364 lines
42 KiB
XML
364 lines
42 KiB
XML
<document id="96701774C77815D39A49D9FAC9083FFF" ID-DOI="10.11646/zootaxa.3941.1.10" ID-GBIF-Dataset="e56e730e-44e7-4efd-922e-9028c5ab01f8" ID-ISSN="1175-5326" ID-Zenodo-Dep="235956" ID-ZooBank="3AC1F816-A929-478D-85B3-23C26674FE30" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461024732864" checkinUser="plazi" docAuthor="Walther, Frank & Hausdorf, Bernhard" docDate="2015" docId="5F42879A2817FFDA1DC2B9AFFE34F908" docLanguage="en" docName="zt03941p148.pdf" docOrigin="Zootaxa 3941 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leiostyla beatae Walther & Hausdorf, 2015, n. sp." docType="treatment" docVersion="7" lastPageNumber="147" masterDocId="A37BFFE22817FFD91D55BF3AFFF0FFA3" masterDocTitle="Leiostyla beatae n. sp. from eastern Georgia (Gastropoda: Lauriidae)" masterLastPageNumber="148" masterPageNumber="144" pageNumber="144" updateTime="1698571579076" updateUser="plazi">
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<mods:mods id="C65522F5F53FA9B53F4C515922AE0CA8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="06E230DD087D127F87835281CB3315D3">
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<mods:title id="3739B183E5342288E497ECFF11F31F8C">Leiostyla beatae n. sp. from eastern Georgia (Gastropoda: Lauriidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="F66D618C63D0B1AA66246F9D55E44BD3" type="personal">
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<mods:namePart id="0CF13B91D5563329302547E075E076AF">Walther, Frank</mods:namePart>
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<mods:name id="AFBE57E87B7A76FFDF939259504BE85C" type="personal">
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<mods:roleTerm id="2A0EB21EFA23C76788430F9622DDB649">Author</mods:roleTerm>
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<mods:namePart id="392425E79949C765F8967B750AC570E5">Hausdorf, Bernhard</mods:namePart>
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<mods:typeOfResource id="8528FA4461671E9F29BFDD7D5872990E">text</mods:typeOfResource>
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<mods:title id="658ADD3150DFC83BF5CEE2F55951764A">Zootaxa</mods:title>
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<mods:part id="36C7A25A2480C268C8C5C4A7DF591F06">
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<mods:date id="0B2DC182FDE5B00E074B4446813D0492">2015</mods:date>
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<mods:number id="447A328C433A374488E381EFD809DF84">3941</mods:number>
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<mods:number id="C324136316F010C2486EABD6B0D3DC30">1</mods:number>
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<mods:start id="C14E34A5DE4391A7492B0D8B286555DB">144</mods:start>
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<mods:classification id="0FDF6E61CD5D57AB9C6565902D3AAB33">journal article</mods:classification>
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<mods:identifier id="CA583473876B508DC9B1ABBF1CB1D9BD" type="DOI">10.11646/zootaxa.3941.1.10</mods:identifier>
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<mods:identifier id="ADD341E1FB26940BDE42BECAE8E8A442" type="GBIF-Dataset">e56e730e-44e7-4efd-922e-9028c5ab01f8</mods:identifier>
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<mods:identifier id="54E6654302DCFA3C3E78FA9E31EA4112" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0D7AC458563423F8CBC7BD0B87D37A1F" type="Zenodo-Dep">235956</mods:identifier>
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<treatment id="5F42879A2817FFDA1DC2B9AFFE34F908" ID-DOI="http://doi.org/10.5281/zenodo.6095581" ID-GBIF-Taxon="119606016" ID-Zenodo-Dep="6095581" LSID="urn:lsid:plazi:treatment:5F42879A2817FFDA1DC2B9AFFE34F908" httpUri="http://treatment.plazi.org/id/5F42879A2817FFDA1DC2B9AFFE34F908" lastPageId="3" lastPageNumber="147" pageId="0" pageNumber="144">
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<subSubSection id="9FF165072817FFD91DC2B9AFFE52F913" box="[151,418,1685,1712]" pageId="0" pageNumber="144" type="nomenclature">
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<paragraph id="D754368C2817FFD91DC2B9AFFE52F913" blockId="0.[151,418,1685,1712]" box="[151,418,1685,1712]" pageId="0" pageNumber="144">
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<heading id="8C1C81E02817FFD91DC2B9AFFE52F913" bold="true" box="[151,418,1685,1712]" fontSize="11" level="1" pageId="0" pageNumber="144" reason="1">
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<emphasis id="E59FEA9E2817FFD91DC2B9AFFE52F913" bold="true" box="[151,418,1685,1712]" pageId="0" pageNumber="144">
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<taxonomicName id="10EB4D0F2817FFD91DC2B9AFFEA9F90C" ID-CoL="3T2MM" box="[151,345,1685,1711]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="0" pageNumber="144" phylum="Mollusca" rank="species" species="beatae" status="sp. nov.">
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<emphasis id="E59FEA9E2817FFD91DC2B9AFFEA9F90C" bold="true" box="[151,345,1685,1711]" italics="true" pageId="0" pageNumber="144">Leiostyla beatae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FEAC57E52817FFD91C35B9ACFE52F913" box="[352,418,1686,1712]" pageId="0" pageNumber="144" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9FF165072817FFDB1DC2B9E1FF2AFEDC" lastPageId="2" lastPageNumber="146" pageId="0" pageNumber="144" type="diagnosis">
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<paragraph id="D754368C2817FFD91DC2B9E1FE26F8B6" blockId="0.[151,1436,1754,1986]" pageId="0" pageNumber="144">
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<emphasis id="E59FEA9E2817FFD91DC2B9E1FEFCF951" bold="true" box="[151,268,1755,1778]" pageId="0" pageNumber="144">Diagnosis.</emphasis>
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A
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<taxonomicName id="10EB4D0F2817FFD91C78B9E0FE7FF951" box="[301,399,1754,1778]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="0" pageNumber="144" phylum="Mollusca" rank="genus">
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<emphasis id="E59FEA9E2817FFD91C78B9E0FE7FF951" box="[301,399,1754,1778]" italics="true" pageId="0" pageNumber="144">Leiostyla</emphasis>
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</taxonomicName>
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species characterized by a cylindrical-ovate, rib-striated shell with a columellar lamella forming a wide upwards bending arch.
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</paragraph>
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<paragraph id="D754368C2817FFDB1D92B81BFB13FF57" blockId="0.[151,1436,1754,1986]" lastBlockId="2.[151,1437,152,625]" lastPageId="2" lastPageNumber="146" pageId="0" pageNumber="144">
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<emphasis id="E59FEA9E2817FFD91D92B81BFE83F89B" bold="true" box="[199,371,1825,1848]" pageId="0" pageNumber="144">
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Shell (
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<figureCitation id="4FD02A092817FFD91C5AB81BFEA0F89B" box="[271,336,1825,1848]" captionStart="FIGURE 1" captionStartId="1.[151,255,1817,1840]" captionTargetBox="[252,1326,194,1774]" captionTargetId="figure@1.[252,1327,194,1775]" captionTargetPageId="1" captionText="FIGURE 1. Shells of Leiostyla species. A – C. Leiostyla beatae n. sp., Georgia: Lagodekhishevi valley 2 km upstream of Lagodekhi; A. holotype (ZMH 101605); B. paratype (NHMW); C. paratype (ZMH 101606); D. Leiostyla silicea (Schileyko, 1975), Georgia: Tekhuri valley 6.7 km towards Lebarde (ZMH 86618); E. Leiostyla paulinae (Lindholm, 1913), Georgia: Kintrishi valley, Chakhati (ZMH 81306); F. Leiostyla caucasica (L. Pfeiffer, 1857), Georgia: Pasanauri, lower valley of Shavi Aragvi river (ZMH 81326). Scale bar = 1.0 mm for all figures except oblique view into aperture; scale bare = 0.5 mm for oblique view into aperture." httpUri="https://zenodo.org/record/235957/files/figure.png" pageId="0" pageNumber="144">Fig. 1</figureCitation>
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A).
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</emphasis>
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Cylindrical-ovate; perforate; rib-striated (12–15 ribs per mm of the penultimate whorl), corneous; 6.75–7.25 slightly convex whorls separated by a slightly impressed suture; body whorl ascending towards the aperture; outside of the palatal wall with a distinct incision corresponding to the lower palatal fold; aperture cordate; peristome expanded and strengthened by a lip; angular lamella very high, without appendages; parietal lamella low; both are connected with the insertions of the peristome by a more or less thin callus; columellar lamella forming a wide upwards bending arch, inserting in a sharp angle and ending at the middle of the columellar edge; the suprapalatal fold is toothlike and forms a roundish sinulus with the angular lamella; upper palatal fold absent; lower palatal fold long and strong, below the middle of the palatal wall, fused anteriorly with the lip; basal fold long, also reaching the lip.
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</paragraph>
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<caption id="839466042816FFD81DC2B823FD4BF87E" httpUri="https://zenodo.org/record/235957/files/figure.png" pageId="1" pageNumber="145" targetBox="[252,1326,194,1774]" targetPageId="1">
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<paragraph id="D754368C2816FFD81DC2B823FD4BF87E" blockId="1.[151,1436,1816,2013]" pageId="1" pageNumber="145">
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<emphasis id="E59FEA9E2816FFD81DC2B823FEE9F893" bold="true" box="[151,281,1817,1840]" pageId="1" pageNumber="145">FIGURE 1.</emphasis>
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Shells of
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<taxonomicName id="10EB4D0F2816FFD81CD1B822FE16F893" box="[388,486,1816,1840]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="145" phylum="Mollusca" rank="genus">
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<emphasis id="E59FEA9E2816FFD81CD1B822FE16F893" box="[388,486,1816,1840]" italics="true" pageId="1" pageNumber="145">Leiostyla</emphasis>
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</taxonomicName>
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species.
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<emphasis id="E59FEA9E2816FFD81F13B823FD71F893" bold="true" box="[582,641,1817,1840]" pageId="1" pageNumber="145">A–C.</emphasis>
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<taxonomicName id="10EB4D0F2816FFD81FD2B822FCC6F893" box="[647,822,1816,1840]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="145" phylum="Mollusca" rank="species" species="beatae" status="sp. nov.">
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<emphasis id="E59FEA9E2816FFD81FD2B822FCC6F893" box="[647,822,1816,1840]" italics="true" pageId="1" pageNumber="145">Leiostyla beatae</emphasis>
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</taxonomicName>
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<emphasis id="E59FEA9E2816FFD81E69B823FC88F893" bold="true" box="[828,888,1817,1840]" pageId="1" pageNumber="145">
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<taxonomicNameLabel id="FEAC57E52816FFD81E69B823FC88F893" box="[828,888,1817,1840]" pageId="1" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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, Georgia: Lagodekhishevi valley 2 km upstream of Lagodekhi;
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<emphasis id="E59FEA9E2816FFD81C48B806FEC7F8F0" bold="true" box="[285,311,1852,1875]" pageId="1" pageNumber="145">A.</emphasis>
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holotype (ZMH 101605);
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<emphasis id="E59FEA9E2816FFD81F3EB806FD73F8F0" bold="true" box="[619,643,1852,1875]" pageId="1" pageNumber="145">B.</emphasis>
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paratype (NHMW);
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<emphasis id="E59FEA9E2816FFD81E20B806FC7FF8F0" bold="true" box="[885,911,1852,1875]" pageId="1" pageNumber="145">C.</emphasis>
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paratype (ZMH 101606);
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<emphasis id="E59FEA9E2816FFD81994B806FB2BF8F0" bold="true" box="[1217,1243,1852,1875]" pageId="1" pageNumber="145">D.</emphasis>
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<taxonomicName id="10EB4D0F2816FFD819BDB801FEABF8D6" authority="Schileyko, 1975" authorityName="Schileyko" authorityYear="1975" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="145" phylum="Mollusca" rank="species" species="silicea">
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<emphasis id="E59FEA9E2816FFD819BDB801FA6BF8F0" box="[1256,1435,1851,1875]" italics="true" pageId="1" pageNumber="145">Leiostyla silicea</emphasis>
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(Schileyko, 1975)
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</taxonomicName>
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, Georgia: Tekhuri valley 6.7 km towards Lebarde (ZMH 86618);
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<emphasis id="E59FEA9E2816FFD81965B864FBB9F8D6" bold="true" box="[1072,1097,1886,1909]" pageId="1" pageNumber="145">E.</emphasis>
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<taxonomicName id="10EB4D0F2816FFD81907B867FF27F83B" authority="Lindholm, 1913" authorityName="Lindholm" authorityYear="1913" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="145" phylum="Mollusca" rank="species" species="paulinae">
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<emphasis id="E59FEA9E2816FFD81907B867FAEAF8D6" box="[1106,1306,1885,1909]" italics="true" pageId="1" pageNumber="145">Leiostyla paulinae</emphasis>
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(Lindholm, 1913)
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</taxonomicName>
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, Georgia: Kintrishi valley, Chakhati (ZMH 81306);
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<emphasis id="E59FEA9E2816FFD81E51B8BBFCE6F83B" bold="true" box="[772,790,1921,1944]" pageId="1" pageNumber="145">F.</emphasis>
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<taxonomicName id="10EB4D0F2816FFD81E75B8BAFB4FF83B" authority="L. Pfeiffer, 1857" authorityName="L. Pfeiffer" authorityYear="1857" box="[800,1215,1920,1944]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="145" phylum="Mollusca" rank="species" species="caucasica">
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<emphasis id="E59FEA9E2816FFD81E75B8BAFC03F83B" box="[800,1011,1920,1944]" italics="true" pageId="1" pageNumber="145">Leiostyla caucasica</emphasis>
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(L. Pfeiffer, 1857)
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</taxonomicName>
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, Georgia: Pasanauri, lower valley of Shavi Aragvi river (ZMH 81326). Scale bar = 1.0 mm for all figures except oblique view into aperture; scale bare = 0.5 mm for oblique view into aperture.
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</paragraph>
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</caption>
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<paragraph id="D754368C2815FFDB1D92BE3AFF2AFEDC" blockId="2.[151,1437,152,625]" pageId="2" pageNumber="146">
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Measurements (n=27): shell diameter D:
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<quantity id="10139B692815FFDB1FD2BE3AFCD7FEB4" box="[647,807,256,279]" metricMagnitude="-3" metricUnit="m" metricValue="1.98" metricValueMax="2.16" metricValueMin="1.8" pageId="2" pageNumber="146" unit="mm" value="1.98" valueMax="2.16" valueMin="1.8">1.80–2.16 mm</quantity>
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, mean 1.94 +
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<quantity id="10139B692815FFDB1E90BE3AFBD9FEB4" box="[965,1065,256,279]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="146" unit="mm" value="0.09">0.09 mm</quantity>
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; shell diameter at third whorl D3:
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<quantity id="10139B692815FFDB1DC2BE18FEC7FE9A" box="[151,311,290,313]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" metricValueMax="1.46" metricValueMin="1.12" pageId="2" pageNumber="146" unit="mm" value="1.29" valueMax="1.46" valueMin="1.12">1.12–1.46 mm</quantity>
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, mean 1.26 +
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<quantity id="10139B692815FFDB1C8DBE18FDCDFE9A" box="[472,573,290,313]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="146" unit="mm" value="0.09">0.09 mm</quantity>
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; shell height H:
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<quantity id="10139B692815FFDB1FA2BE18FC66FE9A" box="[759,918,290,313]" metricMagnitude="-3" metricUnit="m" metricValue="3.66" metricValueMax="3.92" metricValueMin="3.4" pageId="2" pageNumber="146" unit="mm" value="3.66" valueMax="3.92" valueMin="3.4">3.40–3.92 mm</quantity>
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, mean 3.64 +
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<quantity id="10139B692815FFDB1914BE18FB57FE9A" box="[1089,1191,290,313]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="146" unit="mm" value="0.17">0.17 mm</quantity>
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; aperture diameter ad: 1.06–1.32, mean 1.19 +
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<quantity id="10139B692815FFDB1CCDBE7FFE0AFEFF" box="[408,506,325,348]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="2" pageNumber="146" unit="mm" value="0.07">0.07 mm</quantity>
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; aperture height ah: 1.14–1.44, mean 1.29 +
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<quantity id="10139B692815FFDB1E87BE7FFBC3FEFF" box="[978,1075,325,348]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="146" unit="mm" value="0.08">0.08 mm</quantity>
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; D/H: 0.484–0.818, mean 0.535 + 0.031.
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</paragraph>
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</subSubSection>
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<subSubSection id="9FF165072815FFDA1D92BEB0FE10FEFF" lastPageId="3" lastPageNumber="147" pageId="2" pageNumber="146" type="discussion">
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<paragraph id="D754368C2815FFDB1D92BEB0FBDBFDD2" blockId="2.[151,1437,152,625]" pageId="2" pageNumber="146">
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<emphasis id="E59FEA9E2815FFDB1D92BEB0FEC5FE02" bold="true" box="[199,309,394,417]" pageId="2" pageNumber="146">Remarks.</emphasis>
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<taxonomicName id="10EB4D0F2815FFDB1C68BEB3FE1EFE02" box="[317,494,393,417]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="146" phylum="Mollusca" rank="species" species="beatae">
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<emphasis id="E59FEA9E2815FFDB1C68BEB3FE1EFE02" box="[317,494,393,417]" italics="true" pageId="2" pageNumber="146">Leiostyla beatae</emphasis>
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</taxonomicName>
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has been identified with
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<taxonomicName id="10EB4D0F2815FFDB1E57BEB3FB8AFE02" authority="Schileyko, 1975" authorityName="Schileyko" authorityYear="1975" box="[770,1146,393,417]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="146" phylum="Mollusca" rank="species" species="silicea">
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<emphasis id="E59FEA9E2815FFDB1E57BEB3FC41FE02" box="[770,945,393,417]" italics="true" pageId="2" pageNumber="146">Leiostyla silicea</emphasis>
|
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(
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<bibRefCitation id="B37A4B7D2815FFDB1E94BEB0FB81FE02" author="Schileyko" box="[961,1137,394,417]" pageId="2" pageNumber="146" refString="Schileyko, A. A. (1975) Mollyuski podsemeystva Lauriinae fauny SSSR (Pulmonata, Pupillidae). Zoologicheskiy Zhurnal, 54, 1767 - 1782." type="journal article" year="1975">Schileyko, 1975</bibRefCitation>
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)
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</taxonomicName>
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(
|
||
<figureCitation id="4FD02A092815FFDB19DCBEB0FB37FE02" box="[1161,1223,394,417]" captionStart="FIGURE 1" captionStartId="1.[151,255,1817,1840]" captionTargetBox="[252,1326,194,1774]" captionTargetId="figure@1.[252,1327,194,1775]" captionTargetPageId="1" captionText="FIGURE 1. Shells of Leiostyla species. A – C. Leiostyla beatae n. sp., Georgia: Lagodekhishevi valley 2 km upstream of Lagodekhi; A. holotype (ZMH 101605); B. paratype (NHMW); C. paratype (ZMH 101606); D. Leiostyla silicea (Schileyko, 1975), Georgia: Tekhuri valley 6.7 km towards Lebarde (ZMH 86618); E. Leiostyla paulinae (Lindholm, 1913), Georgia: Kintrishi valley, Chakhati (ZMH 81306); F. Leiostyla caucasica (L. Pfeiffer, 1857), Georgia: Pasanauri, lower valley of Shavi Aragvi river (ZMH 81326). Scale bar = 1.0 mm for all figures except oblique view into aperture; scale bare = 0.5 mm for oblique view into aperture." httpUri="https://zenodo.org/record/235957/files/figure.png" pageId="2" pageNumber="146">Fig. 1</figureCitation>
|
||
B) by
|
||
<bibRefCitation id="B37A4B7D2815FFDB1845BEB0FEF7FE67" pageId="2" pageNumber="146" refString="Pokryszko, B. M., Cameron, R. A. D., Mumladze, L. & Tarkhnishvili, D. (2011) Forest snail faunas from Georgian Transcaucasia: patterns of diversity in a Pleistocene refugium. Biological Journal of the Linnean Society, 102, 239 - 250. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.2010.01575. x" type="journal article">
|
||
Pokryszko
|
||
<emphasis id="E59FEA9E2815FFDB18DCBEB0FF43FE67" italics="true" pageId="2" pageNumber="146">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
. It differs from that species in the more cylindrical, lighter shell, the columellar lamella that is bending upwards and not running horizontally, and the lack of an upper palatal fold. Pilsbry (
|
||
<date id="A355104C2815FFDB197ABEEAFB59FE44" box="[1071,1193,464,487]" pageId="2" pageNumber="146" value="1922" valueMax="1923">1922–1923</date>
|
||
) classified the species with a steeply ascending columellar lamella, namely
|
||
<taxonomicName id="10EB4D0F2815FFDB1F8DBECBFB98FDAA" authority="Lindholm, 1913" authorityName="Lindholm" authorityYear="1913" box="[728,1128,497,521]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="146" phylum="Mollusca" rank="species" species="paulinae">
|
||
<emphasis id="E59FEA9E2815FFDB1F8DBECBFC6FFDAA" box="[728,927,497,521]" italics="true" pageId="2" pageNumber="146">Leiostyla paulinae</emphasis>
|
||
(
|
||
<bibRefCitation id="B37A4B7D2815FFDB1EE5BEC8FB90FDAA" author="Lindholm" box="[944,1120,498,521]" pageId="2" pageNumber="146" refString="Lindholm, W. A. (1913) Beschreibung neuer Arten und Formen aus dem Kaukasus-Gebiete. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 45, 17 - 23, 62 - 69." type="journal article" year="1913">Lindholm, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="10EB4D0F2815FFDB19CABECBFEC6FD8F" authority="L. Pfeiffer, 1857" authorityName="L. Pfeiffer" authorityYear="1857" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="146" phylum="Mollusca" rank="species" species="caucasica">
|
||
<emphasis id="E59FEA9E2815FFDB19CABECBFA84FDAA" box="[1183,1396,497,521]" italics="true" pageId="2" pageNumber="146">Leiostyla caucasica</emphasis>
|
||
(
|
||
<bibRefCitation id="B37A4B7D2815FFDB18D1BEC8FEDEFD8F" author="Pfeiffer" pageId="2" pageNumber="146" refString="Pfeiffer, L. (1857) Neue Landschnecken. Malakozoologische Blatter, 4, 85 - 89." type="journal article" year="1857">L. Pfeiffer, 1857</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, in a separate section. The shape of the columellar lamella of
|
||
<taxonomicName id="10EB4D0F2815FFDB1E8EBD2FFBB0FD8F" box="[987,1088,532,556]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="146" phylum="Mollusca" rank="species" species="beatae">
|
||
<emphasis id="E59FEA9E2815FFDB1E8EBD2FFBB0FD8F" box="[987,1088,532,556]" italics="true" pageId="2" pageNumber="146">L. beatae</emphasis>
|
||
</taxonomicName>
|
||
resembles that of these species. However,
|
||
<taxonomicName id="10EB4D0F2815FFDB1C50BD02FE9AFDEC" box="[261,362,567,591]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="146" phylum="Mollusca" rank="species" species="beatae">
|
||
<emphasis id="E59FEA9E2815FFDB1C50BD02FE9AFDEC" box="[261,362,567,591]" italics="true" pageId="2" pageNumber="146">L. beatae</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="10EB4D0F2815FFDB1CACBD02FD83FDEC" box="[505,627,567,591]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="146" phylum="Mollusca" rank="species" species="paulinae">
|
||
<emphasis id="E59FEA9E2815FFDB1CACBD02FD83FDEC" box="[505,627,567,591]" italics="true" pageId="2" pageNumber="146">L. paulinae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="4FD02A092815FFDB1FD7BD02FD4FFDEC" box="[642,703,568,591]" captionStart="FIGURE 1" captionStartId="1.[151,255,1817,1840]" captionTargetBox="[252,1326,194,1774]" captionTargetId="figure@1.[252,1327,194,1775]" captionTargetPageId="1" captionText="FIGURE 1. Shells of Leiostyla species. A – C. Leiostyla beatae n. sp., Georgia: Lagodekhishevi valley 2 km upstream of Lagodekhi; A. holotype (ZMH 101605); B. paratype (NHMW); C. paratype (ZMH 101606); D. Leiostyla silicea (Schileyko, 1975), Georgia: Tekhuri valley 6.7 km towards Lebarde (ZMH 86618); E. Leiostyla paulinae (Lindholm, 1913), Georgia: Kintrishi valley, Chakhati (ZMH 81306); F. Leiostyla caucasica (L. Pfeiffer, 1857), Georgia: Pasanauri, lower valley of Shavi Aragvi river (ZMH 81326). Scale bar = 1.0 mm for all figures except oblique view into aperture; scale bare = 0.5 mm for oblique view into aperture." httpUri="https://zenodo.org/record/235957/files/figure.png" pageId="2" pageNumber="146">Fig. 1</figureCitation>
|
||
C) in the cylindrical, more strongly rib-striated shell and the lack of an upper palatal fold and from
|
||
<taxonomicName id="10EB4D0F2815FFDB1CB4BD60FD97FDD2" box="[481,615,602,625]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="146" phylum="Mollusca" rank="species" species="caucasica">
|
||
<emphasis id="E59FEA9E2815FFDB1CB4BD60FD97FDD2" box="[481,615,602,625]" italics="true" pageId="2" pageNumber="146">L. caucasica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="4FD02A092815FFDB1F23BD60FD42FDD2" box="[630,690,602,625]" captionStart="FIGURE 1" captionStartId="1.[151,255,1817,1840]" captionTargetBox="[252,1326,194,1774]" captionTargetId="figure@1.[252,1327,194,1775]" captionTargetPageId="1" captionText="FIGURE 1. Shells of Leiostyla species. A – C. Leiostyla beatae n. sp., Georgia: Lagodekhishevi valley 2 km upstream of Lagodekhi; A. holotype (ZMH 101605); B. paratype (NHMW); C. paratype (ZMH 101606); D. Leiostyla silicea (Schileyko, 1975), Georgia: Tekhuri valley 6.7 km towards Lebarde (ZMH 86618); E. Leiostyla paulinae (Lindholm, 1913), Georgia: Kintrishi valley, Chakhati (ZMH 81306); F. Leiostyla caucasica (L. Pfeiffer, 1857), Georgia: Pasanauri, lower valley of Shavi Aragvi river (ZMH 81326). Scale bar = 1.0 mm for all figures except oblique view into aperture; scale bare = 0.5 mm for oblique view into aperture." httpUri="https://zenodo.org/record/235957/files/figure.png" pageId="2" pageNumber="146">Fig. 1</figureCitation>
|
||
D) in the smaller, rib-striated shell.
|
||
</paragraph>
|
||
<caption id="839466042815FFDB1DC2B8AAFD3AF869" httpUri="https://zenodo.org/record/235958/files/figure.png" pageId="2" pageNumber="146" targetBox="[181,1416,688,1892]" targetPageId="2">
|
||
<paragraph id="D754368C2815FFDB1DC2B8AAFD3AF869" blockId="2.[151,1436,1935,1994]" pageId="2" pageNumber="146">
|
||
<emphasis id="E59FEA9E2815FFDB1DC2B8AAFEEBF804" bold="true" box="[151,283,1936,1959]" pageId="2" pageNumber="146">FIGURE 2.</emphasis>
|
||
Separation of
|
||
<taxonomicName id="10EB4D0F2815FFDB1C95B8B5FD82F804" box="[448,626,1935,1959]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="146" phylum="Mollusca" rank="species" species="beatae" status="sp. nov.">
|
||
<emphasis id="E59FEA9E2815FFDB1C95B8B5FD82F804" box="[448,626,1935,1959]" italics="true" pageId="2" pageNumber="146">Leiostyla beatae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E59FEA9E2815FFDB1F2EB8AAFD4AF804" bold="true" box="[635,698,1936,1959]" pageId="2" pageNumber="146">
|
||
<taxonomicNameLabel id="FEAC57E52815FFDB1F2EB8AAFD4AF804" box="[635,698,1936,1959]" pageId="2" pageNumber="146" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="10EB4D0F2815FFDB1F9FB8B5FC8AF804" box="[714,890,1935,1959]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="146" phylum="Mollusca" rank="species" species="silicea">
|
||
<emphasis id="E59FEA9E2815FFDB1F9FB8B5FC8AF804" box="[714,890,1935,1959]" italics="true" pageId="2" pageNumber="146">Leiostyla silicea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="10EB4D0F2815FFDB1EE1B8B5FB8CF804" box="[948,1148,1935,1959]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="146" phylum="Mollusca" rank="species" species="paulinae">
|
||
<emphasis id="E59FEA9E2815FFDB1EE1B8B5FB8CF804" box="[948,1148,1935,1959]" italics="true" pageId="2" pageNumber="146">Leiostyla paulinae</emphasis>
|
||
</taxonomicName>
|
||
according to the first two discriminant functions based on shell measurements.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D754368C2814FFDA1D92BFA2FE10FEFF" blockId="3.[151,1436,151,799]" pageId="3" pageNumber="147">
|
||
A discriminant analysis based on the shell measurements (D, D3, H, ad, ah) of
|
||
<taxonomicName id="10EB4D0F2814FFDA194CBFA2FB8DFF0C" box="[1049,1149,151,175]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="147" phylum="Mollusca" rank="species" species="beatae">
|
||
<emphasis id="E59FEA9E2814FFDA194CBFA2FB8DFF0C" box="[1049,1149,151,175]" italics="true" pageId="3" pageNumber="147">L. beatae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="10EB4D0F2814FFDA19DEBFA2FB1CFF0C" box="[1163,1260,151,175]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="147" phylum="Mollusca" rank="species" species="silicea">
|
||
<emphasis id="E59FEA9E2814FFDA19DEBFA2FB1CFF0C" box="[1163,1260,151,175]" italics="true" pageId="3" pageNumber="147">L. silicea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="10EB4D0F2814FFDA1877BFA2FA6CFF0C" box="[1314,1436,151,175]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="147" phylum="Mollusca" rank="species" species="paulinae">
|
||
<emphasis id="E59FEA9E2814FFDA1877BFA2FA6CFF0C" box="[1314,1436,151,175]" italics="true" pageId="3" pageNumber="147">L. paulinae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="4FD02A092814FFDA1DCABF80FF13FF72" box="[159,227,186,209]" captionStart="FIGURE 2" captionStartId="2.[151,255,1936,1959]" captionTargetBox="[181,1416,688,1892]" captionTargetId="figure@2.[180,1416,687,1892]" captionTargetPageId="2" captionText="FIGURE 2. Separation of Leiostyla beatae n. sp., Leiostyla silicea and Leiostyla paulinae according to the first two discriminant functions based on shell measurements." httpUri="https://zenodo.org/record/235958/files/figure.png" pageId="3" pageNumber="147">Fig. 2</figureCitation>
|
||
) indicated that the null hypothesis that the means of all of these variables are equal across the species can significantly be rejected (Wilk's lambda for the first two discriminant functions = 4.4%;
|
||
<emphasis id="E59FEA9E2814FFDA19D6BFE7FB60FF57" box="[1155,1168,221,244]" italics="true" pageId="3" pageNumber="147">p</emphasis>
|
||
<0.001). Based on shell measurements alone, 96.9% of the individuals were correctly classified using the first two discriminant functions (93.8% after cross validation). Note, however, that we consider also the shell sculpture and the apertural armature as essential for the determination of
|
||
<taxonomicName id="10EB4D0F2814FFDA1C20BE7FFE28FEFF" box="[373,472,324,348]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="147" phylum="Mollusca" rank="species" species="beatae">
|
||
<emphasis id="E59FEA9E2814FFDA1C20BE7FFE28FEFF" box="[373,472,324,348]" italics="true" pageId="3" pageNumber="147">L. beatae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9FF165072814FFDA1D92BE52FE7BFD8F" pageId="3" pageNumber="147" type="materials_examined">
|
||
<paragraph id="D754368C2814FFDA1D92BE52FD95FE67" blockId="3.[151,1436,151,799]" pageId="3" pageNumber="147">
|
||
<emphasis id="E59FEA9E2814FFDA1D92BE52FE9FFEDC" bold="true" box="[199,367,360,383]" pageId="3" pageNumber="147">
|
||
<typeStatus id="0850882E2814FFDA1D92BE52FF0FFEDC" box="[199,255,360,383]" pageId="3" pageNumber="147">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<typeStatus id="0850882E2814FFDA1C2DBE52FE10FEDC" box="[376,480,360,383]" pageId="3" pageNumber="147" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<collectingCountry id="AFFC761C2814FFDA1CB9BE52FDB6FEDC" box="[492,582,360,383]" name="Georgia" pageId="3" pageNumber="147">Georgia</collectingCountry>
|
||
: Lagodekhishevi valley
|
||
<quantity id="10139B692814FFDA1E05BE52FC79FEDC" box="[848,905,360,383]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="147" unit="km" value="2.0">2 km</quantity>
|
||
upstream of Lagodekhi (
|
||
<collectionCode id="B1FAAE492814FFDA19CABE52FB2CFEDC" box="[1183,1244,360,383]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="3" pageNumber="147">ZMH</collectionCode>
|
||
<date id="A355104C2814FFDA19B0BE52FAC7FEDC" box="[1253,1335,360,383]" pageId="3" pageNumber="147" value="05-10-16">101605</date>
|
||
, D =
|
||
<quantity id="10139B692814FFDA182FBE52FF33FE02" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="147" unit="mm" value="1.9">1.9 mm</quantity>
|
||
, H =
|
||
<quantity id="10139B692814FFDA1DABBEB0FEA3FE02" box="[254,339,394,417]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="147" unit="mm" value="3.8">3.8 mm</quantity>
|
||
, ad =
|
||
<quantity id="10139B692814FFDA1CC1BEB0FE19FE02" box="[404,489,394,417]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="147" unit="mm" value="1.3">1.3 mm</quantity>
|
||
; ah =
|
||
<quantity id="10139B692814FFDA1F7FBEB0FD71FE02" box="[554,641,394,417]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="147" unit="mm" value="1.4">1.4 mm</quantity>
|
||
).
|
||
<typeStatus id="0850882E2814FFDA1FC6BEB0FD0FFE02" box="[659,767,394,417]" pageId="3" pageNumber="147" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<collectingCountry id="AFFC761C2814FFDA1E5CBEB0FC92FE02" box="[777,866,394,417]" name="Georgia" pageId="3" pageNumber="147">Georgia</collectingCountry>
|
||
: Lagodekhishevi valley
|
||
<quantity id="10139B692814FFDA1933BEB0FB6CFE02" box="[1126,1180,394,417]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="147" unit="km" value="2.0">2 km</quantity>
|
||
upstream of Lagodekhi (FW 7137/28;
|
||
<collectionCode id="B1FAAE492814FFDA1C61BE97FE7FFE67" box="[308,399,429,452]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="147">NHMW</collectionCode>
|
||
/4;
|
||
<collectionCode id="B1FAAE492814FFDA1CFBBE97FE1AFE67" box="[430,490,429,452]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="3" pageNumber="147">ZMH</collectionCode>
|
||
<date id="A355104C2814FFDA1CA5BE97FDB5FE67" box="[496,581,429,452]" pageId="3" pageNumber="147" value="06-10-16">101606</date>
|
||
/3).
|
||
</paragraph>
|
||
<paragraph id="D754368C2814FFDA1D92BEEAFE7BFD8F" blockId="3.[151,1436,151,799]" pageId="3" pageNumber="147">
|
||
<emphasis id="E59FEA9E2814FFDA1D92BEEAFEAEFE44" bold="true" box="[199,350,464,487]" pageId="3" pageNumber="147">
|
||
<typeStatus id="0850882E2814FFDA1D92BEEAFF0FFE44" box="[199,255,464,487]" pageId="3" pageNumber="147">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="AFFC761C2814FFDA1C30BEEAFE4FFE44" box="[357,447,464,487]" name="Georgia" pageId="3" pageNumber="147">Georgia</collectingCountry>
|
||
: Lagodekhishevi valley
|
||
<quantity id="10139B692814FFDA1F96BEEAFD09FE44" box="[707,761,464,487]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="147" unit="km" value="2.0">2 km</quantity>
|
||
upstream of Lagodekhi, dry scree on a steep slope with mixed deciduous forest (
|
||
<taxonomicName id="10EB4D0F2814FFDA1C00BEC8FE48FDAA" box="[341,440,498,521]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="147" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="E59FEA9E2814FFDA1C00BEC8FE48FDAA" box="[341,440,498,521]" italics="true" pageId="3" pageNumber="147">Carpinus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="10EB4D0F2814FFDA1C93BEC8FDD6FDAA" box="[454,550,498,521]" class="Magnoliopsida" family="Oleaceae" genus="Fraxinus" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="147" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="E59FEA9E2814FFDA1C93BEC8FDD6FDAA" box="[454,550,498,521]" italics="true" pageId="3" pageNumber="147">Fraxinus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="10EB4D0F2814FFDA1F61BEC8FD87FDAA" box="[564,631,498,521]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="147" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="E59FEA9E2814FFDA1F61BEC8FD87FDAA" box="[564,631,498,521]" italics="true" pageId="3" pageNumber="147">Fagus</emphasis>
|
||
</taxonomicName>
|
||
) and a dense undergrowth of
|
||
<taxonomicName id="10EB4D0F2814FFDA1EE1BECBFC07FDAA" box="[948,1015,497,521]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="147" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="E59FEA9E2814FFDA1EE1BECBFC07FDAA" box="[948,1015,497,521]" italics="true" pageId="3" pageNumber="147">Rubus</emphasis>
|
||
</taxonomicName>
|
||
and some
|
||
<taxonomicName id="10EB4D0F2814FFDA193CBECBFB4DFDAA" box="[1129,1213,497,521]" class="Magnoliopsida" family="Betulaceae" genus="Corylus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="147" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="E59FEA9E2814FFDA193CBECBFB4DFDAA" box="[1129,1213,497,521]" italics="true" pageId="3" pageNumber="147">Corylus</emphasis>
|
||
</taxonomicName>
|
||
bushes,
|
||
<quantity id="10139B692814FFDA184FBEC8FAADFDAA" box="[1306,1373,498,521]" metricMagnitude="2" metricUnit="m" metricValue="7.05" pageId="3" pageNumber="147" unit="m" value="705.0">705 m</quantity>
|
||
a.s.l.,
|
||
<geoCoordinate id="B2DF504B2814FFDA1DC2BD2FFEFBFD8F" box="[151,267,533,556]" direction="north" orientation="latitude" pageId="3" pageNumber="147" precision="5" value="41.8563">41.8563°N</geoCoordinate>
|
||
<geoCoordinate id="B2DF504B2814FFDA1C47BD2FFE77FD8F" box="[274,391,533,556]" direction="east" orientation="longitude" pageId="3" pageNumber="147" precision="5" value="46.2996">46.2996°E</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9FF165072814FFDA1D92BD02FBC5FD7A" pageId="3" pageNumber="147" type="distribution">
|
||
<paragraph id="D754368C2814FFDA1D92BD02FBC5FD7A" blockId="3.[151,1436,151,799]" pageId="3" pageNumber="147">
|
||
<emphasis id="E59FEA9E2814FFDA1D92BD02FE47FDEC" bold="true" box="[199,439,568,591]" pageId="3" pageNumber="147">Distribution (Fig. 3).</emphasis>
|
||
<taxonomicName id="10EB4D0F2814FFDA1C95BD0DFD82FDEC" box="[448,626,567,591]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="147" phylum="Mollusca" rank="species" species="beatae">
|
||
<emphasis id="E59FEA9E2814FFDA1C95BD0DFD82FDEC" box="[448,626,567,591]" italics="true" pageId="3" pageNumber="147">Leiostyla beatae</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="0850882E2814FFDA1EDCBD02FC47FDEC" box="[905,951,568,591]" pageId="3" pageNumber="147">type</typeStatus>
|
||
locality. Despite the land snail fauna of the Lagodekhi Nature Reserve is relatively well known (
|
||
<bibRefCitation id="B37A4B7D2814FFDA1F82BD60FB40FDD2" author="Lezhawa" box="[727,1200,602,625]" pageId="3" pageNumber="147" refString="Lezhawa, G. I. (1983) Nazemnye mollyuski Lagodekhskogo zapovednika. In: Fauna i ekologiya bespozvonochnykh zhivotnykh Gruzii. Tbilisi, Metsniereba, pp. 147 - 151." type="book chapter" year="1983">Lezhawa 1983; L. Mumladze, pers. comm.</bibRefCitation>
|
||
), the new species has been detected only recently (
|
||
<bibRefCitation id="B37A4B7D2814FFDA1C92BD47FD42FD37" author="Pokryszko" box="[455,690,636,660]" pageId="3" pageNumber="147" refString="Pokryszko, B. M., Cameron, R. A. D., Mumladze, L. & Tarkhnishvili, D. (2011) Forest snail faunas from Georgian Transcaucasia: patterns of diversity in a Pleistocene refugium. Biological Journal of the Linnean Society, 102, 239 - 250. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.2010.01575. x" type="journal article" year="2011">
|
||
Pokryszko
|
||
<emphasis id="E59FEA9E2814FFDA1F15BD47FD86FD37" box="[576,630,636,660]" italics="true" pageId="3" pageNumber="147">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
) because of its special microhabitat, which is difficult to access due to steepness and thorny vegetation. Only empty shells were found in the leaf litter. A systematic survey of suitable habitats in eastern
|
||
<collectingCountry id="AFFC761C2814FFDA1C51BDF8FEA9FD7A" box="[260,345,706,729]" name="Georgia" pageId="3" pageNumber="147">Georgia</collectingCountry>
|
||
and western
|
||
<collectingCountry id="AFFC761C2814FFDA1CB2BDF8FDACFD7A" box="[487,604,706,729]" name="Azerbaijan" pageId="3" pageNumber="147">Azerbaijan</collectingCountry>
|
||
will probably reveal additional populations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9FF165072814FFDA1D92BDDFFE34F908" pageId="3" pageNumber="147" type="etymology">
|
||
<paragraph id="D754368C2814FFDA1D92BDDFFEFDFCBC" blockId="3.[151,1436,151,799]" pageId="3" pageNumber="147">
|
||
<emphasis id="E59FEA9E2814FFDA1D92BDDFFEB7FD5F" bold="true" box="[199,327,741,764]" pageId="3" pageNumber="147">Etymology.</emphasis>
|
||
The species is named in honour of Prof. Dr. Beata Pokryszko, Wrocław, who discovered the first specimens.
|
||
</paragraph>
|
||
<paragraph id="D754368C2814FFDA1DC2B94AFE34F908" blockId="3.[151,1436,1647,1708]" pageId="3" pageNumber="147">
|
||
<emphasis id="E59FEA9E2814FFDA1DC2B94AFEEAF924" bold="true" box="[151,282,1648,1671]" pageId="3" pageNumber="147">FIGURE 3.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="10EB4D0F2814FFDA1C9DB955FD88F924" box="[456,632,1647,1671]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="147" phylum="Mollusca" rank="species" species="beatae" status="sp. nov.">
|
||
<emphasis id="E59FEA9E2814FFDA1C9DB955FD88F924" box="[456,632,1647,1671]" italics="true" pageId="3" pageNumber="147">Leiostyla beatae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E59FEA9E2814FFDA1F2AB94AFD4CF924" bold="true" box="[639,700,1648,1671]" pageId="3" pageNumber="147">
|
||
<taxonomicNameLabel id="FEAC57E52814FFDA1F2AB94AFD4CF924" box="[639,700,1648,1671]" pageId="3" pageNumber="147" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="10EB4D0F2814FFDA1F9EB955FC6EF924" box="[715,926,1647,1671]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="147" phylum="Mollusca" rank="species" species="caucasica">
|
||
<emphasis id="E59FEA9E2814FFDA1F9EB955FC6EF924" box="[715,926,1647,1671]" italics="true" pageId="3" pageNumber="147">Leiostyla caucasica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="10EB4D0F2814FFDA1EF9B955FB83F924" box="[940,1139,1647,1671]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="147" phylum="Mollusca" rank="species" species="paulinae">
|
||
<emphasis id="E59FEA9E2814FFDA1EF9B955FB83F924" box="[940,1139,1647,1671]" italics="true" pageId="3" pageNumber="147">Leiostyla paulinae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="10EB4D0F2814FFDA19FCB955FAA8F924" box="[1193,1368,1647,1671]" class="Gastropoda" family="Lauriidae" genus="Leiostyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="147" phylum="Mollusca" rank="species" species="silicea">
|
||
<emphasis id="E59FEA9E2814FFDA19FCB955FAA8F924" box="[1193,1368,1647,1671]" italics="true" pageId="3" pageNumber="147">Leiostyla silicea</emphasis>
|
||
</taxonomicName>
|
||
in the Caucasus region (0.5° grid).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |