548 lines
55 KiB
XML
548 lines
55 KiB
XML
<document id="87D0CB8650B5712FB922BA0152EE2186" ID-CLB-Dataset="305027" ID-DOI="10.3897/zse.100.132111" ID-GBIF-Dataset="6717c3bc-c84e-4604-ab36-042a63c45eac" ID-ZooBank="B39508EB-6EB2-4CC0-88D8-B3E87BDCED02" ID-publisher-id="132111" URI-arpha="6D90141D-221E-5A1C-82D3-04B8878C6781" URI-zoobank="http://zoobank.org/B39508EB-6EB2-4CC0-88D8-B3E87BDCED02" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1731000560999" checkinUser="pensoft" docAuthor="Nguyen, Anh D., Nguyen, Lien T. P. & Korsós, Zoltán" docDate="2024" docId="92A4751707F158C499FD3B94C2D1B998" docLanguage="en" docName="Zoosystematics and Evolution 100(4): 1515-1541" docOrigin="Zoosystematics and Evolution 100 (4)" docSource="https://zse.pensoft.net/article/132111/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Pseudodesmus irregularis Nguyen, Nguyen & Korsós, 2024, sp. nov." docType="treatment" docVersion="4" dtd-version="3.0" lastPageNumber="1541" masterDocId="6D90141D221E5A1C82D304B8878C6781" masterDocTitle="A review of the millipede genus Pseudodesmus Pocock, 1887 (Diplopoda, Platydesmida, Andrognathidae) from Vietnam, with descriptions of five new species and notes on its phylogeny" masterLastPageNumber="1541" masterPageNumber="1515" pageNumber="1515" updateTime="1731004247642" updateUser="ExternalLinkService">
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<mods:titleInfo id="2E42E3C396AF46555BAE53C3AD915DEE">
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<mods:title id="3A752025103862C895FF2F44C39C7ECB">A review of the millipede genus Pseudodesmus Pocock, 1887 (Diplopoda, Platydesmida, Andrognathidae) from Vietnam, with descriptions of five new species and notes on its phylogeny</mods:title>
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<mods:name id="1F6B599555DB8C01D72C0FDD8D76988A" type="personal">
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<mods:roleTerm id="9381B317E031465E49CE1BF240891EB3">Author</mods:roleTerm>
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<mods:namePart id="8FA2977C66EF796A36B4538AC6A4A533">Nguyen, Anh D.</mods:namePart>
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<mods:nameIdentifier id="401C15BB175792C3BF7DFAB975E8D148" type="ORCID">0000-0001-9273-0040</mods:nameIdentifier>
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<mods:affiliation id="B2B958CF7401AFF934721FB1705302BB">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
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</mods:name>
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<mods:name id="87A6B435570F279D6C57EE285C477B9B" type="personal">
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<mods:roleTerm id="A975DA460ADFD843CE1B88B65BD1F98A">Author</mods:roleTerm>
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<mods:namePart id="107B8EDD8DC00011321A303E0FB1F3D7">Nguyen, Lien T. P.</mods:namePart>
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<mods:affiliation id="637EB34E480401C6356E52B3549761D7">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
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</mods:name>
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<mods:name id="88676384D3519CD957D99D2AEED602F8" type="personal">
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<mods:role id="B6BB28BF04A705B4780FB0B18A79A9A8">
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<mods:roleTerm id="757A5FA91122F176FAFC3406D729FA52">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BF277558C55137C8CCEA7800A7987648">Korsós, Zoltán</mods:namePart>
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<mods:nameIdentifier id="95F94A1F3E09E62EE4A8D1285E56BA94" type="ORCID">0000-0003-1545-5086</mods:nameIdentifier>
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<mods:affiliation id="D0694A41D9AFE398130CDE427E4F120F">University of Veterinary Medicine Budapest, István u. 2, 1078 Budapest, Hungary</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="ECB8F5E62964ADFC138470B4095E5936">text</mods:typeOfResource>
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<mods:title id="C6B0A4D5931BDFAE160E02B3BB60B5CF">Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part id="3AF729D6306586AEBB781DACC65435D4">
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<mods:date id="C01F28197B77C4AB04DCB8F4CE5FF904">2024</mods:date>
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<mods:detail id="DADD9956FDD689232FFB5E6C68F4143B" type="pubDate">
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<mods:number id="94FCC5CEB27CCFD552FBAAFBC4C83222">2024-11-07</mods:number>
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<mods:detail id="D07ABCDF6B2D6A02F9D77F44BF210695" type="volume">
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<mods:number id="A5648947FACB5D4933C93B487C798CE7">100</mods:number>
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</mods:detail>
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<mods:detail id="0F674EE586652A7563BBF66FF16CE00A" type="issue">
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<mods:number id="0E73A11C072971FBA9489AB8B52192F7">4</mods:number>
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<mods:extent id="C3CB6A28778AB6FB41C32702AAA02EC2" unit="page">
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<mods:start id="036BAD0B5E117509D34D21A98ACCE340">1515</mods:start>
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<mods:end id="47AE44DE68D858102DFF71FA1CEC863A">1541</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="F37985AE94922906B19A6C28F7FAB47D">journal article</mods:classification>
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<mods:identifier id="ED1A9B023970CA36D9D32636483A5801" type="DOI">10.3897/zse.100.132111</mods:identifier>
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<treatment id="92A4751707F158C499FD3B94C2D1B998" ID-DOI="http://doi.org/10.5281/zenodo.14052233" ID-Zenodo-Dep="14052233" ID-arpha="92A47517-07F1-58C4-99FD-3B94C2D1B998" ID-zoobank="https://zoobank.org/2FD3A321-E3F2-419E-A200-4446A132B10A" LSID="urn:lsid:plazi:treatment:92A4751707F158C499FD3B94C2D1B998" LSID-ZBK="urn:lsid:zoobank.org:act:2FD3A321-E3F2-419E-A200-4446A132B10A" httpUri="http://treatment.plazi.org/id/92A4751707F158C499FD3B94C2D1B998">
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||
<subSubSection id="76FFCDCF45C694B1C77CAF291391CD99" type="nomenclature">
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||
<paragraph id="F69C680C1A19D4EA43488545D256D60E">
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<taxonomicName id="835A0541C3EE713BE6866B3C0A338FB7" authorityName="Nguyen & Nguyen & Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="irregularis" status="sp. nov.">
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<emphasis id="2F3CCCF33AE2DEDA2944C06802B9771A" italics="true">Pseudodesmus irregularis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="624BB255AAB10403A82A1E3DECAB7E8E" rank="species">sp. nov.</taxonomicNameLabel>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="DB7883FBE81A28543D9CDA4D31C65830" type="description">
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||
<paragraph id="95AC61DB027DCA6DBB9D8FFBD79631B9">
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||
<figureCitation id="D4C05D747F6C78DBC36975B8FC042950" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">Figs 21</figureCitation>
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,
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<figureCitation id="C1E3904937A970D761D1BE3B011D60C4" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22</figureCitation>
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,
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<figureCitation id="FC3E325D6CA5D60EBF2DB13101800BEE" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. A. Left anterior gonopod, anterior view; B. Left posterior gonopod, sublateral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure23" httpUri="https://binary.pensoft.net/fig/1169913">23</figureCitation>
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,
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<figureCitation id="250B03AF395B4166D6693C28E8F05619" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. SEM. A, B. Left anterior gonopod, anterior view (A), ventral view (B); C. Distal part of left anterior gonopod, anterior view; D, E. Posterior gonopod and leg 11, ventral view (D), sublateral view (E); F. Distal part of posterior gonopod, ventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure24" httpUri="https://binary.pensoft.net/fig/1169914">24</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0ED1BG" type="material">
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||
<paragraph id="E11A1E99A6914A3E50FB95B9151D4077">
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||
<heading id="04E2C7C7DBB1B22490B74AA6B10EFB8B" reason="title">Material examined.</heading>
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||
</paragraph>
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||
<paragraph id="070712A5CAC703A52ACE3488530D78ED">
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||
<materialsCitation id="49374A688EF41B262508D28DDAF83464" collectedFrom="primary forest" collectingDate="2006-04-15" collectingDateMax="2006-04-24" collectingDateMin="2006-04-15" collectorName="A. D. Nguyen" country="Vietnam" elevation="1500" latitude="12.11124" location="Hon Ba Mt." longLatPrecision="1" longitude="108.98426" specimenCode="IEBR-Myr 958 H" specimenCount="1" specimenCount-male="1" stateProvince="Khanh Hoa Province" typeStatus="holotype">
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||
<emphasis id="A961241B165D41FC56ADF56525C9BFDC" bold="true" italics="true">
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||
<typeStatus id="35D62E9349500536C4BF59B88FD87AEE" type="holotype">Holotype</typeStatus>
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||
.
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||
</emphasis>
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||
<collectingCountry id="A5BE041FB1BEBA45DDDFBEA7B722B2E0" name="Vietnam">Vietnam</collectingCountry>
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||
•
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||
<specimenCount id="807AE7A2380DD8BF4FB54413BA991D2C" count="1" type="male">1 male</specimenCount>
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||
;
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||
<collectingRegion id="A2FA515AC9FC2A786BF659810B33B23C" country="Vietnam" name="Khanh Hoa">Khanh Hoa Province</collectingRegion>
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||
,
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||
<location id="2911AB3C3CBA63534BAEE12E6BFC04C7" LSID="urn:lsid:plazi:treatment:92A4751707F158C499FD3B94C2D1B998:2911AB3C3CBA63534BAEE12E6BFC04C7" country="Vietnam" latitude="12.11124" longLatPrecision="1" longitude="108.98426" name="Hon Ba Mt." stateProvince="Khanh Hoa Province">Hon Ba Mt.</location>
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||
;
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||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ER1BG" specific-use="{"type":"Point","coordinates":[108.984260,12.111240]}">
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||
<geoCoordinate id="68A2A8F4ADD3E45DFFF2C077A971C5AB" degrees="12.11124" direction="north" orientation="latitude" precision="1" value="12.11124">12.11124 ° N</geoCoordinate>
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||
,
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||
<geoCoordinate id="146C429CDD142418F932E22A0BA2E39C" degrees="108.98426" direction="east" orientation="longitude" precision="1" value="108.98426">108.98426 ° E</geoCoordinate>
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||
</named-content>
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||
;
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||
<elevation id="D5DA4A84E2526B21F85E753982BF56C4" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">
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||
<quantity id="FB840B0B50D716B5937F761DB1A103E7" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1,500 m</quantity>
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||
a. s. l.
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||
</elevation>
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||
;
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||
<date id="FFBF79BCDB3C6D0CC1D4FF7F6C9CCA7D" value="2006-04-15" valueMax="2006-04-24" valueMin="2006-04-15">
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<collectingDate id="11FCAAEC64C2A118AB6F239BA6AC6B13" value="2006-04-15" valueMax="2006-04-24" valueMin="2006-04-15">15–24 Apr. 2006</collectingDate>
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||
</date>
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;
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<collectorName id="F13C63EC4814B88798A72C32792A026B">A. D. Nguyen</collectorName>
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leg.;
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<collectedFrom id="9D1000EAF9C6DE200DD18BC92F63B273">primary forest</collectedFrom>
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;
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<specimenCode id="0343CD70B21CB78DECF732E70D0A29B3">
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<abbrev content-type="institution" id="ABBRID0EW1BG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
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958 H
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</specimenCode>
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||
.
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</materialsCitation>
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</paragraph>
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<paragraph id="06C116DDDAF7C37FC70A739CAF5E2F75">
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<materialsCitation id="6E30D8536BA40EA05C38D1130B3E062F" collectedFrom="primary forest" collectingDate="2006-04-15" collectingDateMax="2006-04-24" collectingDateMin="2006-04-15" collectorName="A. D. Nguyen" country="Vietnam" elevation="1500" latitude="12.11124" location="Vietnam" longLatPrecision="1" longitude="108.98426" specimenCode="IEBR-Myr 958 P" specimenCount="1" specimenCount-male="1" stateProvince="Khanh Hoa" typeStatus="paratype">
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||
<emphasis id="7060CC674070ECD3B6744AF8CE37EDA4" bold="true" italics="true">
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||
<typeStatus id="C738A765EA0B3A282D7F483030322DA8" type="paratype">Paratypes</typeStatus>
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||
.
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</emphasis>
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||
<collectingCountry id="BFCDC57F6CD2A522E8A8D9FC7AEED731" name="Vietnam">Vietnam</collectingCountry>
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||
•
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<specimenCount id="CE61F84AE3502B9C432E4BDA437D3E39" count="1" type="male">1 male</specimenCount>
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||
; same data as for holotype;
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||
<specimenCode id="ABB77F1615EBB72516385C6845CDC296">
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<abbrev content-type="institution" id="ABBRID0EC2BG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
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||
958 P
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||
</specimenCode>
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||
</materialsCitation>
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||
•
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||
<materialsCitation id="E405F93C2297CC10E20F1B975F2517D6" collectedFrom="regenerated forest" collectingDate="2004-03-25" collectingDateMax="2004-04-11" collectingDateMin="2004-03-25" collectorName="A. D. Nguyen" country="Vietnam" elevation="1700" latitude="15.07071" location="Ngoc Linh Mt" longLatPrecision="1" longitude="107.97142" specimenCode="IEBR-Myr 960" specimenCount="1" specimenCount-female="1" stateProvince="Kon Tum Province" typeStatus="paratype">
|
||
<specimenCount id="8946CDE967036F0CAE997D145AF0FC3B" count="1" type="female">1 female</specimenCount>
|
||
;
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||
<collectingRegion id="1CDFB8321203ACB28BE752F502365F24" country="Vietnam" name="Kon Tum">Kon Tum Province</collectingRegion>
|
||
,
|
||
<location id="EE6923B08EF0375718027810DD839802" LSID="urn:lsid:plazi:treatment:92A4751707F158C499FD3B94C2D1B998:EE6923B08EF0375718027810DD839802" country="Vietnam" latitude="15.07071" longLatPrecision="1" longitude="107.97142" name="Ngoc Linh Mt" stateProvince="Kon Tum Province">Ngoc Linh Mt</location>
|
||
;
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EK2BG" specific-use="{"type":"Point","coordinates":[107.971420,15.070710]}">
|
||
<geoCoordinate id="37304AC4EF3B7FDF574ABE44C3627B4A" degrees="15.07071" direction="north" orientation="latitude" precision="1" value="15.07071">15.07071 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5DF52E8F526DA0F90972FB31B258FB8D" degrees="107.97142" direction="east" orientation="longitude" precision="1" value="107.97142">107.97142 ° E</geoCoordinate>
|
||
</named-content>
|
||
;
|
||
<elevation id="36F52CABC211C1A33B938D7A6FB4C77C" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
|
||
<quantity id="382D83E3F908B2165542E7DD8A133B40" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1,700 m</quantity>
|
||
a. s. l.
|
||
</elevation>
|
||
;
|
||
<date id="37F78344CF7B80F99D2B6B8BCD6C3CDE" value="2004-03-25" valueMax="2004-04-11" valueMin="2004-03-25">
|
||
<collectingDate id="AC5AEC19A877DAF7E871DE9B98C72677" value="2004-03-25" valueMax="2004-04-11" valueMin="2004-03-25">25 Mar. – 11 Apr. 2004</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="5C5C0D75F98A4BAEE3943D7FCA8C4209">A. D. Nguyen</collectorName>
|
||
leg.;
|
||
<collectedFrom id="8DF9F10FDE1440A4D188BEEFCF222A1D">regenerated forest</collectedFrom>
|
||
;
|
||
<specimenCode id="85650FF782B12BAA08F62AE7B2EB2853">
|
||
<abbrev content-type="institution" id="ABBRID0EP2BG" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
|
||
960
|
||
</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EU2BG" type="diagnosis">
|
||
<paragraph id="06517F8696C6FB2254B27AD57219AD7D">
|
||
<heading id="C126A4C0EDDF84F9BDAD812F6F89C025" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
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<paragraph id="94D844B2511758CEF3916D654EB3F4E6">Head narrower than collum. Collum bilobed, caudal margin highly elevated with 3 + 3 tubercles with 3 + 3 tubercles, and with a row of 1 + 1 tubercles. Midbody ring with 2 distinct rows of tubercles, anterior row of 6–9 + 6 – 9 and posterior of 3 + 3. Paratergal margins with small / tiny dentations or notches. Posterior gonopods 6 - segmented, with three apical stylets.</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="SECID0EZ2BG" type="diagnosis">
|
||
<paragraph id="CE4B11782FCBD379D35E5E666E660167">
|
||
<heading id="18820DD803CAD07441F65C76CD3A88C8" reason="title">Diagnosis remarks.</heading>
|
||
</paragraph>
|
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<paragraph id="722EB33E483565CCAEDDE3E954AB1012">
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The new species is distinguished from
|
||
<taxonomicName id="652CBEC50E862E58F72A343B60A597A7" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="camptotrichus">
|
||
<emphasis id="C05AAAECEEE1B4BA209DA72CC12C3F95" italics="true">P. camptotrichus</emphasis>
|
||
</taxonomicName>
|
||
by coloration (irregularly yellow with brown spots vs earth brown), shorter length (
|
||
<quantity id="89A3544D888F2E903FD039C0C4A8439C" metricMagnitude="-2" metricUnit="m" metricValue="1.292" unit="mm" value="12.92">12.92 mm</quantity>
|
||
vs
|
||
<quantity id="4A5C382AEEAB2BB5A361494DBC2CDC7E" metricMagnitude="-2" metricUnit="m" metricValue="1.4" unit="mm" value="14.0">14 mm</quantity>
|
||
), slightly wider (
|
||
<quantity id="D97A0F4656D6E6741AD8BC174E121F72" metricMagnitude="-3" metricUnit="m" metricValue="2.39" unit="mm" value="2.39">2.39 mm</quantity>
|
||
vs 2.0 mm), having less number of tubercles in posterior metazonal row (3 + 3 vs 6–9 + 6 – 9). The new species is similar to
|
||
<taxonomicName id="5C5A78FBB7818C0839E643EDBCFD903F" baseAuthorityName="Attems" baseAuthorityYear="1938" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="variegatus">
|
||
<emphasis id="0BD96D7B51E5AEF0536DC1F60885CB4A" italics="true">P. variegatus</emphasis>
|
||
</taxonomicName>
|
||
regarding body coloration (irregularly yellow with brown spots), but two species can be separated by body size (length:
|
||
<quantity id="7D952364108AC7B99192574B47C36B04" metricMagnitude="-2" metricUnit="m" metricValue="1.296" unit="mm" value="12.96">12.96 mm</quantity>
|
||
vs
|
||
<quantity id="3E15586925CECEBFA805A75AF8F37CA0" metricMagnitude="-2" metricUnit="m" metricValue="3.2" unit="mm" value="32.0">32 mm</quantity>
|
||
; width:
|
||
<quantity id="4D65BE99F37388C508CFA54EDF571392" metricMagnitude="-3" metricUnit="m" metricValue="2.39" unit="mm" value="2.39">2.39 mm</quantity>
|
||
vs
|
||
<quantity id="074CFAA1668693BC1224CC76EF249B70" metricMagnitude="-3" metricUnit="m" metricValue="5.8" unit="mm" value="5.8">5.8 mm</quantity>
|
||
) and and number of metazonal tubercles (two rows of 6–9 + 6 – 9 and 3 + 3 vs two rows of 8–15 + 8 – 15 and 5–7 + 5 – 7).
|
||
</paragraph>
|
||
<paragraph id="1C892FB59ADEB79C529BCA58835B36C5">
|
||
<taxonomicName id="EAB86053E8D4CBCAD58C2BD0C1DD9A38" authorityName="Nguyen & Nguyen & Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="irregularis" status="sp. nov.">
|
||
<emphasis id="40B771AB9AB5B7E40470A8CBFFE4E4D7" italics="true">Pseudodesmus irregularis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FD9E428DD9EA076D0B15E1DBFD8A3EF0" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is differentiated from above four new species,
|
||
<taxonomicName id="16EEB83CE5DD57F1D0A8FE4B43E6414F" authorityName="Nguyen & Nguyen & Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="bidoup" status="sp. nov.">
|
||
<emphasis id="BC6DF077DE9D394887EE3DD0700D5242" italics="true">P. bidoup</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D371B82A1D43F3B9EC0CCBED22A193DC" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="731F6AE1D2B08DCE66676921EB4334D4" authorityName="Nguyen & Nguyen & Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="ngoclinh" status="sp. nov">
|
||
<emphasis id="7D70FCD230DF420E8F99048E841F03C0" italics="true">P. ngoclinh</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F315AC10B4878F3738FD53BF8464A279" rank="species">sp. nov</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="F846FF89687F032752455F90310984BC" authorityName="Nguyen & Nguyen & Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="karstomus" status="sp. nov.">
|
||
<emphasis id="BE464DDA84334345A325614109E60492" italics="true">P. karstomus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="615082F07DE9CF68EA08BDF7F5777DBF" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="004E05020971C4C347BC4832B4BDC73A" authorityName="Nguyen & Nguyen & Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="condao" status="sp. nov.">
|
||
<emphasis id="3E9680D2AD3F9C98D5FD49F6B8702C75" italics="true">P. condao</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BBCB4E7D938FAA794C7C65DCA4240426" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in body coloration of irregularly yellow with brown spots, body size and number of stylets (three vs four, two, four, and two, respectively).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EO5BG" type="description">
|
||
<paragraph id="76D01D1670F676C1B800E82F0971F42C">
|
||
<heading id="83CF53C61049AC080554EAB234C88C39" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="1F233A94A28D4C1A640E79250D21E4E8">
|
||
<emphasis id="802FBEF239FA916488C992619A42BDAF" bold="true">
|
||
Male
|
||
<typeStatus id="34BE8C29AB1BA1CC548889EE8D3588CB" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="718B01DF5C3446917972955859EAC0A2">
|
||
<emphasis id="6ADCAB57E55781C4DDB41DA84C4C7EA2" bold="true" italics="true">Measurements</emphasis>
|
||
: 42 body rings plus telson, length about
|
||
<quantity id="388D46933AC9FF62AA59147669AFF5B9" metricMagnitude="-2" metricUnit="m" metricValue="1.296" unit="mm" value="12.96">12.96 mm</quantity>
|
||
, length of midbody metazona
|
||
<quantity id="50F4B5503AD3F6A48069A7CF6A8B6AA3" metricMagnitude="-4" metricUnit="m" metricValue="2.4" unit="mm" value="0.24">0.24 mm</quantity>
|
||
, width of midbody metazona
|
||
<quantity id="D527EAEC5F1A162EFD8779AF9375066D" metricMagnitude="-3" metricUnit="m" metricValue="2.39" unit="mm" value="2.39">2.39 mm</quantity>
|
||
, width of midbody prozona
|
||
<quantity id="BBDBE19BCA836217F1713B77B8007AB3" metricMagnitude="-3" metricUnit="m" metricValue="1.08" unit="mm" value="1.08">1.08 mm</quantity>
|
||
; ratio of width of head and width of collum = 0.74.
|
||
</paragraph>
|
||
<paragraph id="6C77CCFDBD860387FBF2D8A05F336273">
|
||
<emphasis id="8A5AF910AFED73F5128B8C7DA9628A06" bold="true" italics="true">Coloration</emphasis>
|
||
of ethanol-preserved specimens (Fig.
|
||
<figureCitation id="066538CBDE51A864F4F1D9D1FEC522A0" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudodesmus condao sp. nov. Holotype (IEBR-Myr 982 H), gonopods. SEM. A, B. Left posterior gonopod, subventral view (A), anterior view (B); C. Anterior gonopods, anterior view; D. Left anterior gonopod, anterior view; E. Distal part of left posterior gonopod, anterior view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure20" httpUri="https://binary.pensoft.net/fig/1169910">20</figureCitation>
|
||
): complex and irregular: body rings 1–4 with none, body rings 5–7 with 2 brown spots, body ring 8 brown on the right, body ring 9 with two brown spots, body ring 10 brown on the right, body ring 11 brown on the left, body rings 12–13 with two brown spots, body ring 14 none, body ring 15 brown on the left, body rings 16–17 brown on the right, body ring 18 brown on left half, body ring 19 brown on right half, body ring 20 brown on left half, body ring 21 none, body ring 22 with left spot, body ring 23 with two brown spots, body rings 24–26 brown on left half, body rings 27–28 full dark, body ring 29 none, body rings 30–32 brown left, body rings 33–34 brown right, body rings 35–36 full, body rings 37–38 none, body ring 39 brown right, body rings 40–42 light brown.
|
||
</paragraph>
|
||
<paragraph id="ED3E6A3206B9DA399DF7BE47AEE77000">
|
||
Head (Figs
|
||
<figureCitation id="25532EFAABB1847EAA99A669B5F71D9A" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21 B</figureCitation>
|
||
,
|
||
<figureCitation id="9C6E26D0D9B008F4BC60FF0D35991E41" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 B</figureCitation>
|
||
) narrower than collum in width, smooth, round-shaped. Antenna stout, clavate,
|
||
<emphasis id="EE8D3BB145792A81C740C0BB39CB50B2" italics="true">in situ</emphasis>
|
||
extending to body ring 6; antennomere 6> 5> 3> 4 = 2> 7 =
|
||
<quantity id="B5384444DC325EAF128AF6FC4BC27622" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
|
||
length (Figs
|
||
<figureCitation id="3D18ADC72CAB9BE5426AF7F2BB7972A5" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21 B</figureCitation>
|
||
,
|
||
<figureCitation id="3649B38DC65B84E3C75ADECE6E4848BD" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 B</figureCitation>
|
||
). Collum (Figs
|
||
<figureCitation id="67F4AAF36335D3C4E713C84E2AC87AF1" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21 B</figureCitation>
|
||
,
|
||
<figureCitation id="D12254901AE1D727C6AC66A41E4F6E9C" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 B</figureCitation>
|
||
) bilobuled, caudal margin highly elevated with 3 + 3 tubercles, and with a row of 1 + 1 tubercles, paramedian one much larger than lateral one.
|
||
</paragraph>
|
||
<caption id="6FC915BBC12E5A9CC34CC0A5F0534755" ID-DOI="10.3897/zse.100.132111.figure21" ID-arpha="3BFBCC07-E890-575A-A50D-59E368F11E98" httpUri="https://binary.pensoft.net/fig/1169911" startId="F21">
|
||
<paragraph id="44D73A148AEDAFD9C86C84D3AF41E68E">
|
||
<label id="96BACA2E26B33F210F0AF1F3AC707BF7">Figure 21.</label>
|
||
</paragraph>
|
||
<paragraph id="618CD4BE16461E273AECDAEBC3535B3C">
|
||
<taxonomicName id="215E79C3E519AFF5629E953B51360736" authorityName="Nguyen & Nguyen & Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="irregularis" status="sp. nov.">
|
||
<emphasis id="DBA25AFBAADD4DA4F93100307B869172" italics="true">Pseudodesmus irregularis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E523AD0F377784AAD24FC3895419141A" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype (
|
||
<specimenCode id="0746A23CB779922999F41C46871939F7">
|
||
<abbrev content-type="institution" id="ABBRID0EYAAI" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
|
||
958
|
||
</specimenCode>
|
||
H), under normal light.
|
||
<emphasis id="00A5D2704D0D9E9FBAB8583EF0CAEA20" bold="true">A, B.</emphasis>
|
||
Anterior part of body, dorsal view (
|
||
<emphasis id="F0942817E5530F410077E12BE2E86A67" bold="true">A</emphasis>
|
||
), ventral view (
|
||
<emphasis id="9A7C467E75C5C005353130A49F88776F" bold="true">B</emphasis>
|
||
);
|
||
<emphasis id="D5CD85EE7A85F6D9A07B092ACE5ECF29" bold="true">C, D.</emphasis>
|
||
Midbody body rings, dorsal view (
|
||
<emphasis id="C3B86C2D89B869BF56315E29A1CBAED8" bold="true">C</emphasis>
|
||
), ventral view (
|
||
<emphasis id="4DA344CA524C5DB67F647C1071A154DC" bold="true">D</emphasis>
|
||
);
|
||
<emphasis id="20A5A46E1E4C988AE8ADDF9E3AB137ED" bold="true">E, F.</emphasis>
|
||
Posterior part of body, dorsal view (
|
||
<emphasis id="C1255ED4102F79F8C9BC2BAB69725CF9" bold="true">E</emphasis>
|
||
), ventral view (
|
||
<emphasis id="9BEF0D801ECA9FDBED17F4718980202B" bold="true">F</emphasis>
|
||
). Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="343AE333CA58A3DCE1A2A46C9ABD55A7" ID-DOI="10.3897/zse.100.132111.figure22" ID-arpha="4F60D10F-21F1-5FCB-BB11-8B1451742D6B" httpUri="https://binary.pensoft.net/fig/1169912" startId="F22">
|
||
<paragraph id="F0AB55AD1C680939A7F05674229CC5B6">
|
||
<label id="CE1DC4B2568988F15749CC123AE06880">Figure 22.</label>
|
||
</paragraph>
|
||
<paragraph id="8A3BB8394EC26187117BC8ACCF936DE7">
|
||
<taxonomicName id="6DE6B1850173235F10F1E04F6F0481F9" authorityName="Nguyen & Nguyen & Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="irregularis" status="sp. nov.">
|
||
<emphasis id="9607A79F7F2F4234ADEA155ACAFD37A6" italics="true">Pseudodesmus irregularis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BA5A5752F747CFD28BD229D254A16B26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype (
|
||
<specimenCode id="6F0A870AF1236FD3C434B228068CACB6">
|
||
<abbrev content-type="institution" id="ABBRID0EHCAI" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
|
||
958
|
||
</specimenCode>
|
||
H), under UV light.
|
||
<emphasis id="DF183C5F1C02B11C17AF5CB628E7DA10" bold="true">A, B.</emphasis>
|
||
Anterior part of body, dorsal view (
|
||
<emphasis id="B7B8E3DA76BF84A066943B7F6312F08B" bold="true">A</emphasis>
|
||
), ventral view (
|
||
<emphasis id="A14E54B8A6039FF4C353C496782D30C5" bold="true">B</emphasis>
|
||
);
|
||
<emphasis id="7A2F7716B5DB0F88F9AE2403DA59CC90" bold="true">C.</emphasis>
|
||
Cross section of midbody body ring, posterior view;
|
||
<emphasis id="5544A4493A9FC70FCFD25A8B2B905743" bold="true">D.</emphasis>
|
||
Midbody body rings, dorsal view;
|
||
<emphasis id="8394E9391FAE743A072D6ED1DAD7C1D3" bold="true">E, F.</emphasis>
|
||
Posterior part of body, dorsal view (
|
||
<emphasis id="0BF3E1DC0FE54A09321F74ADE6646BB0" bold="true">E</emphasis>
|
||
), ventral view (
|
||
<emphasis id="B58579B50D9247C2FCF40D4A66A0EFA9" bold="true">F</emphasis>
|
||
). Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="98C8C486D36A52F6501B51D3D83CC9C6">
|
||
Body (Figs
|
||
<figureCitation id="0384762FF27C72577566968EC1D4002D" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21</figureCitation>
|
||
,
|
||
<figureCitation id="2061633F32DEB4D557BBF762A9134164" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22</figureCitation>
|
||
): ring 2 with two rows of 3 + 3 and 1 + 1 tubercles; ring 3 with two rows of 3 + 3 and 3 + 3 tubercles; other rings with 2 distinct rows of tubercles, anterior row of 6 + 6 (increasing to (9 + 9) on midbody rings), and posterior of 2 + 4 (body ring 4), 3 + 3 (ring 5), 3 + 2 (ring 6), 3 + 3 (on other rings). The last 5 rings with two rows of 3 + 3 and 3 + 3 tubercles (Fig.
|
||
<figureCitation id="A6255C721A6218C6106C5B222DDE0B02" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 E</figureCitation>
|
||
). Anterior rows extending to 2 / 3 paraterga. Paramedian tubercles on posterior rows much larger than ones on anterior rows.
|
||
</paragraph>
|
||
<paragraph id="693091F54DBA47DD9B1BD5A84D1A9CDF">
|
||
Paraterga well developed, slightly curved anteriad on body rings 2–3, lateral margins smooth (Figs
|
||
<figureCitation id="1762B618EC8C477FB9E8F20967D4B863" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21 A, B</figureCitation>
|
||
,
|
||
<figureCitation id="6A004220E042E333674958B00228B343" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 A, B</figureCitation>
|
||
); increasingly less so on the following body rings, slightly curved caudad on body rings 25–27, and strongly curved caudad on body rings 37–40; paraterga of penultimate body ring produced strictly caudad and flanking telson (Figs
|
||
<figureCitation id="99BAED6CD845F7C238F587366F3289CF" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21 E, F</figureCitation>
|
||
,
|
||
<figureCitation id="4853AF22948D0A0B5AEB5DE2B21FBB6F" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 E, F</figureCitation>
|
||
). Lateral margin of paraterga with small / tiny dentations or notches (Figs
|
||
<figureCitation id="9411B93EE652B8260EB37AC3DBAE8355" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21</figureCitation>
|
||
,
|
||
<figureCitation id="743952980C718A396521DF7E53447497" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="157D70479DDFEA81B40F70AC56A1EF2E">
|
||
Telson (Figs
|
||
<figureCitation id="F6304C1F2868C2D8CF96A0C5C5D033D3" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under normal light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C, D. Midbody body rings, dorsal view (C), ventral view (D); E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure21" httpUri="https://binary.pensoft.net/fig/1169911">21 E, F</figureCitation>
|
||
,
|
||
<figureCitation id="645B1610D76EEEA2906A3B7812CC3A87" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 E, F</figureCitation>
|
||
) short, caudal margin rounded; epiproct with 2 + 2 small / tiny tubercles at caudal margin. Paraprocts and hypoproct semi-circular.
|
||
</paragraph>
|
||
<paragraph id="B7D5ED95B2906C12A0CE122BBE54228A">
|
||
Legs slender, shorter than body ring width together with paraterga, terminating before lateral paratergal margins. Claws normal. Coxal sacs strong, present on body rings 3–32 (Fig.
|
||
<figureCitation id="A45C65EFC0F7A11606D3999279D56E68" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), under UV light. A, B. Anterior part of body, dorsal view (A), ventral view (B); C. Cross section of midbody body ring, posterior view; D. Midbody body rings, dorsal view; E, F. Posterior part of body, dorsal view (E), ventral view (F). Scale bars: 1 mm." figureDoi="10.3897/zse.100.132111.figure22" httpUri="https://binary.pensoft.net/fig/1169912">22 F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FD4D1A73AD25C48BEEDD16E1CD1A40A8">
|
||
<emphasis id="71358AAE26BE6B9CA49C13B0E1F88881" bold="true" italics="true">Gonopods</emphasis>
|
||
(Figs
|
||
<figureCitation id="0CF3380E799F78809638D68B2F5DD685" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. A. Left anterior gonopod, anterior view; B. Left posterior gonopod, sublateral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure23" httpUri="https://binary.pensoft.net/fig/1169913">23</figureCitation>
|
||
,
|
||
<figureCitation id="5A316F33988032BB255A4CF8FAA3F3D3" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. SEM. A, B. Left anterior gonopod, anterior view (A), ventral view (B); C. Distal part of left anterior gonopod, anterior view; D, E. Posterior gonopod and leg 11, ventral view (D), sublateral view (E); F. Distal part of posterior gonopod, ventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure24" httpUri="https://binary.pensoft.net/fig/1169914">24</figureCitation>
|
||
): Two pairs of gonopods directed anteriad. Anterior gonopods (Figs
|
||
<figureCitation id="E42736B52FCDAA425767618DBE69C676" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. A. Left anterior gonopod, anterior view; B. Left posterior gonopod, sublateral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure23" httpUri="https://binary.pensoft.net/fig/1169913">23 A</figureCitation>
|
||
,
|
||
<figureCitation id="27CC88799850710E6DEF20EB79F013DB" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. SEM. A, B. Left anterior gonopod, anterior view (A), ventral view (B); C. Distal part of left anterior gonopod, anterior view; D, E. Posterior gonopod and leg 11, ventral view (D), sublateral view (E); F. Distal part of posterior gonopod, ventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure24" httpUri="https://binary.pensoft.net/fig/1169914">24 A – C</figureCitation>
|
||
) 6 - segmented, covered with long setae; coxite (
|
||
<emphasis id="18C9E09C5ED8B1E9A307DFC58529B498" italics="true">co</emphasis>
|
||
) broad, basal part sparsely covered with microgranulations; podomeres 2–5, short and stout; podomere 6 slightly longer than other podomeres, distally carrying macrosetae (
|
||
<emphasis id="C194DD37F0BCBC703C5F6C5A954C166F" italics="true">mse</emphasis>
|
||
). Posterior gonopods (Figs
|
||
<figureCitation id="1A3589A9A5982CEAAC35BF2061E6E56A" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. A. Left anterior gonopod, anterior view; B. Left posterior gonopod, sublateral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm." figureDoi="10.3897/zse.100.132111.figure23" httpUri="https://binary.pensoft.net/fig/1169913">23 B</figureCitation>
|
||
,
|
||
<figureCitation id="ABED24E3CA8C4256C57501F56A056498" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Pseudodesmus irregularis sp. nov. Holotype (IEBR-Myr 958 H), gonopods. SEM. A, B. Left anterior gonopod, anterior view (A), ventral view (B); C. Distal part of left anterior gonopod, anterior view; D, E. Posterior gonopod and leg 11, ventral view (D), sublateral view (E); F. Distal part of posterior gonopod, ventral view. Abbreviations: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet." figureDoi="10.3897/zse.100.132111.figure24" httpUri="https://binary.pensoft.net/fig/1169914">24 D – F</figureCitation>
|
||
) 6 - segmented, covered with sparse long setae; coxite (
|
||
<emphasis id="2E71C5A71B095DA17B2D23A0E58BD7BD" italics="true">co</emphasis>
|
||
) broad, basal part sparsely covered with microgranulations; podomeres 2–5 short and stout; ultimate podomere (
|
||
<emphasis id="E927E08799E0082881A62B3B0C6516EF" italics="true">po 6</emphasis>
|
||
) longest, with three apical stylets (
|
||
<emphasis id="B69FC763E72295105C58AA4F7F416337" italics="true">sty</emphasis>
|
||
).
|
||
</paragraph>
|
||
<caption id="0A9E9451888D3018FEB7EFC98A7B3701" ID-DOI="10.3897/zse.100.132111.figure23" ID-arpha="D64B6B27-019F-54C0-BD85-2EE09DADBA95" httpUri="https://binary.pensoft.net/fig/1169913" startId="F23">
|
||
<paragraph id="169493012E1D3E5A2E220BC8CB922CAA">
|
||
<label id="FAED3053614D83ACD178D56E7EF70E3C">Figure 23.</label>
|
||
</paragraph>
|
||
<paragraph id="24B7FEE99E669F454AA47E2D70DEAF16">
|
||
<taxonomicName id="41DAC60934412D93DAB6BF0DAC9EF8D9" authorityName="Nguyen & Nguyen & Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="irregularis" status="sp. nov.">
|
||
<emphasis id="90588EE9C819FC6FDD456EEF990341BF" italics="true">Pseudodesmus irregularis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C15E21951480CFB98C7752064190DBC5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype (
|
||
<specimenCode id="B6589DCF2C5729B5FDDCDC022CFA5ED2">
|
||
<abbrev content-type="institution" id="ABBRID0EVGAI" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
|
||
958
|
||
</specimenCode>
|
||
H), gonopods.
|
||
<emphasis id="8AD1AC9DA8BD065DD1FE38BD48CDFBA3" bold="true">A.</emphasis>
|
||
Left anterior gonopod, anterior view;
|
||
<emphasis id="2C91F3C24C7C0FDF3BB83664DFD11C41" bold="true">B.</emphasis>
|
||
Left posterior gonopod, sublateral view.
|
||
<emphasis id="74A386EE11228B0D631B0AA9F424F409" italics="true">Abbreviations</emphasis>
|
||
: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="65CF122E46A0129575630D8B84033AE9" ID-DOI="10.3897/zse.100.132111.figure24" ID-arpha="A4B4F4D5-C781-5AC6-8AD4-108C2D0F22D0" httpUri="https://binary.pensoft.net/fig/1169914" startId="F24">
|
||
<paragraph id="9DB5F4399FF8C0C96415BE7B0538E34C">
|
||
<label id="0BF3673C9F181185B3A8B92AA687CA99">Figure 24.</label>
|
||
</paragraph>
|
||
<paragraph id="6FAD5032C28F18B8573F5FDAB4B9B01B">
|
||
<taxonomicName id="1D6E3BB144488C22CFE65F21098A9520" authorityName="Nguyen & Nguyen & Korsós" authorityYear="2024" class="Diplopoda" family="Andrognathidae" genus="Pseudodesmus" kingdom="Animalia" order="Platydesmida" phylum="Arthropoda" rank="species" species="irregularis" status="sp. nov.">
|
||
<emphasis id="47CB23D8B9085AC4FE2EA3405A508519" italics="true">Pseudodesmus irregularis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6207951DE2B1C91A920EBBD6A2EE87C5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype (
|
||
<specimenCode id="5B259262030F275B0DBD29ED716979F9">
|
||
<abbrev content-type="institution" id="ABBRID0EYHAI" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
|
||
958
|
||
</specimenCode>
|
||
H), gonopods.
|
||
<abbrev id="ABBRID0E4HAI" xlink_title="Scanning electron microscope">SEM</abbrev>
|
||
.
|
||
<emphasis id="AEB12D2C8E6F9A676EB99B706EA822C4" bold="true">A, B.</emphasis>
|
||
Left anterior gonopod, anterior view (
|
||
<emphasis id="3C6CD6C7B689E34D35CB36D3EBD8B60C" bold="true">A</emphasis>
|
||
), ventral view (
|
||
<emphasis id="3CBFC0CD2816C906B4E9BB00BE977B31" bold="true">B</emphasis>
|
||
);
|
||
<emphasis id="D069726F37FF74BA24DDA483840EDD30" bold="true">C.</emphasis>
|
||
Distal part of left anterior gonopod, anterior view;
|
||
<emphasis id="A395C70ED5417056DC26EA1CF47254EB" bold="true">D, E.</emphasis>
|
||
Posterior gonopod and leg 11, ventral view (
|
||
<emphasis id="CAFBA0D61C770A92EC8489D92B5774CF" bold="true">D</emphasis>
|
||
), sublateral view (
|
||
<emphasis id="44C70B8573E45103262994307FB5B288" bold="true">E</emphasis>
|
||
);
|
||
<emphasis id="F38311C5CD238CEE91FD1BF534BE73F8" bold="true">F.</emphasis>
|
||
Distal part of posterior gonopod, ventral view.
|
||
<emphasis id="4DF3BEF8A65C2E41F906B17DB1E28EBB" italics="true">Abbreviations</emphasis>
|
||
: co = coxite = podomere 1; po 2 = podomere 2; po 3 = podomere 3; po 4 = podomere 4; po 5 = podomere 5; po 6 = podomere 6 (ultimate podomere); mse = macrosetae; sty = stylet.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EYIAI" type="variation">
|
||
<paragraph id="7CA18905F3F2EFE44AEF85327AC6E5AA">
|
||
<heading id="674365FE2F049A5BF622AD541AECC742" reason="title">Variation.</heading>
|
||
</paragraph>
|
||
<paragraph id="7AB555462AEEAC565A2B9C3670E18295">
|
||
<emphasis id="279C0EABE33C2CECED678AE70A3BC6DA" bold="true" italics="true">
|
||
Male
|
||
<typeStatus id="82179F7139BA2F95B45D06AB265B6E12" type="paratype">paratype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<emphasis id="C31713F87114B948F6A4BF2741AD9D25" bold="true">
|
||
<specimenCode id="69BED6A856F6319B33943B1514819FE1">
|
||
<abbrev content-type="institution" id="ABBRID0EEJAI" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
|
||
958
|
||
</specimenCode>
|
||
P
|
||
</emphasis>
|
||
): 36 body rings plus telson, length about
|
||
<quantity id="3FF6D7CB536994CE66883865AC7B2B14" metricMagnitude="-3" metricUnit="m" metricValue="9.75" unit="mm" value="9.75">9.75 mm</quantity>
|
||
, length of midbody metazoan ca.
|
||
<quantity id="DBE0294014BC6B6F769ED43BA5A42C96" metricMagnitude="-4" metricUnit="m" metricValue="1.4" unit="mm" value="0.14">0.14 mm</quantity>
|
||
, width of midbody metazona ca.
|
||
<quantity id="D1346F46D89806C6E83A593722453E90" metricMagnitude="-3" metricUnit="m" metricValue="1.72" unit="mm" value="1.72">1.72 mm</quantity>
|
||
, width of midbody prozona ca.
|
||
<quantity id="B1BE29736A3B6FDE7D07EB569E47AEEF" metricMagnitude="-4" metricUnit="m" metricValue="8.2" unit="mm" value="0.82">0.82 mm</quantity>
|
||
; ratio of width of head and width of collum = 0.85.
|
||
</paragraph>
|
||
<paragraph id="EB0D102C15C2FBCBF5B44CBF3B09B284">
|
||
The female from Ngoc Linh Mts (
|
||
<emphasis id="F8E5FE013525FBE1BCB91C98649FCCB8" bold="true">
|
||
<specimenCode id="E62928C03AB3B2DC2054FD95242BD988">
|
||
<abbrev content-type="institution" id="ABBRID0ENJAI" xlink_title="Institute of Ecology and Biological Resources, Myriapod collection">IEBR-Myr</abbrev>
|
||
960
|
||
</specimenCode>
|
||
</emphasis>
|
||
) has 41 body rings plus telson; midbody width
|
||
<quantity id="32BF92EC18ABF8E97352406E0E521A8E" metricMagnitude="-3" metricUnit="m" metricValue="2.1" unit="mm" value="2.1">2.1 mm</quantity>
|
||
, length about
|
||
<quantity id="111A9C5A1043F49C515C259DBA9DB702" metricMagnitude="-2" metricUnit="m" metricValue="1.011" unit="mm" value="10.11">10.11 mm</quantity>
|
||
. Coloration with widening pattern repeated between 7–12, 13–17, 18–24, 25–31, 32–36 body rings (see also
|
||
<bibRefCitation id="001F83566BAA4F219CEAC754D11CE1CF" DOI="10.3897/ijm.6.1949" author="Enghoff" firstAuthor="Enghoff" journalOrPublisher="International Journal of Myriapodology" pagination="1-27" refId="B10" refString="Enghoff H (2011) Trans-segmental serial colour patterns in millipedes and their developmental interpretation (Diplopoda). International Journal of Myriapodology 6: 1–27. https://doi.org/10.3897/ijm.6.1949" title="Trans-segmental serial colour patterns in millipedes and their developmental interpretation (Diplopoda)." volume="6" year="2011">Enghoff 2011</bibRefCitation>
|
||
). Tubercles are small: on collum only one row, on 2–4 two weak rows (reaching only one row on paraterga), from body ring 5 onwards two rows, each 1 + 1 large tubercles in the middle, then anterior row 10–13 small tubercles reaching edge of paraterga, in posterior row only 3–4 small tubercles, on the last five body rings anterior row is weaker but still with 8–10 small tubercles, in posterior row the median tubercles larger, with 4 small tubercles laterally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EXJAI" type="etymology">
|
||
<paragraph id="59292F5DDA7E4E6C03B4BD1FFBC6D850">
|
||
<heading id="B43539290021D2B67EC8ABD0BC834EAA" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="55BF7134A4914D8D860ACB97AC6E1E1F">To emphasize the irregular pattern of terga. Noun in apposition.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E3JAI" type="distribution">
|
||
<paragraph id="68BD8110F705332DDEF24B5691E9FB8F">
|
||
<heading id="A81E860EB629FF31AA584E90816B503C" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="82349D04E083289A86E7E4E20DB3DF52">
|
||
The species has been only known from
|
||
<collectingRegion id="44A9A4EF17F489B324C39A30933F2625" country="Vietnam" name="Khanh Hoa">Khanh Hoa</collectingRegion>
|
||
and
|
||
<collectingRegion id="62020231B084070407464CB77E6BDF8E" country="Vietnam" name="Kon Tum">Kon Tum Province</collectingRegion>
|
||
, southcentral
|
||
<collectingCountry id="60FB2A2F77CDA04F014C0088F027E93E" name="Vietnam">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EBKAI" type="DNA barcoding">
|
||
<paragraph id="929C51424D8580806C148122061687E1">
|
||
<heading id="7352C28E3BF4D9DF47D0F6F4D7E121FC" reason="title">DNA barcoding.</heading>
|
||
</paragraph>
|
||
<paragraph id="A9A7A0D2975CBEC53CA47E51BDD4BF0E">
|
||
Sequencing a fragment of the
|
||
<abbrev id="ABBRID0EHKAI" xlink_title="cytochrome c oxidase subunit I">COI</abbrev>
|
||
gene failed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |