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<mods:namePart>Matalin, Andrey V.</mods:namePart>
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<mods:affiliation>Moscow State Pedagogical University, Department of Zoology &amp; Ecology, Kibalchicha str. 6, build. 3, Moscow 129164, Russia. andrei-matalin @ yandex. ru; https: // orcid. org / 0000 - 0002 - 7790 - 8709 &amp; Pirogov National Research Medical University, Department of Biology, Ostrovitianova str. 1, Moscow 117997, Russia.</mods:affiliation>
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<taxonomicName ID-CoL="8LNYT" authority="Tan, 1981" authorityName="Tan" authorityYear="1981" box="[151,514,366,393]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="motoensis">
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<emphasis bold="true" box="[151,385,366,393]" italics="true" pageId="20" pageNumber="421">Therates motoensis</emphasis>
<bibRefCitation author="Tan, J. - J." box="[392,514,366,393]" pageId="20" pageNumber="421" pagination="331 - 335" refId="ref27058" refString="Tan, J. - J. (1981) Cicindelidae. In: Insects of Xizang (Tibet). Vol. I. Comprehensive Scientific Expedition to the Qinghai-Xizang Plateau, Chinese Academy of Sciences. Science Press, Beijing, pp. 331 - 335. [in Chinese, with English summary]" type="book chapter" year="1981">Tan, 1981</bibRefCitation>
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<figureCitation box="[151,278,403,429]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="20" pageNumber="421">Figs 13, 14</figureCitation>
, 
<figureCitation box="[290,362,403,429]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="20" pageNumber="421">40, 41</figureCitation>
, 
<figureCitation box="[374,446,403,429]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="20" pageNumber="421">62, 63</figureCitation>
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<figureCitation box="[458,530,403,429]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="20" pageNumber="421">97, 98</figureCitation>
, 
<figureCitation box="[542,587,403,429]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="20" pageNumber="421">172</figureCitation>
.
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<emphasis box="[151,368,439,465]" italics="true" pageId="20" pageNumber="421">Therates motoensis</emphasis>
<bibRefCitation author="Tan, J. - J." box="[375,544,439,465]" pageId="20" pageNumber="421" pagination="331 - 335" refId="ref27058" refString="Tan, J. - J. (1981) Cicindelidae. In: Insects of Xizang (Tibet). Vol. I. Comprehensive Scientific Expedition to the Qinghai-Xizang Plateau, Chinese Academy of Sciences. Science Press, Beijing, pp. 331 - 335. [in Chinese, with English summary]" type="book chapter" year="1981">Tan, 1981: 332</bibRefCitation>
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, Mêdog).
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<taxonomicName ID-CoL="8MKSW" authority="W. Horn, 1913" authorityName="W. Horn" authorityYear="1913" box="[151,598,507,531]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="vitalisi">
<emphasis box="[151,442,507,531]" italics="true" pageId="20" pageNumber="421">Therates fruhstorferi vitalisi</emphasis>
W. 
<bibRefCitation author="Horn, W." box="[485,598,507,531]" pageId="20" pageNumber="421" pagination="362 - 366" refId="ref25772" refString="Horn, W. (1913) Materiaux pour servir a l'etude de la faune entomologique de l'Indochine. Cicindelinae. Annales de la Societe Entomologique de Belgique, 57, 362 - 366. https: // doi. org / 10.5962 / bhl. part. 4597" type="journal article" year="1913">Horn, 1913</bibRefCitation>
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<bibRefCitation author="Wiesner, J." box="[824,918,507,531]" pageId="20" pageNumber="421" refId="ref27575" refString="Wiesner, J. (1992) Verzeichnis der Sandlaufkafer der Welt, Checklist of the Tiger Beetles of the World. Vol. 27. Beitrag zur Kenntnis der Cicindelidae. Verlag Erna Bauer, Keltern, 364 pp." type="book" year="1992">1992: 92</bibRefCitation>
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; 
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<bibRefCitation author="Lorenz, W." box="[930,1103,507,531]" pageId="20" pageNumber="421" refId="ref26095" refString="Lorenz, W. (1998) Systematic list of extant ground beetles of the word (Insects Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). First Edition. Published by the author, Tutzing, Germany, 503 pp." type="book" year="1998">Lorenz 1998: 38</bibRefCitation>
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; 
<treatmentCitation author="Puchkov, A. V. &amp; Matalin, A. V." page="115" pageId="20" pageNumber="421" year="2003">
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<taxonomicName ID-CoL="8LNYT" authority="Tan, 1981" authorityName="Tan" authorityYear="1981" box="[151,451,568,593]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="motoensis">
<emphasis box="[151,347,568,592]" italics="true" pageId="20" pageNumber="421">Therates motoensis</emphasis>
<bibRefCitation author="Tan, J. - J." box="[354,451,569,593]" pageId="20" pageNumber="421" pagination="331 - 335" refId="ref27058" refString="Tan, J. - J. (1981) Cicindelidae. In: Insects of Xizang (Tibet). Vol. I. Comprehensive Scientific Expedition to the Qinghai-Xizang Plateau, Chinese Academy of Sciences. Science Press, Beijing, pp. 331 - 335. [in Chinese, with English summary]" type="book chapter" year="1981">Tan, 1981</bibRefCitation>
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<bibRefCitation author="Wu, X. - Q." box="[478,612,569,593]" pageId="20" pageNumber="421" pagination="21 - 43" refId="ref27836" refString="Wu, X. - Q. (2011) A revised list of the tiger beetles of China (Coleoptera: Cicindelidae). Cicindela, 43 (2), 21 - 43." type="journal article" year="2011">Wu 2011: 31</bibRefCitation>
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, 37; 
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<bibRefCitation author="Wiesner, J. &amp; Bi, W. - X." box="[661,912,568,593]" pageId="20" pageNumber="421" pagination="233 - 236" refId="ref27747" refString="Wiesner, J. &amp; Bi, W. - X. (2014) Taxonomical notes on some species of Therates Latreille, 1817 from Xizang, China (Coleoptera: Cicindelidae). Entomologische Zeitschrift, Schwanfeld, 124 (4), 233 - 236." type="journal article" year="2014">Wiesner &amp; Bi 2014: 233</bibRefCitation>
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; Putchkov &amp; Matalin 2017: 245; 
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3ÔÔ 
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<collectingCountry box="[486,562,676,703]" name="China" pageId="20" pageNumber="421">China</collectingCountry>
, 
<collectingRegion box="[584,668,675,701]" country="China" name="Xizang" pageId="20" pageNumber="421">Xizang</collectingRegion>
, 
<collectingMunicipality box="[688,766,675,701]" pageId="20" pageNumber="421">Motuo</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:050B87D78D54247AE1B81818FC917184:887D601A8D54247AE23C1BD5FCA274BC" box="[787,866,675,701]" country="China" municipality="Motuo" name="Hanmi" pageId="20" pageNumber="421" stateProvince="Xizang">Hanmi</location>
, 
<quantity box="[886,983,675,701]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="20" pageNumber="421" unit="m" value="2100.0">
<elevation box="[886,983,675,701]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="20" pageNumber="421" unit="m" value="2100.0">2100 m</elevation>
</quantity>
, 
<date box="[1005,1186,674,701]" pageId="20" pageNumber="421" value="2013-07-15" valueMax="2013-07-30" valueMin="2013-07-15">
<collectingDate box="[1005,1186,674,701]" pageId="20" pageNumber="421" value="2013-07-15" valueMax="2013-07-30" valueMin="2013-07-15">15–30.VII.2013</collectingDate>
</date>
, leg. 
<collectorName box="[1265,1432,675,701]" pageId="20" pageNumber="421">Wen-Xuan Bi</collectorName>
</materialsCitation>
; 
<materialsCitation box="[151,558,711,739]" collectingDate="2013-07-12" collectingDateMax="2013-07-24" collectingDateMin="2013-07-12" collectorName="Wen-Xuan Bi" country="China" elevation="2100" location="Hanmi" municipality="Motuo" pageId="20" pageNumber="421" specimenCount="17" specimenCount-female="17" stateProvince="Xizang">
19ÔÔ 
<specimenCount box="[226,285,712,739]" pageId="20" pageNumber="421" type="female">17♀♀</specimenCount>
— 
<emphasis box="[319,363,711,737]" italics="true" pageId="20" pageNumber="421">ibid</emphasis>
, 
<date box="[376,558,711,737]" pageId="20" pageNumber="421" value="2013-07-12" valueMax="2013-07-24" valueMin="2013-07-12">
<collectingDate box="[376,558,711,737]" pageId="20" pageNumber="421" value="2013-07-12" valueMax="2013-07-24" valueMin="2013-07-12">12–24.VII.2013</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation box="[569,953,710,739]" collectingDate="2013-07-25" collectingDateMax="2013-07-31" collectingDateMin="2013-07-25" collectorName="Wen-Xuan Bi" country="China" elevation="2100" location="Hanmi" municipality="Motuo" pageId="20" pageNumber="421" specimenCount="8" specimenCount-female="8" stateProvince="Xizang">
6ÔÔ 
<specimenCount box="[630,674,712,739]" pageId="20" pageNumber="421" type="female">8♀♀</specimenCount>
— 
<emphasis box="[709,753,711,737]" italics="true" pageId="20" pageNumber="421">ibid</emphasis>
, 
<date box="[765,953,710,737]" pageId="20" pageNumber="421" value="2013-07-25" valueMax="2013-07-31" valueMin="2013-07-25">
<collectingDate box="[765,953,710,737]" pageId="20" pageNumber="421" value="2013-07-25" valueMax="2013-07-31" valueMin="2013-07-25">25–31. VII.2013</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation box="[964,1342,711,739]" collectingDate="2011-07-23" collectingDateMax="2011-07-28" collectingDateMin="2011-07-23" collectorName="Wen-Xuan Bi" country="China" elevation="2100" location="Hanmi" municipality="Motuo" pageId="20" pageNumber="421" specimenCount="2" specimenCount-female="2" stateProvince="Xizang">
6ÔÔ 
<specimenCount box="[1024,1070,712,739]" pageId="20" pageNumber="421" type="female">2♀♀</specimenCount>
— 
<emphasis box="[1104,1148,711,737]" italics="true" pageId="20" pageNumber="421">ibid</emphasis>
, 
<date box="[1160,1342,711,737]" pageId="20" pageNumber="421" value="2011-07-23" valueMax="2011-07-28" valueMin="2011-07-23">
<collectingDate box="[1160,1342,711,737]" pageId="20" pageNumber="421" value="2011-07-23" valueMax="2011-07-28" valueMin="2011-07-23">23–28.VII.2011</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation collectingDate="2013-08-18" collectorName="Wen-Xuan Bi" country="China" elevation="1850" location="Motuo" pageId="20" pageNumber="421" specimenCount="1" stateProvince="Xizang">
2ÔÔ— 
<collectingCountry box="[151,221,747,773]" name="China" pageId="20" pageNumber="421">China</collectingCountry>
, 
<collectingRegion box="[232,316,747,773]" country="China" name="Xizang" pageId="20" pageNumber="421">Xizang</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:050B87D78D54247AE1B81818FC917184:887D601A8D54247AE0681B9DFE557504" box="[327,405,747,773]" country="China" name="Motuo" pageId="20" pageNumber="421" stateProvince="Xizang">Motuo</location>
, 
<quantity box="[416,573,747,773]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="20" pageNumber="421" unit="m" value="1850.0" valueMax="2000.0" valueMin="1700.0">
<elevation box="[416,573,747,773]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="20" pageNumber="421" unit="m" value="1850.0" valueMax="2000.0" valueMin="1700.0">1700–2000 m</elevation>
</quantity>
, 
<date box="[584,733,747,773]" pageId="20" pageNumber="421" value="2013-08-18">
<collectingDate box="[584,733,747,773]" pageId="20" pageNumber="421" value="2013-08-18">18.VIII.2013</collectingDate>
</date>
, leg. 
<collectorName box="[792,950,747,773]" pageId="20" pageNumber="421">Wen-Xuan Bi</collectorName>
</materialsCitation>
; 
<materialsCitation collectingDate="2012-08-03" collectorName="Chao Wu" country="China" elevation="2100" location="Motuo" pageId="20" pageNumber="421" specimenCount="1" specimenCount-female="1" stateProvince="Xizang">
2ÔÔ 
<specimenCount box="[1020,1043,748,775]" pageId="20" pageNumber="421" type="female">1♀</specimenCount>
— 
<collectingCountry box="[1075,1150,748,775]" name="China" pageId="20" pageNumber="421">China</collectingCountry>
, 
<collectingRegion box="[1161,1245,747,773]" country="China" name="Xizang" pageId="20" pageNumber="421">Xizang</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:050B87D78D54247AE1B81818FC917184:887D601A8D54247AE5C71B9DFAF67504" box="[1256,1334,747,773]" country="China" name="Motuo" pageId="20" pageNumber="421" stateProvince="Xizang">Motuo</location>
, 
<quantity box="[1344,1431,747,773]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="20" pageNumber="421" unit="m" value="2100.0">
<elevation box="[1344,1431,747,773]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="20" pageNumber="421" unit="m" value="2100.0">2100 m</elevation>
</quantity>
, 
<date box="[151,286,783,809]" pageId="20" pageNumber="421" value="2012-08-03">
<collectingDate box="[151,286,783,809]" pageId="20" pageNumber="421" value="2012-08-03">3.VIII.2012</collectingDate>
</date>
, leg. 
<collectorName box="[347,456,783,809]" pageId="20" pageNumber="421">Chao Wu</collectorName>
</materialsCitation>
; 
<materialsCitation box="[468,1298,782,809]" collectingDate="2005-08" collectorName="Hao Huang" country="China" location="Motuo" pageId="20" pageNumber="421" specimenCount="1" specimenCount-female="1" stateProvince="Xizang">
<specimenCount box="[468,491,783,809]" pageId="20" pageNumber="421" type="female">1♀</specimenCount>
— 
<collectingCountry box="[524,600,783,809]" name="China" pageId="20" pageNumber="421">China</collectingCountry>
, 
<collectingRegion box="[613,697,783,809]" country="China" name="Xizang" pageId="20" pageNumber="421">Xizang</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:050B87D78D54247AE1B81818FC917184:887D601A8D54247AE3EB1A79FCD27528" box="[708,786,783,809]" country="China" name="Motuo" pageId="20" pageNumber="421" stateProvince="Xizang">Motuo</location>
, Hanmi-A’niqiao, 
<date box="[994,1108,782,809]" pageId="20" pageNumber="421" value="2005-08">
<collectingDate box="[994,1108,782,809]" pageId="20" pageNumber="421" value="2005-08">VIII.2005</collectingDate>
</date>
, leg. 
<collectorName box="[1169,1298,783,809]" pageId="20" pageNumber="421">Hao Huang</collectorName>
</materialsCitation>
(
<bibRefCitation author="Wiesner, J. &amp; Bi, W. - X." pageId="20" pageNumber="421" pagination="233 - 236" refId="ref27747" refString="Wiesner, J. &amp; Bi, W. - X. (2014) Taxonomical notes on some species of Therates Latreille, 1817 from Xizang, China (Coleoptera: Cicindelidae). Entomologische Zeitschrift, Schwanfeld, 124 (4), 233 - 236." type="journal article" year="2014">Wiesner &amp; Bi 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="421" type="diagnosis">
<paragraph blockId="20.[151,1437,638,1925]" pageId="20" pageNumber="421">
<emphasis bold="true" box="[199,322,854,880]" pageId="20" pageNumber="421">Diagnosis.</emphasis>
From all related species 
<taxonomicName authorityName="Tan" authorityYear="1981" box="[605,746,855,880]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="motoensis">
<emphasis box="[605,746,855,880]" italics="true" pageId="20" pageNumber="421">T. motoensis</emphasis>
</taxonomicName>
is readily distinguished by the bluish-green topside (
<figureCitation captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="20" pageNumber="421">Figs 13, 14</figureCitation>
); by the colour of labrum (
<figureCitation box="[483,612,891,917]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="20" pageNumber="421">Figs 62, 63</figureCitation>
); as well as by the aedeagus clearly dilated in apical third (
<figureCitation box="[1272,1371,891,917]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="20" pageNumber="421">Fig. 172</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="421" type="description">
<paragraph blockId="20.[151,1437,638,1925]" box="[199,898,926,953]" pageId="20" pageNumber="421">
<emphasis bold="true" box="[199,373,926,952]" pageId="20" pageNumber="421">Redescription.</emphasis>
TL = 
<quantity box="[442,604,927,953]" metricMagnitude="-2" metricUnit="m" metricValue="1.205" metricValueMax="1.29" metricValueMin="1.1199999999999999" pageId="20" pageNumber="421" unit="mm" value="12.05" valueMax="12.9" valueMin="11.2">11.2–12.9 mm</quantity>
(mean = 
<quantity box="[710,812,927,953]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="20" pageNumber="421" unit="mm" value="12.1">12.1 mm</quantity>
, n = 6).
</paragraph>
<paragraph blockId="20.[151,1437,638,1925]" pageId="20" pageNumber="421">
Head shining bluish-green with light golden reflection; orbital plates striated throughout or finely striated in posterior third; frons convex, semi-circular, smooth or slightly rugose, frontal sulci deep especially in anterior third, convergent in anterior 2/3, shallow and distinctly divergent in posterior third. Mandibles light brown underside, pale-yellow topside with light-brown teeth and apical molar. Labial and maxillary palpi yellow with slightly darker basal palpomeres. Antennae lanceolate, extending posteriorly nearly to the elytral shoulders in males, somewhat shorter in the females, scape yellow or yellow-brown on anterior side and brown-black on posterior side, remaining antennomeres black, antennomeres 2–5 with light blue tinge, antennomeres 6–11 approximately equal width without protruding downward anterior lower margins (
<figureCitation box="[690,827,1215,1241]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="20" pageNumber="421">Figs 40, 41</figureCitation>
). Labrum indistinctly transverse, LW/LL = 1.0–1.14 (mean = 1.07, n = 4), with six–eight apical teeth and seven–nine setae between them, yellowish with black margins and large basal spot in males (
<figureCitation box="[489,572,1287,1313]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="20" pageNumber="421">Fig. 62</figureCitation>
), black with yellowish oval apical spot in females (
<figureCitation box="[1147,1230,1287,1313]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="20" pageNumber="421">Fig. 63</figureCitation>
).
</paragraph>
<paragraph blockId="20.[151,1437,638,1925]" pageId="20" pageNumber="421">Pronotum slightly transverse, PW/PL = 1.0–1.1 (mean = 1.03, n = 4), shining bluish-green with light golden tinge, apical lobe slightly wider in middle portion; thorax greenish-blue with light golden tinge.</paragraph>
<paragraph blockId="20.[151,1437,638,1925]" pageId="20" pageNumber="421">Fore and middle femora pale-yellow along posterior side and black-brown with bluish tinge along anterior side, hind femora pale-yellow in basal half and black with blue tinge in apical half; fore and middle tibiae pale-yellow except narrow brown-black apices, hind tibiae pale-yellow except brown-black basal third, distinctly wider than fore and middle tibiae; tarsomeres 1 and 2 of fore- and middle legs pale-yellow or light brown with darker apices, other tarsomeres dark-brown, all hind tarsomeres pale-yellow except dark-brown apices; HTbL/HTaL = 1.09–1.15 (mean = 1.12, n = 2).</paragraph>
<paragraph blockId="20.[151,1437,638,1925]" pageId="20" pageNumber="421">
Elytra prolonged, indistinctly divergent toward apex, EL/EW = 2.04–2.13 (mean = 2.08, n = 4); longer in females—EL/PL = 4.0–4.25 (mean = 4.13, n = 2) than in males—3.7–3.8 (mean = 3.75, n = 2); shining green with golden reflection; punctuation deep and regular between shoulders and central dot, shallower behind, in apical quarter virtually indistinct; scutellum dark blue with golden-green tinge; apical margin gradually rounded. Elytral pattern presented by very small invisible dorsally yellow basal portion of humeral lunule and small trapezoid yellow central dot (
<figureCitation box="[285,414,1791,1817]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="20" pageNumber="421">Figs 13, 14</figureCitation>
, 
<figureCitation box="[426,500,1791,1817]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="20" pageNumber="421">97, 98</figureCitation>
).
</paragraph>
<paragraph blockId="20.[151,1437,638,1925]" pageId="20" pageNumber="421">
Aedeagus clearly dilated in apical third, with gradually sinuate, prolonged apex (
<figureCitation box="[1105,1202,1827,1853]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="20" pageNumber="421">Fig. 172</figureCitation>
), AL = 
<quantity box="[1288,1437,1826,1853]" metricMagnitude="-3" metricUnit="m" metricValue="2.825" metricValueMax="2.9" metricValueMin="2.75" pageId="20" pageNumber="421" unit="mm" value="2.825" valueMax="2.9" valueMin="2.75">2.75–2.9 mm</quantity>
(mean = 2.8, n = 2), EL/AL = 2.69.
</paragraph>
</subSubSection>
<subSubSection box="[199,849,1898,1925]" pageId="20" pageNumber="421" type="distribution">
<paragraph blockId="20.[151,1437,638,1925]" box="[199,849,1898,1925]" pageId="20" pageNumber="421">
<emphasis bold="true" box="[199,353,1898,1924]" pageId="20" pageNumber="421">Distribution.</emphasis>
<collectingCountry box="[359,437,1899,1925]" name="China" pageId="20" pageNumber="421">CHINA</collectingCountry>
: 
<collectingRegion box="[447,528,1899,1925]" country="China" name="Xizang" pageId="20" pageNumber="421">Xizang</collectingRegion>
(Mêdog County) (
<figureCitation box="[738,836,1899,1925]" captionStart="FIGURES 184" captionStartId="28.[152,270,1579,1603]" captionTargetBox="[151,1435,597,1556]" captionTargetId="figure-226@28.[151,1435,597,1556]" captionTargetPageId="28" captionText="FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png." figureDoi="http://doi.org/10.5281/zenodo.7758916" httpUri="https://zenodo.org/record/7758916/files/figure.png" pageId="20" pageNumber="421">Fig. 184</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>