF927051A8AAF947FA54F912" bold="true" box="[1412,1436,1695,1719]" pageId="22" pageNumber="173">A.</emphasis>
Male (MUSENUV-Ar 438).
<emphasis id="09FB224FFF927051ACEDF91BFE12F97E" bold="true" box="[451,474,1731,1755]" pageId="22" pageNumber="173">B.</emphasis>
Female (MUSENUV-Ar 436).
<emphasis id="09FB224FFF927051AE09F91BFCAAF97E" bold="true" box="[807,866,1731,1755]" pageId="22" pageNumber="173">C, D.</emphasis>
<taxonomicName id="FC8F85DEFF927051AE45F91CFBB4F97E" authority="Pinto-Da-Rocha &amp; Cabra-García, 2022" authorityName="Pinto-Da-Rocha &amp; Cabra-García" authorityYear="2022" box="[875,1148,1731,1755]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="moralensis" status="sp. nov." subGenus="Atreus">
<emphasis id="09FB224FFF927051AE45F91CFC60F97E" box="[875,936,1732,1755]" italics="true" pageId="22" pageNumber="173">Tityus</emphasis>
(
<emphasis id="09FB224FFF927051AE94F91BFC33F97F" box="[954,1019,1731,1754]" italics="true" pageId="22" pageNumber="173">Atreus</emphasis>
)
<emphasis id="09FB224FFF927051A923F91BFBB4F97E" box="[1037,1148,1731,1755]" italics="true" pageId="22" pageNumber="173">moralensis</emphasis>
</taxonomicName>
<emphasis id="09FB224FFF927051A9A8F91BFB34F97E" bold="true" box="[1158,1276,1731,1755]" pageId="22" pageNumber="173">
<taxonomicNameLabel id="12C89F34FF927051A9A8F91BFB12F97E" box="[1158,1242,1731,1755]" pageId="22" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
C.
</emphasis>
Male holotype (MUSENUV-Ar 2104).
<emphasis id="09FB224FFF927051ACA2F93FFE6DF95A" bold="true" box="[396,421,1767,1791]" pageId="22" pageNumber="173">D.</emphasis>
Female paratype (ICN-1402). Scale bars= 5 mm.
</paragraph>
</caption>
<paragraph id="3B30FE5DFF927051ADE9F8F5FC47F816" blockId="22.[151,1436,1837,1972]" pageId="22" pageNumber="173">
<emphasis id="09FB224FFF927051ADE9F8F5FE55F8E2" bold="true" box="[199,413,1837,1863]" pageId="22" pageNumber="173">Doubtful records.</emphasis>
<materialsCitation id="8BE7F400FF927051AC88F8F5FC76F8E2" ID-GBIF-Occurrence="3816447320" box="[422,958,1837,1864]" country="Colombia" location="Caldas Department" pageId="22" pageNumber="173" specimenCount="1" stateProvince="Caldas">
<emphasis id="09FB224FFF927051AC88F8F5FD86F8E2" bold="true" box="[422,590,1837,1863]" pageId="22" pageNumber="173">
<collectingCountry id="4398BECDFF927051AC88F8F5FD81F8E2" box="[422,585,1837,1863]" name="Colombia" pageId="22" pageNumber="173">COLOMBIA</collectingCountry>
:
</emphasis>
<emphasis id="09FB224FFF927051AF79F8F5FCFEF8E2" box="[599,822,1837,1863]" italics="true" pageId="22" pageNumber="173">
<collectingRegion id="F94B30BFFF927051AF79F8F5FCFEF8E2" box="[599,822,1837,1863]" country="Colombia" name="Caldas" pageId="22" pageNumber="173">Caldas Department</collectingRegion>
</emphasis>
: Manizales
</materialsCitation>
.
<materialsCitation id="8BE7F400FF927051AEE4F8F5FF37F8C9" ID-GBIF-Occurrence="3816447301" collectorName="La Vega &amp; La Reserva" country="Colombia" location="Cundinamarca Department" pageId="22" pageNumber="173" specimenCount="1" stateProvince="Cundinamarca">
<collectingRegion id="F94B30BFFF927051AEE4F8F5FB35F8E2" box="[970,1277,1837,1863]" country="Colombia" name="Cundinamarca" pageId="22" pageNumber="173">
Cundinamarca
<emphasis id="09FB224FFF927051A959F8F6FB35F8E2" box="[1143,1277,1838,1863]" italics="true" pageId="22" pageNumber="173">Department</emphasis>
</collectingRegion>
:
<collectorName id="967A9B8BFF927051A820F8F6FAA4F8ED" box="[1294,1388,1838,1864]" pageId="22" pageNumber="173">La Vega</collectorName>
(
<collectorName id="967A9B8BFF927051A853F8F5FF3EF8C9" pageId="22" pageNumber="173">La Reserva</collectorName>
)
</materialsCitation>
.
<emphasis id="09FB224FFF927051AC25F889FE3FF8CE" box="[267,503,1873,1899]" italics="true" pageId="22" pageNumber="173">
<collectingRegion id="F94B30BFFF927051AC25F889FE3FF8CE" box="[267,503,1873,1899]" country="Colombia" name="Quindio" pageId="22" pageNumber="173">Quindío Department</collectingRegion>
</emphasis>
: Armenia and Salento.
<emphasis id="09FB224FFF927051AE27F889FC2DF8CE" box="[777,997,1873,1899]" italics="true" pageId="22" pageNumber="173">
<collectingRegion id="F94B30BFFF927051AE27F889FC2DF8CE" box="[777,997,1873,1899]" country="Colombia" name="Tolima" pageId="22" pageNumber="173">Tolima Department</collectingRegion>
</emphasis>
: Ibagué (Cay Pathway).
<materialsCitation id="8BE7F400FF927051A829F889FE78F82A" ID-GBIF-Occurrence="3816447309" country="Ecuador" location="Azuay province" pageId="22" pageNumber="173" specimenCount="1" stateProvince="Azuay">
<emphasis id="09FB224FFF927051A829F889FA5DF8CE" bold="true" box="[1287,1429,1873,1899]" pageId="22" pageNumber="173">
<collectingCountry id="4398BECDFF927051A829F889FA5DF8CE" box="[1287,1429,1873,1899]" name="Ecuador" pageId="22" pageNumber="173">ECUADOR</collectingCountry>
</emphasis>
:
<emphasis id="09FB224FFF927051ADB9F8ADFE8EF82A" box="[151,326,1909,1935]" italics="true" pageId="22" pageNumber="173">
<collectingRegion id="F94B30BFFF927051ADB9F8ADFE8EF82A" box="[151,326,1909,1935]" country="Ecuador" name="Azuay" pageId="22" pageNumber="173">Azuay province</collectingRegion>
</emphasis>
: Cuenca
</materialsCitation>
.
<emphasis id="09FB224FFF927051AC92F8ADFD59F82A" box="[444,657,1909,1935]" italics="true" pageId="22" pageNumber="173">
<collectingRegion id="F94B30BFFF927051AC92F8ADFD59F82A" box="[444,657,1909,1935]" country="Ecuador" name="Cotopaxi" pageId="22" pageNumber="173">Cotopaxi Province</collectingRegion>
</emphasis>
: Las Pampas.
<emphasis id="09FB224FFF927051AE12F8ADFBD3F82A" box="[828,1051,1909,1935]" italics="true" pageId="22" pageNumber="173">
<collectingRegion id="F94B30BFFF927051AE12F8ADFBD3F82A" box="[828,1051,1909,1935]" country="Ecuador" name="Pichincha" pageId="22" pageNumber="173">Pichincha Province</collectingRegion>
</emphasis>
: Alluriquin and Chiriboga.
<emphasis id="09FB224FFF927051A872F8ADFEE8F816" italics="true" pageId="22" pageNumber="173">Santo Domingo de</emphasis>
<materialsCitation id="8BE7F400FF927051AC06F841FC43F816" ID-GBIF-Occurrence="3816447308" box="[296,907,1945,1972]" country="Ecuador" location="Santo Domingo de Los Colorados" pageId="22" pageNumber="173" specimenCount="1" stateProvince="Tsachilas Province">
<emphasis id="09FB224FFF927051AC06F841FE34F816" box="[296,508,1945,1971]" italics="true" pageId="22" pageNumber="173">
<collectingRegion id="F94B30BFFF927051AC06F841FE34F816" box="[296,508,1945,1971]" inferredBySuffix="true" pageId="22" pageNumber="173">Tsáchilas Province</collectingRegion>
</emphasis>
:
<location id="3E50A886FF927051AF24F841FC43F816" LSID="urn:lsid:plazi:treatment:B3264F4BFF8D7051ADB9FB73FC47F816:3E50A886FF927051AF24F841FC43F816" box="[522,907,1945,1972]" country="Ecuador" name="Santo Domingo de Los Colorados" pageId="22" pageNumber="173" stateProvince="Tsachilas Province">Santo Domingo de Los Colorados</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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