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<document id="C382F047258E8D180AE215147498A934" ID-CLB-Dataset="3376" ID-DOI="10.5852/ejt.2021.767.1481" ID-GBIF-Dataset="9b3e85f1-1856-49bc-8cde-b2d197897698" ID-ISSN="2118-9773" ID-Zenodo-Dep="5715526" ID-ZooBank="32E080F8-A27A-4CBB-BA48-044DF82335A9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637362086142" checkinUser="admin" docAuthor="Liang, Zulong, Angus, Robert B. &amp; Jia, Fenglong" docDate="2021" docId="56557D6EFFEFFFF0557ACCC73391F9C1" docLanguage="en" docName="ejt-767_1-39.pdf.imf" docOrigin="European Journal of Taxonomy 767" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Patrus jiangxiensis Liang &amp; Angus &amp; Jia 2021, sp. nov." docType="treatment" docUuid="74887BE8-29AD-43F6-8E93-84770844D59C" docUuidSource="ZooBank" docVersion="7" lastPageNumber="10" masterDocId="AA6C0516FFE8FFF95709CD48375DFFC8" masterDocTitle="Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="8" updateTime="1699469142537" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FA45EB67F5D970883F888FAC7B2E50D7">Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae)</mods:title>
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<mods:name id="2FED1376F9B583474880F6A5C6713E66" type="personal">
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<mods:namePart id="AF910FDF0F7C754B9A850D5EC4880643">Liang, Zulong</mods:namePart>
<mods:nameIdentifier id="A0EF21750E2C72F379DF04FC05BBDFF8" type="LSID">75479591-435D-45B4-9821-234D76B89043</mods:nameIdentifier>
<mods:affiliation id="DDD4120EB353C4977949629B1EC81281">Institute of Entomology, Life Science School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China. Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK.</mods:affiliation>
<mods:nameIdentifier id="55F964E3E4C9C0B3CC071A77F3672831" type="email">liangzl3@mail2.sysu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="C66B5F3B83B6A993BC4703C6719BDB83">Angus, Robert B.</mods:namePart>
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<mods:namePart id="2921DE6C9BCDBF37D60DDE4D2A0560AE">Jia, Fenglong</mods:namePart>
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<mods:title id="3AEF975368F9AECA1260ADD1220B5098">European Journal of Taxonomy</mods:title>
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<mods:part id="2EEC080EE103BB06C799924AEFFD1851">
<mods:date id="86DC4E2EC3E24F7E6C1F739D195C7BB5">2021</mods:date>
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<mods:number id="271607ED18B293406A73D46EAFC7F949">2021-09-02</mods:number>
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<treatment id="56557D6EFFEFFFF0557ACCC73391F9C1" ID-DOI="http://doi.org/10.5281/zenodo.5715635" ID-GBIF-Taxon="190335231" ID-Zenodo-Dep="5715635" LSID="urn:lsid:zoobank.org:act:74887BE8-29AD-43F6-8E93-84770844D59C" httpUri="http://treatment.plazi.org/id/56557D6EFFEFFFF0557ACCC73391F9C1" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
<subSubSection id="96E69FF3FFEFFFFE557ACCC733E0FE05" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="DE43CC78FFEFFFFE557ACCC7349DFE62" blockId="7.[374,1213,399,496]" box="[627,960,399,426]" pageId="7" pageNumber="8">
<heading id="850B7B14FFEFFFFE557ACCC7349DFE62" box="[627,960,399,426]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<taxonomicName id="19FCB7FBFFEFFFFE557ACCC73402FE61" authority="Liang &amp; Angus &amp; Jia, 2021" authorityName="Liang &amp; Angus &amp; Jia" authorityYear="2021" box="[627,863,399,425]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="jiangxiensis" status="sp. nov.">
<emphasis id="EC88106AFFEFFFFE557ACCC73402FE61" bold="true" box="[627,863,399,425]" italics="true" pageId="7" pageNumber="8">Patrus jiangxiensis</emphasis>
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<taxonomicNameLabel id="F7BBAD11FFEFFFFE546FCCD8349DFE62" box="[870,960,400,426]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="96E69FF3FFEFFFFE55BCCC9D3423FE27" box="[693,894,469,496]" pageId="7" pageNumber="8" type="description">
<paragraph id="DE43CC78FFEFFFFE55BCCC9D3423FE27" blockId="7.[374,1213,399,496]" box="[693,894,469,496]" pageId="7" pageNumber="8">
<figureCitation id="46C7D0FDFFEFFFFE55BCCC9D346FFE38" box="[693,818,469,496]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[189,1398,359,1889]" captionTargetId="figure-21@4.[189,1398,359,1889]" captionTargetPageId="4" captionText="Fig. 1. Habitus. AC. Patrus hainanensis sp. nov., ♂, paratype (SYSU). DF. Patrus jiangxiensis. sp. nov., ♂, paratypes (SYSU). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5715528" httpUri="https://zenodo.org/record/5715528/files/figure.png" pageId="7" pageNumber="8">Figs 1DF</figureCitation>
,
<figureCitation id="46C7D0FDFFEFFFFE5437CC9E3412FE38" box="[830,847,470,496]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="7" pageNumber="8">4</figureCitation>
,
<figureCitation id="46C7D0FDFFEFFFFE5455CC9D3423FE27" box="[860,894,469,495]" captionStart="Fig" captionStartId="11.[189,232,1857,1883]" captionTargetBox="[204,1379,1111,1818]" captionTargetId="figure-216@11.[189,1398,1111,1818]" captionTargetPageId="11" captionText="Fig. 6. Female reproductive tract of new species, paratype with the same collection data as for holotype (SYSU). A. Patrus hainanensis sp. nov. B. Patrus jiangxiensis sp. nov. C. Patrus shangchuanensis sp. nov.Abbreviations: AG = accessory gland; BC = bursa copulatrix; BS = bursa sclerite; CO = common oviduct; FD = fertilization duct; GC = gonocoxa; LT = laterogtergite; SP = spermatheca; VA = vagina. Scale bars = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.5715542" httpUri="https://zenodo.org/record/5715542/files/figure.png" pageId="7" pageNumber="8">6B</figureCitation>
</paragraph>
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<subSubSection id="96E69FF3FFEFFFF157B4CF5F35D1FCCE" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="DE43CC78FFEFFFFE57B4CF5F366AFDF9" blockId="7.[189,311,535,561]" box="[189,311,535,561]" pageId="7" pageNumber="8">
<heading id="850B7B14FFEFFFFE57B4CF5F366AFDF9" bold="true" box="[189,311,535,561]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="EC88106AFFEFFFFE57B4CF5F366AFDF9" bold="true" box="[189,311,535,561]" pageId="7" pageNumber="8">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DE43CC78FFEFFFF157B4CF0D33D4FEA1" blockId="7.[188,1399,580,784]" lastBlockId="8.[188,1399,265,361]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
This species can be diagnosed by the following characters combined: 1) labrum less than ⅓ as long as wide, with anterior margin straight; 2) pronotal setose region anteriorly expanded inward onto pronotal disc to external third of dorsal eye; 3) dorsal surface with strong and dense micro-punctation, much stronger on elytron (
<figureCitation id="46C7D0FDFFEFFFFE56A3CFE73574FD01" box="[426,553,687,713]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="7" pageNumber="8">Fig. 4EG</figureCitation>
); 4) pronoto-elytral glabrous region regular oval, terminating posteriorly with an obtuse but not very wide angle, slightly wider in female (
<figureCitation id="46C7D0FDFFEFFFFE54E8CF9A331EFD24" box="[993,1091,722,748]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="7" pageNumber="8">Fig. 4A</figureCitation>
); 5) elytral apex straight, epipleural angle obtuse; 6) male protarsus as in
<figureCitation id="46C7D0FDFFEFFFFE55EBCFBD3403FCD8" box="[738,862,757,784]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="7" pageNumber="8">Fig. 4BC</figureCitation>
, adhesive pallet with dense adhesive setae and small suction discs, distinctly longer than protibia; 7) aedeagus as in
<figureCitation id="46C7D0FDFFE0FFF154D0CC413368FEEB" box="[985,1077,265,291]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="8" pageNumber="9">Fig. 4D</figureCitation>
, median lobe broad, shorter than parameres, extending
<superScript id="29896130FFE0FFF156F7CC64355BFEF2" attach="right" box="[510,518,300,314]" fontSize="6" pageId="8" pageNumber="9">4</superScript>
/5 of their length; 10) female reproductive tract as in
<figureCitation id="46C7D0FDFFE0FFF15372CC64338BFE8E" box="[1147,1238,300,326]" captionStart="Fig" captionStartId="11.[189,232,1857,1883]" captionTargetBox="[204,1379,1111,1818]" captionTargetId="figure-216@11.[189,1398,1111,1818]" captionTargetPageId="11" captionText="Fig. 6. Female reproductive tract of new species, paratype with the same collection data as for holotype (SYSU). A. Patrus hainanensis sp. nov. B. Patrus jiangxiensis sp. nov. C. Patrus shangchuanensis sp. nov.Abbreviations: AG = accessory gland; BC = bursa copulatrix; BS = bursa sclerite; CO = common oviduct; FD = fertilization duct; GC = gonocoxa; LT = laterogtergite; SP = spermatheca; VA = vagina. Scale bars = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.5715542" httpUri="https://zenodo.org/record/5715542/files/figure.png" pageId="8" pageNumber="9">Fig. 6B</figureCitation>
, spermatheca elongate, C-shaped; fertilization duct slender, forming several loose hairpin hoops.
</paragraph>
<caption id="8A839CF0FFEFFFFE57B5CA2C355DF821" ID-DOI="http://doi.org/10.5281/zenodo.5715536" ID-Zenodo-Dep="5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="7" pageNumber="8" startId="7.[188,231,1892,1918]" targetBox="[189,1398,847,1855]" targetPageId="7">
<paragraph id="DE43CC78FFEFFFFE57B5CA2C355DF821" blockId="7.[187,1399,1892,2025]" pageId="7" pageNumber="8">
<emphasis id="EC88106AFFEFFFFE57B5CA2C3655F8B6" bold="true" box="[188,264,1892,1918]" pageId="7" pageNumber="8">Fig. 4.</emphasis>
<taxonomicName id="19FCB7FBFFEFFFFE5606CA2D36AFF8B6" authority="Liang &amp; Angus &amp; Jia, 2021" authorityName="Liang &amp; Angus &amp; Jia" authorityYear="2021" box="[271,498,1893,1918]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="jiangxiensis" status="sp. nov.">
<emphasis id="EC88106AFFEFFFFE5606CA2D36AFF8B6" box="[271,498,1893,1918]" italics="true" pageId="7" pageNumber="8">Patrus jiangxiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7BBAD11FFEFFFFE56FECA2D350FF8B7" box="[503,594,1893,1919]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ (SYSU).
<emphasis id="EC88106AFFEFFFFE55EACA2C35A5F8B6" bold="true" box="[739,760,1892,1918]" pageId="7" pageNumber="8">A</emphasis>
. Left elytron, paratype.
<emphasis id="EC88106AFFEFFFFE5302CA2D3342F8B7" bold="true" box="[1035,1055,1893,1919]" pageId="7" pageNumber="8">B</emphasis>
. Front leg, paratype.
<emphasis id="EC88106AFFEFFFFE521ACA2C3275F8B6" bold="true" box="[1299,1320,1892,1918]" pageId="7" pageNumber="8">C</emphasis>
. Front tarsus, paratype.
<emphasis id="EC88106AFFEFFFFE5682CAC036C3F86A" bold="true" box="[395,414,1928,1954]" pageId="7" pageNumber="8">D</emphasis>
. Genitalia, holotype.
<emphasis id="EC88106AFFEFFFFE55ACCAC035EBF86A" bold="true" box="[677,694,1928,1954]" pageId="7" pageNumber="8">E</emphasis>
. Surface sculpture on head, paratype.
<emphasis id="EC88106AFFEFFFFE538ECAC033CAF86A" bold="true" box="[1159,1175,1928,1954]" pageId="7" pageNumber="8">F</emphasis>
. Surface sculpture on pronotum, paratype.
<emphasis id="EC88106AFFEFFFFE56DCCAE336B1F80D" bold="true" box="[469,492,1963,1989]" pageId="7" pageNumber="8">G</emphasis>
. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm.
</paragraph>
</caption>
<paragraph id="DE43CC78FFE0FFF157B4CCD93377FDB6" blockId="8.[189,1399,401,639]" pageId="8" pageNumber="9">
This species is also a member of Group
<quantity id="1904619DFFE0FFF15598CCD935E0FE63" box="[657,701,401,427]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="9" unit="in" value="2.0">2 in</quantity>
Vaziranis (1984) key. It can be further assigned to the
<taxonomicName id="19FCB7FBFFE0FFF15248CCDA3231FE63" baseAuthorityName="Ochs" baseAuthorityYear="1932" box="[1345,1388,402,427]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="wui">
<emphasis id="EC88106AFFE0FFF15248CCDA3231FE63" box="[1345,1388,402,427]" italics="true" pageId="8" pageNumber="9">wui</emphasis>
</taxonomicName>
- subgroup proposed by
<bibRefCitation id="BA6DB189FFE0FFF156D8CCFC35C5FE06" author="Mazzoldi" box="[465,664,436,462]" firstAuthor="Mazzoldi" pageId="8" pageNumber="9" pagination="137 - 146" refId="ref16803" refString="Mazzoldi P. 1998. Gyrinidae: New species of Orectochilus Dejean, 1833 subgenus Patrus Aube, 1838 (Coleoptera). In: Jach M. A. &amp; Ji L. (eds) Water Beetles of China: 137 - 146. Zoologisch-Botanische Gesellschaft in Osterreichs and Wiener Coleopterologenverein, Vienna." type="book chapter" year="1998">Mazzoldi (1998)</bibRefCitation>
by the very large protarsus in the male. This species closely resembles
<taxonomicName id="19FCB7FBFFE0FFF15633CC9035F0FE39" authority="(Mazzoldi, 1998)" baseAuthorityName="Mazzoldi" baseAuthorityYear="1998" box="[314,685,471,497]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="wangi">
<emphasis id="EC88106AFFE0FFF15633CC903684FE39" box="[314,473,472,497]" italics="true" pageId="8" pageNumber="9">Patrus wangi</emphasis>
(
<bibRefCitation id="BA6DB189FFE0FFF156E0CC9F35F8FE39" author="Mazzoldi" box="[489,677,471,497]" firstAuthor="Mazzoldi" pageId="8" pageNumber="9" pagination="137 - 146" refId="ref16803" refString="Mazzoldi P. 1998. Gyrinidae: New species of Orectochilus Dejean, 1833 subgenus Patrus Aube, 1838 (Coleoptera). In: Jach M. A. &amp; Ji L. (eds) Water Beetles of China: 137 - 146. Zoologisch-Botanische Gesellschaft in Osterreichs and Wiener Coleopterologenverein, Vienna." type="book chapter" year="1998">Mazzoldi, 1998</bibRefCitation>
)
</taxonomicName>
described from
<collectingRegion id="1C38029AFFE0FFF15464CC9F3488FE39" box="[877,981,471,497]" country="China" name="Zhejiang" pageId="8" pageNumber="9">Zhejiang</collectingRegion>
and
<collectingRegion id="1C38029AFFE0FFF15307CC9F3306FE39" box="[1038,1115,471,497]" country="China" name="Anhui" pageId="8" pageNumber="9">Anhui</collectingRegion>
. These two species both have a well-developed protarsus which is distinctly longer than the protibia. However, it can be easily distinguished from the latter by the shape of the protarsus in the male (
<figureCitation id="46C7D0FDFFE0FFF15338CF5633CDFDF0" box="[1073,1168,542,568]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="8" pageNumber="9">Fig. 4B</figureCitation>
) and of the elytral glabrous region in the female. Besides, it is larger in size, the truncature of the protibia is more oblique and the median lobe of the aedeagus lacks the hook-like process (
<figureCitation id="46C7D0FDFFE0FFF154B7CF2C3340FDB6" box="[958,1053,612,638]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="8" pageNumber="9">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="DE43CC78FFE0FFF157B4CFED35D1FCCE" blockId="8.[188,1399,677,774]" pageId="8" pageNumber="9">
The new species is also similar to
<taxonomicName id="19FCB7FBFFE0FFF15540CFEE3468FD08" authority="(Ochs, 1932)" baseAuthorityName="Ochs" baseAuthorityYear="1932" box="[585,821,677,704]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="wui">
<emphasis id="EC88106AFFE0FFF15540CFEE35CCFD77" box="[585,657,678,703]" italics="true" pageId="8" pageNumber="9">P. wui</emphasis>
(
<bibRefCitation id="BA6DB189FFE0FFF155A8CFED3476FD08" author="Ochs" box="[673,811,677,704]" firstAuthor="Ochs" pageId="8" pageNumber="9" pagination="57 - 58" refId="ref17264" refString="Ochs G. 1932. Note on species of Gyrinidae from China. Peking Natural History Bulletin 6: 57 - 58." type="journal article" year="1932">Ochs, 1932</bibRefCitation>
)
</taxonomicName>
. Compared to
<taxonomicName id="19FCB7FBFFE0FFF154D4CFEE3378FD77" baseAuthorityName="Ochs" baseAuthorityYear="1932" box="[989,1061,678,703]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="wui">
<emphasis id="EC88106AFFE0FFF154D4CFEE3378FD77" box="[989,1061,678,703]" italics="true" pageId="8" pageNumber="9">P. wui</emphasis>
</taxonomicName>
, this species is larger in size, the protarsus is also larger, longer than the protibia, the glabrous region is less broad posteriorly, and the median lobe of the aedeagus is broader.
</paragraph>
</subSubSection>
<subSubSection id="96E69FF3FFE0FFF157B4CE6534FBFCBE" box="[189,934,813,886]" pageId="8" pageNumber="9" type="etymology">
<paragraph id="DE43CC78FFE0FFF157B4CE653618FC8F" blockId="8.[189,325,813,839]" box="[189,325,813,839]" pageId="8" pageNumber="9">
<heading id="850B7B14FFE0FFF157B4CE653618FC8F" bold="true" box="[189,325,813,839]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EC88106AFFE0FFF157B4CE653618FC8F" bold="true" box="[189,325,813,839]" pageId="8" pageNumber="9">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="DE43CC78FFE0FFF157B4CE1434FBFCBE" blockId="8.[189,934,860,886]" box="[189,934,860,886]" pageId="8" pageNumber="9">
This species is named after the
<typeStatus id="014772DAFFE0FFF15524CE14353DFCBE" box="[557,608,860,886]" pageId="8" pageNumber="9">type</typeStatus>
locality ʻJiangxi Provinceʼ.
</paragraph>
</subSubSection>
<subSubSection id="96E69FF3FFE0FFF157B4CED53374FA39" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="DE43CC78FFE0FFF157B4CED536F0FC7F" blockId="8.[189,429,925,951]" box="[189,429,925,951]" pageId="8" pageNumber="9">
<heading id="850B7B14FFE0FFF157B4CED536F0FC7F" bold="true" box="[189,429,925,951]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EC88106AFFE0FFF157B4CED536F0FC7F" bold="true" box="[189,429,925,951]" pageId="8" pageNumber="9">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DE43CC78FFE0FFF157E3CE843600FC2E" blockId="8.[189,1399,972,1069]" box="[234,349,972,998]" pageId="8" pageNumber="9">
<emphasis id="EC88106AFFE0FFF157E3CE843600FC2E" bold="true" box="[234,349,972,998]" pageId="8" pageNumber="9">
<typeStatus id="014772DAFFE0FFF157E3CE843600FC2E" box="[234,349,972,998]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DE43CC78FFE0FFF157B4CEA73390FBE4" blockId="8.[189,1399,972,1069]" pageId="8" pageNumber="9">
<materialsCitation id="6E94C625FFE0FFF157B4CEA73390FBE4" ID-GBIF-Occurrence="3409120352" collectingDate="2016-09-22" collectingDateMax="2016-09-24" collectingDateMin="2016-09-22" collectionCode="SYSU" collectorName="Ruijuan Zhang &amp; Yudan Tang" country="China" county="Luxi County" elevation="520" latitude="27.798056" location="Wugongshan" municipality="Pingxiang City" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" stateProvince="Jiangxi" typeStatus="holotype">
<collectingCountry id="A6EB8CE8FFE0FFF157B4CEA73645FBC1" box="[189,280,1007,1033]" name="China" pageId="8" pageNumber="9">CHINA</collectingCountry>
<emphasis id="EC88106AFFE0FFF1564CCEA736FCFBC1" bold="true" box="[325,417,1007,1033]" pageId="8" pageNumber="9">
<collectingRegion id="1C38029AFFE0FFF1564CCEA736FCFBC1" box="[325,417,1007,1033]" country="China" name="Jiangxi" pageId="8" pageNumber="9">Jiangxi</collectingRegion>
</emphasis>
<specimenCount id="C8FA07F1FFE0FFF156C3CEB836BEFBC2" box="[458,483,1008,1034]" pageId="8" pageNumber="9" type="male"></specimenCount>
;
<collectingRegion id="1C38029AFFE0FFF156FECEA73511FBC1" box="[503,588,1007,1033]" pageId="8" pageNumber="9">Jiangxi</collectingRegion>
Prov.
<collectingMunicipality id="3E275602FFE0FFF155AFCEA7343DFBC1" box="[678,864,1007,1033]" pageId="8" pageNumber="9">Pingxiang City</collectingMunicipality>
,
<collectingCounty id="3722B4F4FFE0FFF1547ACEA7334FFBC1" box="[883,1042,1007,1033]" pageId="8" pageNumber="9">Luxi County</collectingCounty>
,
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;
<geoCoordinate id="BBC8AABFFFE0FFF153DCCEA5322FFBC2" box="[1237,1394,1005,1034]" degrees="27" direction="north" minutes="47" orientation="latitude" pageId="8" pageNumber="9" precision="15" seconds="53" value="27.798056">27°4753″ N</geoCoordinate>
,
<geoCoordinate id="BBC8AABFFFE0FFF157B4C9593600FBE5" box="[189,349,1041,1069]" degrees="114" direction="east" minutes="14" orientation="longitude" pageId="8" pageNumber="9" seconds="83">114°1483″ E</geoCoordinate>
; alt.
<quantity id="1904619DFFE0FFF1569FC95A36BEFBE5" box="[406,483,1042,1069]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="8" pageNumber="9" unit="m" value="520.0">
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</quantity>
;
<date id="AA42EAB8FFE0FFF156E6C95A35E4FBE4" box="[495,697,1042,1068]" pageId="8" pageNumber="9" value="2016-09-22" valueMax="2016-09-24" valueMin="2016-09-22">
<collectingDate id="BA061350FFE0FFF156E6C95A35E4FBE4" box="[495,697,1042,1068]" pageId="8" pageNumber="9" value="2016-09-22" valueMax="2016-09-24" valueMin="2016-09-22">2224 Sep. 2016</collectingDate>
</date>
;
<collectorName id="7309A9AEFFE0FFF155CCC95A3429FBE4" box="[709,884,1042,1068]" pageId="8" pageNumber="9">Ruijuan Zhang</collectorName>
and
<collectorName id="7309A9AEFFE0FFF154A5C95A3365FBE5" box="[940,1080,1042,1069]" pageId="8" pageNumber="9">Yudan Tang</collectorName>
leg.;
<collectionCode id="B8ED54BDFFE0FFF15373C95A3394FBE4" box="[1146,1225,1042,1068]" pageId="8" pageNumber="9">SYSU</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="DE43CC78FFE0FFF157E3C91C3634FBA6" blockId="8.[189,1399,1108,1521]" box="[234,361,1108,1134]" pageId="8" pageNumber="9">
<emphasis id="EC88106AFFE0FFF157E3C91C3634FBA6" bold="true" box="[234,361,1108,1134]" pageId="8" pageNumber="9">
<typeStatus id="014772DAFFE0FFF157E3C91C3634FBA6" box="[234,361,1108,1134]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DE43CC78FFE0FFF157B4C93F3374FA39" blockId="8.[189,1399,1108,1521]" pageId="8" pageNumber="9">
<materialsCitation id="6E94C625FFE0FFF157B4C93F34A6FB59" ID-GBIF-Occurrence="3409120356" box="[189,1019,1143,1169]" collectingDate="2016-09-22" collectingDateMax="2016-09-24" collectingDateMin="2016-09-22" collectorName="Ruijuan Zhang &amp; Yudan Tang" country="China" county="Luxi County" elevation="520" latitude="27.798056" location="Wugongshan" municipality="Jiangxi Prov." pageId="8" pageNumber="9" specimenCount="30" stateProvince="Jiangxi" typeStatus="paratype">
<collectingCountry id="A6EB8CE8FFE0FFF157B4C93F3645FB59" box="[189,280,1143,1169]" name="China" pageId="8" pageNumber="9">CHINA</collectingCountry>
<emphasis id="EC88106AFFE0FFF1563AC93F36D2FB59" bold="true" box="[307,399,1143,1169]" pageId="8" pageNumber="9">
<collectingRegion id="1C38029AFFE0FFF1563AC93F36D2FB59" box="[307,399,1143,1169]" country="China" name="Jiangxi" pageId="8" pageNumber="9">Jiangxi</collectingRegion>
</emphasis>
<specimenCount id="C8FA07F1FFE0FFF156A1C93F36AFFB59" box="[424,498,1143,1169]" pageId="8" pageNumber="9" type="generic">30 ex.</specimenCount>
; same collection data as for holotype;
<collectionCode id="B8ED54BDFFE0FFF154B9C93F34A6FB59" box="[944,1019,1143,1169]" pageId="8" pageNumber="9">SYSU</collectionCode>
</materialsCitation>
<materialsCitation id="6E94C625FFE0FFF1531BC93F345BFB1F" ID-GBIF-Occurrence="3409120335" collectingDate="2016-09-19" collectingDateMax="2016-09-21" collectingDateMin="2016-09-19" collectorName="Ruijuan Zhang &amp; Yudan Tang" country="China" county="Jiangxi Prov. Shangyou County" elevation="183" latitude="25.915277" location="Nature Reserve" longLatPrecision="20" longitude="114.052505" municipality="Guanggushan" pageId="8" pageNumber="9" specimenCount="2" specimenCount-female="2" stateProvince="Jiangxi" typeStatus="paratype">
<specimenCount id="C8FA07F1FFE0FFF1531BC93F3308FB59" box="[1042,1109,1143,1169]" pageId="8" pageNumber="9" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="1C38029AFFE0FFF1536BC93F33EAFB59" box="[1122,1207,1143,1169]" pageId="8" pageNumber="9">Jiangxi</collectingRegion>
Prov.
<collectingCounty id="3722B4F4FFE0FFF15208C93F3648FB7C" pageId="8" pageNumber="9">Shangyou County</collectingCounty>
,
<collectingMunicipality id="3E275602FFE0FFF1562BC9D2369EFB7C" box="[290,451,1178,1204]" pageId="8" pageNumber="9">Guanggushan</collectingMunicipality>
(光姑山)
<location id="DB239AA3FFE0FFF1554AC9D335A3FB7D" LSID="urn:lsid:plazi:treatment:56557D6EFFEFFFF0557ACCC73391F9C1:DB239AA3FFE0FFF1554AC9D335A3FB7D" box="[579,766,1179,1205]" country="China" county="Jiangxi Prov. Shangyou County" latitude="25.915277" longLatPrecision="20" longitude="114.052505" municipality="Guanggushan" name="Nature Reserve" pageId="8" pageNumber="9" stateProvince="Jiangxi">Nature Reserve</location>
;
<geoCoordinate id="BBC8AABFFFE0FFF15405C9D034F9FB7D" box="[780,932,1176,1205]" degrees="25" direction="north" minutes="54" orientation="latitude" pageId="8" pageNumber="9" precision="15" seconds="55" value="25.915277">25°5455″ N</geoCoordinate>
,
<geoCoordinate id="BBC8AABFFFE0FFF154BBC9D03308FB7D" box="[946,1109,1176,1205]" degrees="114" direction="east" minutes="03" orientation="longitude" pageId="8" pageNumber="9" precision="15" seconds="09" value="114.052505">114°0309″ E</geoCoordinate>
; alt.
<quantity id="1904619DFFE0FFF1539BC9D233BFFB7D" box="[1170,1250,1178,1205]" metricMagnitude="2" metricUnit="m" metricValue="1.83" pageId="8" pageNumber="9" unit="m" value="183.0">
<elevation id="55D12B4BFFE0FFF1539BC9D233BFFB7D" box="[1170,1250,1178,1205]" metricMagnitude="2" metricUnit="m" metricValue="1.83" pageId="8" pageNumber="9" unit="m" value="183.0">183 m</elevation>
</quantity>
;
<date id="AA42EAB8FFE0FFF153F9C9D237A6FB1F" pageId="8" pageNumber="9" value="2016-09-19" valueMax="2016-09-21" valueMin="2016-09-19">
<collectingDate id="BA061350FFE0FFF153F9C9D237A6FB1F" pageId="8" pageNumber="9" value="2016-09-19" valueMax="2016-09-21" valueMin="2016-09-19">1921 Sep. 2016</collectingDate>
</date>
;
<collectorName id="7309A9AEFFE0FFF1560EC9F536EBFB1F" box="[263,438,1213,1239]" pageId="8" pageNumber="9">Ruijuan Zhang</collectorName>
and
<collectorName id="7309A9AEFFE0FFF156E7C9F53524FB10" box="[494,633,1213,1240]" pageId="8" pageNumber="9">Yudan Tang</collectorName>
leg.;
<collectionCode id="B8ED54BDFFE0FFF155B2C9F5345BFB1F" box="[699,774,1213,1239]" pageId="8" pageNumber="9">SYSU</collectionCode>
</materialsCitation>
<materialsCitation id="6E94C625FFE0FFF15416C9F533D6FB33" ID-GBIF-Occurrence="3409120384" collectingDate="2014-07-20" collectorName="Lin Renchao" country="China" elevation="690" latitude="29.04" location="Jiangxi" longLatPrecision="737" longitude="115.14" municipality="Guanyinyan" pageId="8" pageNumber="9" specimenCount="14" stateProvince="Jiangxi" typeStatus="paratype">
<specimenCount id="C8FA07F1FFE0FFF15416C9F53434FB10" box="[799,873,1213,1240]" pageId="8" pageNumber="9" type="generic">14 ex.</specimenCount>
; 江西靖安县观音岩 [
<collectingMunicipality id="3E275602FFE0FFF15366C9F53259FB1F" box="[1135,1284,1213,1239]" pageId="8" pageNumber="9">Guanyinyan</collectingMunicipality>
,
<location id="DB239AA3FFE0FFF15219C9F53236FB10" LSID="urn:lsid:plazi:treatment:56557D6EFFEFFFF0557ACCC73391F9C1:DB239AA3FFE0FFF15219C9F53236FB10" box="[1296,1387,1213,1240]" country="China" latitude="29.04" longLatPrecision="737" longitude="115.14" municipality="Guanyinyan" name="Jiangxi" pageId="8" pageNumber="9" stateProvince="Jiangxi">
<collectingRegion id="1C38029AFFE0FFF15219C9F53236FB10" box="[1296,1387,1213,1240]" pageId="8" pageNumber="9">Jiangxi</collectingRegion>
</location>
];
<geoCoordinate id="BBC8AABFFFE0FFF157B4C9A93642FB33" box="[189,287,1249,1275]" degrees="29.04" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="555" value="29.04">29.04 N</geoCoordinate>
,
<geoCoordinate id="BBC8AABFFFE0FFF15625C9A836C4FB33" box="[300,409,1248,1275]" degrees="115.14" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="555" value="115.14">115.14 E</geoCoordinate>
; alt.
<quantity id="1904619DFFE0FFF156DDC9A8357FFB33" box="[468,546,1248,1275]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="8" pageNumber="9" unit="m" value="690.0">
<elevation id="55D12B4BFFE0FFF156DDC9A8357FFB33" box="[468,546,1248,1275]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="8" pageNumber="9" unit="m" value="690.0">690 m</elevation>
</quantity>
;
<date id="AA42EAB8FFE0FFF15539C9A93598FB33" box="[560,709,1248,1275]" pageId="8" pageNumber="9" value="2014-07-20">
<collectingDate id="BA061350FFE0FFF15539C9A93598FB33" box="[560,709,1248,1275]" pageId="8" pageNumber="9" value="2014-07-20">20 Jul. 2014</collectingDate>
</date>
; 林仁超采 [
<collectorName id="7309A9AEFFE0FFF1545EC9A834AFFB32" box="[855,1010,1248,1274]" pageId="8" pageNumber="9">Lin Renchao</collectorName>
leg.];
<collectionCode id="B8ED54BDFFE0FFF15349C9A933D6FB33" box="[1088,1163,1249,1275]" pageId="8" pageNumber="9">SYSU</collectionCode>
</materialsCitation>
<materialsCitation id="6E94C625FFE0FFF153AFC9A934E4FA89" ID-GBIF-Occurrence="3409120427" collectingDate="2014-07-22" collectorName="Lin Renchao" country="China" county="Jiangxi" elevation="660" latitude="29.04" location="Sanzhualun" longLatPrecision="737" longitude="115.11" municipality="Jing'an" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" stateProvince="Jiangxi" typeStatus="paratype">
<specimenCount id="C8FA07F1FFE0FFF153AFC9A93389FB33" box="[1190,1236,1249,1275]" pageId="8" pageNumber="9" type="female">1 ♀</specimenCount>
; 江西靖安县 三爪仑乡白水洞景区 [
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<collectingRegion id="1C38029AFFE0FFF156DEC84C356CFAD6" box="[471,561,1284,1310]" pageId="8" pageNumber="9">Jiangxi</collectingRegion>
</collectingCounty>
,
<collectingMunicipality id="3E275602FFE0FFF15549C84C35C4FAD6" box="[576,665,1284,1310]" pageId="8" pageNumber="9">Jingan</collectingMunicipality>
,
<location id="DB239AA3FFE0FFF155A0C84C3468FAD6" LSID="urn:lsid:plazi:treatment:56557D6EFFEFFFF0557ACCC73391F9C1:DB239AA3FFE0FFF155A0C84C3468FAD6" box="[681,821,1284,1310]" country="China" county="Jiangxi" latitude="29.04" longLatPrecision="737" longitude="115.11" municipality="Jing'an" name="Sanzhualun" pageId="8" pageNumber="9" stateProvince="Jiangxi">Sanzhualun</location>
,
<location id="DB239AA3FFE0FFF1544AC84C3324FAD6" LSID="urn:lsid:plazi:treatment:56557D6EFFEFFFF0557ACCC73391F9C1:DB239AA3FFE0FFF1544AC84C3324FAD6" box="[835,1145,1283,1310]" country="China" county="Jiangxi" latitude="29.04" longLatPrecision="737" longitude="115.11" municipality="Jing'an" name="Baishuidong Scenic Area" pageId="8" pageNumber="9" stateProvince="Jiangxi">Baishuidong Scenic Area</location>
];
<geoCoordinate id="BBC8AABFFFE0FFF15386C84C33A9FAD6" box="[1167,1268,1284,1310]" degrees="29.04" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="555" value="29.04">29.04 N</geoCoordinate>
,
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; alt.
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;
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</date>
; 林仁超采 [
<collectorName id="7309A9AEFFE0FFF15577C86F3441FA89" box="[638,796,1319,1345]" pageId="8" pageNumber="9">Lin Renchao</collectorName>
leg.];
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<specimenCount id="C8FA07F1FFE0FFF154D0C86F3356FA89" box="[985,1035,1319,1345]" pageId="8" pageNumber="9" type="male">1 ♂</specimenCount>
; 江西井冈山白银湖 [
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,
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,
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]; alt.
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;
<date id="AA42EAB8FFE0FFF1558BC802347FFAAC" box="[642,802,1354,1380]" pageId="8" pageNumber="9" value="2011-04-23">
<collectingDate id="BA061350FFE0FFF1558BC802347FFAAC" box="[642,802,1354,1380]" pageId="8" pageNumber="9" value="2011-04-23">23 Apr. 2011</collectingDate>
</date>
; 采集人: 赵爽 [
<collectorName id="7309A9AEFFE0FFF154EFC80233DAFAAC" box="[998,1159,1354,1380]" pageId="8" pageNumber="9">Zhao Shuang</collectorName>
leg.];
<collectionCode id="B8ED54BDFFE0FFF153D1C802327EFAAC" box="[1240,1315,1354,1380]" pageId="8" pageNumber="9">SYSU</collectionCode>
</materialsCitation>
<materialsCitation id="6E94C625FFE0FFF15248C8023696FA63" ID-GBIF-Occurrence="3409120437" collectingDate="2011-04-28" collectorName="Jia Fenglong" country="China" county="Jiangxi" location="Huyangta" municipality="Jinggangshan" pageId="8" pageNumber="9" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Jiangxi" typeStatus="paratype">
<specimenCount id="C8FA07F1FFE0FFF15248C802322CFAAD" box="[1345,1393,1354,1381]" pageId="8" pageNumber="9" type="male">1 ♂</specimenCount>
,
<specimenCount id="C8FA07F1FFE0FFF157B4C8253658FA40" box="[189,261,1389,1416]" pageId="8" pageNumber="9" type="female">3 ♀♀</specimenCount>
; 江西井冈山湖羊塔 [
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,
<collectingMunicipality id="3E275602FFE0FFF1558BC8253478FA4F" box="[642,805,1389,1415]" pageId="8" pageNumber="9">Jinggangshan</collectingMunicipality>
,
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];
<date id="AA42EAB8FFE0FFF154CCC8253337FA4F" box="[965,1130,1389,1415]" pageId="8" pageNumber="9" value="2011-04-28">
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</date>
; 采集人: 贾凤龙 [
<collectorName id="7309A9AEFFE0FFF1525AC8253671FA62" pageId="8" pageNumber="9">Jia Fenglong</collectorName>
leg.];
<collectionCode id="B8ED54BDFFE0FFF15689C8D93696FA63" box="[384,459,1425,1451]" pageId="8" pageNumber="9">SYSU</collectionCode>
</materialsCitation>
<materialsCitation id="6E94C625FFE0FFF156E4C8D93613FA06" ID-GBIF-Occurrence="3409120382" collectingDate="2011-04-01" collectorName="Li Yun" country="China" location="Jinggang Shan" municipality="Jiangxi" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" stateProvince="Jiangxi" typeStatus="paratype">
<specimenCount id="C8FA07F1FFE0FFF156E4C8D9357CFA63" box="[493,545,1425,1451]" pageId="8" pageNumber="9" type="male">1 ♂</specimenCount>
; 江西井冈山 [
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<collectingRegion id="1C38029AFFE0FFF155DEC8D8346CFA62" box="[727,817,1424,1450]" pageId="8" pageNumber="9">Jiangxi</collectingRegion>
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,
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];
<date id="AA42EAB8FFE0FFF15319C8D933F9FA63" box="[1040,1188,1425,1451]" pageId="8" pageNumber="9" value="2011-04-01">
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</date>
; 李韵采 [
<collectorName id="7309A9AEFFE0FFF15229C8D8322BFA63" box="[1312,1398,1424,1451]" pageId="8" pageNumber="9">Li Yun</collectorName>
leg.];
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<specimenCount id="C8FA07F1FFE0FFF15662C8FC36C1FA06" box="[363,412,1460,1486]" pageId="8" pageNumber="9" type="male">1 ♂</specimenCount>
; W.
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,
<collectingMunicipality id="3E275602FFE0FFF15531C8FC35F3FA06" box="[568,686,1460,1486]" pageId="8" pageNumber="9">Jian City</collectingMunicipality>
,
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,
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;
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,
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; alt.
<quantity id="1904619DFFE0FFF157E0C89F3619FA39" box="[233,324,1495,1521]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="8" pageNumber="9" unit="m" value="1130.0">
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;
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<collectingDate id="BA061350FFE0FFF15659C89F36B9FA39" box="[336,484,1495,1521]" pageId="8" pageNumber="9" value="2014-08-02">2 Aug. 2014</collectingDate>
</date>
;
<collectorName id="7309A9AEFFE0FFF156F9C89F356CFA39" box="[496,561,1495,1521]" pageId="8" pageNumber="9">Chen</collectorName>
,
<collectorName id="7309A9AEFFE0FFF15535C89F353EFA39" box="[572,611,1495,1521]" pageId="8" pageNumber="9">Hu</collectorName>
,
<collectorName id="7309A9AEFFE0FFF15566C89F35CDFA39" box="[623,656,1495,1521]" pageId="8" pageNumber="9">Lv</collectorName>
and
<collectorName id="7309A9AEFFE0FFF155C1C89F35B4FA39" box="[712,745,1495,1521]" pageId="8" pageNumber="9">Yu</collectorName>
leg.; in the stream;
<collectionCode id="B8ED54BDFFE0FFF154D8C89F3378FA39" box="[977,1061,1495,1521]" pageId="8" pageNumber="9">SHNU</collectionCode>
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.
</paragraph>
</subSubSection>
<subSubSection id="96E69FF3FFE0FFF057B4CB503591FA51" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
<paragraph id="DE43CC78FFE0FFF157B4CB503612F9FA" blockId="8.[189,335,1560,1586]" box="[189,335,1560,1586]" pageId="8" pageNumber="9">
<heading id="850B7B14FFE0FFF157B4CB503612F9FA" bold="true" box="[189,335,1560,1586]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EC88106AFFE0FFF157B4CB503612F9FA" bold="true" box="[189,335,1560,1586]" pageId="8" pageNumber="9">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DE43CC78FFE0FFF157B4CB0F34C9F94C" blockId="8.[188,1399,1607,1669]" pageId="8" pageNumber="9">
MEASUREMENTS.
<typeStatus id="014772DAFFE0FFF15675CB0F36B3F9A9" box="[380,494,1607,1633]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
:
<quantity id="1904619DFFE0FFF156F1CB0F350FF9A9" box="[504,594,1607,1634]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="9" unit="mm" value="8.5">8.5 mm</quantity>
in length,
<quantity id="1904619DFFE0FFF155C5CB0F347BF9A9" box="[716,806,1607,1633]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="8" pageNumber="9" unit="mm" value="3.9">3.9 mm</quantity>
in width; male:
<quantity id="1904619DFFE0FFF154EACB0F3332F9A9" box="[995,1135,1607,1633]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" metricValueMax="9.8" metricValueMin="8.4" pageId="8" pageNumber="9" unit="mm" value="9.1" valueMax="9.8" valueMin="8.4">8.49.8 mm</quantity>
in length, 3.94.0 mm in width; female: 7.0
<quantity id="1904619DFFE0FFF156C9CB223544F94D" box="[448,537,1642,1669]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="8" pageNumber="9" unit="mm" value="8.6">8.6 mm</quantity>
in length,
<quantity id="1904619DFFE0FFF1559FCB223479F94D" box="[662,804,1642,1669]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.9" metricValueMin="3.1" pageId="8" pageNumber="9" unit="mm" value="3.5" valueMax="3.9" valueMin="3.1">3.13.9 mm</quantity>
in width.
</paragraph>
<paragraph id="DE43CC78FFE0FFF157B4CBE434CEF8E7" blockId="8.[189,1399,1707,1840]" pageId="8" pageNumber="9">
HABITUS AND COLOUR (
<figureCitation id="46C7D0FDFFE0FFF156C6CBE3350CF90E" box="[463,593,1707,1734]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[189,1398,359,1889]" captionTargetId="figure-21@4.[189,1398,359,1889]" captionTargetPageId="4" captionText="Fig. 1. Habitus. AC. Patrus hainanensis sp. nov., ♂, paratype (SYSU). DF. Patrus jiangxiensis. sp. nov., ♂, paratypes (SYSU). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5715528" httpUri="https://zenodo.org/record/5715528/files/figure.png" pageId="8" pageNumber="9">Fig. 1DF</figureCitation>
). Medium-sized species, oblong-oval in shape, widest just before middle, moderately convex in lateral view. Dorsal side black with strong bronze sheen. Pronotum and elytra with yellow lateral borders. Ventral side mostly reddish brown to dark brown, with legs and epipleura yellow to reddish yellow, last abdomen ferruginous.
</paragraph>
<paragraph id="DE43CC78FFE0FFF057B4CA1F3673FEA1" blockId="8.[186,1398,1877,2011]" lastBlockId="9.[189,1399,265,361]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
HEAD. Labrum wide and short, ⅓ as long as wide, with anterior margin straight; anterior surface smooth; posterior surface with strong punctate-tomentose. Clypeus slightly shorter than labrum, with anterior margin straight. Fronto-clypeal suture distinct.Surface of clypeus with weak micro-reticulation anteriorly, while distinct posteriorly, among meshes of micro-reticulation with dense irregular punctation. Surface on frons and vertex with strong micro-reticulation formed by isodiametric polygonal meshes, gradually obliterated backwards. Amongst meshes of the micro-reticulation strong irregular-shaped punctation (
<figureCitation id="46C7D0FDFFE1FFF057CFCC07367FFEA1" box="[198,290,335,361]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="9" pageNumber="10">Fig. 4E</figureCitation>
).
</paragraph>
<paragraph id="DE43CC78FFE1FFF057B4CCD9322FFCBC" blockId="9.[188,1400,401,885]" pageId="9" pageNumber="10">
THORAX. Pronotum regularly attenuated from base to apex, with margins weakly convex. Surface of pronotal glabrous region with distinct micro-reticulation formed by isodiametric polygonal meshes and strong irregular punctation (
<figureCitation id="46C7D0FDFFE1FFF05500CC9F353FFE39" box="[521,610,471,497]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="9" pageNumber="10">Fig. 4F</figureCitation>
), much of which merges in rows to form distinct wrinkles. Lateral pubescence anteriorly expanded inward onto pronotal disc to external third of eye, posteriorly weakly reduced backwards in concave line. Scutellum wide and short, less than ½ as long as wide, with surface smooth. Pronoto-elytral glabrous region regular oval, terminating posteriorly with obtuse but not very wide angle (
<figureCitation id="46C7D0FDFFE1FFF05646CF2C36F3FDB6" box="[335,430,612,638]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="9" pageNumber="10">Fig. 4A</figureCitation>
). Lateral pubescent band in male anteriorly as wide as pronotal one, subparallel to elytral margin backwards until middle of elytron, then regularly expanded, reaching suture at about onetenth of distance from truncature; in female anteriorly as wide as pronotal one, parallel to elytral margin backwards until ⅔ of elytron, then regularly expanded, reaching suture at about
<superScript id="29896130FFE1FFF0538FCF8533D2FD13" attach="right" box="[1158,1167,717,731]" fontSize="6" pageId="9" pageNumber="10">1</superScript>
/
<subScript id="4278CE3DFFE1FFF0539FCF9E33FAFD2C" attach="left" box="[1174,1191,726,740]" fontSize="6" pageId="9" pageNumber="10">12</subScript>
of distance from truncature, so that elytral glabrous region in female slightly wider. Surface of elytral glabrous region covered by weak micro-reticulation formed by transverse or oblique and slightly elongate polygonal meshes and by strong punctation formed by short transverse or oblique strioles (
<figureCitation id="46C7D0FDFFE1FFF0536CCE7F3399FC99" box="[1125,1220,823,849]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="9" pageNumber="10">Fig. 4G</figureCitation>
). Truncature of elytra straight; epipleural angle obtuse, narrowly rounded, sutural angle almost 90°, narrowly rounded.
</paragraph>
<paragraph id="DE43CC78FFE1FFF057B4CED43692FC34" blockId="9.[188,1398,923,1020]" pageId="9" pageNumber="10">
MALE GENITALIA (
<figureCitation id="46C7D0FDFFE1FFF0568ECED436BEFC7E" box="[391,483,924,950]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="9" pageNumber="10">Fig. 4D</figureCitation>
). Median lobe shorter than parameres extending
<superScript id="29896130FFE1FFF0532CCED33370FC61" attach="right" box="[1061,1069,923,937]" fontSize="6" pageId="9" pageNumber="10">4</superScript>
/5 of their length, basal third parallel, then slightly narrowed until apical third, sub-apex parallel, apical part regularly narrowed, apex point shortly narrowed.
</paragraph>
<paragraph id="DE43CC78FFE1FFF057B4C96C34C8FB6F" blockId="9.[187,1398,1060,1192]" pageId="9" pageNumber="10">
FEMALE REPRODUCTIVE TRACT (
<figureCitation id="46C7D0FDFFE1FFF05506C96C3537FBF7" box="[527,618,1060,1087]" captionStart="Fig" captionStartId="11.[189,232,1857,1883]" captionTargetBox="[204,1379,1111,1818]" captionTargetId="figure-216@11.[189,1398,1111,1818]" captionTargetPageId="11" captionText="Fig. 6. Female reproductive tract of new species, paratype with the same collection data as for holotype (SYSU). A. Patrus hainanensis sp. nov. B. Patrus jiangxiensis sp. nov. C. Patrus shangchuanensis sp. nov.Abbreviations: AG = accessory gland; BC = bursa copulatrix; BS = bursa sclerite; CO = common oviduct; FD = fertilization duct; GC = gonocoxa; LT = laterogtergite; SP = spermatheca; VA = vagina. Scale bars = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.5715542" httpUri="https://zenodo.org/record/5715542/files/figure.png" pageId="9" pageNumber="10">Fig. 6B</figureCitation>
). Gonocoxosternite elongate, apically narrowly protruding, anterior process short; laterotergite elongate, slightly narrowed towards apex; vagina elongate; spermatheca elongate, C-shaped, strongly curved; fertilization duct slender, forming several loose hairpin hoops; accessory gland short and slender, inserting dorsally on bursa.
</paragraph>
<paragraph id="DE43CC78FFE1FFF057B4C9873591FA51" blockId="9.[189,1401,1231,1433]" pageId="9" pageNumber="10">
SEXUAL DIMORPHISM. Male slightly larger in size than female; protibia in male short and wide, short triangular, strongly expanded exteriorly towards apex, apex of protibia obliquely truncate, anteroexternal angle obtuse, rounded (
<figureCitation id="46C7D0FDFFE1FFF05534C85D35C6FAE7" box="[573,667,1301,1327]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="9" pageNumber="10">Fig. 4B</figureCitation>
), while shorter and less expanded, and apex straight-truncate in female. Protarsus of male large, strongly expanded, distinctly longer than protibia, slightly narrowed towards apex, ventral side with dense adhesive setae and small suction discs (
<figureCitation id="46C7D0FDFFE1FFF05344C81333F7FABE" box="[1101,1194,1371,1398]" captionStart="Fig" captionStartId="7.[188,231,1892,1918]" captionTargetBox="[189,1398,847,1855]" captionTargetId="figure-129@7.[189,1398,847,1855]" captionTargetPageId="7" captionText="Fig. 4. Patrus jiangxiensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg, paratype.C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715536" httpUri="https://zenodo.org/record/5715536/files/figure.png" pageId="9" pageNumber="10">Fig. 4C</figureCitation>
); elytral glabrous region slightly broader posteriorly in female.
</paragraph>
</subSubSection>
<subSubSection id="96E69FF3FFE1FFF057B4C8883391F9C1" box="[189,1228,1472,1545]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="DE43CC78FFE1FFF057B4C8883605FA12" blockId="9.[189,344,1472,1498]" box="[189,344,1472,1498]" pageId="9" pageNumber="10">
<heading id="850B7B14FFE1FFF057B4C8883605FA12" bold="true" box="[189,344,1472,1498]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="EC88106AFFE1FFF057B4C8883605FA12" bold="true" box="[189,344,1472,1498]" pageId="9" pageNumber="10">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DE43CC78FFE1FFF057B4C8A73391F9C1" blockId="9.[189,1228,1519,1545]" box="[189,1228,1519,1545]" pageId="9" pageNumber="10">
Known from Pingxiang, Shangyou, Jingxi and Jinggangshan in
<collectingRegion id="1C38029AFFE1FFF054A2C8A7332FF9C1" box="[939,1138,1519,1545]" country="China" name="Jiangxi" pageId="9" pageNumber="10">Jiangxi Province</collectingRegion>
,
<collectingCountry id="A6EB8CE8FFE1FFF05377C8A73395F9C1" box="[1150,1224,1519,1545]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>