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<document ID-DOI="10.11646/zootaxa.5256.5.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7758872" ID-ZooBank="E5B9708E-D626-4923-825A-A9FD59AE4024" approvalRequired="3" approvalRequired_for_textStreams="2" approvalRequired_for_treatments="1" checkinTime="1679468565563" checkinUser="plazi" docAuthor="Matalin, Andrey V. &amp; Wiesner, Jürgen" docDate="2023" docId="050B87D78D482466E1B81BD1FD3671CC" docLanguage="en" docName="zootaxa.5256.5.1.pdf" docOrigin="Zootaxa 5256 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.5.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Therates ziyardamensis Matalin et Wiesner 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="409" masterDocId="F932FFAF8D40246EE12F1976FFC07601" masterDocTitle="Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)" masterLastPageNumber="433" masterPageNumber="401" pageNumber="409" updateTime="1679469078842" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Matalin, Andrey V.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-7790-8709</mods:nameIdentifier>
<mods:affiliation>Moscow State Pedagogical University, Department of Zoology &amp; Ecology, Kibalchicha str. 6, build. 3, Moscow 129164, Russia. andrei-matalin @ yandex. ru; https: // orcid. org / 0000 - 0002 - 7790 - 8709 &amp; Pirogov National Research Medical University, Department of Biology, Ostrovitianova str. 1, Moscow 117997, Russia.</mods:affiliation>
<mods:nameIdentifier type="email">andrei-matalin@yandex.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wiesner, Jürgen</mods:namePart>
<mods:affiliation>Dresdener Ring 11, D- 38444 Wolfsburg, Germany.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5256</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>5</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>401</mods:start>
<mods:end>433</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5256.5.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5256.5.1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7758872</mods:identifier>
<mods:identifier type="ZooBank">E5B9708E-D626-4923-825A-A9FD59AE4024</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7758898" ID-Zenodo-Dep="7758898" LSID="urn:lsid:plazi:treatment:050B87D78D482466E1B81BD1FD3671CC" httpUri="http://treatment.plazi.org/id/050B87D78D482466E1B81BD1FD3671CC" lastPageNumber="409" pageId="8" pageNumber="409">
<subSubSection box="[151,761,679,706]" pageId="8" pageNumber="409" type="nomenclature">
<paragraph blockId="8.[151,761,679,743]" box="[151,761,679,706]" pageId="8" pageNumber="409">
<heading bold="true" box="[151,761,679,706]" fontSize="11" level="1" pageId="8" pageNumber="409" reason="1">
<emphasis bold="true" box="[151,761,679,706]" pageId="8" pageNumber="409">
<taxonomicName authority="Matalin et Wiesner" authorityName="Matalin et Wiesner" authorityYear="2023" box="[151,682,679,706]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="ziyardamensis" status="sp. n.">
<emphasis bold="true" box="[151,435,679,706]" italics="true" pageId="8" pageNumber="409">Therates ziyardamensis</emphasis>
Matalin et Wiesner
</taxonomicName>
,
<taxonomicNameLabel box="[695,761,680,706]" pageId="8" pageNumber="409" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,424,716,743]" pageId="8" pageNumber="409" type="description">
<paragraph blockId="8.[151,761,679,743]" box="[151,424,716,743]" pageId="8" pageNumber="409">
<figureCitation box="[151,236,716,742]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="8" pageNumber="409">Figs 12</figureCitation>
,
<figureCitation box="[248,278,716,743]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="8" pageNumber="409">35</figureCitation>
,
<figureCitation box="[290,320,716,742]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="8" pageNumber="409">64</figureCitation>
,
<figureCitation box="[332,362,716,742]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="8" pageNumber="409">96</figureCitation>
,
<figureCitation box="[374,419,716,742]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="8" pageNumber="409">131</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[151,1106,784,809]" pageId="8" pageNumber="409" type="reference_group">
<paragraph blockId="8.[151,1106,784,809]" box="[151,1106,784,809]" pageId="8" pageNumber="409">
<treatmentCitationGroup box="[151,1106,784,809]" pageId="8" pageNumber="409">
<taxonomicName ID-CoL="8MKSW" authority="W. Horn, 1913" authorityName="W. Horn" authorityYear="1913" box="[151,590,784,808]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="vitalisi">
<emphasis box="[151,438,784,808]" italics="true" pageId="8" pageNumber="409">Therates fruhstorferi vitalisi</emphasis>
W.
<bibRefCitation author="Horn, W." box="[478,590,784,808]" pageId="8" pageNumber="409" pagination="362 - 366" refId="ref25772" refString="Horn, W. (1913) Materiaux pour servir a l'etude de la faune entomologique de l'Indochine. Cicindelinae. Annales de la Societe Entomologique de Belgique, 57, 362 - 366. https: // doi. org / 10.5962 / bhl. part. 4597" type="journal article" year="1913">Horn, 1913</bibRefCitation>
</taxonomicName>
—
<treatmentCitation author="Wiesner, J." box="[617,814,784,809]" page="372" pageId="8" pageNumber="409" year="1999">
<bibRefCitation author="Wiesner, J." box="[617,814,784,809]" pageId="8" pageNumber="409" pagination="369 - 372" refId="ref27616" refString="Wiesner, J. (1999) Zwei neue Therates Latreille 1817 from Burma (Coleoptera: Cicindelidae). Entomologische Zeitschrift, 109 (9), 369 - 372." type="journal article" year="1999">Wiesner 1999: 372</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Cassola, F. &amp; Klicha, M." box="[824,1101,784,809]" page="39" pageId="8" pageNumber="409" year="2002">
<bibRefCitation author="Cassola, F. &amp; Klicha, M." box="[824,1101,784,809]" pageId="8" pageNumber="409" pagination="31 - 43" refId="ref25300" refString="Cassola, F. &amp; Klicha, M. (2002) Full specific status of Calochroa corbetti (W. Horn, 1899), and notes on the tiger beetle fauna of Myanmar (Burma) (Coleoptera: Cicindelidae). Mitteilungen des Internationalen Entomologischen Vereins, 27 (1 / 2), 31 - 43." type="journal article" year="2002">Cassola &amp; Klícha 2002: 39</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="409" type="materials_examined">
<paragraph blockId="8.[151,1437,854,1997]" pageId="8" pageNumber="409">
<emphasis bold="true" box="[151,322,854,881]" pageId="8" pageNumber="409">Type material.</emphasis>
<materialsCitation collectingDate="1998-05-21" collectionCode="V, JW" collectorName="S. Murzin &amp; Sinyaev" country="Myanmar" elevation="1250" location="Putao" pageId="8" pageNumber="409" specimenCount="1" typeStatus="holotype">
<typeStatus box="[326,444,855,881]" pageId="8" pageNumber="409">HOLOTYPE</typeStatus>
, Ô—
<collectingCountry box="[502,651,856,883]" name="Myanmar" pageId="8" pageNumber="409">MYANMAR</collectingCountry>
N (Burma),
<quantity box="[785,853,854,881]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="409" unit="km" value="65.0">65 km</quantity>
NW
<location LSID="urn:lsid:plazi:treatment:050B87D78D482466E1B81BD1FD3671CC:887D601A8D482466E2A31A21FC0F7570" box="[908,975,855,881]" country="Myanmar" name="Putao" pageId="8" pageNumber="409">Putao</location>
,
<quantity box="[983,1067,854,881]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="409" unit="m" value="1250.0">
<elevation box="[983,1067,854,881]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="409" unit="m" value="1250.0">1250 m</elevation>
</quantity>
, Zi Yar Dam vill., 18–
<date box="[1301,1432,854,881]" pageId="8" pageNumber="409" value="1998-05-21">
<collectingDate box="[1301,1432,854,881]" pageId="8" pageNumber="409" value="1998-05-21">21.05.1998</collectingDate>
</date>
, leg.
<collectorName box="[199,311,891,917]" pageId="8" pageNumber="409">S. Murzin</collectorName>
&amp;
<collectionCode box="[347,367,891,917]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="409" type="Museum">V</collectionCode>
.
<collectorName box="[377,467,891,917]" pageId="8" pageNumber="409">Sinyaev</collectorName>
(
<collectionCode box="[483,526,891,917]" pageId="8" pageNumber="409">JW</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="409" type="diagnosis">
<paragraph blockId="8.[151,1437,854,1997]" pageId="8" pageNumber="409">
<emphasis bold="true" box="[151,274,926,952]" pageId="8" pageNumber="409">Diagnosis.</emphasis>
This new species is reliably distinguished from
<taxonomicName authorityName="W. Horn" authorityYear="1913" box="[817,921,927,953]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis box="[817,921,927,953]" italics="true" pageId="8" pageNumber="409">T. vitalisi</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Tan, Mo, Liang" authorityYear="1991" box="[936,1078,927,953]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="biserratus">
<emphasis box="[936,1078,927,953]" italics="true" pageId="8" pageNumber="409">T. biserratus</emphasis>
</taxonomicName>
,
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[1091,1245,927,953]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="hunanensis" status="sp. n.">
<emphasis box="[1091,1245,927,953]" italics="true" pageId="8" pageNumber="409">T. hunanensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1253,1317,927,953]" pageId="8" pageNumber="409">
<taxonomicNameLabel box="[1253,1317,927,953]" pageId="8" pageNumber="409" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName authorityName="W. Horn" authorityYear="1912" box="[1331,1436,927,952]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis box="[1331,1436,927,952]" italics="true" pageId="8" pageNumber="409">T. sauteri</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Tan" authorityYear="1981" box="[199,339,963,988]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="motoensis">
<emphasis box="[199,339,963,988]" italics="true" pageId="8" pageNumber="409">T. motoensis</emphasis>
</taxonomicName>
by apex of aedeagus without lateral carinae (
<figureCitation box="[848,942,963,989]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="8" pageNumber="409">Fig. 131</figureCitation>
<emphasis box="[949,980,963,988]" italics="true" pageId="8" pageNumber="409">vs.</emphasis>
<figureCitation box="[987,1145,962,989]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="8" pageNumber="409">Figs 152–174</figureCitation>
). From
<taxonomicName authorityName="W. Horn" authorityYear="1902" box="[1232,1388,963,989]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="fruhstorferi">
<emphasis box="[1232,1388,963,989]" italics="true" pageId="8" pageNumber="409">T. fruhstorferi</emphasis>
</taxonomicName>
and
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[151,334,999,1025]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="pseudovitalisi" status="sp. n.">
<emphasis box="[151,334,999,1025]" italics="true" pageId="8" pageNumber="409">T. pseudovitalisi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[341,404,999,1025]" pageId="8" pageNumber="409">
<taxonomicNameLabel box="[341,404,999,1025]" pageId="8" pageNumber="409" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
this new species is recognized by thin apex of aedeagus (
<figureCitation box="[1045,1139,999,1025]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="8" pageNumber="409">Fig. 131</figureCitation>
<emphasis box="[1145,1176,999,1024]" italics="true" pageId="8" pageNumber="409">vs.</emphasis>
<figureCitation box="[1182,1334,998,1025]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="8" pageNumber="409">Figs 115–130</figureCitation>
and
<figureCitation captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="8" pageNumber="409">Figs 132–135</figureCitation>
, respectively) as well as by the rough punctuation of the elytra (
<figureCitation box="[969,1055,1035,1061]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="8" pageNumber="409">Figs 12</figureCitation>
,
<figureCitation box="[1067,1095,1035,1061]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="8" pageNumber="409">96</figureCitation>
<emphasis box="[1102,1133,1035,1060]" italics="true" pageId="8" pageNumber="409">vs.</emphasis>
<figureCitation box="[1140,1280,1035,1061]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="8" pageNumber="409">Figs 1, 3–11</figureCitation>
,
<figureCitation box="[1292,1366,1034,1061]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="8" pageNumber="409">81–95</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="409" type="description">
<paragraph blockId="8.[151,1437,854,1997]" pageId="8" pageNumber="409">
<emphasis bold="true" box="[199,345,1070,1096]" pageId="8" pageNumber="409">Description.</emphasis>
TL =
<quantity box="[427,534,1071,1097]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="8" pageNumber="409" unit="mm" value="10.6">10.6 mm</quantity>
. Head black with distinct violet reflection; orbital plates finely but distinctly striated in posterior third; frons semi-circular, with distinct impression and transverse rugoses (see above), slightly downward anteriorly (see at side), frontal sulci very deep, distinctly convergent in anterior half and shallow and sharply divergent posteriorly; occiput flat with shallow impression anteriorly. Mandibles dark brown underside, yellow-pale topside, with brown teeth and apical molar. Labial palpi pale with dark-brown apical palpomeres; maxillary palpomeres 1 and 2 light brown, palpomeres 3 and 4 dark brown. Antennae not extend shoulders, black, antennomeres 1–5 with light violet tinge and brown apices, antennomeres 9–11 club-shaped, antennomeres 9 and 10 serrated, with downward anterior lower margins, antennomere 11 clearly dilated with abrupt apical margin (
<figureCitation captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="8" pageNumber="409">Fig. 35</figureCitation>
). Labrum as wide as long (LL = LW—
<quantity box="[619,706,1359,1385]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="409" unit="mm" value="1.4">1.4 mm</quantity>
), black-brown with large elongate apical yellow spot unseparated from central apical teeth (
<figureCitation box="[439,524,1395,1421]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="8" pageNumber="409">Fig. 64</figureCitation>
).
</paragraph>
<paragraph blockId="8.[151,1437,854,1997]" pageId="8" pageNumber="409">
Pronotum shining black with blue-violet reflection; as wide as long (PL = PW—
<quantity box="[1114,1202,1431,1457]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="409" unit="mm" value="1.9">1.9 mm</quantity>
), apical lobe slightly wider in medial part; thorax black with distinct blue reflection.
</paragraph>
<paragraph blockId="8.[151,1437,854,1997]" pageId="8" pageNumber="409">Fore femora pale along anterior margin and black with violet tinge along posterior margin; middle femora pale in basal half and black with violet tinge in apical half; hind femora pale in basal third and black with violet tinge in apical 2/3; tibiae black with distinct violet lustre; tarsi black with light violet tinge except brown apices of all apical tarsomeres; HTbL/HTaL = 1.22.</paragraph>
<paragraph blockId="8.[151,1437,854,1997]" pageId="8" pageNumber="409">
Elytra black-blue with bright purple-violet reflection; indistinctly divergent in apical third, EL/EW = 1.94 (EL =
<quantity box="[174,262,1683,1709]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="8" pageNumber="409" unit="mm" value="6.6">6.6 mm</quantity>
, EW =
<quantity box="[345,435,1683,1709]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="409" unit="mm" value="3.4">3.4 mm</quantity>
); punctuation deep and regular in basal 3/4 except humeral area, in apical quarter shallow and tenuous; scutellum black with violet tinge, apical margin rounded but shortly top-cut. Elytral pattern presented by small pale basal portion of humeral lunule (see in front) and relatively short practically transverse central dot indistinctly dilated in lateral portion (
<figureCitation box="[569,656,1791,1817]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="8" pageNumber="409">Figs 12</figureCitation>
,
<figureCitation box="[668,700,1791,1817]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="8" pageNumber="409">96</figureCitation>
).
</paragraph>
<paragraph blockId="8.[151,1437,854,1997]" pageId="8" pageNumber="409">
Aedeagus (only apical half) with thin elongate slightly sinuate rounded apex, without lateral carinae (
<figureCitation captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="8" pageNumber="409">Fig. 131</figureCitation>
).
</paragraph>
<paragraph blockId="8.[151,1437,854,1997]" box="[199,401,1899,1925]" pageId="8" pageNumber="409">Female unknown.</paragraph>
</subSubSection>
<subSubSection box="[199,1135,1935,1961]" pageId="8" pageNumber="409" type="etymology">
<paragraph blockId="8.[151,1437,854,1997]" box="[199,1135,1935,1961]" pageId="8" pageNumber="409">
<emphasis bold="true" box="[199,334,1935,1961]" pageId="8" pageNumber="409">Etymology.</emphasis>
This new species is named after the
<typeStatus box="[745,793,1935,1961]" pageId="8" pageNumber="409">type</typeStatus>
locality—Zi Yar Dam village.
</paragraph>
</subSubSection>
<subSubSection box="[199,758,1970,1997]" pageId="8" pageNumber="409" type="distribution">
<paragraph blockId="8.[151,1437,854,1997]" box="[199,758,1970,1997]" pageId="8" pageNumber="409">
<emphasis bold="true" box="[199,353,1970,1996]" pageId="8" pageNumber="409">Distribution.</emphasis>
<collectingCountry box="[359,478,1971,1997]" name="Myanmar" pageId="8" pageNumber="409">MYANMAR</collectingCountry>
:
<collectingRegion box="[488,632,1971,1997]" country="Myanmar" name="Kachin" pageId="8" pageNumber="409">Kachin State</collectingRegion>
(
<figureCitation box="[647,745,1971,1997]" captionStart="FIGURES 184" captionStartId="28.[152,270,1579,1603]" captionTargetBox="[151,1435,597,1556]" captionTargetId="figure-226@28.[151,1435,597,1556]" captionTargetPageId="28" captionText="FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png." figureDoi="http://doi.org/10.5281/zenodo.7758916" httpUri="https://zenodo.org/record/7758916/files/figure.png" pageId="8" pageNumber="409">Fig. 184</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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